identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0388F8254C36CA74FE90FF68FC01FAF0.text	0388F8254C36CA74FE90FF68FC01FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helcogramma inclinata (Fowler 1946)	<div><p>Helcogramma inclinata (Fowler, 1946)</p> <p>[Standard Japanese name: Aya-hebigimpo] (Figs 5E–F, 6C; Table 2)</p> <p>Enneapterygius inclinatus Fowler, 1946: 191, fig. 55 (type locality: Aguni-jima island, Ryukyu Islands).</p> <p>Helcogramma habena Williams and McCormick, 1990: 1020, figs 2, 6A–B, 8 (type locality: Batan Island, Batan Islands, Philippines).</p> <p>Helcogramma ellioti (not of Herre, 1944): Hayashi 1993: 944 (Ryukyu Islands); Hayashi 2000: 1079 (Ryukyu Islands); Kato 2011: 222, unnumbered fig. (Yaku-shima island, Ryukyu Islands).</p> <p>Helcogramma fuscopinna (not of Holleman, 1982): Aizawa 1997: 560, unnumbered fig. (Iriomote-jima island, Ryukyu Islands).</p> <p>Helcogramma inclinatum: Fricke 1997: 446, fig. 91 (Ishigaki-jima island, Ryukyu Islands); Hayashi 2013: 1282, unnumbered fig. (Yaku-shima and Aguni-jima islands, Ryukyu Islands).</p> <p>Helcogramma inclinata: Williams and Howe 2003: 164 (Aguni-jima island, Ryukyu Islands); Senou et al. 2006: 84 (Ie-jima island, Ryukyu Islands); Motomura et al. 2006: 107, figs 2–4 (Satsuma Peninsula, Kyushu; Yaku-shima island, Ryukyu Islands); Yoshigou and Nakamura 2008: 70 (in part; HMNH-P 5739, 7425, 7426, 8625, and 9023 from Kume-jima and Okinawa-jima islands, Ryukyu Islands); Motomura et al. 2010: 188, fig. 434 (Yaku-shima island, Ryukyu Islands); Tashiro 2013: 294, unnumbered figs (Iou-jima and Take-shima islands, Ryukyu Islands).</p> <p>Helcogramma nesion (not of Williams and Howe, 2003): Yoshigou and Nakamura 2008: 71 (in part; HMNH-P 7945 from Okinawa-jima island, Ryukyu Islands).</p> <p>Helcogramma sp.: Yoshigou and Nakamura 2008: 71 (in part; HMNH-P 7910 from Okinoerabu-jima island, Ryukyu Islands).</p> <p>Type material examined. Enneapterygius inclinatus: ANSP 72067 (holotype), male, 26.1 mm SL, Aguni-jima island, Ryukyu Islands, E. Tinkham, 27 July 1945.</p> <p>Non-type material examined. 107 specimens, 13.5–51.5 mm SL. SATSUMA PENINSULA, KYUSHU: KAUM –I. 4246, female, 50.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.16917&amp;materialsCitation.latitude=31.374722" title="Search Plazi for locations around (long 130.16917/lat 31.374722)">Kurose Beach</a>, Kasasa, Minamisatsuma, 31°22′29″N, 130°10′09″E, 2 m depth, M. Ito, 28 May 2007; KAUM –I. 6522, male, 50.9 mm SL, KAUM –I. 6576, male, 43.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.16917&amp;materialsCitation.latitude=31.374722" title="Search Plazi for locations around (long 130.16917/lat 31.374722)">Kurose Beach</a>, Kasasa, Minamisatsuma, 31°22′29″N, 130°10′09″E, 2 m depth, M. Ito, 9 September 2007. TAKE-SHIMA ISLAND: KAUM –I. 29668, male, 43.9 mm SL, KAUM –I. 29669, male, 42.5 mm SL, KAUM –I. 29683, male, 13.8 mm SL, KAUM –I. 29684, female, 15.2 mm SL, off south coast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.40916&amp;materialsCitation.latitude=30.808887" title="Search Plazi for locations around (long 130.40916/lat 30.808887)">Ombozaki</a>, 30°48′32″N, 130°24′33″E, 5–20 m depth, KAUM Fish Team, 29 May 2010; KAUM –I. 37989, male, 45.6 mm SL, off south coast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.40916&amp;materialsCitation.latitude=30.808887" title="Search Plazi for locations around (long 130.40916/lat 30.808887)">Ombozaki</a>, 30°48′32″N, 130°24′33″E, 5–20 m depth, KAUM Fish Team, 18 May 2011. IOU-JIMA ISLAND: KAUM –I. 29529, male, 13.5 mm SL, off west coast, 30°47′04″N, 130°15′42″E, 5–20 m depth, KAUM Fish Team, 25 May 2010; KAUM –I. 29592, male, 17.9 mm SL, off west coast, 30°47′04″N, 130°15′42″E, 5–20 m depth, KAUM Fish Team, 28 May 2010; KAUM –I. 30625, female, 49.2 mm SL, KAUM –I. 30626, male, 31.3 mm SL, off west coast, 30°47′04″N, 130°15′42″E, 2–22 m depth, KAUM Fish Team, 27 June 2010; KAUM –I. 37834, male, 44.2 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.28082&amp;materialsCitation.latitude=30.771944" title="Search Plazi for locations around (long 130.28082/lat 30.771944)">Harumase</a>, 30°46′19″N, 130°16′51″E, 10–15 m depth, KAUM Fish Team, 16 May 2011. YAKU-SHIMA ISLAND: KAUM –I. 34, male, 44.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.48834&amp;materialsCitation.latitude=30.457224" title="Search Plazi for locations around (long 130.48834/lat 30.457224)">Isso</a>, 30°27′26″N, 130°29′18″E, 3 m depth, S. Harazaki, 20 January 2006; KAUM –I. 11189, male, 43.1 mm SL, KAUM –I. 11190, female, 42.7 mm SL, KAUM –I. 11242, female, 44.0 mm SL, KAUM –I. 11243, female, 41.0 mm SL, KAUM –I. 11763, male, 32.3 mm SL, south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.41444&amp;materialsCitation.latitude=30.2675" title="Search Plazi for locations around (long 130.41444/lat 30.2675)">Kamazenohana</a>, Kurio, 30°16′03″N, 130°24′52″E, 0–3 m depth, KAUM Fish Team, 10 August 2008; KAUM –I. 11424, male, 50.8 mm SL, KAUM –I. 11425, female, 50.3 mm SL, KAUM –I. 11426, male, 34.4 mm SL, KAUM –I. 11427, female, 36.5 mm SL, KAUM –I. 11428, female, 24.9 mm SL, KAUM –I. 11731, female, 28.0 mm SL, KAUM –I. 11732, female, 46.6 mm SL, KAUM –I. 11733, female, 36.5 mm SL, KAUM –I. 11734, female, 32.2 mm SL, KAUM –I. 11735, female, 42.7 mm SL, KAUM –I. 11736, female, 51.5 mm SL, KAUM –I. 11737, female, 26.5 mm SL, KAUM –I. 11738, female, 25.5 mm SL, KAUM –I. 24443, male, 21.7 mm SL, west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.41306&amp;materialsCitation.latitude=30.265833" title="Search Plazi for locations around (long 130.41306/lat 30.265833)">Kamazenohana</a>, Kurio, 30°15′57″N, 130°24′47″E, 0–3 m depth, KAUM Fish Team, 12 August 2008; KAUM –I. 11472, female, 26.1 mm SL, KAUM –I. 11473, female, 22.9 mm SL, KAUM –I. 11474, male, 29.9 mm SL, KAUM –I. 11478, male, 23.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.49973&amp;materialsCitation.latitude=30.45639" title="Search Plazi for locations around (long 130.49973/lat 30.45639)">Isso</a>, 30°27′23″N, 130°29′59″E, 1–5 m depth, KAUM Fish Team, 13 August 2008; KAUM –I. 20192, male, 28.8 mm SL, KAUM –I. 20209, male, 34.8 mm SL, tide pool at west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.47194&amp;materialsCitation.latitude=30.232779" title="Search Plazi for locations around (long 130.47194/lat 30.232779)">Yudomari Port</a>, Yudomari, 30°13′58″N, 130°28′19″E, 0–3 m depth, KAUM Fish Team, 29 October 2008; KAUM –I. 20348, male, 28.0 mm SL, KAUM –I. 20350, male, 30.6 mm SL, KAUM –I. 20351, male, 39.7 mm SL, southeast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.50055&amp;materialsCitation.latitude=30.4575" title="Search Plazi for locations around (long 130.50055/lat 30.4575)">Cape Yahazu</a>, Isso, 30°27′27″N, 130°30′02″E, 1–3 m depth, KAUM Fish Team, 31 October 2008; KAUM –I. 21857, male, 24.8 mm SL, KAUM –I. 21858, female, 23.3 mm SL, west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.41306&amp;materialsCitation.latitude=30.265833" title="Search Plazi for locations around (long 130.41306/lat 30.265833)">Kamazenohana</a>, Kurio, 30°15′57″N, 130°24′47″E, 0–4 m depth, KAUM Fish Team, 30 July 2009; KAUM –I. 38696, male, 41.9 mm SL, south of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.47194&amp;materialsCitation.latitude=30.231945" title="Search Plazi for locations around (long 130.47194/lat 30.231945)">Yudomari Port</a>, Yudomari, 30°13′55″N, 130°28′19″E, 1–5 m depth, KAUM Fish Team, 11 June 2011. NAKANO-SHIMA ISLAND: YCM-P 26990, tidepool at Funakura, M. Hayashi, 3 May 1992. TAKARA- JIMA ISLAND: KAUM – I. 2037, male, 39.4 mm SL, 25 March 1965. AMAMI-OSHIMA ISLAND: KAUM –I. 4126, male, 35.1 mm SL, 4 May 1965; KAUM –I. 17495, male, 44.1 mm SL, KAUM –I. 17496, female, 44.1 mm SL, tidepool at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.67528&amp;materialsCitation.latitude=28.409445" title="Search Plazi for locations around (long 129.67528/lat 28.409445)">Tsuchihama</a>, Kasari, 28°24′34″N, 129°40′31″E, 0.5–1 m depth, M. Meguro and T. Yoshida, 27 March. 2009. OKINOERABU-JIMA ISLAND: HMNH-P 7910, male, 41.1 mm SL, female, 37.8 mm SL, Okidomari Beach, Tina, 2 May 2004; KAUM – I. 2002, male, 49.1 mm SL, July 1962. YORON-JIMA ISLAND: KAUM –I. 40069, male, 20.1 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.061943" title="Search Plazi for locations around (long 128.41722/lat 27.061943)">Udonosu Beach</a>, 27°03′43″N, 128°25′02″E, 0.3–1 m depth, M. Matsunuma, 14 August 2011; KAUM –I. 40103, male, 20.9 mm SL, KAUM –I. 40397, male, 41.5 mm SL, KAUM –I. 44370, male, 32.3 mm SL, KAUM –I. 44371, male, 26.1 mm SL, KAUM –I. 44372, female, 24.9 mm SL, KAUM –I. 44373, male, 24.6 mm SL, KAUM –I. 44374, female, 21.8 mm SL, off southwest coast, 27°01′48″N, 128°24′37″E, 2–10 m depth, M. Meguro and E. Katayama, 17 August 2011; KAUM –I. 45757, male, 33.1 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.031666" title="Search Plazi for locations around (long 128.40805/lat 27.031666)">Tomori Fishing Port</a>, 27°01′54″N, 128°24′29″E, 5–10 m depth, M. Yamashita and T. Yoshida, 15 April 2012; KAUM –I. 45832, male, 34.8 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 5 m depth, H. Motomura, 16 April 2012; KAUM –I. 45861, male, 36.8 mm SL, KAUM –I. 45895, male, 35.1 mm SL, KAUM –I. 45896, male, 31.3 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 3–10 m depth, KAUM Fish Team, 17 April 2012; KAUM –I. 47823, male, 42.2 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 2–8 m depth, KAUM Fish Team, 13 August 2012; KAUM –I. 47878, male, 39.8 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 2 m depth, KAUM Fish Team, 13 August 2012; KAUM –I. 48144, male, 28.5 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 1 m depth, KAUM Fish Team, 19 August 2012; KAUM –I. 50844, male, 22.8 mm SL, KAUM –I. 50848, male, 26.7 mm SL, KAUM –I. 50850, female, 23.0 mm SL, KAUM –I. 50851, female, 23.0 mm SL, KAUM –I. 50852, male, 20.9 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 2–3 m depth, KAUM Fish Team, 20 August 2012; KAUM –I. 50863, male, 38.0 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 15–17 m depth, KAUM Fish Team, 21 August 2012; KAUM –I. 51239, female, 30.1 mm SL, KAUM –I. 51243, female, 31.3 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 3–5 m depth, KAUM Fish Team, 23 October 2012; KAUM –I. 51241, male, 37.6 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 2 m depth, KAUM Fish Team, 23 October 2012; KAUM –I. 51339, male, 27.4 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.42973&amp;materialsCitation.latitude=27.022223" title="Search Plazi for locations around (long 128.42973/lat 27.022223)">Ritcho</a>, 27°01′20″N, 128°25′47″E, 10 m depth, KAUM Fish Team, 24 October 2012; KAUM –I. 51376, female, 29.7 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 8 m depth, KAUM Fish Team, 25 October 2012; KAUM –I. 51399, male, 34.1 mm SL, KAUM –I. 51400, male, 25.5 mm SL, KAUM –I. 51401, male, 24.1 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 8 m depth, KAUM Fish Team, 26 October 2012; KAUM –I. 51455, female, 31.4 mm SL, KAUM –I. 51456, female, 28.6 mm SL, KAUM –I. 51457, female, 27.8 mm SL, KAUM –I. 51458, female, 23.5 mm SL, KAUM –I. 51558, male, 32.8 mm SL, KAUM –I. 51559, male, 36.7 mm SL, KAUM –I. 51560, male, 34.5 mm SL, KAUM –I. 51586, male, 39.0 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.40805&amp;materialsCitation.latitude=27.020277" title="Search Plazi for locations around (long 128.40805/lat 27.020277)">Maehama Beach</a>, 27°01′13″N, 128°24′29″E, 2–8 m depth, KAUM Fish Team, 29 October 2012; KAUM –I. 51470, female, 30.5 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Chabana</a>, 27°03′40″N, 128°25′02″E, 8 m depth, KAUM Fish Team, 27 October 2012. OKINAWA-JIMA ISLAND: HMNH-P 5515, male, 32.6 mm SL, Giizabanda, Asato, Yaese, 10 August 2002; HMNH-P 7425, female, 42.7 mm SL, HMNH-P 7426, male, 30.0 mm SL, Komesu Beach, Itoman, 27 December 2003; HMNH-P 8625, 2 males, 36.4–45.7 mm SL, 3 females, 32.0– 40.5 mm SL, Odo Beach, Itoman, 12 December 2004; HMNH-P 9023, 3 females, 34.3–38.2 mm SL, Akasaki Beach, Kunigami, 31 December 2005. KUME-JIMA ISLAND: HMNH-P 5739, male, 22.1 mm SL, beach near Kume Airport, 13 August 2002.</p> <p>Diagnosis. A species of Helcogramma with the following combination of characters: 13–16 (rarely 13; usually 15) second dorsal-fin spines; 22–35 pored lateral-line scales; 5–10 symphyseal mandibular pores; anterior nostril tentacle unbranched; orbital tentacle simple; nape scaled; lower half of head (area bounded by snout, ventral margin of orbit, and preopercular margin) black; bright blue stripe extending from anterior part of upper lip, through ventral margin of orbit, to preopercular margin; all fins and body black (all color descriptions for mature males only).</p> <p>Distribution. Helcogramma inclinata is distributed in the western Pacific Ocean, where it ranges from southern Japan to the Philippines (Fricke 1997; Williams and Howe 2003). In Japan, this species has been collected from the Satsuma Peninsula in southern Kyushu and from Takeshima, Iou-jima, Yaku-shima, Nakano-shima, Takara-jima, Yoron-jima, Okinawa-jima, and Kume-jima islands in the Ryukyu Islands. In addition, this species has been recorded elsewhere in the central and southern Ryukyu Islands: i.e., Ie-jima (Senou et al. 2006), Aguni-jima (Fowler 1946), Ishigaki-jima (Fricke 1997), and Iriomote-jima (Aizawa 1997) islands. Helcogramma inclinata usually inhabits rocky reefs at depths of 3– 8 m.</p> <p>Remarks. Helcogramma inclinata, belonging to the H. fuscopinna species group (sensu Fricke 1997), is distinguished from all other members of the group by its usually having 15 second dorsal-fin rays, 5–10 symphyseal mandibular pores, and a scaled nape, and by its mature male coloration (e.g., lower head, body, and all fins black).</p> <p>Motomura et al. (2006) reported H. inclinata from the southern end of the Satsuma Peninsula, on the basis of an underwater photograph only, as the northernmost record for the species. Two of our specimens (KAUM–I. 4246, 50.0 mm SL; KAUM–I. 6522, 50.9 mm SL) collected from Kasasa, Minamisatsuma, on the west coast of Satsuma Peninsula, represent a northward distributional range extension for H. inclinata.</p> </div>	https://treatment.plazi.org/id/0388F8254C36CA74FE90FF68FC01FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Tashiro, Satokuni;Motomura, Hiroyuki	Tashiro, Satokuni, Motomura, Hiroyuki (2014): The Validity of Helcogramma ishigakiensis (Aoyagi, 1954) and a Synopsis of Species of Helcogramma from the Ryukyu Islands, Southern Japan (Perciformes: Tripterygiidae). Species Diversity 19: 97-110, DOI: 10.12782/sd.19.2.097
0388F8254C35CA75FC0BFA34FDC4F8D8.text	0388F8254C35CA75FC0BFA34FDC4F8D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helcogramma rhinoceros Hansen 1986	<div><p>Helcogramma rhinoceros Hansen, 1986</p> <p>[Standard Japanese name: Tengu-hebigimpo] (Figs 5G–H, 6E; Table 2)</p> <p>Helcogramma rhinoceros Hansen, 1986: 332, fig. 7 (left), 15 (type locality: Putic Island, Philippines); Aizawa 1997: 560, unnumbered fig. (Iriomote-jima island, Ryukyu Islands); Suzuki et al. 1998: 2, figs 1–7 (Nakano-ugan-jima and Iriomote-jima islands, Ryukyu Islands); Motomura et al. 2010: 189, fig. 436 (Yaku-shima island, Ryukyu Islands); Kato 2011: 221, unnumbered fig. (Yaku-shima island, Ryukyu Islands); Tashiro 2013: 298, unnumbered figs (Take-shima island, Ryukyu Islands); Hayashi 2013; 1283, 2098 (Yaku-shima, Amami-oshima, and Okinawajima islands, Ryukyu Islands).</p> <p>Non-type material examined. 12 specimens, 21.1– 30.3 mm SL. TAKE-SHIMA ISLAND: KAUM –I. 32332, male, 26.9 mm SL, off south coast of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.40916&amp;materialsCitation.latitude=30.808887" title="Search Plazi for locations around (long 130.40916/lat 30.808887)">Ombozaki</a>, 30°48′32″N, 130°24′33″E, 5–10 m depth, KAUM Fish Team, 19 September 2010. NAKANO-SHIMA ISLAND: KAUM –I. 63252, female, 30.3 mm SL, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.84888&amp;materialsCitation.latitude=29.837223" title="Search Plazi for locations around (long 129.84888/lat 29.837223)">Nakano-shima Port</a>, 29°50′14″N, 129°50′56″E, 17–19 m depth, S. Tashiro, 30 August 2014; KAUM –I. 63467, female, 29.4 mm SL, south side of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=129.85085&amp;materialsCitation.latitude=29.836945" title="Search Plazi for locations around (long 129.85085/lat 29.836945)">Nakano-shima Port</a>, 29°50′13″N, 129°51′03″E, 5 m depth, S. Tashiro, 2 September 2014. AMAMI-OSHIMA ISLAND: KAUM –I. 57683, female, 21.2 mm SL, KAUM –I. 57729, female, 21.1 mm SL, 28°07′57″N, 129°21′07″E, 3–15 m depth, Sokaru, Setouchi, S. Tashiro et al., 13 December 2013. YORON-JIMA ISLAND: KAUM –I. 51381, 27.2 mm SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Yoron Port</a>, 27°3′40″N, 128°25′02″E, 8 m depth, S. Tashiro, 26 October 2012; KAUM –I. 51428, male, 26.8 mm SL, KAUM –I. 51465, male, 26.7 mm SL, KAUM –I. 51466, male, 23.8 SL, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.41722&amp;materialsCitation.latitude=27.06111" title="Search Plazi for locations around (long 128.41722/lat 27.06111)">Yoron Port</a>, 27°3′40″N, 128°25′02″E, 8 m depth, S. Tashiro, 27 October 2012. ISHIGAKI-JIMA ISLAND: NSMT-P 55449, female, 25.2 mm SL, 23°26′60″N, 124°04′80″E, Uganzaki, K. Matsuura et al., 24 February 1998. NAKANOUGAN-JIMA ISLAND: KPM-NI 4341, male, 26.1 mm SL, KPM-NI 4342, female, 23.6 mm SL, K. Yano, 28 July 1997.</p> <p>Diagnosis. A species of Helcogramma with the following combination of characters: 14–15 (modally 15) second dorsal-fin spines; 19–24 pored lateral-line scales; 10–11 third dorsal-fin rays; 4+1+4 mandibular pores; nape without scales; anterior nostril tentacle unbranched; orbital tentacle present; males with dermal prolongation on tip of snout; lower half of head (area below level of upper lip) black; narrow blue stripe running along upper lip, through ventral margin of orbit, to preopercular margin; pelvic fin, pectoral-fin base, and lower 7 pectoral-fin rays black (all color descriptions for mature males only).</p> <p>Distribution. Helcogramma rhinoceros is distributed in the eastern Indian Ocean and western Pacific Ocean, where it ranges from the Andaman Sea to southern Japan, Vietnam, Indonesia, New Caledonia, and Vanuatu (Hansen 1986; Fricke 1997; Suzuki et al. 1998). In the Ryukyu Islands, this species was collected from Take-shima, Nakanoshima, Amami-oshima, Yoron-jima, Ishigaki-jima, and Nakano-ugan-jima islands during this study and had been previously recorded from Yaku-shima (Motomura et al. 2010) and Iriomote-jima (Aizawa 1997) islands. It was also confirmed from Kume-jima island (KPM-NR 36216, 64513, 64515) on the basis of underwater photographs. Helcogramma rhinoceros has been usually observed near the upper or lateral surfaces of rocks, at depths of 6– 8 m.</p> <p>Remarks. Mature males of Helcogramma rhinoceros can be distinguished easily from all other species of Helcogramma by their having a dermal prolongation on the tip of the snout. Females of H. rhinoceros are similar to females of H. fuscipectoris in coloration, but H. rhinoceros differs from the latter in having a simple anterior nostril tentacle (a branched tentacle in H. fuscipectoris).</p> </div>	https://treatment.plazi.org/id/0388F8254C35CA75FC0BFA34FDC4F8D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Tashiro, Satokuni;Motomura, Hiroyuki	Tashiro, Satokuni, Motomura, Hiroyuki (2014): The Validity of Helcogramma ishigakiensis (Aoyagi, 1954) and a Synopsis of Species of Helcogramma from the Ryukyu Islands, Southern Japan (Perciformes: Tripterygiidae). Species Diversity 19: 97-110, DOI: 10.12782/sd.19.2.097
0388F8254C34CA72FE8FF82CFE66FA91.text	0388F8254C34CA72FE8FF82CFE66FA91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helcogramma striata Hansen 1986	<div><p>Helcogramma striata Hansen, 1986</p> <p>[Standard Japanese name: Tatejima-hebigimpo] (Figs 5I–J, 6E; Table 2)</p> <p>Helcogramma striata Hansen, 1986: 350, fig. 18 (type locality: Miyake-jima island, Izu Islands, Japan; paratypes from Tokomasari Reef, Okinawa-jima island, Ryukyu Islands); Aizawa 1997: 559, unnumbered fig. (Iriomote-jima island, Ryukyu Islands); Hayashi 2000: 945 (south of Amami-oshima island, Ryukyu Islands); Sakai et al. 2005: 10 (Nakano-shima island, Ryukyu Islands); Watai et al. 2009: 129 (Tokashiki-jima island, Ryukyu Islands); Motomura et al. 2010: 189, fig. 437 (Yaku-shima island, Ryukyu Islands); Tashiro 2013: 297, unnumbered figs (Iou-jima and Take-shima islands, Ryukyu Islands).</p> <p>Helcogramma striatum: Fricke 1997: 481, fig. 98 (Ryukyu Islands); Senou et al. 2006: 84 (Ie-jima island, Ryukyu Islands); Senou et al. 2007: 64 (Miyako-jima island, Ryukyu Islands); Hayashi 2013: 1281, unnumbered fig. (Yakushima island and south of Amami-oshima island in the Ryukyu Islands).</p> <p>Material examined. 29 specimens, 11.4–38.5 mm SL, from the Ryukyu Islands. TAKE-SHIMA ISLAND: KAUM –I. 29720, male, 18.9 mm SL, KAUM –I. 29721, 11.4 mm SL, off south coast of Ombozaki, 30°48′32″N, 130°24′33″E, 5–40 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.40916&amp;materialsCitation.latitude=30.808887" title="Search Plazi for locations around (long 130.40916/lat 30.808887)">Fish Team</a>, 29 May 2010; KAUM –I. 30110, male, 37.1 mm SL, KAUM –I. 30111, male, 31.2 mm SL, KAUM –I. 30112, female, 20.3 mm SL, KAUM –I. 30113, male, 14.8 mm SL, off south coast of Ombozaki, 30°48′32″N, 130°24′33″E, 5–40 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.40916&amp;materialsCitation.latitude=30.808887" title="Search Plazi for locations around (long 130.40916/lat 30.808887)">Fish Team</a>, 27 May 2010; KAUM –I. 37748, male, 38.5 mm SL, KAUM –I. 37749, male, 36.7 mm SL, KAUM –I. 37750, male, 35.8 mm SL, KAUM –I. 37751, 2 males, 30.7– 38.5 mm SL, 2 females, 28.8–29.3 mm SL, Take-shima Port, 30°48′59″N, 130°25′06″E, 3–5 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.41833&amp;materialsCitation.latitude=30.816387" title="Search Plazi for locations around (long 130.41833/lat 30.816387)">Fish Team</a>, 11 May 2011. IOU-JIMA ISLAND: KAUM –I. 29515, female, 33.5 mm SL, KAUM –I. 29516, male, 20.8 mm SL, off south coast, 30°46′32″N, 130°16′43″E, 10–60 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.27861&amp;materialsCitation.latitude=30.775555" title="Search Plazi for locations around (long 130.27861/lat 30.775555)">Fish Team</a>, 25 May 2010; KAUM –I. 30049, male, 13.6 mm SL, off west coast, 30°47′04″N, 130°15′42″E, 5–20 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.26167&amp;materialsCitation.latitude=30.784445" title="Search Plazi for locations around (long 130.26167/lat 30.784445)">Fish Team</a>, 28 May 2010; KAUM –I. 31713, male, 27.6 mm SL, KAUM –I. 31714, male, 25.7 mm SL, off west coast, 30°47′04″N, 130°15′42″E, 10–15 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.26167&amp;materialsCitation.latitude=30.784445" title="Search Plazi for locations around (long 130.26167/lat 30.784445)">Fish Team</a>, 14 August 2010. YAKU-SHIMA ISLAND: KAUM –I. 20174, male, 20.4 mm SL, tide pool at west side Yudomari Port, 30°13′58″N, 130°28′19″E, 0–3 m depth, KAUM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.47194&amp;materialsCitation.latitude=30.232779" title="Search Plazi for locations around (long 130.47194/lat 30.232779)">Fish Team</a>, 29 October 2008. NAKANO- SHIMA ISLAND: KAUM –I. 63256, male, 24.4 mm SL, KAUM –I. 63289, female, 21.7 mm SL, KAUM –I. 63290, male, 20.1 mm SL, KAUM –I. 63291, male, 13.8 mm SL, Nakano-shima Port, 29°50′14″N, 129°50′56″E, 17–19 m depth, S. Tashiro et al., 30 August 2014; KAUM –I. 63265, male, 23.6 mm SL, south side of Nakano-shima Port, 29°50′13″N, 129°51′03″E, 5 m depth, S. Tashiro et al., 2 September 2014. AMAMI-OSHIMA ISLAND: YCM –P 26162, male, 25.5 mm SL, Sakinome, Setouchi, Amami Fish Team, 27 August 1991. IE-JIMA ISLAND: KPM-NI 4110, male, 34.7 mm SL, east of Nakanose, T. Nomura, 28 July 1997; KPM-NI 27124, male, 26.2 mm SL, male, 36.9 mm SL, Nakanose, T. Nomura, 24 October 2010. TOKASHIKI-JIMA ISLAND: KPM-NI 22043, female, 35.4 mm SL, off Tokashiku Beach, H. Gunji, 10 March. 2004; KPM-NI 22057,</p> <p>female, 33.6 mm SL, off Tokashiku Beach, A. Murase, 10 March 2004.</p> <p>Diagnosis. A species of Helcogramma with the following combination of characters: 13–15 (modally 14) second dorsal-fin spines; 3–4+2+3–4 mandibular pores; anterior nostril tentacle unbranched; orbital tentacle absent; nape scaled; three longitudinal stripes running from snout to caudal peduncle; row of 6 or 7 spots between middle and ventral stripes.</p> <p>Distribution. Helcogramma striata is distributed in the Indo-West Pacific, where it ranges from Japan to Australia and from the Andaman Sea to Fiji and the Line Islands (Hansen 1986; Fricke 1997; Holleman 2007). In Japan, this species occurs from Shizuoka Prefecture to the Ryukyu Islands, where it was collected from Take-shima, Iou-jima, Yaku-shima, Nakano-shima, Amami-oshima, Ie-jima, and Tokashiki-jima islands during the present study and had been recorded previously from Nakano-shima (Sakai et al. 2005), Okinawa-jima (Hansen 1986), Miyako-jima (Senou et al. 2007), and Iriomote-jima (Aizawa 1997) islands. It was also confirmed from Yonaguni-jima island (KPM-NR 12127) based on underwater photographs. Helcogramma striata usually inhabits the upper surface of rocks or corals, at depths of 3– 20 m.</p> <p>Remarks. Helcogramma striata is similar to H. maldivensis Fricke and Randall, 1992 in coloration and in lacking both the orbital tentacle and scales on the nape anterior to the dorsal fin. Helcogramma striata can be easily distinguished from H. maldivensis by its having usually 14 second dorsal-fin spines (vs usually 13 in H. maldivensis) and three longitudinal stripes in both sexes (vs three dotted lines). Mature males of H. striata have a dark-reddish body whereas females have a pale red body with a pale yellow area below the lowermost stripe.</p> </div>	https://treatment.plazi.org/id/0388F8254C34CA72FE8FF82CFE66FA91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Tashiro, Satokuni;Motomura, Hiroyuki	Tashiro, Satokuni, Motomura, Hiroyuki (2014): The Validity of Helcogramma ishigakiensis (Aoyagi, 1954) and a Synopsis of Species of Helcogramma from the Ryukyu Islands, Southern Japan (Perciformes: Tripterygiidae). Species Diversity 19: 97-110, DOI: 10.12782/sd.19.2.097
0388F8254C33CA72FF6DFAF6FA4AFE01.text	0388F8254C33CA72FF6DFAF6FA4AFE01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helcogramma McCulloch and Waite 1918	<div><p>Key to the species of Helcogramma in the Ryukyu Islands</p> <p>1a Lateral surface of body with three white (dusky in mature males) longitudinal stripes; orbital tentacles absent............................................................................... H. striata</p> <p>1b Lateral surface of body without longitudinal stripes; orbital tentacles present........................................................ 2</p> <p>2a Symphyseal mandibular pores 5 or more....................... 3</p> <p>2b Symphyseal mandibular pore 1........................................ 4</p> <p>3a Nape without scales; second dorsal-fin spines usually 13........................................................................... H. aquila</p> <p>3b Nape with scales; second dorsal-fin spines usually 15............................................................................... H. inclinata</p> <p>4a Pored lateral-line scales 28–39; no longitudinal row of reddish or dark blotches on middle of lateral surface of body; largest specimen of more than 50 mm SL.................................................................................. H. ishigakiensis</p> <p>4b Pored lateral-line scales 17–25; longitudinal row of reddish or dark blotches in middle of lateral surface of body (indistinct in mature males); largest specimen of less than 35 mm SL............................................................ 5</p> <p>5a Males with dermal prolongation on tip of snout; second dorsal-fin spines usually 15; anterior nostril tentacle unbranched.......................................... H. rhinoceros</p> <p>5b No dermal prolongation on tip of snout; second dorsalfin spins usually 14; anterior nostril tentacle branched.................................................................... H. fuscipectoris</p></div> 	https://treatment.plazi.org/id/0388F8254C33CA72FF6DFAF6FA4AFE01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Tashiro, Satokuni;Motomura, Hiroyuki	Tashiro, Satokuni, Motomura, Hiroyuki (2014): The Validity of Helcogramma ishigakiensis (Aoyagi, 1954) and a Synopsis of Species of Helcogramma from the Ryukyu Islands, Southern Japan (Perciformes: Tripterygiidae). Species Diversity 19: 97-110, DOI: 10.12782/sd.19.2.097
