taxonID	type	description	language	source
0388878031080547FF6198C2FA51789B.taxon	type_taxon	Type-species: Chelonus aculeatus Ashmead, 1890.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080547FF6198C2FA51789B.taxon	diagnosis	Diagnosis. Head transverse (Figs 2 H, 4 G); antenna slender, filiform (Figs 2 I, 4 I); face and vertex behind ocelli smooth to densely granulate, clypeus flat and densely punctate (Figs 2 G, 4 F); eyes setose (Figs 2 J, 4 H); mesoscutum evenly convex, transverse suture distinct; scutellar suture comparatively deep; propodeum coarsely reticulate-rugose with transversal carina (Figs 2 C, 4 C); metasomal carapace with the straight elongate apex (Figs 2 D, 4 D).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080547FF6198C2FA51789B.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080547FF6198C2FA51789B.taxon	distribution	Distribution. Oriental.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080547FF6198C2FA51789B.taxon	discussion	Key to species of subgenus Arichelonus	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	description	Figs 1 – 2	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	materials_examined	Material examined. Holotype: female, China, Zhejiang Prov., Mt. Jiulong, IV. 1995, Hanlin Chen, No. 95507 (ZJUH).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	diagnosis	Diagnosis. This new species is similar to C. (A.) sternalis Tobias, 1964, but can be separated from the latter by the following characters: length of metasoma 2.3 × its height (4.0 × in C. (A.) sternalis Tobias, 1964); scutellum coarsely reticulate rugose and punctate medially (smooth medially); antennomeres of female 18, length of antenna distinctly shorter than body length (antenna very slender and filiform, antennomeres 38, as long as body).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	description	Description. Holotype, female, length of body 3.4 mm, of fore wing 2.2 mm. Head. Antennomeres 18, basal half of left antenna distinctly broadened; third antennomere equal in length to fourth; third, fourth and terminal antennomeres 3.2 ×, 3.2 × and 2.3 × longer than wide (Fig. 2 I); head transverse, width of head 2.6 × its dorsal length roundly contracted behind eyes; OOL: OD: POL = 9: 4: 9; longitudinal diameter of eye 1.1 × height of temples; vertex behind ocelli and temple evenly convex, transversely striate; frons distinctly concave, smooth (Fig. 2 H); face coarsely striate-rugose; clypeus shiny, finely punctate with dense setae (Fig. 2 G). Mesosoma. Length of mesosoma 1.4 × its height; pronotum finely reticulate-rugose; mesoscutum evenly convex, reticulate-rugose; scutellum hardly convex, more strongly and coarsely rugose than mesoscutum, punctate medially; scutellar suture comparatively deep, narrow, with carinae; apical transverse carina of propodeum short, near posterior margin of propodeum (Fig. 2 C); mesopleuron evenly convex, coarsely reticulate-rugose (Fig. 2 B). Wings. Length of fore wing 2.6 × its width; length of marginal cell 1.25 × pterostigma; vein r straight, r: 3 - SR: SR 1 = 7: 7: 45; 1 - CU 1: 2 - CU 1 = 6: 17 (Fig. 2 A). Legs. Hind coxa mainly smooth and shallowly punctate; hind coxa, femur and tibia all distinctly robust; length of hind femur, tibia and basitarsus 3.0 ×, 3.75 × and 3.8 × their maximum width, respectively (Fig. 2 E). Metasoma. Carapace elongate-oval, but distinctly contracted at the one-third of apex, two thirds of it longitudinally striate, and reticulate by weaker and transverse rugae basally; length of carapace 2.3 × its maximum width in dorsal view (Fig. 2 D); ventrally metasoma completely open to apex of carapace; length of ovipositor projecting part 0.5 × of carapace length, and the ovipositor sheath 0.13 × as long as fore wing, 0.38 × as long as hind tibia (Fig. 2 E). Colour. Black; antennomeres dark brown; palp blackish-brown; pterostigma and veins yellowish-brown; pterostigma and veins light brown; wing membrane hyaline; legs almost entirely yellowish-brown, basally hind tibia light yellow and apically black; ovipositor sheath brown. Male. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031080544FF619C05FB7E7ECB.taxon	etymology	Etymology. Named after its special abdomen: allokotos is Greek for unusual and strange.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	description	Figs 3 – 4	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	materials_examined	Material examined. Holotype: female, China, Zhejiang Prov., Deqing, 27. V. 1995, Junhua He, No. 965092 (ZJUH); Paratypes, 2 females, Deqing, Zhejiang province, 27. V. 1995, Junhua He, Nos. 954349, 954350.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	diagnosis	Diagnosis. This new species is similar to C. (A.) olgae Kokujev, 1895, but can be separated from the latter by the following characters: ovipositor obviously extended beyond apex of metasomal carapace and covered with a semi-round and semi-hyaline lamella (the ovipositor not or weakly projecting beyond apex of metasoma in C. (A.) olgae Kokujev, 1895); metasoma entirely brown (metasoma with pair of yellow basal spots); about 22 antennomeres of female, distinctly shorter than body length (antennomeres of female about 45, equal to body length).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	description	Description. Holotype, female, length of body 4.6 mm, of fore wing 3.3 mm. Head. Antennomeres 22 (Fig. 4 I); third antennomere 1.1 × longer than fourth; third, fourth and terminal antennomeres 4.0 ×, 3.5 × and 1.75 × longer than wide; head transverse, in dorsal view roundly contracted behind eyes; OOL: OD: POL = 19: 15: 9; longitudinal diameter of eye 2.1 × height of temples; vertex behind ocelli and temple transversely striate; frons distinctly concave (Fig. 4 G); face evenly convex, coarsely transversely striate; clypeus hardly convex and densely punctate, slightly shiny; length of malar space 1.5 × basal width of mandible (Fig. 4 F); face and eyes densely setose in lateral view (Fig. 4 H). Mesosoma. Length of mesosoma 1.3 × its height; mesonotum evenly convex, irregular areolate-rugose and more strongly rugose postero-medially; scutellum hardly convex, punctate medially, rugose laterally; scutellar suture comparatively deep; propodeum coarsely reticulaterugose (Fig. 4 C); mesopleuron evenly convex, coarsely reticulate-rugose (Fig. 4 B). Wings. Length of fore wing 2.9 × its width; length of marginal cell 1.3 × pterostigma; r: 3 - SR: SRl = 9: 9: 43; SRl straight; parastigma distinctly swollen; 1 - CU 1: 2 - CU 1 = 10: 27 (Fig. 4 A). Legs. Hind coxa completely smooth and punctate; hind femur pretty robust; length of hind femur, tibia and basitarsus 3.1 ×, 5.2 × and 4.75 × their maximum width, respectively. Metasoma. Carapace elongate-oval, a narrow lamella dorsally of apical carapace in dorsal view, posterior part of carapace with transversely curved deepened furrow; length of carapace 2.6 × its maximum width in dorsal view (Fig. 4 D); ventral cavity completely open to apex; ovipositor extended beyond apex of metasomal carapace and covered with a semi-round and semi-hyaline short lamella, ovipositor slender and the ovipositor sheath 0.11 × as long as fore wing, 0.32 × as long as hind tibia (Fig. 4 E). Colour. Black; antennomeres blackish-brown; pterostigma and veins blackish-brown; fore wing more or less darkened; metasoma entirely brown; legs almost entirely yellowish-brown; ovipositor sheath brown. Variation. Body length 3.7 – 4.6 mm, and length of ovipositor 0.2 – 0.4 × of carapace length. Male. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310D0540FF619ADAFE9A7F53.taxon	etymology	Etymology. From kenos (Greek for empty) and etron (Greek for belly) because its ventral cavity completely open to the apex.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F0540FF6198C3FA517A49.taxon	type_taxon	Type-species: Megachelonus bidentatus Baker, 1926.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F0540FF6198C3FA517A49.taxon	diagnosis	Diagnosis. Head transverse (Figs 6 G, 8 B, 10 B, 12 E); antennomeres of apical half of antenna distinctly flattened and broadened; face striate to reticulate rugose, clypeus semi-elliptical (Figs 6 F, 8 A, 10 A, 12 D); eyes long elliptical or ovoid in lateral view (Figs 6 I, 8 G, 10 G, 12 F); mesosoma coarsely reticulate-rugose, transverse suture distinct; scutellum distinctly protruding above level of mesoscutum, with same sculpture as mesosoma; propodeum with a transverse carina and distinct tooth-like tubercles (Figs 6 C, 6 J, 8 F, 10 F, 12 C); metasomal carapace with two ventroapical tooth-like protuberances (Figs 6 E, 8 C, 10 C, 12 H).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F0540FF6198C3FA517A49.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F0540FF6198C3FA517A49.taxon	distribution	Distribution. Oriental.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F0540FF6198C3FA517A49.taxon	discussion	Key to species of subgenus Megachelonus	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	description	Figs 5 – 6	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	materials_examined	Material examined. Holotype: female, China, Yunnan Prov., Mt. Gaoligong, 16 - 17. VII. 2006, Jie Zeng, No. 200611349 (ZJUH). Paratypes: 1 female, China, Guangdong Prov., Mt. Guanyin, 15. IX. 2007, Jie Zeng, No. 200807931.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	diagnosis	Diagnosis. This new species is similar to C. (M.) macros Zhang, Chen et He, but can be separated from the latter by the following characters: carapace without lamella (carapace with long and wide lamella in C. macros); apex of carapace narrowed in dorsal view (apex of carapace rounded); apex of propodeum strongly concaved with two pairs of blunt tubercles, lateral tubercles distinctly bigger and longer than medial tubercles (apex of propodeum weakly concaved with four comparably average size blunt tubercles).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	description	Description. Holotype, female, length of body 4.4 mm, of fore wing 3.0 mm. Head. Antennomeres 16 in female, average length of 3 - 7 th antennomeres 3.3 × their width, 8 - 11 th antennomeres 1.5 × longer than wide; head transverse, width of head 1.9 × its dorsal length, roundly contracted behind eyes; temple rounded posteriorly; OOL: OD: POL = 20: 6: 19, vertex evenly convex and with transverse rugae behind ocelli, temple finely and longitudinally striate- rugose, frons concave, striate-rugose around antennal sockets; longitudinal diameter of eye 2.1 × height of temples (Fig. 6 G); face striate-rugose, clypeus densely setose, shiny (Fig. 6 F). Mesosoma. Length of mesosoma 1.4 × its height; pronotum finely reticulate-rugose; mesoscutum evenly convex, reticulate-rugose, more strongly and coarsely rugose postero-medially than anteriorly; scutellum punctate medially, rugose laterally (Fig. 6 B); scutellar suture comparatively deep with 4 carinae (Fig. 6 C); propodeum coarsely reticulate-rugose, strongly concave with two pairs of blunt tubercles apically, lateral tubercles distinctly bigger and longer than medial ones (Fig. 6 J). Wings. Length of fore wing 3.9 × its width; length of marginal cell 2.9 × pterostigma; vein 2 - Rl just present; r: 3 - SR: SRl = 1: 1: 7; SRl complete and straight; parastigma not swollen; 1 - CUl: 2 - CUl = 1: 4; vein CUlb present (Fig. 6 A). Legs. Hind соха completely smooth and setose, hind femur robust, length of hind femur, tibia and basitarsus 3.7 ×, 4.3 × and 6.2 × their maximum width, respectively; length of hind tibial spurs 0.5 × hind basitarsus. Metasoma. Carapace elongate-oviform, length of metasoma 2.2 × its maximum width in dorsal view; metasomal carapace much more strongly narrowed towards apex than towards base, with two ventro-apical tooth-like protuberances, coarsely longitudinally rugulose, connected by transverse elements (Fig. 6 E), carapace without lamella around ventral opening (Fig. 6 D); ovipositor long, slender and curved, the ovipositor sheath 0.19 × as long as fore wing, 0.59 × as long as hind tibia (Fig. 6 H). Colour. Black; 3 basal antennomeres yellowish-brown, remainder of antennomeres dark brown; palp light yellow; mandible brown but basally blaсk, pterostigma and most of veins blackish brown, veins of basal half of wing light yellow; parastigma light yellow; margination of metasomal carapace yellowish; legs yellowish-brown but hind tibia apically and base of hind coxa dark brown. Male. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
03888780310F054FFF619D22FF3C7E7B.taxon	etymology	Etymology. From ir (Latin for without) and lamina (Latin for thin plate), because metasoma lacks the lateral lamella.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	description	Figs 9 – 10	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	materials_examined	Material examined. Holotype: female, China, Yunnan Prov., Tongbiguan, 23. VI. 2005, Jinxian Liu, No. 200609605 (ZJUH).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	diagnosis	Diagnosis. This species differs from the other species in the subgenus Megachlonus by having a pair of long and large teeth at apex of metasomal carapace (Figs 10 G, 10 H).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	description	Description. Holotype, female, length of body 5.3 mm, of fore wing 3.9 mm. Head. Antennomeres 28 (remaining, broken in holotype), basal half of antenna distinctly broadened, then rapidly tapering to apex; third antennomere 1.3 × longer than fourth; third, fourth and terminal antennomeres 4.7 ×, 3.7 × and 3.2 × longer than wide, respectively; 9 - 20 th antennomeres flattened, distinctly shorter than wide; head transverse, in dorsal view roundly contracted behind eyes; OOL: OD: POL = 15: 6: 10; longitudinal diameter of eye 2.0 × height of temples; vertex coarsely striate-rugose; frons distinctly concave, reticulate-rugose but smooth near antennal sockets; temple irregularly reticulate-rugose (Fig. 10 E); face coarsely reticulate-rugose with longitudinal elements, clypeus flat and densely punctate, smooth and shining, length of malar space 1.6 × basal width of mandible (Fig. 10 D); face densely setose in lateral view (Fig. 10 F). Mesosoma. Length of mesosoma 1.4 × its height; mesoscutum areolate-rugose, but smooth and shiny medially; scutellum punctate medially, rugose laterally; scutellar suture comparatively deep; mesopleuron evenly convex, coarsely reticulate-rugose (Fig. 10 B); propodeum coarsely reticulaterugose, with a distinct transversal carina connected with 4 blunt tubercles, lateral tubercles larger than medial ones (Fig. 10 C). Wings. Length of fore wing 2.7 × its width; length of marginal cell 0.8 × pterostigma; vein 2 - Rl just present, vein r shorter than 3 - SR; r: 3 - SR: SRl = 1: 1: 5; SRl more or less curved; parastigma distinctly swollen; 1 - CU 1: 2 - CU 1 = 1: 2; vein CUlb present (Fig. 10 A). Legs. Hind coxa densely short setose and striate-rugose; hind femur robust; length of hind femur, tibia and basitarsus 3.2 ×, 4.5 × and 6.4 × their maximum width, respectively. Metasoma. Carapace elongate-oval in dorsal view, coarsely reticulaterugose basally, longitudinal striate-rugose medially, densely reticulaterugose apically; length of carapace 2.4 × its maximum width in dorsal view (Fig. 10 H); ventral cavity (as a narrow channel) open to apex, with a pair of long and large teeth; ovipositor long, slender and the ovipositor sheath 0.16 × as long as fore wing, 0.44 × as long as hind tibia (Figs 10 G, 10 H). Colour. Black; antennomeres blackish-brown; basal half of clypeus reddish-brown; pterostigma and veins blackish-brown; fore wing more or less infuscate; basal third of carapace yellowish; legs almost entirely yellowishbrown, hind tibia light yellow basally and black apically; ovipositor sheath brown. Male. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
038887803102054BFF619ADAFDD07B3C.taxon	etymology	Etymology. From macro (Greek for long) and odontos (Greek for teeth), because of the apex of the metasomal carapace bears a pair of long teeth.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031040549FF619CE9FDEA7F76.taxon	type_taxon	Type-species: Chelonus (Scabrichelonus) sinensis He, Chen & van Achterberg, 1997 (Monotype). Microchelonus (Scabrichelonus): Tobias, 2000: 481. Scabrichelonus: Chen & Ji, 2003: 111 (as syn. of Microchelonus Szépligeti).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031040549FF619CE9FDEA7F76.taxon	diagnosis	Diagnosis. Head triangular in anterior view (Figs 14 F, 16 G), trapezoid in dorsal view (Figs 14 G, 16 H); temple distinctly swollen, with 3 strong ridge-like transverse carinae; face evenly convex, distinctly transversely rugose, with a longitudinal carina dorso-medially, mandibles robust, not twisted apically, upper tooth longer than lower tooth; (Figs 14 F, 16 G); occipital carina thin, arched dorso-medially, ventrally connected to hypostomal carina above base of mandible (Figs 14 G, 16 H); mesoscutum evenly convex, transverse suture distinct; notauli superficially impressed; propodeum short, near posterior margin of propodeum (Figs 14 C, 16 C); metapleuron coarsely rugose, without metapleural flange; apical transverse carina of (Figs 14 B, 16 B) length of metasoma equal to combined length of head and mesosoma; metasomal carapace elongate, without sutures, only slightly widened apically; ventral metasomal opening close to posterior margin of metasomal carapace; metasoma of male with an apical opening.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031040549FF619CE9FDEA7F76.taxon	biology_ecology	Biology. Unknown.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031040549FF619CE9FDEA7F76.taxon	distribution	Distribution. Oriental (China).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	description	Figs 13 – 16	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	materials_examined	Material examined. Holotype: female, China, Hainan Prov., Mt. Wuzhi, 16 – 20. V. 2007, Liqiong Wen, No. 200705133 (ZJUH). Paratype, 1 male, China, Hainan Prov., Mt. Wuzhi, 17. V. 2007, Bin Xiao, No. 200705132 (ZJUH).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	diagnosis	Diagnosis. This new species is similar to C. (S.) sinensis He, Chen & van Achterberg, but can be separated from the latter by the following characters: hind coxa and hind femur black in both sexes (hind coxa apically and hind femur basally more or less yellowish-brown in C. (S.) sinensis); apical opening of male carapace is relatively large and close to lateral margins of carapace (apical opening of male carapace relatively small and remaining removed from lateral margins of carapace).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	description	Description. Holotype, female, length of body 4.5 mm, of fore wing 3.4 mm. Head. Antennomeres more than 8 (broken in holotype), scapus robust, length of third antennomere 1.2 × fourth, length of third and fourth antennomeres 3.8 × and 3.3 × their width, respectively; in dorsal view its maximum width and width between eyes 1.2 × and 1.8 × its median length, respectively; temple strongly swollen posteriorly; frons slightly concave; temple and vertex distinctly transversely rugose; OOL: OD: POL = 21: 9: 11; longitudinal diameter of eye 0.8 × height of temples (Fig. 14 G); face as long as its width, evenly convex; clypeus flat, sparsely and finely punctate; length of malar space 0.5 × basal width of mandible (Fig. 14 F); face and eyes with densely setose in lateral view (Fig. 14 I). Mesosoma. Length of mesosoma 2.2 × its height; pronotum densely irregularly reticulate; mesoscutum evenly convex, irregularly finely reticulate-rugose, more strongly and coarsely rugose postero-medially; scutellum finely punctate-rugose; scutellar sulcus narrow, with 7 carinae; propodeum densely reticulate-rugose (Fig. 14 C); mesopleuron evenly convex, densely finely reticulate-rugose, more strongly and coarsely so antero-dorsally; prepectal carina just extending to level of postero-ventral margin of pronotum; metapleuron coarsely rugose (Fig. 14 B). Wings. Fore wing (Fig. 14 A): length of fore wing 3.0 × its width; length of vein 1 - R 1 0.9 × length of pterostigma; vein 2 - R 1 just present; r: 3 - SR: SRl = 11: 19: 70; parastigma slightly swollen; 1 - CU 1: 2 - CU 1 = 13: 46; vein CUlb present. Legs. Hind coxa shallowly punctate-rugose; hind femur robust, its length 3.1 × its width; hind tibia rather thick apically, its length 1.2 × length of tarsus; hind tarsus narrowed towards apex; length of basitarsus 0.4 × length of tarsus, 2.0 × length of inner tibial spur (Fig. 14 E). Metasoma. Length of metasoma 2.8 × its maximum width in dorsal view; metasomal carapace longitudinally densely reticulate-rugose (Fig. 14 H); ovipositor is hidden under the ventral opening of metasomal carapace, the ovipositor sheath 0.12 × as long as fore wing, 0.31 × as long as hind tibia (Fig. 14 D). Colour. Black; antennomeres blackish-brown; palp yellowish brown; scapus (except for base) and mandibles basally reddish-brown; pterostigma and most of veins blackish-brown; basal third of carapace yellowish; legs almost yellowish-brown totally; hind соха blасk but apically yellowish-brown; hind femur blackish-brown; basal hind tibia light yellow and apically brown. male. Antennomeres 30; similar to female, but differs in entirely black carapace (Figs 16 D, 16 E); carapace with a small apical opening which transverse elongate-oval in posterior view, 2.3 × wider than high, and height of opening 0.5 × ventral height of carapace (Fig. 16 K).	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	biology_ecology	Biology. Unknown	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
0388878031180555FF619ADAFAB67EA6.taxon	etymology	Etymology. Named after its blackish-brown hind femur: niger is Latin for black, femora is Latin for femur.	en	Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: 10.11646/zootaxa.5165.4.5
