taxonID	type	description	language	source
4C1E29C51D40F94610B877CAF7301DD2.taxon	materials_examined	Material examined. Greece: 2 ♂♂, Ionian Islands, Kerkyra, Chalikiopoulos [lagoon], 22. iv. 1935 (NHMW); 1 ♂, Eastern Macedonia and Thrace, Evros Distr., plain of Evros river, 26. vii. 1988, M. Jaech leg. (NHMW); 1 ♀, Central Macedonia, Khalkidhiki Distr., Sithonia, 2 km S of Kalamitsion, 12. viii. 2000, J. Hotovy leg. (NMPC); 5 ♂♂ 5 ♀♀, NW Peloponnese, 3 km S Kalogria, 38.1213 N, 21.3810 E, ca. 3 m, shallow seasonal swamp, 17. v. 2010, H. Fery & L. Hendrich leg. (NMPC). Hungary: 1 ♂ 1 ♀, Hungary 46 36 (BMNH); 1 ♀, Bacs-Kiskun, Kiskunmajsa env., 11. viii. 1999, J. Hajek leg. (NMPC). Montenegro: 1 ♂, Vranjina env., Skadarsko jezero, 20. ix. 2001, J. Hajek leg. (NMPC). Russia: 1 ♀, Orenburg reg., Totskoye, 1917, S. Jurecek leg. (NMPC); 2 ♂♂ 2 ♀♀, Samara, K. Fausta leg. (ZMAS); 1 ♀, Stavropol reg., Kuma river, 20. iv. 1911 (ZMAS); 1 ♂ 2 ♀♀, Volgograd reg., 2 km south of Zryanin village, 48 ° 36 ' 60 '' N, 43 ° 10 ' 10 '' E, small lakes near Liska river, incl silty open bay with grasses, Alisma and Juncus, 29 - 30. iv. 2002, J. Bergsten & A. Nilsson leg. (NHRS); 1 ♂ 1 ♀, Volgograd reg., Archeda-Don rivers alluvial sandy plain, 16 km ESE of Terkin village, 49.6861 N, 43.3333 E, different lakes, grassy ponds, fens and stream, 2 - 3. v. 2002, J. Bergsten & A. Nilsson leg. (NHRS); 3 ♂♂ 2 ♀♀, Volgograd reg., Kretskiy, 48.6083 N, 44.7061 E, river-arm, newly flooded grassland, 5. v. 2002, J. Bergsten & A. Nilsson leg. (NHRS). Turkey: 2 spec., Aydin vil. [= province], S of Aydin, ditch, 4. iv. 14987, H. Fery leg. (HFCB); 4 ♂♂ 1 ♀, Mugla vil. [= province], Koeycegiz, 27. v. 1991, S. Schoedl leg. (NHMW, NMPC). Slovakia: 1 ♂, 1 km N of Tvrdosovce, 24. iv. 2000, T. Kopecky leg. (NMPC). Ukraine: 1 ♂, Kherson distr., monast. Korsunskij, cursus inf. fl. Dnjepr, 3. vi. 1927, S. Medvedev leg. (ex coll. Zakharenko, ZMAS).	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
4C1E29C51D40F94610B877CAF7301DD2.taxon	diagnosis	Diagnosis. Habitus as depicted in Figs 4 c, 5 c. Clypeus with anterior margin rounded (Fig. 6 a). Reticulation of dorsal surface confined to head, more distinct and impressed anteriorly. Punctation of head fine, visible on whole surface; punctures sparse, distance between them usually equal or bigger than their diameter (Fig. 6 a). Punctation of pronotum double, fine, distance between larger punctures bigger than their diameter. Punctation of elytra double, diameter of small puncture less than half of diameter of large punctures; distance between large punctures bigger than their diameter. Epipleura smooth with fine punctures. Metatibia with sinuous outer margin. Male. Longer metatibial spur long, nearly as long as metatarsomere I-II combined (Fig. 7 a); spur bisinuate with only indistinct serration basally (Fig. 7 a). Male genitalia as in Fig. 8 a-d, median lobe in ventral view slightly narrowing from base to apex. Female. Both shiny and matt forms known of females of H. anatolicus. Shiny form agreeing well with male; matt form with whole surface densely reticulated, meshes somewhat elongate on elytra. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur shorter than in male; broad and with serration in basal two thirds, narrowed, slightly curved and without serration in apical third. Female genitalia as in Fig. 8 e-g.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
4C1E29C51D40F94610B877CAF7301DD2.taxon	distribution	Distribution. The species is distributed in the Eastern Mediterranean and in south-eastern Europe. It occurs in Italy, southernmost Slovakia, Hungary, the Balkan Peninsula, Turkey, southern Ukraine and Russia up to latitude 55 ° and east to the Ural Mountains (Fig. 9). First record from Slovakia and Ukraine.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
7EF60A8A6FAA9238C99941F20F2166A0.taxon	materials_examined	Material examined. We have examined more than 600 specimens from the Czech Republic, Finland, France, Germany, Great Britain, Russia, Slovakia, Sweden, and Ukraine, deposited in NHRS and NMPC.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
7EF60A8A6FAA9238C99941F20F2166A0.taxon	diagnosis	Diagnosis. Habitus as depicted in Figs 4 b, 5 b. Clypeus with anterior margin medially nearly straight (Fig. 6 b). Reticulation of dorsal surface confined to head, more distinct and impressed anteriorly (Fig. 6 b). Punctation of head fine, visible only in posterior half, punctures on clypeus imperceptible due to strong reticulation (Fig. 6 b); punctures dense, distance between them smaller than their diameter (Fig. 6 b). Punctation of pronotum double, coarse, distance between larger punctures smaller than their diameter. Punctation of elytra double, diameter of small puncture about half of diameter of large punctures; distance between large punctures, at least basally, smaller than their diameter. Epipleura smooth with fine punctures. Metatibia with outer margin nearly straight. Male. Longer metatibial spur short, only slightly longer than metatarsomere I (Fig. 7 b); spur nearly straight, broad with distinct serration (Fig. 7 b). Male genitalia as in Fig. 8 h-k, median lobe in ventral view parallel-sided in most of its length. Female. Both shiny and matt forms are known for females of H. ovatus. Shiny form agreeing well with male; matt form with whole surface densely reticulated, meshes distinctly elongate on elytra. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur similar to that of male. Female genitalia as in Fig. 8 l-n.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
7EF60A8A6FAA9238C99941F20F2166A0.taxon	distribution	Distribution. Widely distributed Palaearctic species. With the exception of the Iberian Peninsula, it occurs in most of territory of Europe and temperate Asia east to the Baikal Lake (east Siberia).	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
2A5ECDD7849177F8C1EC6A3C83D17A90.taxon	materials_examined	Material examined. Israel: 2 ♂♂, 1 ♀, Hula reserve, 21. iii. 1985; 4 ♂♂, 8 ♀♀, same locality, but 13. iv. 1986; 1 ♂, 3 ♀♀, Talme Elazar, 21. iv. 1986; 1 ♂, 6 ♀♀, Magan Michael, 21. iv. 1986, all M. Jaech leg. (NHMW, NMPC). Turkey: 3 ♂♂, 13 ♀♀, Mugla vil. [= province], Koeycegiz, 27. v. 1991, S. Schoedl leg. (NHMW, NMPC); 3 ♂♂, 2 ♀♀, same data, but M. Jaech leg. (NHMW, NMPC).	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
2A5ECDD7849177F8C1EC6A3C83D17A90.taxon	diagnosis	Diagnosis. Habitus as depicted in Figs 4 d, 5 d. Clypeus with anterior margin medially nearly straight (Fig. 6 c). Reticulation of dorsal surface confined to head and more distinct and impressed anteriorly (Fig. 6 c), and to sides of pronotum. Punctation of head fine, visible on whole surface (Fig. 6 c); punctures dense, distance between them smaller than their diameter (Fig. 6 c). Punctation of pronotum double, fine, distance between larger punctures bigger than their diameter. Punctation of elytra double, diameter of small punctures less than half of diameter of large punctures; distance between large punctures bigger than their diameter. Epipleura reticulated with very fine punctures. Metatibia with outer margin nearly straight. Male. Longer metatibial spur long, nearly as long as metatarsomere I-II combined (Fig. 7 c); spur broad and straight in basal two thirds with small but distinct serration, attenuated and curved apically (Fig. 7 c). Male genitalia as in Fig. 8 o-r, median lobe in ventral view slightly narrowing from base to apex. Female. Only matt females of H. sanctus are known so far. Whole surface densely reticulated, meshes on elytra somewhat elongate. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur similar to that of male, but almost straight in apical third. Female genitalia as in Fig. 8 s-u.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
2A5ECDD7849177F8C1EC6A3C83D17A90.taxon	distribution	Distribution. A species distributed in the Levant region of the Near East. So far recorded from several localities in Israel, Jordan and Syria (Fig. 9). First record from Turkey.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
D33CB305D2B2DBCFF23DCE4577DF4B1F.taxon	materials_examined	Material examined. Turkey: 1 ♂, 2 ♀♀, Mugla vil. [= province], Koeycegiz, 27. v. 1991, S. Schoedl leg. (NHMW, NMPC).	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
D33CB305D2B2DBCFF23DCE4577DF4B1F.taxon	distribution	Distribution. Predominantly a Mediterranean species. First record from Turkey.	en	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
