identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4C1E29C51D40F94610B877CAF7301DD2.text	4C1E29C51D40F94610B877CAF7301DD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus anatolicus Guignot 1957	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hyphydrus anatolicus Guignot, 1957</p>
            <p> Hyphydrus anatolicus Guignot, 1957: 91 (orig. descr.; type locality:  “Angora” [Ankara, Turkey]). </p>
            <p> Hyphydrus carrarai Sanfilippo, 1963: 77 (orig. descr.; type locality: "Macchia di Migliarino, Torre del Lago (Toscana)" [Italy]); synonymy by Pederzani 1976: 166. </p>
            <p> Hyphydrus sanctus :  Biström 1982: 39 (partim, misidentification). </p>
            <p>Published records.</p>
            <p> Bosnia and Hercegovina:  Biström (1982: 39 as  H. sanctus ). Croatia:  Guéorguiev (1971: 8 as  H. carrarai );  Biström (1982: 39 as  H. sanctus );  Ádám (1992: 194 as  Hyphydrus sanctus );  Temunović et al. (2007: 17);  Krčmar (2014: 20). Greece:  Biström (1982: 39 as  H. sanctus ); Wewalka (1984: 131). Hungary:  Ádám (1992: 194 as  H. sanctus ); Csabai et al. (1999: 148 as  H. sanctus );  Móra et al. (2004: 153); Csabai and Nosek (2006: 73);  Kálmán et al. (2008: 76);  Molnár (2008:110);  Sóos et al. (2008: 223);  Lőkkös (2010:161). Italy: Sanfilippo (1963: 77 as  H. carrarai ); Angelini (1972: 182 as  H. carrarai ; 1984: 54); Pederzani (1976: 166);  Biström (1982: 39 as  H. sanctus ); Rocchi (1991: 68); Pederzani &amp; Campadelli (1996: 21); Nardi (1997: 132); Bordoni et al. (2006: 87). Macedonia:  Biström (1982: 39 as  H. sanctus ). Montenegro: Scheers (2016: 209). Russia:  Biström (1982: 39 as  H. sanctus ). Serbia:  Mesaroš (2015: 50). Turkey: Guignot (1957: 91). </p>
            <p>Material examined.</p>
            <p> Greece: 2♂♂, Ionian Islands, Kerkyra, Chalikiopoulos [lagoon], 22.iv.1935 (NHMW); 1♂, Eastern Macedonia and Thrace,  Évros Distr., plain of  Évros river, 26.vii.1988, M.  Jäch leg. (NHMW); 1♀, Central Macedonia, Khalkidhiki Distr., Sithonia, 2 km S of  Kalamítsion , 12.viii.2000, J.  Hotový leg. (NMPC); 5♂♂ 5♀♀, NW Peloponnese, 3 km S Kalogria, 38.1213N, 21.3810E, ca. 3 m, shallow seasonal swamp, 17.v.2010, H. Fery &amp; L. Hendrich leg. (NMPC). Hungary: 1♂ 1♀, Hungary 46 36 (BMNH); 1♀,  Bács-Kiskun , Kiskunmajsa env., 11.viii.1999, J.  Hájek leg. (NMPC). Montenegro: 1♂, Vranjina env., Skadarsko jezero, 20.ix.2001, J.  Hájek leg. (NMPC). Russia: 1♀, Orenburg reg., Totskoye, 1917,  Š .  Jureček leg. (NMPC); 2♂♂ 2♀♀, Samara, K. Fausta leg. (ZMAS); 1♀, Stavropol reg., Kuma river, 20.iv.1911 (ZMAS); 1♂ 2♀♀, Volgograd reg., 2 km south of Zryanin village, 48°36  ‘60‘‘ N, 43°10  ‘10‘‘ E, small lakes near Liska river, incl silty open bay with grasses,  Alisma and  Juncus , 29-30.iv.2002, J. Bergsten &amp; A. Nilsson leg. (NHRS); 1♂ 1♀, Volgograd reg., Archeda-Don rivers alluvial sandy plain, 16 km ESE of Terkin village, 49.6861N, 43.3333E, different lakes, grassy ponds, fens and stream, 2-3.v.2002, J. Bergsten &amp; A. Nilsson leg. (NHRS); 3♂♂ 2♀♀, Volgograd reg., Kretskiy, 48.6083N, 44.7061E, river-arm, newly flooded grassland, 5.v.2002, J. Bergsten &amp; A. Nilsson leg. (NHRS). Turkey: 2 spec., Aydin vil. [= province], S of Aydin, ditch, 4.iv.14987, H. Fery leg. (HFCB); 4♂♂ 1♀,  Muğla vil. [= province],  Köyçeğiz , 27.v.1991, S.  Schödl leg. (NHMW, NMPC). Slovakia: 1♂, 1 km N of  Tvrdošovce , 24.iv.2000, T.  Kopecký leg. (NMPC). Ukraine: 1♂, Kherson distr., monast. Korsunskij, cursus inf. fl. Dnjepr, 3.vi.1927, S. Medvedev leg. (ex coll. Zakharenko, ZMAS). </p>
            <p>Diagnosis.</p>
            <p>Habitus as depicted in Figs 4c, 5c. Clypeus with anterior margin rounded (Fig. 6a). Reticulation of dorsal surface confined to head, more distinct and impressed anteriorly. Punctation of head fine, visible on whole surface; punctures sparse, distance between them usually equal or bigger than their diameter (Fig. 6a). Punctation of pronotum double, fine, distance between larger punctures bigger than their diameter. Punctation of elytra double, diameter of small puncture less than half of diameter of large punctures; distance between large punctures bigger than their diameter. Epipleura smooth with fine punctures. Metatibia with sinuous outer margin.</p>
            <p> Male. Longer metatibial spur long, nearly as long as metatarsomere I-II combined (Fig. 7a); spur bisinuate with only indistinct serration basally (Fig. 7a). Male genitalia as in Fig. 8  a–d , median lobe in ventral view slightly narrowing from base to apex. </p>
            <p> Female. Both shiny and matt forms known of females of  H. anatolicus . Shiny form agreeing well with male; matt form with whole surface densely reticulated, meshes somewhat elongate on elytra. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur shorter than in male; broad and with serration  in basal two thirds, narrowed, slightly curved and without serration in apical third. Female genitalia as in Fig. 8  e–g . </p>
            <p>Habitat.</p>
            <p> The species inhabits various types of standing water, predominantly densely vegetated pools, ditches and small ponds.  H. anatolicus tolerates also saline habitats. </p>
            <p>Distribution.</p>
            <p>The species is distributed in the Eastern Mediterranean and in south-eastern Europe. It occurs in Italy, southernmost Slovakia, Hungary, the Balkan Peninsula, Turkey, southern Ukraine and Russia up to latitude 55° and east to the Ural Mountains (Fig. 9). First record from Slovakia and Ukraine.</p>
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	https://treatment.plazi.org/id/4C1E29C51D40F94610B877CAF7301DD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bergsten, Johannes;Weingartner, Elisabeth;Hajek, Jiri	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
7EF60A8A6FAA9238C99941F20F2166A0.text	7EF60A8A6FAA9238C99941F20F2166A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus ovatus (Linnaeus 1760) Linnaeus 1760	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hyphydrus ovatus (Linnaeus, 1760)</p>
            <p> Dytiscus ovatus Linnaeus, 1760: 547 (type locality: Svecia [Sweden]). For full list of synonymy, see Nilsson &amp;  Hájek (2017a: 199). </p>
            <p>Material examined.</p>
            <p>We have examined more than 600 specimens from the Czech Republic, Finland, France, Germany, Great Britain, Russia, Slovakia, Sweden, and Ukraine, deposited in NHRS and NMPC.</p>
            <p>Diagnosis.</p>
            <p>Habitus as depicted in Figs 4b, 5b. Clypeus with anterior margin medially nearly straight (Fig. 6b). Reticulation of dorsal surface confined to head, more distinct and impressed anteriorly (Fig. 6b). Punctation of head fine, visible only in posterior half, punctures on clypeus imperceptible due to strong reticulation (Fig. 6b); punctures dense, distance between them smaller than their diameter (Fig. 6b). Punctation of pronotum double, coarse, distance between larger punctures smaller than their diameter. Punctation of elytra double, diameter of small puncture about half of diameter of large punctures; distance between large punctures, at least basally, smaller than their diameter. Epipleura smooth with fine punctures. Metatibia with outer margin nearly straight.</p>
            <p> Male. Longer metatibial spur short, only slightly longer than metatarsomere I (Fig. 7b); spur nearly straight, broad with distinct serration (Fig. 7b). Male genitalia as in Fig. 8  h–k , median lobe in ventral view parallel-sided in most of its length. </p>
            <p> Female. Both shiny and matt forms are known for females of  H. ovatus . Shiny form agreeing well with male; matt form with whole surface densely reticulated, meshes distinctly elongate on elytra. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur similar to that of male. Female genitalia as in Fig. 8  l–n . </p>
            <p>Habitat.</p>
            <p>The species inhabits various types of standing and slowly flowing water bodies with at least some vegetation. The typical habitats represent (frequently eutrophic) ponds, densely vegetated pools, ditches, oxbows or open swamps.</p>
            <p>Distribution.</p>
            <p>Widely distributed Palaearctic species. With the exception of the Iberian Peninsula, it occurs in most of territory of Europe and temperate Asia east to the Baikal Lake (east Siberia).</p>
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	https://treatment.plazi.org/id/7EF60A8A6FAA9238C99941F20F2166A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bergsten, Johannes;Weingartner, Elisabeth;Hajek, Jiri	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
2A5ECDD7849177F8C1EC6A3C83D17A90.text	2A5ECDD7849177F8C1EC6A3C83D17A90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus sanctus Sharp 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hyphydrus
sanctus Sharp, 1882
</p>
            <p> Hyphydrus sanctus Sharp, 1882: 380. </p>
            <p>Published records.</p>
            <p> Israel: Sharp (1882: 380);  Biström (1982: 39); Wewalka (1984: 131). Jordan:  Biström (1982: 39); Wewalka (1984: 131). Syria:  Biström (1982: 39); Wewalka (1984: 131). </p>
            <p>Material examined.</p>
            <p> Israel: 2♂♂, 1♀, Hula reserve, 21.iii.1985; 4♂♂, 8♀♀, same locality, but 13.iv.1986; 1♂, 3♀♀, Talme Elazar, 21.iv.1986; 1♂, 6♀♀, Magan Michael, 21.iv.1986, all M.  Jäch leg. (NHMW, NMPC). Turkey: 3♂♂, 13♀♀,  Muğla vil. [=province],  Köyçeğiz , 27.v.1991, S.  Schödl leg. (NHMW, NMPC); 3♂♂, 2♀♀, same data, but M.  Jäch leg. (NHMW, NMPC). </p>
            <p>Diagnosis.</p>
            <p>Habitus as depicted in Figs 4d, 5d. Clypeus with anterior margin medially nearly straight (Fig. 6c). Reticulation of dorsal surface confined to head and more distinct and impressed anteriorly (Fig. 6c), and to sides of pronotum. Punctation of head fine, visible on whole surface (Fig. 6c); punctures dense, distance between them smaller than their diameter (Fig. 6c). Punctation of pronotum double, fine, distance between larger punctures bigger than their diameter. Punctation of elytra double, diameter of small punctures less than half of diameter of large punctures; distance between large punctures bigger than their diameter. Epipleura reticulated with very fine punctures. Metatibia with outer margin nearly straight.</p>
            <p> Male. Longer metatibial spur long, nearly as long as metatarsomere I-II combined (Fig. 7c); spur broad and straight in basal two thirds with small but distinct serration, attenuated and curved apically (Fig. 7c). Male genitalia as in Fig. 8  o–r , median lobe in ventral view slightly narrowing from base to apex. </p>
            <p> Female. Only matt females of  H. sanctus are known so far. Whole surface densely reticulated, meshes on elytra somewhat elongate. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur similar to that of male, but almost straight in apical third. Female genitalia as in Fig. 8  s–u . </p>
            <p>Habitat.</p>
            <p> Similarly to the other two species,  H. sanctus inhabits various types of standing and slowly flowing water bodies with at least some vegetation. Wewalka (1984) reported several specimens from a densely vegetated pool and single occurrences from an artificial pool with clear water, an irrigation ditch and from a stream. </p>
            <p>Distribution.</p>
            <p>A species distributed in the Levant region of the Near East. So far recorded from several localities in Israel, Jordan and Syria (Fig. 9). First record from Turkey.</p>
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	https://treatment.plazi.org/id/2A5ECDD7849177F8C1EC6A3C83D17A90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bergsten, Johannes;Weingartner, Elisabeth;Hajek, Jiri	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
D33CB305D2B2DBCFF23DCE4577DF4B1F.text	D33CB305D2B2DBCFF23DCE4577DF4B1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyphydrus aubei Ganglbauer 1891	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hyphydrus aubei Ganglbauer, 1891</p>
            <p>Note.</p>
            <p> Hyphydrus aubei is the fourth European species in the  Hyphydrus ovatus species group sensu  Biström (1982). It does not belong to the  Hyphydrus ovatus complex as here defined and it is easily separated from the preceding three species based on colouration (Figures 4-5). </p>
            <p> Material examined. </p>
            <p> Turkey: 1♂, 2♀♀,  Muğla vil. [=province],  Köyçeğiz , 27.v.1991, S.  Schödl leg. (NHMW, NMPC). </p>
            <p>Distribution.</p>
            <p>Predominantly a Mediterranean species. First record from Turkey.</p>
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	https://treatment.plazi.org/id/D33CB305D2B2DBCFF23DCE4577DF4B1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Bergsten, Johannes;Weingartner, Elisabeth;Hajek, Jiri	Bergsten, Johannes, Weingartner, Elisabeth, Hajek, Jiri (2017): Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae). ZooKeys 678: 73-96, DOI: http://dx.doi.org/10.3897/zookeys.678.12886, URL: http://dx.doi.org/10.3897/zookeys.678.12886
