taxonID	type	description	language	source
038D927EFFE9FF9145EA78E2FEADFE96.taxon	description	(Figs 1, 4 – 5, 9)	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	materials_examined	Material studied. Holotype: male (PHILIPPINES, LAGUNA PROVINCE): three labels: " (PHILIPPINES) / Mt. Maquiling / alt. 400 m / Laguna Prov. / Luzon, 12. IX. / 1985, K. Ishikawa " [white, printed], " Tullgren sample / of leaf litter " [white, printed], " PENICILLIDMUS / maquilingensis m. / P. Jałoszyński, 2018 / HOLOTYPUS " [red, printed] (EUMJ).	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	diagnosis	Diagnosis. Male: BL ~ 1 mm; frons and vertex with punctures much more distinct than those on pronotum and elytra; pronotal base with only one pair of small and shallow pits connected by a distinct transverse groove; aedeagus with broadly subtriangular apex.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	description	Description. Body of male (Fig. 1) elongate but moderately slender, strongly convex, with short and robust appendages, BL 1.08 mm; cuticle glossy, pigmentation reddish-brown with elytra slightly darker than remaining body parts, vestiture of setae yellowish. Head slightly flattened and transverse, broadest at eyes, HL 0.20 mm, HW 0.25 mm; tempora in dorsal view distinctly shorter than eyes; vertex weakly and evenly convex; frons confluent with vertex, weakly convex at middle and slightly flattened at eyes; supraantennal tubercles small and feebly elevated. Eyes large, weakly emarginate posteriorly, each composed of about 18 ommatidia, distinctly projecting laterally from the head silhouette. Punctures on vertex and frons distinct and dense, with sharply marked margins and separated by spaces subequal to diameters of punctures; setae short and dense, nearly recumbent. Antennae slender but short, AnL 0.45 mm; antennomere I about as long as broad, II distinctly elongate, III – VIII each indistinctly transverse, IX strongly transverse, much broader than VIII, X slightly transverse, distinctly broader and longer than IX, XI about as long as X but slightly narrower, about as long as broad. Pronotum broadest near middle, PL 0.30 mm, PW 0.35 mm; anterior margin weakly arcuate, lateral margins rounded in anterior half, weakly angulate at insertions of lateral setal brushes and behind them nearly straight and distinctly convergent posteriorly; posterior pronotal corners obtuse-angled and blunt; posterior margin arcuate; base with only one pair of small and shallow pits connected by a distinct transverse groove. Sides of pronotum lacking longitudinal carinae. Punctures on pronotal disc fine, superficial, inconspicuous; setae dense, longer than those on vertex and suberect. Elytra oval, as convex as pronotum, broadest in front of middle, EL 0.58 mm, EW 0.50 mm, EI 1.15; basal impressions short; elytral apices separately rounded. Punctures on elytral disc more distinct than those on pronotum, nearly as dense as those on frons but superficial and diffuse; setae short, moderately dense and suberect, additionally sides of elytra with sparse, long and erect setae. Hind wings long, functional. Legs moderately long and slender, unmodified. Aedeagus (Figs 4 – 5) moderately slender, AeL 0.13 mm; basal half nearly parallel-sided, apical third gradually narrowed distad, subtrapezoidal, with triangular, obtuse-angled apex; endophallus with a symmetrical pair of curved bunches of elongate sclerites in subapical region and a pair of rod-like structures in sub-basal area. Parameres slender, short, each with one long apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	distribution	Distribution (Fig. 9). Philippines, southern Luzon, Laguna Province.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	etymology	Etymology. Locotypical, after Mt. Maquiling.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFE9FF9145EA78E2FEADFE96.taxon	discussion	Remarks. The aedeagus of P. maquilingensis differs from that of all known congeners in a distinctly stouter median lobe, and especially its apical portion, which forms an obtuse apical angle, whereas it is sharp-angled in all remaining species.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	description	(Figs 2, 6 – 7, 9)	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	materials_examined	Material studied. Holotype: male (PHILIPPINES, LAGUNA PROVINCE): three labels: " (PHILIPPINES) / Mt. Maquiling / alt. 400 m / Laguna Prov. / Luzon, 12. IX. / 1985, K. Ishikawa " [white, printed], " Tullgren sample / of fallen fruit / of Ficus sp. " [white, printed], " PENICILLIDMUS / lagunensis m. / P. Jałoszyński, 2018 / HOLOTYPUS " [red, printed] (EUMJ).	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	diagnosis	Diagnosis. Male: BL distinctly below 1 mm; frons and vertex with punctures only slightly more distinct than those on pronotum and elytra; pronotal base with one pair of large submedian and one pair of smaller lateral pits not connected by groove; aedeagus with very narrow, slender subtriangular apex.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	description	Description. Body of male (Fig. 2) elongate but moderately slender, strongly convex, with short and robust appendages, BL 0.89 mm; cuticle glossy, pigmentation uniformly brown, vestiture of setae yellowish. Head weakly and evenly convex, transverse, broadest at eyes, HL 0.19 mm, HW 0.23 mm; tempora in dorsal view distinctly shorter than eyes; vertex and frons confluent, weakly and evenly convex; supraantennal tubercles small and feebly elevated. Eyes large, weakly emarginate posteriorly, each composed of 14 – 15 ommatidia, distinctly projecting laterally from the head silhouette. Punctures on vertex and frons distinct and dense, but with diffuse margins, separated by spaces subequal to diameters of punctures; setae short and dense, nearly recumbent. Antennae slender but short, AnL 0.28 mm; antennomere I about as long as broad, II distinctly elongate, III – X each transverse, XI indistinctly longer than X but slightly narrower, about as long as broad. Pronotum broadest near middle, PL 0.23 mm, PW 0.23 mm; anterior margin weakly arcuate, lateral margins rounded in anterior half, distinctly angulate at papillate insertions of lateral setal brushes and behind them weakly concave and strongly convergent posteriorly; posterior pronotal corners obtuse-angled and blunt; posterior margin arcuate; base with two pairs of antebasal pits, of which submedian pits are large and lateral ones are much smaller, transverse groove absent. Sides of pronotum lacking longitudinal carinae. Punctures on pronotal disc similarly dense as those on frons and vertex but indistinctly shallower; setae dense, longer than those on vertex and suberect. Elytra oval, as convex as pronotum, broadest indistinctly in front of middle, EL 0.48 mm, EW 0.35 mm, EI 1.36; basal impressions short but deep; elytral apices separately rounded; narrow stripe at each side of suture distinctly raised in anterior half of elytra. Punctures on elytral disc less distinct than those on pronotum, superficial; setae short, moderately dense and suberect. Hind wings long, functional. Legs moderately long and slender, unmodified. Aedeagus (Figs 6 – 7) slender, AeL 0.10 mm; median lobe drop-shaped, broadest near base and gradually narrowing up to subapical third, where it narrows more rapidly to form a conspicuously long and narrow apical portion, which is subtriangular and its tip is sharp-angled; endophallus with an asymmetrical pair of long rod-like structures in proximal half. Parameres were damaged during preparation. Female. Unknown.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	distribution	Distribution (Fig. 9). Philippines, southern Luzon, Laguna Province.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	etymology	Etymology. Locotypical, after the Laguna Province.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEAFF9145EA7BE4FED4F860.taxon	discussion	Remarks. The aedeagus of P. lagunensis differs from that of all congeners in the conspicuously narrow and long apical portion of the median lobe; it also has a dorsal apical plate similar in shape, but much shorter than the ventral apical projection; in all remaining species the dorsal apical plate is either short or indistinctly defined (P. masseyensis, P. unicolor and P. lavongai), much shorter and broader (P. maquilingensis), or truncated at apex (P. luzonicus).	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	description	(Figs 3, 8, 9)	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	materials_examined	Material studied. Holotype: male (PHILIPPINES, LAGUNA PROVINCE): three labels: " (PHILIPPINES) / Mt. Maquiling / alt. 400 m / Laguna Prov. / Luzon, 12. IX. / 1985, K. Ishikawa " [white, printed], " Tullgren sample / of fallen fruit / of Ficus sp. " [white, printed], " PENICILLIDMUS / luzonicus m. / P. Jałoszyński, 2018 / HOLOTYPUS " [red, printed] (EUMJ).	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	diagnosis	Diagnosis. Male: BL distinctly less than 1 mm; frons and vertex with punctures as inconspicuous as those on pronotum and elytra; pronotal base with one pair of large submedian and one pair of much smaller lateral pits not connected by groove; aedeagus with moderately narrow subtriangular apex.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	description	Description. Body of male (Fig. 3) elongate but moderately slender, strongly convex, with short and robust appendages, BL 0.75 mm; cuticle glossy, pigmentation uniformly light brown, vestiture of setae yellowish. Head weakly and evenly convex, broadest at eyes, HL 0.15 mm, HW 0.18 mm; tempora in dorsal view indistinctly shorter than eyes; vertex and frons confluent and weakly, evenly convex; supraantennal tubercles small and feebly elevated. Eyes large, only indistinctly emarginate posteriorly, each composed of about 16 ommatidia, distinctly projecting laterally from the head silhouette. Punctures on vertex and frons dense but very shallow and diffuse, superficial and inconspicuous; setae short and dense, nearly recumbent. Antennae slender but short, AnL 0.24 mm; antennomere I about as long as broad, II distinctly elongate, III – X each transverse, XI indistinctly longer than X but slightly narrower, about as long as broad. Pronotum broadest near middle, PL 0.23 mm, PW 0.23 mm; anterior margin weakly arcuate, lateral margins rounded in anterior half, weakly angulate at papillate insertions of lateral setal brushes and behind them nearly straight and distinctly convergent posteriorly; posterior pronotal corners obtuse-angled and blunt; posterior margin arcuate; base with two pairs of antebasal pits, of which submedian pits are large and lateral ones much smaller, transverse groove absent. Sides of pronotum lacking longitudinal carinae. Punctures on pronotal disc fine and inconspicuous; setae short, moderately dense, suberect. Elytra oval, as convex as pronotum, broadest indistinctly in front of middle, EL 0.38 mm, EW 0.28 mm, EI 1.36; basal impressions short but deep; elytral apices separately rounded. Punctures on elytral disc fine and inconspicuous; setae short, moderately dense and suberect. Hind wings long, functional. Legs moderately long and slender, unmodified. Aedeagus (Fig. 8) moderately slender, AeL 0.13 mm; median lobe drop-shaped, nearly parallel-sided in proximal half and rapidly narrowing distad in apical half, with its apical portion forming a triangular projection with sharp-angled apex; endophallus weakly sclerotized, with a symmetrical median elongate structure. Parameres broad and short, each with one long apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	distribution	Distribution (Fig. 9). Philippines, southern Luzon, Laguna Province.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	etymology	Etymology. Locotypical, after the island of Luzon.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
038D927EFFEBFF9645EA7C11FED0FEB3.taxon	discussion	Remarks. The aedeagus of P. luzonicus differs from that of all congeners in the subtrapezoidal and truncated ventral subapical plate; see remarks for P. lagunensis. The aedeagus of the holotype was studied and illustrated only in ventral view to avoid damage during manipulations; it is particularly weakly sclerotized and prone to distortions.	en	Jałoszyński, Paweł (2019): Three new species of Penicillidmus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa 4563 (1): 185-190, DOI: 10.11646/zootaxa.4563.1.11
