identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038D8112A02DD67AFBE0FC78374226AF.text	038D8112A02DD67AFBE0FC78374226AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travisia brevis Moore 1923	<div><p>Travisia brevis Moore, 1923</p><p>(Figs 1‒3)</p><p>Travisia brevis Moore, 1923: 220‒221 .— Berkeley &amp; Berkeley, 1952: 90‒91, fig. 183.— Hartman, 1961: 34; 1966: 409; 1969: 343‒344, fig. 1.— Imajima, 1963: 361; 1964: 247; 2009: 138–139 — Reish, 1965: 145.— Fauchald, 1972: 237.— Hobson &amp; Banse, 1981: 62, fig. 13h.— Blake, 2000: 161‒162, fig. 7.7.— Hendrickx, 2005: 83.— Méndez, 2006: 777; 2007: 609; 2012: 184.— Mercado-Santiago et al. 2021: 388.— Hernández-Alcántara et al. 2023: 4.— Hernández-Alcántara et al. 2025: 6.</p><p>Material examined. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-100.47&amp;materialsCitation.latitude=16.795" title="Search Plazi for locations around (long -100.47/lat 16.795)">All</a> off Guerrero, Mexico. ICML-EMU-14077-A, TALUD XI, St. 2, BC, 16º47’42”N, 100º28’12”W, 07 June 2007, 880 m, 1 specimen . ICML-EMU-14077-B, TALUD XI, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-100.79278&amp;materialsCitation.latitude=16.876389" title="Search Plazi for locations around (long -100.79278/lat 16.876389)">St.</a> 5, BC, 16º52’35”N, 100º47’34”W, 07 June 2007, 1550 m, 1 specimen . ICML-EMU-14078-A, TALUD XII, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-101.657776&amp;materialsCitation.latitude=17.071112" title="Search Plazi for locations around (long -101.657776/lat 17.071112)">St.</a> 8, BC, 17º04’16”N, 101º39’28”W, 29 March 2008, 1928 m, 3 specimens . ICML-EMU-14078-B, TALUD XII, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-104.5975&amp;materialsCitation.latitude=18.674444" title="Search Plazi for locations around (long -104.5975/lat 18.674444)">St.</a> 27, BS, 18º40’28”N, 104º35’51”W, 02 April 2008, 1040‒1095 m, 3 specimens .</p><p>Diagnosis. Travisia with annulate branchiae from chaetiger 2, 18‒21 pairs of branchia and 19‒22 chaetigers. Mid-ventral groove absent. Nuchal organs small, C-shaped. Parapodial lappets well-developed, present along body.</p><p>Description. Specimens 20‒23 mm long, 3.5‒6.0 mm wide with 19‒22 chaetigers. Body brownish in alcohol; robust, grub-like (Fig. 1A‒B) with rugose epidermis (Figs 1‒3). Mid-ventral groove absent.</p><p>Prostomium small, conical, pointed, smooth, iridescent (Figs 1A, C‒E, 2A, 3A‒B). Nuchal organs small, Cshaped, dorsolateral (Figs 1C‒D, 3A‒B). Eyes absent. Mouth between chaetigers 1 and 2 (Figs 1E, 3B). Pharynx not everted in any specimens.</p><p>Peristomium uniannulated. Chaetiger 1 biannulated. Chaetigers 2 to 19 triannulated. Following chaetigers uniannulated to pygidium. Segments from first two body quarters with annulation visible along entire segment width (from parapodium to parapodium); segments from third body quarter toward posterior end with annulation visible only on middorsal and midventral zones (annulation not reaching parapodia).</p><p>Epithelium of peristomium and first three chaetigers fully covered with small, spherical papillae (dorsal, ventral and laterally) (Figs 1A, C‒E, 2A, 3A‒B). In chaetigers 4 to 13, dorsal and ventral surfaces with three narrow ridges, each ridge with 1‒2 rows of tightly packed, small, spherical papillae with lateral-most papillae enlarged (Figs 1A, 2B, E); lateral sides with epithelium fully covered with small, rounded papillae (Fig. 2B‒C). Mid-body segments with five broad ridges dorsally and ventrally, each composed of many tight papillae, unequal in size (Fig. 2F). An intersegmental package of papillae ventrally along last body quarter, oval-shaped to rim-shaped, depending on fixation (Fig. 3C‒D); papillae diminishing gradually in size toward posterior segments.</p><p>Interamal sensory pores small, present between notochaetae and neurochaetae from chaetiger 2 to near last segments (Figs 1B, 2C). Lateral nephridial pores not seen.</p><p>Branchiae annulate (Fig. 2A‒D), 18‒21 pairs, starting in chaetiger 2. First pair short, increasing in size towards medium chaetigers, decreasing in size gradually to posterior end (Figs 1A‒B, 2A‒D). Last branchia small, digitate.</p><p>Parapodial lappets conical, present from chaetiger 17 continued to posterior segments, covered by clusters of dense, small spherical papillae (Fig. 2G‒I). From chaetiger 20 to end, distal-most papilla of parapodial lappets remarkably larger than basal ones (Fig. 2G‒I).</p><p>Parapodia biramous. All noto- and neurochaetae iridescent, spinulose capillaries. Last 4‒6 segments achaetous.</p><p>Pygidium short, thick, with one ring of 12 lobes, without anal cirri (Figs 1B, 2D, 3C‒E).</p><p>Methyl green staining. Prostomium and branchiae unstained (Fig. 3A, C). Rest of the body is uniformly stained.</p><p>Remarks. Travisia brevis has been reported in many ecological surveys and technical reports (e.g., Méndez 2006, 2007, 2012; Carson &amp; Hentschel 2006; Henkel et al. 2020; Mercado-Santiago et al. 2021), but apart from the original description there are few illustrations of this species. Unfortunately, the original establishment by Moore (1923) did not include drawings. Berkeley &amp; Berkeley (1952: fig. 183) included a drawing of the body in dorsal view which was later replicated by Hartman (1969: fig. 1) and Blake (2000: fig. 7.7A). A chaetiger in lateral view was illustrated by Hobson &amp; Banse (1981: fig. 13h) and then replicated by Blake (2000: fig. 7.7B). Digital photographs of T. brevis are available in technical sheets of the Southern California Association of Marine Invertebrate Taxonomists (SCAMIT 2016) and Burgess (2018a). In the present study, a full description and additional digital photographs of body structures are provided (see Figs 1–3) in order to help further identifications of T. brevis in western Mexico or elsewhere along the eastern Pacific.</p><p>Species of Travisia have been classified in two groups based on the absence or presence of branchiae by Rizzo &amp; Salazar-Vallejo (2020, 2021 taxonomic identification key) and Plathong et al. (2023). Group I includes four species lacking branchiae whereas group II consists of species with branchiae. Group II is also separated into two subgroups based on the type of branchiae: subgroup A with branched branchiae, and subgroup B with cirriform branchiae. Subgroup B is further divided into B1 based on the presence of a ventral groove, and B2 lacking a ventral groove. Following this recognition of groups and subgroups, B2 is currently composed of 33 species when the three species described by Avery et al. (2023) are considered.</p><p>A comprehensive comparison of species of Travisia is a complicated matter, mostly because after their establishment they have seldom been recorded and are often included in lists of species or ecological studies without being thoroughly examined from a morphological point of view. Revisions based on type material, redescriptions, and appropriate illustrations are not available. The majority of the original descriptions do not include adequate description of the morphological features that the current study of Travisia demands (see Rizzo &amp; Salazar 2020, Yang et al. 2022, Avery et al. 2023, Plathong et al. 2023). Some do not even include figures of body structures or, if present, these can be interpreted erroneously or in a speculative manner. In addition, the size of worms, number of chaetigers, achaetous segments and branchia, and the beginning of interamal sensory pores and lateral nephridiopores may be features with discrete or overlapped variations that do not allow a precise distinction of taxa to species level. Besides, intrasegmental divisions and patterns of papillation are features that demand a very detailed and specific study, sometimes difficult to perform due to body contractions or to a dense papillation above the body epithelium.</p><p>Abiotic conditions. The specimens of T. brevis were collected from 880–1928 m depth, under the following environmental conditions: temperature, 2.3–5.1°C; salinity, 34.53–34.63 ppm; dissolved oxygen, 0.24– 1.49 O 2 mg / L; %CO, 1.37–1.96; sediments composition variable, including silty clay, mixed, and muddy sand (Table 1).</p></div>	https://treatment.plazi.org/id/038D8112A02DD67AFBE0FC78374226AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tovar-Hernández, María Ana;Burgess, Dany E.;León-González, Jesús Angel De;Hendrickx, Michel E.	Tovar-Hernández, María Ana, Burgess, Dany E., León-González, Jesús Angel De, Hendrickx, Michel E. (2025): Travisiid worms from deep water off Western Mexico with the establishment of a new species (Annelida: Travisiidae). Zootaxa 5729 (2): 349-369, DOI: 10.11646/zootaxa.5729.2.7, URL: https://doi.org/10.11646/zootaxa.5729.2.7
038D8112A029D676FBE0FA52360121C3.text	038D8112A029D676FBE0FA52360121C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travisia pupa Moore 1906	<div><p>Travisia pupa Moore, 1906</p><p>(Figs 4‒5)</p><p>Travisia pupa Moore, 1906: 228‒231, Pl. XI, fig. 23; 1923: 221.— Uschakov, 1950: 209; 1955: 324, fig. 120F‒G.— Berkeley &amp; Berkeley, 1952: 89, figs 181‒182.— Levenstein, 1970: 214.— Dauvin &amp; Bellan, 1994: 173.— Hobson, 1976: 139.— Hobson &amp; Banse, 1981: 62, fig. 13j. — Blake, 2000: 163, fig. 7.8.— Imajima, 2009: 139‒140, fig. 45A‒E.— Alalykina, 2020: 193.</p><p>Material examined. Washington State Department of Ecology. Puget Sound, Washington, USA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.35889&amp;materialsCitation.latitude=47.60722" title="Search Plazi for locations around (long -122.35889/lat 47.60722)">Elliott Bay</a>, Sta. 187, 103 m, 47°36’26”N, 122°21’32”W, coll. PSEMP Urban Bays Monitoring, 16 June 2021 (1 specimen) . <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.81473&amp;materialsCitation.latitude=47.678333" title="Search Plazi for locations around (long -122.81473/lat 47.678333)">Hood Canal</a>, north of Seabeck, Sta. 222, 126 m, 47°40’42”N, 122°48’53”W, coll. PSEMP Long-term Monitoring, 01 June 2022 (3 specimens) . <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.47139&amp;materialsCitation.latitude=48.098053" title="Search Plazi for locations around (long -122.47139/lat 48.098053)">Saratoga Passage</a>, Sta. 19, 126 m, 48°5’53”N, 122°28’17”W, coll. PSEMP Long-term Monitoring, 01 May 2023 (1 specimen) .</p><p>Diagnosis. Travisia with annulate branchiae from chaetiger 2, 24‒25 pairs of branchiae and 26‒27 chaetigers. Mid-ventral groove present. Nuchal organs small. Parapodial lappets present along body except for the last body quarter.</p><p>Description. Specimens 23‒75 mm long, 9‒18 mm wide with 26‒27 chaetigers (n: 5). Body pale to cream-colored in alcohol; robust, grub-like with rugose epidermis (Fig. 4A‒B). Prostomium small, with rounded margin, smooth (Figs 4C, 5B). Nuchal organs dorsolateral, small narrow splits (Fig. 5B). Eyes absent. Mouth ventral between chaetigers 1 and 2: upper lip with elongate annulations of chaetiger 1, lower lip with short annulation of chaetiger 2 (Figs 4B, 5A, D). Mid-ventral groove present along body, more visible in anterior and posterior segments (Figs 4B, 5A, D, F).</p><p>Peristomium uniannulate. Chaetiger 1 biannulate. Chaetigers 2 to 13‒14 triannulate. Chaetigers 14‒15 to 17‒ 18 biannulate. Following chaetigers to the end uniannulate. Peristomium triangular shaped dorsally, lateral sides oblique, distal margin rounded, basal margin four times wider than anterior margin, covered with small, spherical papillae (Figs 4C, 5B).</p><p>First chaetiger with dorsal and lateral sides covered by rounded packages of equal size spherical papillae (Figs 4C, 5B). From chaetiger 2 towards posterior end, each segment with spherical papillae around body as follows: chaetigers 2‒16 with spherical papillae dorsally (Fig. 5B), large papillae only on posterior edges of annulations (Fig. 5C), anterior papillae to posterior edges considerably smaller; chaetigers 17‒18 to body end with epithelium fully covered with spherical papillae, increasing gradually in size from anterior margin of each segment to posterior margin (Fig. 5E). Last 11 segments with inferior margins flap-like, with ventral margin longer than dorsal margin (Fig. 5E).</p><p>Branchiae slightly annulate, 24‒25 pairs starting at chaetiger 2 and extending to chaetigers 24‒26. First pair short, increasing gradually in size towards chaetiger 12, decreasing in size gradually to posterior end (Figs 4A‒B, 5A).</p><p>Parapodia biramous. Parapodial lappets present on neuropodia of chaetigers 2‒15 (Figs 4A‒B, D, 5A, D, F). Each neuropodial lappet triangular, composed of aglutinated papillae with largest papillae on top (Figs 4D, 5D). Notopodial lappets absent (Fig. 5C). Chaetigers 15 or 16 with a gradual transition of parapodial lappets rather than abrupt transition to end without parapodial lappets.All noto- and neurochaetae iridescent, spinulose capillaries. Last 4‒5 segments achaetous.</p><p>Interamal sensory pores small, between notochaetae and neurochaetae from chaetiger 1 until near last segments. Lateral neuropodial nephridiopores present along chaetigers 3 to 14.</p><p>Pygidium short, terminal or shifted to side, thick, with 10 lobes (Figs 4A‒B, 5G).</p><p>Methyl green staining. Branchiae and mid-ventral groove unstained. Rest of the body is uniformly stained (Fig. 5).</p><p>Remarks. Travisia pupa was originally described by Moore (1906) from the Gulf of Georgia, the Behm Canal, and Kasaan Bay (Alaska, USA), in sediments from 32‒438 m depth. Unfortunately, Moore (1906) did not include illustrations of body structures or papillation, except for a short, anterior area of a chaeta (Moore 1906: pl. XI, fig. 23). Posteriorly, T. pupa has been reported in Alaska, British Columbia, California, Sea of Okhotsk, Bering Sea, Japan Trench, and Kuril-Kamchatka Trench, in shelf and slope depths (Moore 1923; Uschakov 1950, 1955; Berkeley &amp; Berkeley 1952; Levenstein 1970; Hobson 1976, Hobson &amp; Banse 1981; Blake 2000; Imajima 2009), though in all cases poorly illustrated. Among all these records, that of Imajima (2009: fig. 45A‒E, Honshu/ Japan) includes detailed drawings regarding body papillation.</p><p>Moore (1906) noted that the size of T. pupa was 24‒82 mm long, 8‒32 mm width, 31‒32 segments, stating “... the number is not correlated with the size of the worm, the largest two having thirty-one and the smallest thirty-two”, 22‒25 pairs of branchiae, and interamal sensory pores starting at chaetiger 1. While examining the type material, Hobson (1976) reported the following features: worms 40‒85 mm long; 30‒32 segments: 1 achaetous anterior segment, 25‒27 chaetigerous segments, and 3‒6 achaetous pre-anal segments; and 21‒25 pairs of branchiae. Specimens from Puget Sound examined herein measure 23‒75 mm long, 9‒18 mm width, 31‒32 segments, 26‒27 chaetigerous, 4‒5 achaetous pre-anal segments, and 24‒25 pairs of branchia.</p><p>In her contribution, Hobson (1976) declared T. foetida Hartman, 1969, a species described from Redondo Canyon, California to 256 m depth, a junior synonym of T. pupa, a synonym followed by Blake (2000) in his report of T. pupa to Santa Maria Basin and Western Santa Barbara Channel (California, USA) in 91‒197 m depth; however, T. foetida was still included as a valid name in Blake &amp; Maciolek (2019), Rizzo &amp; Salazar-Vallejo (2021) and Read &amp; Fauchald (2025).</p><p>Fauchald (1972) reported the presence of T. foetida in many localities from western Mexico (off Baja California to Acapulco, and in the southern part of the Gulf of California, from 1985‒3383 m depth), and it was therefore suggested that the distribution of T. pupa reaches western Mexico (Blake 2000; Imajima 2009). Fauchald (1972) reported a uniform number of 27 chaetigers in all specimens he reviewed and identified as T. foetida (47 specimens), and the presence of a dorsally and anteriorly directed peristomial lappet. In his drawing (Fauchald 1972: pl. 49, fig. d) this structure is triangular, as in T. pupa here reported from Puget Sound, but the position and size of nuchal organs are peculiar in Fauchald´s specimens: circular, large (as long as prostomium) and placed on the base of the peristomium versus nuchal organs small, as narrow splits located on the anterior margin of the peristomium in T. pupa . This difference is significant and provides support to reject the synonymy proposed by Hobson (1976). It is desirable, however, to carefully revise the types of T. foetida and T. pupa and Fauchald´s material to prove the validity of T. foetida and its presence in Western Mexico.</p><p>Specimens here identified from Puget Sound as T. pupa have a mid-ventral groove running along the body (also see Burgess 2018b). Travisia pupa was included in the subgroup B2 (species without mid-ventral groove) by Plathong et al. (2023), but the presence of a mid-ventral groove clearly indicates that T. pupa belongs to subgroup B1 (species with mid-ventral groove). However, we believe that the use of this feature as a diagnostic character demands further study. In the case of Travisia amoyanus Yang, Wu, Wang, Zhao, Hwang &amp; Cai, 2022, described from the intertidal-subtidal, in Xiamen, China (Yang et al. 2022), the original description suggests that some specimens lack a midventral groove: “Midventral groove absent, if have, present from last four segments”. In Travisia pupa from Puget Sound the mid-ventral groove is clearly visible in anterior and posterior segments only. This raises the question about the presence or absence of a midventral groove. It might represent a variable feature according to ontogeny or fixation, making this character questionable for its current use in classification or recognition of taxa. A further argument in favor of using this character with caution is provided by Santos (1977), who stated that “The relative development of the midventral groove in T. hobsonae [Santos, 1977] appears to be related to the preparation of the specimens. Those that are incompletely narcotized before fixation show a pronounced groove, whereas the specimens that are properly narcotized exhibit a very slight midventral groove”.</p></div>	https://treatment.plazi.org/id/038D8112A029D676FBE0FA52360121C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tovar-Hernández, María Ana;Burgess, Dany E.;León-González, Jesús Angel De;Hendrickx, Michel E.	Tovar-Hernández, María Ana, Burgess, Dany E., León-González, Jesús Angel De, Hendrickx, Michel E. (2025): Travisiid worms from deep water off Western Mexico with the establishment of a new species (Annelida: Travisiidae). Zootaxa 5729 (2): 349-369, DOI: 10.11646/zootaxa.5729.2.7, URL: https://doi.org/10.11646/zootaxa.5729.2.7
038D8112A025D677FBE0FCFE305424EF.text	038D8112A025D677FBE0FCFE305424EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travisia hernandezalcantarai Tovar-Hernández & Burgess & León-González & Hendrickx 2025	<div><p>Travisia hernandezalcantarai sp. nov.</p><p>Zoobank: urn:lsid:zoobank.org:act: 42EA1216-FE10-4C05-9541-E14E38DC390E</p><p>(Figs 6‒10)</p><p>Type Material. Holotype, ICML-EMU-14079, TALUD XII. Sta. 29, 02 April 2008, 19º19’37”N, 105º26’20”W, off <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.43889&amp;materialsCitation.latitude=19.326946" title="Search Plazi for locations around (long -105.43889/lat 19.326946)">Jalisco</a>, Mexico,BS, 1609‒1643m . Paratypes,ICML-EMU-14080-A, TALUD XI. Sta. 26, 10June2007, 18º33’12”N, 104º32’12”W, off Colima, Mexico, BC, 1918 m, 1 specimen . ICML-EMU-14080-B, TALUD XI. Sta. 27, 11 June 2007, 18º41’00”N, 104º33’07”W, off Colima, Mexico, BC, 1305 m, 1 specimen . ICML-EMU-14080-C, TALUD XI. Sta. 28, 11 June 2007, 18º48’05”N, 104º32’36”W, off Colima, Mexico, BC, 1050 m, 1 specimen . UAA, TALUD XII. Sta. 5, 28 March 2008, 16º58’28”N, 100º55’20”W, off Guerrero, Mexico, BC, 1849 m, 1 specimen mounted in gold for SEM. ICML-EMU-14082-A, TALUD XII. Sta. 10, 28 March 2008, 17º11’03”N, 101º28’05”W, off Guerrero, Mexico, BC, 1290 m, 1 specimen . ICML-EMU-14082-B, TALUD XII. Sta. 14, 30 March 2008, 17º36’20”N, 102º01’59”W, off Guerrero, Mexico, BC, 1245 m, 1 specimen . ICML-EMU-14083-A, TALUD XII. Sta. 20, 31 March 2008, 17º54’20”N, 103º10’45”W, off Guerrero, Mexico, BC, 1992 m, 1 specimen . ICML-EMU-14083-B, TALUD XII. Sta. 27, 02 April 2008, 18º40’28”N, 104º35’51”W, off Colima, Mexico, BC, 1158 m, 1 specimen . UANL-8282, TALUD XII. Sta. 28, 02 April 2008, 18º50’19”N, 104º34’14”W, off Colima, Mexico, BC, 1080 m, 1 specimen .</p><p>Diagnosis. Travisia with annulate branchiae from chaetiger 2, 22‒25 pairs of branchiae and 26‒28 chaetigers. Mid-ventral groove present. Nuchal organs remarkably large. Parapodial lappets absent.</p><p>Description. Holotype 32 mm long (20‒39 mm long in paratypes), 8 mm width (7‒9 mm wide in paratypes) with 26 chaetigers (27‒28 chaetigers in paratypes). Body yellowish to cream-colored in alcohol; robust, grub-like (Fig. 8A‒B) with rugose epidermis and remarkable, transverse rows of spherical papillae (Fig. 6A‒B). Prostomium small, conical, pointed, smooth (Figs 6A, C‒D, F, 8A‒D, 9A‒B). Nuchal organs dorsolateral, teardrop-shaped splits, remarkably large (Figs 6A, C‒D, F, 8C, 9B). Eyes absent. Mouth ventral between chaetigers 1 and 2, upper lip with short elongate annulations of chaetiger 1, lower lip with short annulation of chaetiger 2 (Fig. 8A‒D). Pharynx not everted in all specimens.</p><p>Peristomium uniannulated. Chaetiger 1 biannulated (Figs 8C‒D, 9A‒B). Chaetigers 2 to 13‒14 triannulated (Figs 8A‒B, D, 9A). Chaetigers 15‒17 biannulated (Fig. 8A‒B). Chaetigers 18 to the end uniannulated (Fig. 8A‒B).</p><p>Peristomium covered with spherical papillae (Fig. 8C); dorsal side with distal part squared (lateral and distal sides forming right angles), narrower than prostomium width (Fig. 6C‒D, F). Basal part with rounded lateral margins, three times broader than distal part (Fig. 6C‒D, F). First, second and third chaetigers with lateral sides covered by sub-triangular packages of equal-sized spherical papillae (Figs 6A, C‒D, F, 8C, 9A‒B), dorsal side with posterior margin of packages festooned (Fig 6A, C‒D, F). From chaetiger 2 towards posterior end, each segment with rows of spherical papillae around body as follows: A, anterior chaetigers (chaetigers 2‒13) with three rows of spherical papillae (Figs 7E, 8A); B, median chaetigers (chaetigers 14 to 17) with two rows of spherical papillae: median row with large papillae, external row with small papillae (Fig. 7F). C, Posterior chaetigers (from chaetiger 18 to the end) without rows of papillae, all integument covered in full by iminute papillae (Fig. 9F), increasing gradually in size from the anterior margin of each segment to posterior margin (Fig. 7A, I). Posterior margin of each segment flap-like (festooned), with ventral sides longer than dorsal sides (Fig. 7I). Papillae located basally in lateral borders of each segment twice as long as internal papillae, spherical to oval-shaped (Figs 7I, 9A, D‒E). In last 10 segments, lateral-most papillae from within basal margin spherical, +3 times the size of internal papillae (Fig. 7J).</p><p>Interamal sensory pores C-shaped, large (almost as wide as branchial basis) (Fig. 9E), located between notochaetae and neurochaetae from chaetiger 1 until near last segments. Lateral neuropodial nephridiopores not seen under light or SEM microscopy.</p><p>Branchiae annulate (Figs 9C‒E, 10C, E), 24 pairs in holotype (22‒25 pairs in paratypes) starting at chaetiger 2 (Fig. 9A‒B) and extending to chaetiger 25 (chaetiger 22 in six paratypes, chaetiger 24 in one paratype). First pair short, increasing gradually in size towards chaetiger 10 (Figs 7G‒J, 8B), decreasing in size gradually to posterior end (Figs 7J, 8B).</p><p>Parapodia biramous (Figs 9D‒E, 10C, E). All noto- and neurochaetae translucent, spinulose capillaries (Figs 9E, 10D, F). Last 2‒4 segments achaetous (Fig. 9F). Noto- and neuropodial lappets absent (Figs 7G‒H, 9D, F). A shallow, mid-ventral groove present only on last 10‒12 abdominal segments (Fig. 6E).</p><p>Pygidium short, terminal (not shifted to side), thick; anus ventral with 12 lobes in a circle, without anal cirri (Figs 6E, 7A‒D, 8A‒B, 9F).</p><p>Etymology. The species is named after Dr. Pablo Hernández-Alcántara, a Mexican polychaete systematist and ecologist, and a long-time friend and colleague. The species is named in recognition of his contribution to the study on spatial and bathymetric trends of polychaetes in Western Mexico. The species-group name is a noun in the genitive case (ICZN, 1999, Art. 31.1.2).</p><p>Remarks. Travisia hernandezalcantarai sp. nov., is unique among the species currently recognized in the genus by having nuchal organs remarkably large, as long as the tip of the prostomium.</p><p>Travisia hernandezalcantarai sp. nov., and T. pupa can be distinguished based on the following features: 1, Travisia hernandezalcantarai sp. nov., does not have parapodial lappets whereas they are present in T. pupa; 2, nuchal organs large, teardrop-shaped in T. hernandezalcantarai sp. nov., versus discrete, narrow splits in T. pupa; 3, peristomium with an inverted T-shaped area dorsally in T. hernandezalcantarai sp. nov., versus a triangular-shaped area in T. pupa; 4, lateralmost papillae on the posterior margin of each terminal segment being up to 3 times larger than internal papillae in T. hernandezalcantarai sp. nov., versus all papillae from the posterior margins less than 2 times larger than internal papillae in T. pupa; and 5, a mid-ventral groove restricted to posterior segments of the body in T. hernandezalcantarai sp. nov., whereas it is present along the entire body in T. pupa .</p><p>Based on the presence of a ventral groove and as stated in the remarks section for T. pupa, T. hernandezalcantarai sp. nov., and T. pupa should be added to the group B1 proposed by Plathong et al. (2023). Group B1 is composed by T. fusiformis Kudenov, 1975 (intertidal, from the Gulf of California, Mexico), T. hobsonae Santos, 1977 (10‒110 m depth, Tampa Bay, Florida, USA), T. thailandensis Plathong, Plathong &amp; Salazar-Vallejo, 2023 (50‒80 m, Gulf of Thailand), T. pupa and T. hernandezalcantarai sp. nov., Travisia hobsonae is characterized by having branchiae starting at chaetiger 3 and prostomium with pustules (Santos 1977), whereas in T. fusiformis, T. thailandensis, T. pupa and T. hernandezalcantarai sp. nov., branchiae appear in chaetiger 2 and the prostomium lacks pustules. In T. fusiformis, T. thailandensis and T. pupa the mid-ventral groove is present along body whereas the mid-ventral groove extents only to posterior segments in T. hernandezalcantarai sp. nov., Travisia fusiformis has 34‒35 chaetigers, T. thailandensis 26‒28 chaetigers and T. hernandezalcantarai sp. nov., 26‒28 chaetigers. Travisia fusiformis, T. thailandensis, and T. pupa have parapodial lappets. In T. fusiformis, neuropodial lappets are small, conspicuous and well-developed from chaetiger 17 to the end of the body; in T. thailandensis, parapodial lappets from the anterior region are small and neuropodial lappets become progressively larger in posterior region; in T. pupa, lappets are well-developed in three quarters of the body segments (anterior); T. hernandezalcantarai sp. nov., lacks lappets altogether.</p><p>Abiotic conditions. The specimens of Travisia hernandezalcantarai sp. nov., were collected in sediments from 1050–1992 m deep, under the following environmental conditions: temperature, 2.3–5.2°C; salinity, 34.53–34.63; dissolved oxygen, 0.03–1.49 ml O 2 /l; %CO, 0.91–4.64; sediments composition variable, including mixed, sandy mud, muddy sand, and mud (Table 1).</p></div>	https://treatment.plazi.org/id/038D8112A025D677FBE0FCFE305424EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tovar-Hernández, María Ana;Burgess, Dany E.;León-González, Jesús Angel De;Hendrickx, Michel E.	Tovar-Hernández, María Ana, Burgess, Dany E., León-González, Jesús Angel De, Hendrickx, Michel E. (2025): Travisiid worms from deep water off Western Mexico with the establishment of a new species (Annelida: Travisiidae). Zootaxa 5729 (2): 349-369, DOI: 10.11646/zootaxa.5729.2.7, URL: https://doi.org/10.11646/zootaxa.5729.2.7
038D8112A03FD66DFBE0F9DA378A2248.text	038D8112A03FD66DFBE0F9DA378A2248.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Travisia Johnston 1840	<div><p>Key to species of Travisia Johnston, 1840 from the North American Pacific</p><p>1 Branchia branched; branchiae present from chaetiger 3........................ T. filamentosa de León-González, 1998</p><p>- Branchia cirriform.................................................................................... 2</p><p>2 Posterior end tapered; body with more than 19 chaetigers..................................................... 3</p><p>- Posterior end with a short cylindrical cone, deeply furrowed dorsally; 18 chaetigers................................................................................................... T. oregonensis Fauchald &amp; Hancock, 1981</p><p>3 Pygidium with cirri; parapodial lappets well-developed, starting at segment 15.................. T. gigas Hartman, 1938</p><p>- Pygidium without cirri; parapodial lappets absent or present in other arrangement.................................. 4</p><p>4 Nuchal organs large, teardrop-shaped; parapodial lappets absent.................. Travisia hernandezalcantarai sp. nov.</p><p>- Nuchal organs small, discrete, as narrow splits or C-shaped splits............................................... 5</p><p>5 Prostomium blunt or truncate, not conical................................................................. 6</p><p>- Prostomium pointed, conical............................................................................ 7</p><p>6 Branchiae present from chaetiger 2; 31 segments (27 chaetigers); epithelium with entire surface pustulated, not strongly granulated....................................................................... T. foetida Hartman, 1969</p><p>- Branchiae present from chaetiger 3; 32 to 50 segments (27‒45 chaetigers); epithelium with large size pustules, crowded over most parts of the body, giving an aspect of close granulation............................... T. granulata Moore, 1923</p><p>7 Body fusiform; epithelium fully covered with papillae, papillae not forming rows............ T. fusiformis Kudenov, 1975</p><p>- Body robust, grub-like; epithelium with well-defined rows or ridges of papillae at least in some body segments.......... 8</p><p>8 Mid-ventral groove present; parapodial lappets well-developed, present in anterior segments......... T. pupa Moore, 1906</p><p>- Mid-ventral groove absent; parapodial lappets well-developed in posterior segments.............. T. brevis Moore, 1923</p></div>	https://treatment.plazi.org/id/038D8112A03FD66DFBE0F9DA378A2248	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tovar-Hernández, María Ana;Burgess, Dany E.;León-González, Jesús Angel De;Hendrickx, Michel E.	Tovar-Hernández, María Ana, Burgess, Dany E., León-González, Jesús Angel De, Hendrickx, Michel E. (2025): Travisiid worms from deep water off Western Mexico with the establishment of a new species (Annelida: Travisiidae). Zootaxa 5729 (2): 349-369, DOI: 10.11646/zootaxa.5729.2.7, URL: https://doi.org/10.11646/zootaxa.5729.2.7
