taxonID	type	description	language	source
038F870AFF9CEA4308FCFDAE24DBFBC8.taxon	materials_examined	Material examined. Two adult specimens were examined by light microscopy and measured with an ocular micrometer. Holotype specimen collected from Tobago Island in 2012 deposited in Smithsonian National Museum of Natural History under accession number USNM 1760563. Paratype specimen collected from Tobago Island in 2012 deposited in Smithsonian National Museum of Natural History under accession number USNM 1760564. Both specimens were mounted alive. Holotype was photographed and video recorded; paratype was only video recorded. Type locality: Tobago: Highway to Heaven dive spot, 31 m depth: 11 º 20 ’ 154 ” N, 60 º 38 ’ 539 ” W. Temperature: 27 º C. Grain Size: Mean 0.74 mm ± 0.63 SD; Median 0.58 mm.	en	Araújo, Thiago Quintão, Hochberg, Rick (2025): First record and a new species of Dendrodasys (Gastrotricha: Macrodasyida: Dactylopodolidae) from Tobago Island. Zootaxa 5696 (3): 398-408, DOI: 10.11646/zootaxa.5696.3.5, URL: https://doi.org/10.11646/zootaxa.5696.3.5
038F870AFF9CEA4308FCFDAE24DBFBC8.taxon	etymology	Etymology. The species epithet is named after the dive location on the island of Tobago: Highway to Heaven, located in the northwest corner of the island.	en	Araújo, Thiago Quintão, Hochberg, Rick (2025): First record and a new species of Dendrodasys (Gastrotricha: Macrodasyida: Dactylopodolidae) from Tobago Island. Zootaxa 5696 (3): 398-408, DOI: 10.11646/zootaxa.5696.3.5, URL: https://doi.org/10.11646/zootaxa.5696.3.5
038F870AFF9CEA4308FCFDAE24DBFBC8.taxon	diagnosis	Diagnosis Specimens with body lengths of 240 μm – 250 μm long (= 245 μm, n = 2). Maximum body width at head (U 5), neck (U 16), U 50 and caudal end of body (U 65) is 32.1 / 24 / 33 / 22 μm. Pharynx to 50 μm long (U 20). Triangular head with a blunt terminal mouth and prominent conical cephalic lobes on each side of head. A pair of very small presumed sensorial tubes were present dorsally at U 04. A pair of horn-like (triangular) organs medial to cephalic lobes at U 05, and a pair of knob-like pestle organs with narrow basis inserted posterior to the cephalic lobes. One pair of TbA arising from a common base per side; TbD, TbV, TbL and TbVL absent. TbP three per side; one tube (per side) at the anterior constriction of the caudal pedicle and one pair at the tip of each pedicle. Eleven pairs of epidermal glands along the body and a single epidermal gland at the base of the bifurcation of the caudal pedicle. Single large mature egg above intestine. Frontal organ with ventral opening present at U 51. Putative caudal organ present at U 62.	en	Araújo, Thiago Quintão, Hochberg, Rick (2025): First record and a new species of Dendrodasys (Gastrotricha: Macrodasyida: Dactylopodolidae) from Tobago Island. Zootaxa 5696 (3): 398-408, DOI: 10.11646/zootaxa.5696.3.5, URL: https://doi.org/10.11646/zootaxa.5696.3.5
038F870AFF9CEA4308FCFDAE24DBFBC8.taxon	description	Description The type description is mainly based on an adult specimen of 250 μm total body length (Figs. 2, 3 A). Slightly subterminal mouth positioned in the front of a triangular head. One pair of small tubes (3 μm long), presumed to be sensorial, on the dorsal side of the head at U 02. A pair of horn-like (triangular / conical) organs (6 μm) are present at U 05 and dorsally to the cephalic lobes (Figs. 2 A, C, 3 C, D). Knob-like pestle organs with narrow base at U 08 (Figs 2; 3 A) inserted at lateral body surface and posterior to the cephalic lobes. Trunk widened due to the presence of a mature egg (width range: 19 – 36 μm) and narrowed at U 65 where it extended as a caudal pedicle (70 μm long). The caudal pedicle bifurcated at U 88 (Figs. 2, 3 A). Cuticle. The cuticle was smooth across the body and without notable folds or ridges. Epidermal glands. Round to elliptical epidermal glands of 6 – 7 μm diameter. A total of eleven glands were present on each side of the body as follows: a refractile pair at U 05; an opaque pair at U 14; an adjacent pair of refractile / opaque glands at U 22 / U 24, respectively; an adjacent pair of refractile / opaque glands at U 28 / U 30, respectively; an adjacent pair of refractile glands at U 42 / U 44; and three refractile glands at U 54, U 59, and U 64; a single refractile gland present just anterior to the bifurcation of the caudal pedicle at U 89 (Figs. 2 A; 3 A, C). Adhesive tubes. A single pair of TbA per side arises from a common base at U 09; each tube as ca. 11 μm long and completely separated from the other on distal portion (Figs. 2 B, 3 B). Three TbP per side. A single tube to 14 um long was present at the constriction of the caudal pedicle at U 75; a thick fiber extended from each tube into the body. The remaining two tubes were positioned at the tip of each pedicle: both tubes 5 μm long (Figs. 2, 3 A). Cilia. Numerous sensory cilia 10 – 18 μm long projected from the anterior margin of the head and lined the cephalic lobes; cilia ventrally curved over the head margin and were active. The dorsal body wall was also adorned with mobile cilia: some cilia formed a transverse line at U 07 (Fig. 2 A); other cilia were present between the pharynx and lateral body wall (Figs. 2; 3 A – C). Many sensory cilia were adjacent to each other and appeared as pairs (10 – 15 um long) (Fig. 2; 3 A, C). Two columns of ventral locomotory cilia extended from U 03 to U 70. The columns merged as transversal band at U 18. The columns combined into a single field under the caudal pedicle. Digestive tract. The narrow mouth (4 μm wide) was subterminal with a trapezoid-shaped cuticular ring at the tip of the head. The buccal cavity expanded slightly as it continues into the pharynx, which was straight and 50 μm long by 10 μm wide. A pair of dorsolateral pharyngeal pores at U 17 extended inward as cuticle-lined tubes to the pharynx just anterior of the pharyngo-intestinal junction (PhIJ) at U 20. The intestine was 9 – 12 μm wide and contained small refractile contents along its length; it narrowed to 5 μm wide as it approached the ventral anus at U 69. Muscles. All somatic muscles appeared cross-striated (Figs. 2; 3 A). Reproductive anatomy. A single large egg was present dorsal to the intestine; the nucleus was 9 μm in diameter (Figs. 2 A, B; 4 A). Ovary not observed. A frontal organ, somewhat ellipsoidal in shape, was present at U 51 close to the left side of the body; allosperm were present inside the organ. An apparent opening of the frontal organ was observed on the ventral body wall where it was surrounded by four raised bumps (Fig. 2; 4 A – D). No testes were observed but a potential caudal organ (10 μm long) was present on the right side of the body at U 62. The putative caudal organ was pyriform in shape had a potential ventral opening adjacent to the border of the intestine (Figs. 4 F – G).	en	Araújo, Thiago Quintão, Hochberg, Rick (2025): First record and a new species of Dendrodasys (Gastrotricha: Macrodasyida: Dactylopodolidae) from Tobago Island. Zootaxa 5696 (3): 398-408, DOI: 10.11646/zootaxa.5696.3.5, URL: https://doi.org/10.11646/zootaxa.5696.3.5
