identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038F87AEFFBFFFFBFCFAF9A9A4D55DD5.text	038F87AEFFBFFFFBFCFAF9A9A4D55DD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus Simon 1897	<div><p>TITIOTUS SIMON</p> <p>Titiotus Simon, 1897: 113 (type species by original designation Titiotus californicus Simon).</p> <p>DIAGNOSIS: The unusual character combination of three claws plus claw tufts readily separates members of Titiotus and their closest relatives from most other North American spiders. Males of Titiotus differ from those of the other three closely related genera (Liocranoides Keyserling, Anachemmis Chamberlin, and Socalchemmis Platnick and Ubick) by having a tibial apophysis consisting of three or four prongs (rather than just one or two, figs. 9, 14); females differ by having a wide median septum that usually occupies more than two-thirds of the epigynal width (figs. 10, 15).</p> <p>DESCRIPTION: Medium to large spiders, total length of males 8–17, of females 11–21. Carapace oval, widest at rear of coxae II, abruptly narrowed at level of palpi to less than two-thirds of maximum width; thoracic groove long, longitudinal, very deep; surface coated with short recumbent and fewer, longer, erect dark setae, erect setae most numerous along midline and in ocular area; eight eyes in two rows; from above, both eye rows slightly recurved; from front, anterior row recurved, posterior row slightly procurved; anterior median eyes round, smallest; other eyes oval, subequal, with canoe-shaped tapeta; anterior median eyes separated by roughly their diameter, slightly closer to anterior laterals; posterior medians separated by roughly their diameter, much farther from posterior laterals; lateral eyes of each side separated by almost their diameter; median ocular quadrangle wider in back than in front, wider in back than long; clypeal height about twice diameter of anterior median eyes, corners of clypeus with incised margins that overlie cheliceral boss; chilum weakly sclerotized, divided, composed of two triangular sclerites. Chelicerae vertical, anterior surface with few, erect setae; promargin with three teeth situated at proximal end of fang furrow, most proximal tooth smallest, middle tooth largest, retromargin with three larger, more distally situated teeth; very short, narrow, Ishaped posterior sclerite present, separating chelicerae at base. Labium short, only about half as long as endites, distally invaginated at middle, reflexed at almost 90 ° angle relative to sternum. Endites rectangular, distally slightly convergent, with anteromedian scopula and anterolateral serrula consisting of single row of teeth. Sternum rounded, without extensions to coxae, with only slight angular projections between coxae, with erect setae; posterior margin only slightly extended between coxae IV.</p> <p>Leg formula 4123. Typical leg-spination pattern (only surfaces bearing spines listed): femora: I d1-1-1, p0-2-1, r1-1-1; II d1-1-1, p2- 2-1, r0-1-2; III d1-1-1, p2-1-1, r1-2-1; IV d1-1- 1, p1-1-1, r0-0-2; patellae III, IV p0-1-0, r0-1- 0; tibiae: I, II p1-1-0, v4-6-4; III d1-0-1, p0-1-1, v2-2-2, r1-1-1; IV d1-0-1, p0-1-1, v2-2-2, r1-0- 1; metatarsi: I v2-2-2; II p1-0-0, v2-2-2, r1-0-0; III p1-1-2, v2-2-1r, r1-1-2; IV p1-1-2, v2-2-1r, r1-2-2; tarsi with three claws and claw tufts, superior claws with several weak teeth, most distal teeth largest, inferior claws unarmed; all tarsi with strong ventral scopulae, scopular hairs distinct from those of claw tufts; distal segments apparently with trichobothria in two rows; tarsal organ not scanned; all trochanters strongly notched; males without tibial crack; metatarsi without preening combs.</p> <p>Abdomen without anterior or dorsal scutum; anterior lateral spinnerets large, composed of two articles, distal article with two major ampullate gland spigots and about 25 piriform gland spigots (figs. 1, 4); posterior median spinnerets composed of one article, small, triangular, those of female expanded posteriorly, where they bear single large and two smaller cylindrical gland spigots, preceded anteriorly by several smaller aciniform gland spigots and terminally by two minor ampullate gland spigots (figs. 2, 5); posterior lateral spinnerets composed of two articles, distal article about one-fourth as long as proximal article, with numerous long aciniform gland spigots and (in female) three larger cylindrical gland spigots (figs. 3, 6); colulus represented by setae on small lobe.</p> <p>Male palp with patella not widened, tibial apophysis complex, consisting of three or four prongs, of which two or three most retrolateral often share common base (fig. 9); subtegulum and tegulum with interlocking lobes, median apophysis heavily sclerotized at least distally, bearing multiple processes; embolus short, arched, accompanied by hyaline conductor (figs. 7, 8). Female palp with extremely long, dentate claw. Epigynum with wide median septum, usually occupying most of epigynal width (fig. 10); spermathecal bulbs situated posterolaterally.</p> <p>SPECIES GROUPS: Two groups are recognized; the californicus group, containing most of the species, is characterized by having the most ventral prong of the tibial apophysis relatively long and narrow, and situated close to the base of the cymbium. Members of the humboldt group, containing only the new species T. humboldt, T. marin, and T. costa, have the most ventral prong of the tibial apophysis relatively short and wide, and situated close to base of the retrolateral prongs.</p> </div>	https://treatment.plazi.org/id/038F87AEFFBFFFFBFCFAF9A9A4D55DD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFBDFFF1FF79FD76A54D5AAC.text	038F87AEFFBDFFF1FF79FD76A54D5AAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus californicus Simon	<div><p>Titiotus californicus Simon</p> <p>Figures 1–11; map 1</p> <p>Titiotus californicus Simon, 1897: 113, fig. 97 (female holotype from California, no specific locality, in the Muséum National d’Histoire Naturelle, Paris, examined via sketches and detailed description by the late W. Peck).</p> <p>NOTE: Simon did not specify the source of the female holotype, but it may have been provided by George Marx. The Marx collection in the USNM includes two females of this species, labelled with the nomen nudum Cybaeus californicus. Unfortunately, even if this was the source of Simon’s specimen, the type locality remains unspecified, as the USNM females are labelled only as ‘‘occidental Cal.’’.</p> <p>DIAGNOSIS: Males resemble those of the more northern species T. hansii but have a more pronounced dorsal extension on the middle prong of the tibial apophysis (fig. 9); females also resemble those of T. hansii but have a wider epigynal septum that is more narrowed anteriorly (fig. 10).</p> <p>MALE: Total length 10. Carapace light brown, with slightly darker triangular markings radiating from thoracic groove to intercoxal areas; abdomen pale gray; femora light brown, more distal leg segments grading to reddish brown on metatarsi and tarsi. Leg spination: femora: I p0-3-1, r1-2-1; II r1-2-1; III r1-1-2; IV p1-2-1, r0-1-2; tibiae: I, II d1-0- 1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1- 1-0, r1-1-0; III p1-2-2, v2-2-1p, r1-1-1; IV p1- 2-2, v2-2- 1p. Embolus expanded prolaterally, arched distally; median apophysis with subdistal projection; middle prong of tibial apophysis bifid, with enlarged, distally expanded dorsal branch (figs. 7–9).</p> <p>FEMALE: Total length 12. Coloration as in male. Leg spination: femora II p1-2-1; tibiae I, II r1-1-0; metatarsi: I r1-0-0; III p1-2-2, r1-2-2. Epigynal septum wide, occupying most of epigynal width; posterior ducts transverse (figs. 10, 11).</p> <p>MATERIAL EXAMINED: United States: CAL- IFORNIA: Alameda Co.: no specific locality, May (AMNH), 13; Berkeley, May 22, 1945 (R. Smith, AMNH), 1♀, Aug. 1945 (R. Smith, AMNH), 13, July 7, 1969, in basement (J. Smith, CAS), 1♀, May 1976 (UCB), 13, Apr. 15, 1978 (N. Lavrov, CAS), 1♀, July 18, 2001, under pillow in bed (H. Netiv, UCR), 1♀; Berkeley Hills, May 5, 1978, in house (R. McGinley, UCB), 13; Camp Ohlone, Sunol Regional Wilderness, June 30, 1979 (S. Diers, MET), 13; Castro Valley, July 21, 2002, in bathroom (M. Curley, UCR), 13; Oakland, Aug. 22, 1965 (A. Wong, CAS), 1♀, May 4, 1993, in house (L. Alexander, CAS), 13; Pleasant Ridge Regional Park Headquarters, June 2, 1990 (M. Thompson, MET), 13; Strawberry Canyon, Berkeley, May 1981 (UCB), 13; Sunol, July 25, 1979 (A. Crawford, MET), 1♀; Sunol Regional Wilderness, July 21, 1989 (M. Thompson, MET), 13. Amador Co.: Amador Canyon, off Pardee Reservoir, July 7, 1958 (W. Gertsch, V. Roth, AMNH), 43, 9♀; canyon on Highway 49, 2– 3 mi S Jackson, July 7, 1958 (W. Gertsch, V. Roth, AMNH), 1♀; Mokelumne River, S Jackson, July 7, 1958 (W. Gertsch, V. Roth, AMNH), 23, 1♀. Calaveras Co.: Cave City Cave, Feb. 7, 1954, on floor (R. de Saussure, AMNH), 13, June 13, 1960, on open rock (R. Graham, AMNH), 13, 1♀; Cave of the Catacombs, Sept. 1, 1961 (R. Graham, AMNH), 1♀; Hanging Gardens Cave, Sept. 4, 1961 (M. Ivie, R. Graham, AMNH), 1♀; King Tut Cave, Dec. 22, 1990 (D. Cowan, CDU), 1♀, Aug. 24, 1991, elev. 1900 ft (T. Briggs, D. Cowan, G. Malliet, D. Ubick, CDU), 1♀; Mountain Ranch, June 16, 2004 (J. Pargett, UCR), 13; 1 mi NW Murphys, June 11, 1994, under limestone rock (D. Ubick, CDU), 13; Stanislaus River, 5 mi S Dorrington, Aug. 5, 1953 (W., J. Gertsch, AMNH), 3♀; Wilseyville, May 29, 2001 (K. Collins, UCR), 13. Contra Costa Co.: Antioch, May 15, 1956, on sand at night (M. Wasbauer, AMNH), 13, 1♀; Briones Reservoir, July 12– 26, 1980 (J. Fraser, CAS), 33, 2♀; Byron, June 7–8, 2004, in sandblasting building (J. Miller, UCR), 23; Clayton, Apr. 2, 1941 (W. Pearce, UCR), 13, 1♀; El Cerrito, June 1955, in house (R. Black, CAS), 1♀; Kennedy Grove, May 2, 1980, in house (CAS), 1♀; Martinez, June 2001, in house (B. Crumrine, CDU), 13; Shell Ridge, Walnut Creek (J. Allen, CAS), 13. El Dorado Co.: Folsom Lake Campground, 9 mi SW Pilot Hill, May 22, 1993, in restroom at night (D. Ubick, CDU), 23, 1♀. Monterey Co.: Arroyo Seco, Indians Road, 36 ° 13.7 9 N, 121 ° 29.5 9 W, May 19, 2002, under rocks, mixed woods (D., S. Ubick, CDU), 1♀ (spinnerets scanned), May 19–Oct. 13, 2002, pitfalls, oak chaparral, elev. 1,100 ft (D., S. Ubick, CDU), 23, 1♀, July 8, 1995, at night (D. Ubick, S. Fend, W. Savary, CDU), 1♀; Carmel Valley, May 1996 (P. Stadille, CDU), 23; Hastings Natural History State Reserve, 36 ° 22.8 9 N, 121 ° 33.2 9 W, June 18, 1943 (J. Linsdale, CAS), 13, July 7, 1944 (J. Linsdale, CAS), 13, July 8, 1946 (J. Linsdale, JCC), 13, July 10, 1946 (J. Linsdale, AMNH), 13, June 1947 (J. Linsdale, AMNH), 13, June 22, 1948 (J. Linsdale, CAS), 13, June 24, 1948 (J. Linsdale, AMNH), 13, July 31, 1948 (J. Linsdale, AMNH), 23, Aug. 3, 1948, shower room (J. Linsdale, AMNH), 13, July 5,</p> <p>Map 1. Central California, showing distributions of five species of Titiotus.</p> <p>1950, in school at night (J. Linsdale, AMNH), 13, July 1–6, 1955, in house (J. Linsdale, AMNH), 23. Placer Co.: Ruck-A-Chucky Campground, middle fork, American River, May 22, 1993, litter, rocks, mixed forest, elev.</p> <p>800 ft (D. Ubick, CDU), 13. San Francisco Co.: San Francisco, June 1985, in house (N. Ljubic, CDU), 1♀; Yerba Buena Island, Feb. 9, 2006, in house (P. Griffes, CAS), 1♀. San Benito Co.: Hollister, June 18, 1982 (C. Porter,</p> <p>CAS), 13. San Mateo Co.: Atherton, Mar. 2, 1963 (D., R. Dailey, CDU), 13 (spinnerets scanned); Edgewood Park, Dec. 27, 1987, under serpentine rock (D. Ubick, S. Fend, CDU), 1♀; Indian Mortar, Jasper Ridge, Dec. 30, 1988, serpentine grassland (D. Ubick, CDU), 1♀; Pacifica, June 29–30, 1991, in house (D. Pledger, CDU), 13; 0.5 mi W San Bruno, May–Sept. 1962, pitfall (H. Stark, AMNH), 33; Valerga Drive, Belmont, June 26, 2006 (N. Teodoro, CAS), 13. Santa Clara Co.: Alum Rock Park, San Jose, Oct. 23, 1970 (E. Schlinger, CAS), 1♀; Highway 9 near Skyline Boulevard, near Castle Crags State Park, Sept. 1997, in house (E. Spitz-Blum, CAS), 1♀; Los Gatos, Nov. 1979, in house (W. Ferguson, CDU), 1♀; San Jose, Sept. 4, 2002, in house (N. Pepper, UCR), 1♀; Saratoga, July 4, 1981 (D. Ubick, CDU), 13, 1♀; Stanford University environs, Palo Alto, Dec. 25, 1920 (J. Chamberlin, AMNH), 23; vic. Stanford, 1927–1928 (C. Duncan, AMNH), 13. Santa Cruz Co.: Aptos, May 1994, biting human in school (S. Tjosvold, CAS), 13. Stanislaus Co.: Frank Raines State Park, Apr. 17, 1975, on brick wall at night (L. Vincent, UCB), 13.</p> <p>DISTRIBUTION: Widespread in central California (map 1).</p> <p>Titiotus hansii (Schenkel), new combination</p> <p>Figures 12–16; map 1</p> <p>Eusparassus hansii Schenkel, 1950: 70, fig. 24 (female holotype from Monte Rio, Russian River, Sonoma County, California, June 6, 1939, H. Schenkel- Rudin, in Basel, examined via photographs by A. Hänggi).</p> <p>DIAGNOSIS: Males resemble those of the more southern species T. californicus but have a less pronounced dorsal extension on the middle prong of the tibial apophysis (fig. 14); females also resemble those of T. californicus but have a narrower epigynal septum that is wider anteriorly (fig. 15).</p> <p>MALE: Total length 9. Coloration as in T. californicus. Leg spination: femora: I r2-1-1; II p2-1-1, r2-1-1; III r1-1-2; IV p2-1-1; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I p0- 1-0, r0-1-0; II p1-1-0, r1-1-0; III p1-2-2. Embolus not expanded prolaterally, arched, narrowed distally; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis bifid, with dorsal branch smaller than ventral branch (figs. 12–14).</p> <p>FEMALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I r1-2-1; II r1-1-2; IV p1-2-1; tibiae: I p0-0-0, v4-4-4; II p0-1-0, v4-4-4, r0-1-0; III p1-0-1, r1-0-1; metatarsi: II p0-0-0; III p1-2-2, r1-2-2; IV p1-2-2, r1-1-2. Epigynal septum relatively narrow, with almost straight sides; posterior ducts oblique (figs. 15, 16).</p> <p>MATERIAL EXAMINED: United States: CALIFORNIA: Butte Co.: French Creek, 5 mi N, 2.5 mi E Madrone Lake, Plumas National Forest, July 22, 1970 (S. Frommer, UCR), 1♀. Humboldt Co.: Hoopa Valley Indian Reservation, Tish-Tang-a-Tang Campground, 41 ° 01.4 9 N, 123 ° 38.34 9 W, May 28, 2005, leaf litter, rocks, logs, oak forest with madrone and conifer (D. Ubick, CDU), 2♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.7525&amp;materialsCitation.latitude=40.185833" title="Search Plazi for locations around (long -123.7525/lat 40.185833)">Phillipsville</a>, 40 ° 11 9 N, 123 ° 45 9 W, Sept. 2, 1963 (W., J. Ivie, AMNH), 1♀. Mendocino Co.: Anchor Bay, June 28, 1952 (W. Gertsch, AMNH), 2♀; ‘‘ Deerwood’ ’ (Leach family property), July 20, 1923 (E. Leach, CAS), 1♀; Hartsook Inn, Piercy, Sept. 23, 1963 (W. Gertsch, AMNH), 13, 1♀; Longvale, June 30, 1952 (W. Gertsch, AMNH), 103, 36♀; NCCRP, 3 mi N Branscomb, May 21–23, 1982, elev. 1400 ft (C. Griswold, CAS), 23, 4♀; Noyo River, 14.5 air mi E Fort Bragg, 39 ° 25.5 9 N, 123 ° 32 9 W, May 25, 1996 (matured Feb. 1997), redwood forest at night (D. Ubick, CDU), 1♀, May 25–26, 1996 (C. Griswold, CAS), 2♀, redwood forest (D. Ubick, CDU), 3♀, May 24–25, 1997 (C. Griswold, CAS), 13, 1♀; Piercy, S fork of Eel River, July 23, 1953 (W., J. Gertsch, AMNH), 5♀; 2 mi N Piercy, Aug. 19, 1959 (W. Gertsch, V. Roth, AMNH), 4♀; 5 mi N Piercy, Apr. 6, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 53, 13♀; Willits, no date (A. Dahl, UCR), 1♀, Aug. 10, 2000, in home, elev. 3000 ft, redwood forest area (R. Dixon, UCR), 1♀, early Oct. 2000 (R. Dixon, UCR), 13; Willits, Mendocino County Museum, Aug. 2001, sticky trap (E. Hamby, UCR), 1♀. Nevada Co.: Nevada City, Dec. 22, 1993, in house (T. Van Wagner, CDU), 13; White Cloud Campground, May 31, 1964 (P. Arnaud, Jr., CAS), 13. Plumas Co.: Juniper Cave [5 Kloppenberg Cave], Oct. 17, 1959, on small landing at depth of 25 ft (R. Graham, AMNH), 1♀; Kloppenberg Cave, 7 air mi SE Quincy, Sept. 3, 1988, dark zone (D. Ubick, CDU), 1♀; 5 mi W Quincy, Mar. 25, 1977 (J. Dalbec, CAS), 13; Seneca, N fork of Feather River, Aug. 2, 1953 (W., J. Gertsch, AMNH), 5♀; Soda Springs Cave, 8 air mi N Quincy, Sept. 4, 1988, dark zone (D. Ubick, CDU), 13, 1♀; Soda Springs Cave, 10 mi N Quincy (R. Graham, AMNH), 13, 1♀, Sept. 5, 1961 (W. Gertsch, W. Ivie, R. Graham, AMNH), 3♀. Shasta Co.: cave in Low Pass Creek, Jan. 29, 1953, elev. 1600 ft (J. Gorman, AMNH), 13, 1♀. Sierra Co.: Bassetts, Aug. 18, 1993, in culvert, elev. 5400 ft (J. Boutin, CDU), 13; 4.8 mi N Camptonville, Aug. 21, 1993, in culvert, elev. 3000 ft (D. Ubick, CDU), 1♀; 5 mi E Camptonville, Aug. 15, 1993, in culvert, elev. 3000 ft (D. Ubick, CDU), 1♀, Aug. 9, 1999 (matured Sept.), elev. 3000 ft (D. Ubick, CDU), 13; the Cups, Sierra City, Sept. 6, 1959 (W. Gertsch, V. Roth, AMNH), 43, 5♀; Downieville, late July, 2004 (A. Faibish, UCR), 1♀; Downieville, at school, Sept. 24, 1993, elev. 2800 ft (T. Van Wagner, CDU), 1♀, Sept. 27, 1993, elev. 2800 ft (T. Van Wagner, CDU), 13; just E Loganville, no date, roadside rock outcrop at night (D. Ubick, CDU), 1♀; 7 mi W Sierra City, July 8, 1952 (W. Gertsch, AMNH, JCC), 13, 37♀. Siskiyou Co.: 20 mi N Somes Bar, July 20, 1962 (V. Roth, AMNH), 13 (single palp with broken tibial apophysis), 2♀. Sonoma Co.: Bohemian Highway, 2.4 mi SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.316667&amp;materialsCitation.latitude=9.616667" title="Search Plazi for locations around (long -9.316667/lat 9.616667)">Monte Rio</a>, 38 ° 26 9 37 0 N, 122 ° 59 9 19 0 W, Dec. 20, 2001, oakchaparral serpentine (T. Briggs, G. Giribet, D., S. Ubick, CDU), 13, 1♀; 0.2 mi NW Camp Meeker on Bohemian Highway, Jan. 5, 1984, elev. 120 m (D. Kavanaugh, CAS), 1♀; Guerneville, Apr. 4, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 203, 17♀, Aug. 2003, in house (R. Kaspar, UCR), 23; Monte Rio, May 5, 1996 (D. Ubick, CDU), 1♀, June 2003, in house (P. Craig, CDU), 23; 2.8 mi S Monte Rio, July 5, 1958 (W. Gertsch, V. Roth, AMNH), 2♀; Pine Glade Road, 1 air mi NE Monte Rio, Nov. 6, 1999, redwood forest (D., S. Ubick, CDU), 1♀; trail above Pine Glade Road, Monte Rio, June 30, 1996, redwood forest (P. Craig, W. Savary, D. Ubick, CDU), 1♀, Sept. 28, 1996, redwood forest (P. Craig, W. Savary, D. Ubick, CDU), 13, 1♀. Trinity Co.: Indian Valley Creek Cave, Oct. 27, 1990, elev. ca 1800 ft, on walls of cave (D. Ubick, W. Rauscher, CDU), 23, 1♀.</p> <p>DISTRIBUTION: Widespread in northern California (map 1).</p></div> 	https://treatment.plazi.org/id/038F87AEFFBDFFF1FF79FD76A54D5AAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFB7FFF5FF7DF9F0A5E25A84.text	038F87AEFFB7FFF5FF7DF9F0A5E25A84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus shasta Platnick & Ubick 2008	<div><p>Titiotus shasta, new species</p> <p>Figures 17–21; map 1</p> <p>TYPE: Male holotype from McArthur, Shasta Co., California (June 18, 1942; W. Pearce), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition to the genus name taken from the type locality.</p> <p>DIAGNOSIS: Males resemble those of T. californicus and T. hansii but have a much smaller ventral branch on the middle prong of the tibial apophysis (fig. 19); females resemble those of T. californicus but have a posteriorly narrowed epigynal septum (fig. 20).</p> <p>MALE: Total length 10. Coloration as in T. californicus. Leg spination: femora: I, II r1-2- 1; III, IV p1-2-1; tibiae: I, II d1-0-1, v4-4-4, r1- 1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2; IV r1-1-2. Embolus expanded prolaterally, beak-shaped distally; median apophysis with elongate tip, with only tiny subdistal projection; middle prong of tibial apophysis bifid, with ventral branch much smaller than dorsal branch (figs. 17–19).</p> <p>FEMALE: Total length 12. Coloration as in T. californicus. Leg spination: femora: I, II r1- 2-1; III r2-1-1; IV p2-0-1; tibiae: I v4-4-4, r0-1- 0; II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I p0-1-0, r0-1-0; II p1-1-0, r0-1-0; III p1- 2-2. Epigynal septum wide, occupying most of epigynal width, except posteriorly, where narrowed; posterior ducts elbowed (figs. 20, 21).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Colusa Co.: Deafy (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.7025&amp;materialsCitation.latitude=39.352497" title="Search Plazi for locations around (long -122.7025/lat 39.352497)">Dixie</a>) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.7025&amp;materialsCitation.latitude=39.352497" title="Search Plazi for locations around (long -122.7025/lat 39.352497)">Glade Campground on Forest Road</a>, 13 mi W Stonyford, Mendocino National Forest, 39 ° 21 9 N, 122 ° 42 9 W, no date, elev. 1000 m (J. Schweikert, CAS), 13. Glenn Co.: Forest Road, 5.5 mi S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.652504&amp;materialsCitation.latitude=39.5025" title="Search Plazi for locations around (long -122.652504/lat 39.5025)">County Road</a> 308, 9 mi SW Elk Creek, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.652504&amp;materialsCitation.latitude=39.5025" title="Search Plazi for locations around (long -122.652504/lat 39.5025)">Mendocino National Forest</a>, 39 ° 30 9 N, 122 ° 39 9 W, Mar. 30, 1997, elev. 1100 m (J. Schweikert, CAS), 13. Lake Co.: Lucerne, Aug. 17, 1958, molasses bait trap (H. Leech, CAS), 1♀. Mendocino Co.: 7.5 km NE Covelo, 28201 Mendocino Pass Road, 39 ° 50 9 N, 123 ° 10.7 9 W, Sept. 2, 1996, elev. 1680 ft (C. Prince, CAS), 1♀. Shasta Co.: Lake Shasta, June 14, 2006 (C. Abreu, UCR), 13; Palo Cedro, June 17, 2006 (D. McConnel, UCR), 23; Redding, Aug. 26, 2004 (A. Tennett, UCR), 1♀, June 2, 2006, in bathtub (A. Olivas, UCR), 1♀, June 4, 2006 (H. Ray, UCR), 13, June 16, 2006 (G. Snook, UCR), 13, same date (W. Mapes, UCR), 1♀, June 28, 2006 (T. Ressler, UCR), 1♀, June 29, 2006 (M. Wooden, UCR), 1♀. Trinity Co.: Hayfork Ranger Station, May 19, 1973 (M. Bentzien, UCB), 13.</p> <p>DISTRIBUTION: Known only from northwestern California (map 1).</p> <p>Titiotus flavescens (Chamberlin and Ivie)</p> <p>Figures 22–26; map 2</p> <p>Liocranoides flavescens Chamberlin and Ivie, 1941: 24, fig. 18 (female holotype from Ben Lomond, Santa Cruz Co., in AMNH, examined).</p> <p>Titiotus flavescens: Lehtinen, 1967: 271.</p> <p>DIAGNOSIS: Males resemble those of T. shantzi in having the middle prong of the tibial apophysis relatively narrow and the dorsal prong relatively short, but differ in having the middle prong more deeply incised (fig. 24); females have a distinctive epigynal septum, which is greatly narrowed posteriorly (fig. 25).</p> <p>Map 2. Central California, showing distributions of eight species of Titiotus.</p> <p>MALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I, II r1-2- 1; IV p0-2-1, r0-1-1; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I p1-0-0, r0-1-0; II p1-1-0, r1-1-0; III p1-2-2. Embolus with subapical, distally directed projection; median apophysis with subdistal projection; middle prong of tibial apophysis small, bifid, dorsal branch distally invaginated, dorsal prong relatively short (figs. 22–24).</p> <p>FEMALE: Total length 14. Coloration as in T. californicus, except proximal leg segments with dorsal dusky markings. Leg spination: femora: I, II r1-2-1; III, IV p1-2-1; tibiae: I p1- 0-0, v4-4-4, r1-0-0; II v4-4-4, r0-2-0; III, IV p1- 0-1, r1-0-1; metatarsi: III p1-2-2. Posterior quarter of epigynal septum greatly narrowed, forming stalk; posterior ducts oblique (figs. 25, 26).</p> <p>MATERIAL EXAMINED: United States: CALI- FORNIA: Santa Clara Co.: 27950 N Carnegie Ave., Santa Clara, June 8, 1993 (T. Carb, AMNH), 13; San Jose, Oct. 21, 1974, in house (D. Ubick, CDU), 13. Santa Cruz Co.: Aptos, July 10, 1946 (E. Ross, AMNH), 1♀; Bat Cave, Santa Cruz, Apr. 21, 1979 (D. Rudolph, Cowan, Van Ingen, AMNH), 23; Ben Lomond, May 1934 (L. Saylor, AMNH), 1♀ (holotype), June 25, 1952 (W. Gertsch, AMNH, JCC), 13, 21♀, July 4, 1958 (W. Gertsch, V. Roth, AMNH), 12♀, Apr. 3, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 93, 11♀, Sept. 23, 1961 (W. Gertsch, W. Ivie, AMNH), 53, 9♀; Boulder Creek, Aug. 1, 2004, in house (T. Nygard, UCR), 1♀, Oct. 7, 2004, in house (J. Sanchez, UCR), 13; 1 mi NE Boulder Creek, June 29, 1989, dry redwood forest, elev. ca 800 ft (A. Sundman, CDU), 1♀; Cave Gulch, Nov. 16, 1979, redwood forest (D. Ubick, CDU), 1♀; Cave Gulch, cave near Univ. of California, Santa Cruz, Nov. 4, 1972 (T. Stearns, CAS), 1♀; Cave Gulch, IXL Cave, Dec. 6, 1980, on cave wall (T. Briggs, CAS), 13; Felton, Aug. 16, 1959 (W. Gertsch, V. Roth, AMNH), 2♀, Apr. 3, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 1♀, June 1994 (S. Tjosvold, CAS), 1♀, Sept. 2001 (R. Knight, UCR), 13, Oct. 3, 2004, in shoe in bedroom (M. Mister, UCR), 13; N Soquel, Oct. 6, 2004 (A. Fournier, UCR), 13.</p> <p>DISTRIBUTION: Known only from Santa Clara and Santa Cruz counties south of the San Francisco Bay (map 2).</p></div> 	https://treatment.plazi.org/id/038F87AEFFB7FFF5FF7DF9F0A5E25A84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFB3FFE8FF01F9DFA5C55E32.text	038F87AEFFB3FFE8FF01F9DFA5C55E32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus shantzi Platnick & Ubick 2008	<div><p>Titiotus shantzi, new species</p> <p>Figures 27–31; map 2</p> <p>TYPES: Male holotype and female allotype from Santa Barbara, Santa Barbara Co., California (May 1948; H. Shantz), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.</p> <p>DIAGNOSIS: Males resemble those of T. flavescens but have the middle prong of the tibial apophysis short, with a beak-shaped dorsal branch (fig. 29); females have the epigynal septum wider posteriorly than do those of T. flavescens (fig. 30).</p> <p>MALE: Total length 10. Coloration as in T. californicus. Leg spination: femora: I p1-2-1, r2-1-1; II r1-1-2; IV p1-2-1; tibiae: I, II d1-0-1, p0-1-1, v4-4-4, r0-1-1; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0. Embolus with small, subapical, distally directed projection; median apophysis with small, subdistal projection; middle prong of tibial apophysis small, bifid, with beak-shaped dorsal branch, dorsal prong relatively short (figs. 27–29).</p> <p>FEMALE: Total length 14. Coloration as in T. californicus. Leg spination: femora: I, II r1- 1-2; III r2-1-1; IV p1-2-1; tibiae: I, II d1-0-1, p0-1-1, v4-4-4, r0-1-1; III, IV r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Epigynal septum relatively narrow, only slightly narrowed posteriorly; posterior ducts recurved (figs. 30, 31).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Kern Co.: Fort Tejon State Historic Park, June 15, 1975, at lights (M. Muma, AMNH), 23; Tehachapi, June 16, 1975, at light (M. Muma, AMNH), 23. Los Angeles Co.: Acton, May 14, 2000, in house, elev. 2,700 ft (C. Lutz, UCR), 1♀; Agoura, May 6, 2005, in bathroom (D. Trisler, UCR), 13; Agua Dulce, June 2, 1977 (F. Runyan, LACM), 13; Canyon Country, mid-May 2003, in house (R. Fridlund, UCR), 13; Elizabeth Lake, 34 ° 39.7 9 N, 118 ° 21.96 9 W, May 30, 2006, elev. 3,537 ft (C. Becker, UCR), 13; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.36917&amp;materialsCitation.latitude=34.0025" title="Search Plazi for locations around (long -118.36917/lat 34.0025)">Kenneth Hahn State Recreation Area</a>, Baldwin Hills, 34 ° 00 9 N, 118 ° 22 9 W, June 2000, pitfall, elev. 67 m (L. LaPierre, P. Williams, CBH), 13; La Canada, June 24, 1970 (A. Guldaro, LACM), 1♀; Lake Hughes, NW Palmdale, June 2, 2002, elev. 3,200 ft (M. Enright, UCR), 1♀; Lassen Street, Chatsworth, 0.25 mi W Browns Canyon Wash, Apr. 29–May 20, 1966, in house, elev. 950 ft (W. Icenogle, WRI), 2♀, May 23, 1966, in house, elev. 950 ft (W. Icenogle, WRI), 13; Leona Valley, May 9, 2002, elev. 3,200 ft (K. Smith, CAS), 23; Placerita Canyon Park, 3 mi E Newhall, July 26, 1972, elev. 1,550 ft (F.</p> <p>Hovore, WRI), 1♀; Saugas, May 4, 2004 (B. Lytle, UCR), 13; Valencia, Dec. 1, 2003 (M. Pacheco, UCR), 13. Monterey Co.: Paraiso Hot Springs, July 4, 1954, elev. 1,400 ft (O. Bryant, CAS), 1♀. San Luis Obispo Co.: Morro Bay, May 26, 1984 (V. Lee, CAS), 13. Santa Barbara Co.: Lake Cachuma, 18 mi NE Santa</p> <p>Barbara (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 1♀; Santa Barbara, May 1948 (H. Shantz, AMNH), 23; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.06917&amp;materialsCitation.latitude=34.6525" title="Search Plazi for locations around (long -120.06917/lat 34.6525)">Sedgwick Ranch Reserve</a>, May 1997, Malaise trap, elev. 1,800 ft (E., M. Schlinger, CAS), 1♀; Superstar Ranch, Santa Barbara, 34 ° 39 9 N, 120 ° 04 9 W, May–June 1997, elev. 700 ft (E., M. Schlinger, CAS), 13. Ventura Co.: Simi Valley, May 6, 2003, in house (B. Moran, UCR), 13.</p> <p>DISTRIBUTION: Known only from southwestern California (map 2).</p></div> 	https://treatment.plazi.org/id/038F87AEFFB3FFE8FF01F9DFA5C55E32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFAEFFE8FF50FE6DA05E5E58.text	038F87AEFFAEFFE8FF50FE6DA05E5E58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus tahoe Platnick & Ubick 2008	<div><p>Titiotus tahoe, new species</p> <p>Figures 32–36; map 2</p> <p>TYPE: Male holotype taken at an elevation of 6000 ft at Tahoe City, Placer Co., California (July 10, 1952; W. Gertsch), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males and females have not been collected together but are matched here on the basis of geography; males can easily be recognized by the tibial apophysis consisting of four prongs (fig. 34), females by the thumbshaped median lobe situated posteriorly on the epigynal septum (fig. 35).</p> <p>MALE: Total length 8. Coloration as in T. californicus. Leg spination: femora: I r1-2-1; II p2-1-1, r1-1-2; III r1-2-2; IV p1-2-1; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III, IV r0-1-1; metatarsi: I p1-1-0, r0-1-0; II p1-1-0, r1-1-0; III, IV p1-2-2. Embolus expanded prolaterally just beyond base; median apophysis long, with large, subdistal projection; tibial apophysis with four prongs, only two most distal sharing common base (figs. 32–34).</p> <p>FEMALE: Total length 19. Coloration as in T. californicus. Leg spination: femora: II, III r1-1-2; tibiae: I, II r1-1-0; III r0-1-1; IV p1-0-1; metatarsi: II p1-1-0, r1-1-0; III p1-2-2. Epigynal septum with thumb-shaped median lobe occupying posterior fifth of septum length; posterior ducts oblique (figs. 35, 36).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: El Dorado Co.: Placerville, Jan. 1995 (UCR), 13; 5 mi N Pollock Pines, July 6, 1958 (W. Gertsch, V. Roth, AMNH), 13♀; Snowline Camp, July 15, 1948 (P. Hurd, AMNH), 13. Nevada Co.: S fork, Yuba River, 8 mi N Nevada City, Aug. 3, 1953 (W., J. Gertsch, AMNH), 5♀. Placer Co.: Tahoe City, July 10, 1952, elev. 6000 ft (W. Gertsch, AMNH), 13. Yuba Co.: Brown’s Valley, June 28, 2004 (J. Crummett, UCR), 13.</p> <p>DISTRIBUTION: Known only from four counties in the Sierras (map 2).</p></div> 	https://treatment.plazi.org/id/038F87AEFFAEFFE8FF50FE6DA05E5E58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFAEFFEAFCA0FE81A7665EA6.text	038F87AEFFAEFFEAFCA0FE81A7665EA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus gertschi Platnick & Ubick 2008	<div><p>Titiotus gertschi, new species</p> <p>Figures 37–41; map 1</p> <p>TYPES: Male holotype and female allotype from Riverton, El Dorado Co., California (July 11, 1952; W. Gertsch), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.</p> <p>DIAGNOSIS: Males resemble those of T. madera in having the dorsal prong of the median apophysis long and excavated, but differ in having only two projections on the middle prong of the tibial apophysis (fig. 39); females also resemble those of T. madera in having a gradually narrowed posterior projection on the epigynal septum, but have the septum much wider than in that species (fig. 40).</p> <p>MALE: Total length 16. Coloration as in T. flavescens. Leg spination: femora: I r1-2-1; II r1-1-2; IV p2-1-1; tibiae: I, II d1-0-1, r1-1-0; III p1-0-1, r1-0-1; IV p1-0-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus expanded prolaterally just beyond base; median apophysis with large, subdistal projection; middle prong of tibial apophysis bifid, dorsal prong long, ventrally excavated (figs. 37–39).</p> <p>FEMALE: Total length 18. Coloration as in T. flavescens. Leg spination: femora: II p1-1-1, r1-1-1; III, IV p1-2-1; tibiae: I r1-1-0; II d1-0- 1, r1-1-0; III p1-0-1, r1-0-1; metatarsi: I p1-0- 0; II r0-1-0; III p1-2-2. Epigynal septum very wide, occupying most of epigynal width, with gradually narrowing posteromedian ledge; posterior ducts curved (figs. 40, 41).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Amador Co.: Moss Cave, near Volcano, Oct. 17, 1987 (D. Cowan, F. Howarth, D. Ubick, CDU), 23, 1♀, Apr. 24, 1999 (R. Aalbu, CAS), 13, 1♀, Jan. 13, 2002 (R. Aalbu, CAS), 43; Pearl Cave, 0.2 mi NE Volcano, Sept. 13, 1980, on wall (T. Briggs, D. Ubick, CDU), 13; Pearl Cave, 1 mi N Volcano, Feb. 20, 1979 (D. Rudolph, S. Winterath, CAS), 1♀. Calaveras Co.: 7 mi W Angels Camp, Nov. 2, 1979 (R. Laughlin, CDU), 1♀; Brown Deer Cave, Grapevine Gulch, N Columbia, Nov. 27, 1958 (P. Anderson, AMNH), 13; Cave of Skulls, 3 mi N Columbia, Mar. 29, 1979 (D. Rudolph, B. Martin, S. Winterath, W. Elliott, J. Reddell, AMNH), 1♀; Crystal- <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.75&amp;materialsCitation.latitude=9.25" title="Search Plazi for locations around (long -9.75/lat 9.25)">Stanislaus Cave</a>, 38 ° 06 9 15 0 N, 120 ° 24 9 45 0 W, Nov. 25, 1950 (E. Dansky, AMNH), 13; Dublen Cave, Aug. 17, 1963 (R. Graham, AMNH), 2♀; Grapevine Gulch Cave, Oct. 18, 1987 (F. Howarth, D. Cowan, CAS), 13; Pseudoscorpion Cave, Aug. 17, 1963 (R. Graham, AMNH), 2♀; Speleogen Cave, 6 mi NW Columbia, 38 ° 02.85 9 N, 120 ° 28.4 9 W, Feb. 1, 1953, 100 ft in cave (R. de Saussure, AMNH), 13, Dec. 18, 1972 (W. Rauscher, CDU), 13, Mar. 24, 2000 (T. Briggs, J. Ledford, W. Rauscher, D., S. Ubick, CDU), 13; 2.5 mi S West Point, May 2, 1981 (S. Williams, CAS), 1♀; Wood Hollow Cave, Aug. 11, 1962 (R. Graham, AMNH), 1♀. El Dorado Co.: Fallen Leaf Lake, Apr. 9, 1959 (W. Gertsch, V. Roth, AMNH), 13, 1♀, Sept. 9, 1959 (W. Gertsch, V. Roth, AMNH, CAS), 103, 8♀; Riverton, July 11, 1952 (W. Gertsch, AMNH), 63, 17♀. Mariposa Co.: Water Pit Cave, 10 mi E Coulterville, Apr. 7, 1979 (D. Rudolph, B. Martin, S. Winterath, Cowan, Lahner, AMNH), 13. Tuolumne Co.: Bend Cave, 3 mi N Columbia, Feb. 27, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 1♀; mine, 2.5 mi N Columbia, Feb. 22, 1979 (D. Rudolph, S. Winterath, B. Martin, AMNH), 13; Crystal Palace Cave, 3 mi N Columbia, Feb. 4, 1979 (D. Rudolph, S. Winterath, AMNH), 13, Feb. 28, 1979 (D. Rudolph, S. Winterath, B. Martin, AMNH), 13, 1♀; Draper Mine Road, 6 mi E Sonora, Oct. 12, 1973 (matured Feb. 15, 1974), in ravine bank on turned soil, elev. 2,700 ft (W. Icenogle, WRI), 13; Fox Gulch, Transplant Mine, ca 2 mi N Columbia, Dec. 6, 1986, elev. 1,630 ft (D. Ubick, V. Lee, CDU), 13; McLean’s Cave, Marble Quarry Road at South Fork, Stanislaus River, Mar. 5, 1981, on rock face (D. Ubick, CDU), 13, 1♀; McLean’s Cave, 3 mi N Columbia, Mar. 25, 1979 (D. Rudolph, B. Martin, S. Winterath, W. Elliott, J. Reddell, AMNH), 13; Pine Log Cave, 2 mi W Columbia (D. Rudolph, B. Martin, S. Winterath, AMNH), 13, 1♀; Pinnacle Point Cave, 6 mi N Columbia, Feb. 8, 1979 (D. Rudolph, S. Winterath, AMNH), 23, 1♀, Oct. 4, 2004 (R. Aalbu, CAS), 1♀; Porcupine Skull Cave, 2 mi N Columbia, Feb. 11, 1979 (D. Rudolph, S. Winterath, AMNH), 1♀; 5 mi N Priest, July 8, 1959 (W. Gertsch, V. Roth, AMNH, JCC), 13, 23♀; Razorback Grotto Cave, 3 mi N Columbia, Mar. 28, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 13; Scorpion Cave, 12.6 km NNW Sonora, Feb. 1, 1978 (A. Grubbs, AMNH), 13; Sonora, June 2003 (J. Wenz, UCR), 1♀; Strawberry, Sept. 10, 1959 (W. Gertsch, V. Roth, AMNH), 143, 25♀; Tuolumne, Apr. 15, 1934 (CAS), 1♀, June 2005, in or around house (W. Rauscher, CDU), 2♀, June 2006, in house (W. Rauscher, CDU), 1♀; Tuolumne River at N entrance to Yosemite National Park, Aug. 7, 1953 (W., J. Gertsch, AMNH), 2♀; Windeler Cave, 11.3 km NNE Sonora, Jan. 12, 1978 (W. Elliott, A. Grubbs, S. Winterath, AMNH), 1♀.</p> <p>DISTRIBUTION: Known only from the Sierras, generally to the south of T. tahoe (map 1).</p> </div>	https://treatment.plazi.org/id/038F87AEFFAEFFEAFCA0FE81A7665EA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFACFFEBFC9DFDE1A68E595D.text	038F87AEFFACFFEBFC9DFDE1A68E595D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus madera Platnick & Ubick 2008	<div><p>Titiotus madera, new species</p> <p>Figures 42–46; map 2</p> <p>TYPES: Male holotype and female allotype from Bass Lake, Madera Co., California (July 9, 1958; W. Gertsch, V. Roth), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males resemble those of T. gertschi, but can easily be distinguished by the presence of multiple projections on the middle prong of the tibial apophysis (fig. 44); females also resemble those of T. gertschi but have a much narrower epigynal septum (fig. 45).</p> <p>MALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I r1-2-1; II r1-1-2; III r1-1-2; tibiae: I d1-0-1, r1-1-1; II d1- 0-1, r1-1-0; III r0-1-1; metatarsi: p1-0-0, r1-0- 0; III p1-2-2. Embolus expanded prolaterally just beyond base, with short tip; median apophysis with small subdistal projection; middle prong of tibial apophysis bifid, bearing multiple projections, dorsal prong long, ventrally excavated (figs. 42–44).</p> <p>FEMALE: Total length 14. Coloration as in T. californicus. Leg spination: femora: II p1-2- 1, r1-1-2; III r2-1-1; tibiae: I, II p0-1-0, r1-1-0; metatarsi: II p0-0-0, v0-0-0; III p1-2-2. Epigynal septum wide, occupying about twothirds of epigynal width, with gradually narrowing posteromedian ledge; posterior ducts oblique (figs. 45, 46).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Madera Co.: Bass Lake, July 9, 1958 (W. Gertsch, V. Roth, AMNH), 13, 2♀; 5 mi S Fish Camp, just over county line, Sept. 23, 1961 (W. Gertsch, W. Ivie, AMNH), 13. Mariposa Co.: El Portal, Apr. 8, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 1♀; El Portal, outside entrance to Yosemite National Park, July 14, 1952 (W. Gertsch, AMNH), 23, 11♀; 14 mi N Oakhurst, Aug. 7, 1953, elev. 5100 ft (W., J. Gertsch, AMNH), 13, 6♀; Sprinkle Cave, Apr. 7, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 1♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.16917&amp;materialsCitation.latitude=37.53583" title="Search Plazi for locations around (long -119.16917/lat 37.53583)">Wawona Camp</a>, 37 ° 32 9 N, 119 ° 10 9 W, Sept. 17, 1941 (W. Ivie, AMNH), 2♀; Yosemite National Park, July 1935 (AMNH), 1♀; Yosemite National Park, bridge below Nevada Falls, June 26, 1973 (P. Arnaud, Jr., CAS), 13.</p> <p>DISTRIBUTION: Known only from the Sierras of Madera and Mariposa Counties (map 2).</p></div> 	https://treatment.plazi.org/id/038F87AEFFACFFEBFC9DFDE1A68E595D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFADFFEFFD42F996A4C15D96.text	038F87AEFFADFFEFFD42F996A4C15D96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus heberti Platnick & Ubick 2008	<div><p>Titiotus heberti, new species</p> <p>Figures 47–51; map 1</p> <p>TYPES: Male holotype and female allotype from Wilsonia, Kings Canyon National Park, Tulare Co., California (Sept. 13, 1959; W. Gertsch, V. Roth), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a patronym in honor of Mr. Blaine Hebert, assiduous collector of these and other Californian spiders.</p> <p>DIAGNOSIS: Males resemble those of T. tulare in having a large, wide dorsal prong on the tibial apophysis, but differ in having a distally narrower middle prong (fig. 49); females differ from those of T. tulare in having a posteriorly wider epigynal septum (fig. 50).</p> <p>MALE: Total length 12. Coloration as in T. californicus. Leg spination: femora: I p0-3-1, r1-2-1; II r1-2-1; IV p1-2-1; tibiae: I, II d1-0-1, r1-1-0; III r1-0-1; metatarsi: I p1-0-0, r1-1-0; III p1-2-2. Embolus twisted near base, distally wide; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis bifid, with subequal branches, dorsal prong large, wide, flattened (figs. 47– 49).</p> <p>FEMALE: Total length 17. Coloration as in T. californicus. Leg spination: femora: I p0-3- 1, r1-2-1; II r1-2-1; III r1-1-2; IV p1-2-1; patella III p0-0-0; tibiae I, II r1-1-0; metatarsi III, IV p1-2-2. Epigynal septum wide, with posteromedian bulge occupying most of septal width; posterior ducts oblique, arched (figs. 50, 51).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Fresno Co.: Cedar Grove, Kings River Canyon, July 16, 1952, elev. 4633 ft (W. Gertsch, AMNH), 8♀, July 5, 1956, elev. 4633 ft (W., J. Gertsch, AMNH), 23, 6♀; Hume Lake, Dec. 17, 1984 (K. Walker, CAS), 1♀; Lily Pond, 2 mi W Wishon Lake, Oct. 31, 1982, in sandy soil (M. Marshall, CAS), 1♀; Ranger Station, Kings River, Kings Canyon National Park, June 17, 1969, in house (M. Zardus, CAS), 13; Shaver Lake, July 15, 1952 (W. Gertsch, AMNH), 5♀, Sept. 12, 1959 (W. Gertsch, V. Roth, AMNH), 123, 3♀; vicinity Wishon Resv., July 2–6, 1969 (D. Marqua, MET), 13, 2♀, May 28–June 14, 1972 (M. Thompson, MET), 2♀. Kern Co.: Cedar Creek Camp Ground, 0.25 mi N Highway 155, Greenhorn Mountains, 35 ° 44.44 9 N, 118 ° 35.54 9 W, Aug. 25, 2006, on wall of old mine shaft in steep, rocky canyon side near stream, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀, Aug. 26, 2006 (matured Sept. 4, 2006), on ceiling at mouth of old mine shaft in steep, rocky canyon side stream nearby conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀, Aug. 26, 2006 (matured Sept. 8, 2006), on base of tree near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 1♀, Aug. 26, 2006 (matured Dec. 9, 2006), on side of big rock near stream in steep-sided canyon, conifer-oak forest, elev. 5000 ft (B. Hebert, W. Icenogle, WRI), 13. Madera Co.: Cascadel Woods, T 8S,R 23E,S16, July 1–3, 1983 (J. Aotok, CAS), 13; Coarsegold, no date (Oliva, UCR), 1♀, Apr. 21, 2004 (B. Harrigom, UCR), 1♀; Wishon, July 5, 1971 (D. Marqua, MET), 13. Mariposa Co.: Foresta Road, Yosemite National Park, 37 ° 42.242 9 N, 119 ° 43.989 9 W, June 15–18, 2003, pitfalls, elev. 1392 m (A. Leache, C. Conroy, UCB), 63. Tulare Co.: Ash Mountain, Sequoia National Park, Apr. 26, 1951 (E. Schlinger, AMNH), 13; near Ash Mountain, Sequoia National Park, July 9, 1958 (W. Gertsch, V. Roth, AMNH), 13, 5♀; 1 mi below Belnap Springs, near Camp Nelson, July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀; Carmoe Crevice, Nov. 3, 1993 (J. Krejca, CAS), 13; NE Case Mountain Peak, 36.41790 ° N, 118.7933 ° E, July 14–18, 2003, dry pitfall, elev. 1691 m (R. Fisher, AMNH), 13; NE Case Mountain Peak, 36.41870 ° N, 118.79264 ° E, July 14–18, 2003, dry pitfall, elev. 1672 m (R. Fisher, AMNH), 13; NNE Case Mountain Peak, E of E end of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.79264&amp;materialsCitation.latitude=36.4187" title="Search Plazi for locations around (long 118.79264/lat 36.4187)">Salt Creek</a>, 36.42059 ° N, 118.79776 ° E, May 2003, dry pitfall, elev. 1749 m (R. Fisher, AMNH), 13, 36.42083 ° N, 118.79163 ° W, Sept. 2002, dry pitfall, elev. 1609 m (R. Fisher, AMNH), 1♀; NW Case Mountain Peak, S of Salt Creek, 36.41871 ° N, 118.81930 ° W, May 2003, dry pitfall, elev. 1709 m (R. Fisher, AMNH), 13; just S Case Mountain Peak, 36.407 ° N, 118.8016 ° E, July 14–18, 2003, dry pitfall, elev. 1998 m (R. Fisher, AMNH), 13; SE Case Mountain Peak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.8016&amp;materialsCitation.latitude=36.407" title="Search Plazi for locations around (long 118.8016/lat 36.407)">Salt Creek</a> Ridge, 36.39964 ° N, 118.77975 ° W, Sept. 2002, dry pitfall, elev. 1808 m (R. Fisher, AMNH), 13; W Case Mountain Peak, 36.41120 ° N, 118.80909 ° W, July 2002, dry pitfall, elev. 1951 m (R. Fisher, AMNH), 1♀, 36.41139 ° N, 118.80843 ° W, July 2002, dry pitfall, elev. 1932 m (R. Fisher, AMNH), 1♀, July 14–18, 2003, dry pitfall, elev. 1951 m (R. Fisher, AMNH), 1♀; just W Case Mountain Peak, 36.41139 ° N, 118.80843 ° W, July 14–18, 2003, dry pitfall, elev. 1932 m (R. Fisher, AMNH), 13; Cedar Cave, Redwood Canyon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.80843&amp;materialsCitation.latitude=36.41139" title="Search Plazi for locations around (long -118.80843/lat 36.41139)">Kings Canyon National Park</a>, Aug. 16, 1984, under rock outside cave entrance, elev. 1,600 m (D. Ubick, CDU), 1♀; Crystal Sequoia Cave, Sequoia National Park, Apr. 16, 1946 (F. Oberhansley, AMNH), 1♀, July 26, 1991 (D. Ubick, CDU), 1♀; cave in bank on road to Crystal Cave, Sequoia National Park, Nov. 6, 1949, in total darkness (A. Lange, AMNH), 1♀; Harry’s Bend Cave, July 10, 2004 (J. Krejca, A. Gluesenkamp, C. Richard, CAS), 13; Kaweah Power Station #3, Ash Mountain, 40 mi NE Visalia, July 6, 1983 (D. Burdick, CAS, CDU), 23, May 9, 1993 (D. Burdick, CAS), 13, July 2, 1995 (D. Burdick, CAS), 13; Lindcove, July 13, 1976, dead on doorstep (D. Carroll, UCR), 1♀; McIntire Creek, near Camp Nelson, July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13, 1♀; 5–10 mi on Mineking Road from Three Rivers, July 4, 1956, elev. 2000–4000 ft (AMNH), 13, 4♀; Moorhouse Creek, near Camp Nelson, July 11, 1958 (W. Gertsch, V. Roth, AMNH), 13; 1 mi W Camp Nelson, July 11, 1958 (W. Gertsch, V. Roth, AMNH), 2♀; Roads End, Kern River, July 3, 1956 (W. Gertsch, AMNH), 13; Soda Creek, W Camp Nelson, July 11, 1958 (W. Gertsch, V. Roth, AMNH), 23, 2♀; Wilsonia, Kings Canyon National Park, Sept. 13, 1959 (W. Gertsch, V. Roth, AMNH), 23, 3♀; 9 mi N Woodlake, Mar. 22, 1941 (S., D. Mulaik, AMNH), 13. Tuolumne Co.: Tuolumne, May 2004, in house (M. Rauscher, CDU), 13.</p> <p>DISTRIBUTION: Known only from the Sierras, generally to the south of T. gertschi (map 1).</p> </div>	https://treatment.plazi.org/id/038F87AEFFADFFEFFD42F996A4C15D96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA9FFEFFF7AFD31A7945C83.text	038F87AEFFA9FFEFFF7AFD31A7945C83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus tulare Platnick & Ubick 2008	<div><p>Titiotus tulare, new species</p> <p>Figures 52–56; map 2</p> <p>TYPE: Male holotype from near Ash Mountain, Sequoia National Park, Tulare Co., California (July 9, 1958; W. Gertsch, V. Roth), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males and females have not been collected together, and it is possible that the females treated here as T. tulare are actually those of T. icenoglei, and vice versa. Males resemble those of T. heberti but differ in having the tibial apophysis with a distally wider middle prong (fig. 54); females differ from those of T. heberti in having a posteriorly narrowed epigynal septum (fig. 55).</p> <p>MALE: Total length 16. Coloration as in T. californicus. Leg spination: femora: I p0-4-1, r1-2-1; II r1-2-1; III p1-1-2, r1-2-2; IV p1-2-1; tibiae: I v6-6-4, r1-2-0; II d1-0-1, v6-6-4, r1-2- 0; III d1-0-0, p1-1-2, r0-1-1; IV r1-0-2; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus twisted near base, distally narrow; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis bifid but not deeply incised, dorsal prong large, wide, flattened (figs. 52–54).</p> <p>FEMALE: Total length 13. Coloration as in T. californicus. Leg spination: femora: I r1-2-1; II p1-1-2, r1-1-2; tibiae: I d1-0-0, v4-4-4, r1- 1-0; II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: II r1-1-0; III p1-2-2. Epigynal septum wide but short, posterior portion restricted to narrow, triangular bulge; posterior ducts oblique, spermathecae narrow (figs. 55, 56).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Kern Co.: Paradise Valley, Tehachapi Mountains, Aug. 18, 1959, in building, elev. 5000 ft (W. Icenogle, WRI), 1♀; Tehachapi Mountain County Park, Tehachapi Mountains, 35 ° 04.680 9 N, 118 ° 28.857 9 W, Sept. 9, 2004, crawling on road after rain, conifer-oak forest, elev. 5800 ft (W. Icenogle, WRI), 1♀. Tulare Co.: 3 mi W Giant Forest, Sequoia National Park, July 9, 1958 (W. Gertsch, V. Roth, AMNH), 13; Kaweah Power Station #3, Ash Mountain, 40 mi NE Visalia, Mar. 12, 1983 (CDU), 13, July 10, 1983 (D. Burdick, CAS), 13, Apr. 13, 1985 (CAS), 13, Apr. 25, 1996 (D. Burdick, CAS), 13; Kaweah River, 5 mi E Three Rivers, July 17, 1952, elev. 1258 ft (W. Gertsch, AMNH), 1♀; Lake Kaweah, Visalia, May 10, 1994 (D. Bixler, CDB), 13; Three Rivers, July 17, 1952, elev. 1268 ft (W. Gertsch, AMNH), 23.</p> <p>DISTRIBUTION: Known only from Kern and Tulare Counties (map 2).</p> </div>	https://treatment.plazi.org/id/038F87AEFFA9FFEFFF7AFD31A7945C83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA9FFE2FD4EFBD9A7CE5AB2.text	038F87AEFFA9FFE2FD4EFBD9A7CE5AB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus roadsend Platnick & Ubick 2008	<div><p>Titiotus roadsend, new species</p> <p>Figures 57–61; map 2</p> <p>TYPES: Male holotype and female allotype from Roads End, Kern River, Tulare Co., California (July 3, 1956; W. Gertsch), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males of this distinctive species can easily be recognized by the short, sinuous dorsal prong of the tibial apophysis (fig. 59), females by the narrow, posteriorly expanded epigynal septum (fig. 60).</p> <p>MALE: Total length 13. Coloration as in T. californicus. Leg spination: femora: I p0-4-1, r1-2-1; II r1-2-2; III r2-1-1; IV p1-2-2; tibiae: I d1-0-1, p1-2-0, v6-6-4, r1-1-1; II d1-0-1, p1-2- 0, v6-6-4, r1-2-0; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus twisted near base, distally narrow; median apophysis with submedian projection; middle prong of tibial apophysis bifid, incised, dorsal prong short, twisted, situated near cymbium (figs. 57–59).</p> <p>59. Same, retrolateral view. 60. Epigynum, ventral view. 61. Same, dorsal view.</p> <p>FEMALE: Total length 18. Coloration as in T. californicus. Leg spination: femora: I p1-3- 2, r1-2-1; II r1-2-1; III p2-1-2; tibiae: I, II p1-0- 0, v6-6-4, r1-0-0; III r0-1-1; IV d1-0-0, r0-1-1; metatarsi: II p0-1-0; III p1-2-2, r1-2-2; IV r2-2- 2. Epigynal septum narrow, anterior with parallel sides, posteriorly expanded, wrinkled; posterior ducts almost transverse (figs. 60, 61).</p> <p>OTHER MATERIAL EXAMINED: One male and one female taken with the types (AMNH).</p> <p>DISTRIBUTION: Known only from the type locality in Kern County (map 2).</p> <p>69. Same, retrolateral view. 70. Epigynum, ventral view. 71. Same, dorsal view.</p></div> 	https://treatment.plazi.org/id/038F87AEFFA9FFE2FD4EFBD9A7CE5AB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA4FFE3FF54F9C8A5C65CC7.text	038F87AEFFA4FFE3FF54F9C8A5C65CC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus fresno Platnick & Ubick 2008	<div><p>Titiotus fresno, new species</p> <p>Figures 62–66; map 2</p> <p>TYPES: Male holotype and female allotype taken at an elevation of 4633 ft at Cedar Grove, Kings River Canyon, Fresno Co., California (July 16, 1952; W. Gertsch), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males of this distinctive species can easily be recognized by the relatively small embolus (fig. 63) and the relatively wide middle prong of the tibial apophysis (fig. 64), females by the anteriorly rounded epigynal septum (fig. 65).</p> <p>MALE: Total length 17. Coloration as in T. californicus. Leg spination: femora: I p0-4-1, r1-3-1; II p1-2-1, r1-1-3; III p2-2-1, r1-2-2; IV p1-2-1; tibiae: I d1-0-1, v6-6-8, r1-2-1; II p1-2- 1, v6-6-6, r1-2-0; IV r1-1-1; metatarsi: I p2-0-0, r2-0-0; II p2-0-0, r1-2-0; III p1-2-2. Embolus twisted near base, distally very narrow; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis wide, with two incisions, dorsal prong relatively short, blade-shaped (figs. 62–64).</p> <p>FEMALE: Total length 21. Coloration as in T. californicus. Leg spination: femora: I p0-3- 1, r1-2-1; II p1-2-1, r1-2-2; III r2-1-1; IV p1-2- 1; tibiae: I, II p1-2-0, v6-6-6, r1-2-0; IV r1-0-2; metatarsi: I p1-0-0, r1-1-0; II p2-0-0, r1-1-0. Epigynal septum anteriorly with convex lateral margins, posteriorly with straight lateral margins; posterior ducts approximate through much of their length (figs. 65, 66).</p> <p>OTHER MATERIAL EXAMINED: Two males taken with the types (AMNH).</p> <p>DISTRIBUTION: Known only from the type locality in Fresno Co. (map 2).</p> </div>	https://treatment.plazi.org/id/038F87AEFFA4FFE3FF54F9C8A5C65CC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA5FFE3FF0AFC78A68E5D98.text	038F87AEFFA5FFE3FF0AFC78A68E5D98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus icenoglei Platnick & Ubick 2008	<div><p>Titiotus icenoglei, new species</p> <p>Figures 67–71; map 2</p> <p>TYPE: Male holotype taken on the ground below a yellowish floodlight in a conifer-oak forest at an elevation of 5800 ft at Tehachapi Mountain County Park, Tehachapi Mountains, 35 ° 04.068 9 N, 118 ° 28.857 9 W, Kern Co., California (June 26, 2004; W. Icenogle), deposited in AMNH courtesy of Mr. Icenogle.</p> <p>ETYMOLOGY: The specific name is a patronym in honor of the collector of the type (and many other) specimens.</p> <p>DIAGNOSIS: As noted above, the females here placed as T. icenoglei and T. tulare may actually be mismatched. Males of this distinctive species can easily be recognized by the short, deeply incised middle prong of the tibial apophysis (fig. 69), females by the medially expanded epigynal septum (fig. 70).</p> <p>MALE: Total length 10. Coloration as in T. californicus. Leg spination: femora: I r1-2-1; II p1-2-1, r1-2-1; IV p1-2-1; tibiae I, II d1-0-1, v4-4-4, r1-1-0; metatarsi: I p1-1-0, r0-1-0; II p1-1-0, r1-1-0; III p1-2-2. Embolus prolaterally expanded at base, distally blunt; median apophysis with subdistal and retrolateral projections; middle prong of tibial apophysis very short, deeply incised, dorsal prong relatively short, blade-shaped (figs. 67–69).</p> <p>FEMALE: Total length 13. Coloration as in T. californicus. Leg spination: femora: I p0-3- 1, r1-2-1; II p1-3-1, r1-1-3; III p1-2-1, r1-1-2; IV p2-2-1; tibiae: I d0-0-1, v4-4-4, r1-2-0; II d1-0-1, v4-4-4, r1-1-0; III, IV r0-1-1; metatarsi: I p0-1-0, r0-1-0; II p1-1-0, r1-1-0; III p1-2-2. Epigynal septum much narrower anteriorly and posteriorly than at midlength; posterior ducts widely separated (figs. 70, 71).</p> <p>OTHER MATERIAL EXAMINED: One female taken at Bakersfield, Kern Co. in mid-June 1999 (J. Sparks, UCR).</p> <p>DISTRIBUTION: Known only from Kern Co. (map 2).</p></div> 	https://treatment.plazi.org/id/038F87AEFFA5FFE3FF0AFC78A68E5D98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA5FFE5FD50FCD5A61D598A.text	038F87AEFFA5FFE5FD50FCD5A61D598A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus humboldt Platnick & Ubick 2008	<div><p>Titiotus humboldt, new species Figures 72–76; map 3</p> <p>TYPES: Male holotype and female allotype taken 10 mi east of Bridgeville, Humboldt Co., California (Aug. 20, 1959; W. Gertsch), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: This species resembles T. marin and T. costa in having the ventral prong of the tibial apophysis relatively short and wide, and situated close to the base of the middle prong; males can be distinguished by having the third most ventral prong of the tibial apophysis relatively short, only slightly expanded distally, and not incised (fig. 74), females by the relatively wide epigynal septum and protuberant posterolateral epigynal margins (fig. 75).</p> <p>MALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I p1-2-1; II r1-2-1; IV p1-2-1; tibiae: I d1-0-1, v4-4-4, r0-1- 0; II d1-0-1, p1-0-0, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus prolaterally expanded at base, distally narrow; median apophysis with only tiny subdistal projection; two most ventral prongs of tibial apophysis small, third relatively short, square (figs. 72–74).</p> <p>FEMALE: Total length 12. Coloration as in T. californicus. Leg spination: femora: I, II r1- 2-1; III p2-1-2, r1-1-3; IV p2-1-1; tibiae: I v4-4-</p> <p>74. Same, retrolateral view. 75. Epigynum, ventral view. 76. Same, dorsal view.</p> <p>4, r1-0-0; II v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Epigynal septum wide, wider anteriorly than posteriorly, posterolateral epigynal margins protuberant; posterior ducts widely separated (figs. 75, 76).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Humboldt Co.: Five Mile Creek at junction of Highway 96, 4.2 mi S Somes Bar, Oct. 2, 1979, on wall of 6 ft diameter conduit under highway, elev. 450 ft Saussure, AMNH), 1♀, Apr. 9, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 13; Samwell Cave, 40 ° 55 9 N, 122 ° 14 9 2 0 W, June 7, 1995, elev. 1450 ft (D. Ubick, CDU), 1♀, Aug. 30, 1996, elev. 1450 ft (D. Ubick, CDU), 1♀; Shasta Lake Caverns, N side Shasta Lake, 16 mi NNE Redding, Apr. 10, 1979 (D. Rudolph, B. Martin, S. Winterath, AMNH), 1♀. Siskiyou Co.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Ney Springs</a>, 5 mi W Mount Shasta, Sept. 2, 1959 (W. Gertsch, V. Roth, AMNH), 1♀. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Tehama Co.</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Woodson Bridge State Recreation Area</a>, 7 mi E Corning, Aug. 29, 1996, oak camp, at night (D. Ubick, CDU), 1♀. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Trinity Co.</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Big Canyon</a>, 2.6 mi E Camp Trinity, Aug. 12, 1990, under rock (D. Ubick, CDU), 1♀; Del Loma, N fork, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Trinity River</a>, July 25, 1953 (W., J. Gertsch, AMNH), 15♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Forest Glen Cave</a>, on open wall, dry, 75 ft inside cave (R. Graham, AMNH), 23; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.033334&amp;materialsCitation.latitude=40.919167" title="Search Plazi for locations around (long -9.033334/lat 40.919167)">Weaverville</a> to 10 mi E Weaverville, Apr. 6, 1960 (W. Gertsch, W. Ivie, R. Schrammel, AMNH), 23.</p> <p>DISTRIBUTION: Northwestern California (map 3).</p> <p>Map 3. Northern California, showing distributions of three species of Titiotus.</p> <p>(W. Icenogle, WRI), 1♀. Mendocino Co.: near Headquarters, Hopland Field Station, 4 mi E Hopland, Apr. 6, 1972, matured Apr. 28, 1972 (M. Bentzien, UCB), 13. Shasta Co.: Castle Crags State Park, 4 mi W Castella, July 26, 1953 (W., J. Gertsch, AMNH), 1♀; Hazel Creek, July 4, 1952 (W. Gertsch, AMNH), 23, 4♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.75&amp;materialsCitation.latitude=9.0" title="Search Plazi for locations around (long -9.75/lat 9.0)">Potter Creek Cave</a>, 40 ° 47 9 00 0 N, 122 ° 16 9 45 0 W, Nov. 24, 1950 (N. Bostick, AMNH), 13, Sept. 7, 1961 (W. Gertsch, AMNH), 1♀; Samwell Cave, 25 mi N Redding, no date (R. de Saussure, AMNH), 1♀, Apr. 28, 1959, at entrance (AMNH), 1♀, June 5, 1959 (R. de</p> </div>	https://treatment.plazi.org/id/038F87AEFFA5FFE5FD50FCD5A61D598A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFFA3FFD8FCBCFC0DA0D55EC9.text	038F87AEFFA3FFD8FCBCFC0DA0D55EC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus marin Platnick & Ubick 2008	<div><p>Titiotus marin, new species</p> <p>Figures 77–81; map 3</p> <p>TYPE: Male holotype from Mill Valley, Marin Co., California (June 1951; D. Hardy), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males resemble those of T. humboldt and T. costa but can be distinguished by having the third most ventral prong of the tibial apophysis notched (fig. 79); females have a relatively narrow, ovoid epigynal septum (fig. 80).</p> <p>MALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I p0-3-1, r1-2-1; II r0-2-1; III r1-1-2; IV r0-1-2; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2; IV r1-2-1. Embolus prolaterally expanded near tip, distally narrow; median apophysis with only tiny subdistal projection; most ventral prong of tibial apophysis triangular, middle prong relatively short, distally incised, dorsal prong distally narrowed (figs. 77–79).</p> <p>FEMALE: Total length 11. Coloration as in T. californicus. Leg spination: femora: I–III r1-1-2; IV p1-2-1, r0-1-2; tibiae: I p1-0-0, v4-4- 4, r1-0-0; II d1-0-0, p0-1-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi III p1-2-2. Epigynal septum relatively narrow, ovoid, posterolateral epigynal margins protuberant; posterior ducts oblique (figs. 80, 81).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Marin Co.: Burdell Mountain, SE slope, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.15&amp;materialsCitation.latitude=9.8" title="Search Plazi for locations around (long -9.15/lat 9.8)">Novato</a>, 38 ° 07 9 48 0 N, 122 ° 34 9 09 0 W, Mar. 16, 1990, serpentine grassland with adjacent mixed woodland (D.</p> <p>Ubick, T. Briggs, CDU), 13, Mar. 8, 1991, oak woodland, under serpentine (D. Ubick, CDU), 1♀; Fairfax, Nov. 15, 1974 (CAS), 13, 38 ° 00.0 9 N, 122 ° 36.0 9 W, May 6, 1999, in house in oak woodland (J. Schweikert, CAS), 13, Mt. View Road, Apr. 28, 2004 (D. Giampietro, CAS), 13; Marinero Estates, Tiburon Peninsula, Apr. 9, 1994, serpentine grassland (D. Ubick, CDU), 23; Mill Valley, no date, running on floor in house (E. Ross, CAS), 1♀, June 1951 (D. Hardy, AMNH), 13, June 7, 1954 (H. Leech, AMNH), 13, June 17, 1955, at light (H. Leech, AMNH), 13, June 1956 (E. Ross, AMNH), 13, Aug. 1956, in house (E. Ross, AMNH), 1♀; Apr. 1957, in empty bathtub at night (E. Ross, AMNH), 13, June 14, 1957, in house (H. Leech, AMNH), 13, June 1, 1958 (F. Leech, CAS), 13, Sept. 22, 1958 (H. Leech, CAS), 13, Oct. 1, 1958, in house (E. Ross, AMNH), 1♀, Aug. 1, 1963, in house at night (E. Ross, CAS), 1♀, May 1965, house basement (E. Ross, CAS), 23, 1♀, June 6, 1988, in basement of house (C. Hausner, CDU), 1♀, Sept. 5, 1991 (L. Alderson, CDU), 1♀, June 9, 1995 (V. Roth, CAS), 13, June 11, 1996, in house (E. Ross, CAS), 13, Aug. 4, 2002 (S. Hall, UCR), 1♀, May 30, 2003 (E. May, UCR), 1♀; Mount Tamalpais, Mar. 14, 1909 (E. VanDyke, CAS), 13; Novato, May 6, 1973 (B. Kessel, CAS), 13, Mar. 14, 1982, under serpentine (D. Ubick, CDU), 13, Apr. 25, 1982, under serpentine float (D. Ubick, CDU), 13, Jan. 1990, under serpentine (D. Ubick, CDU), 13; Ring Mountain, Tiburon Peninsula, Apr. 15, 1994, serpentine grassland (D. Ubick, J. Boutin, CDU), 13 (penultimate, but palp details visible through cuticle); San Anselmo, May 22, 1998, in house (CAS), 1♀, Aug. 29, 2000, in house (D. Burns, CAS), 1♀; San Geronimo, Jan. 15, 1994 (matured Mar. 24, 1994), serpentine grassland (D. Ubick, J. Boutin, CDU), 13; San Rafael, June 28, 1993, in house (N. Durighello, CDU), 1♀, mid-May 2000, in house (D. Halliday, UCR), 13; N San Rafael, Feb. 6, 1988 (3 matured Mar. 21, 1988, ♀ Apr. 4, 1988), serpentine grassland (D. Ubick, CDU), 13, 1♀; San Rafael Ridge, San Anselmo, May 27, 1996 (L. Freihofer, CAS), 1♀; Sausalito (hilly part), May 23, 2000, in house (A. Weisskopf, CAS), 13; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.1&amp;materialsCitation.latitude=9.083333" title="Search Plazi for locations around (long -9.1/lat 9.083333)">South Tiburon Ridge</a>, 37 ° 53 9 05 0 N, 122 ° 27 9 06 0 W, Feb. 27, 1993, serpentine grassland (D. Ubick, J. Boutin, C. Kellner, CDU), 2♀, Apr. 18, 1993, serpentine grassland (D. Ubick, CDU), 23, 1♀; Terra Linda, San Rafael, 38 ° 01.2 9 N, 122 ° 34.5 9 W, all border of suburb and open space, grassy hills, oaks, laurels, shrubs, elev. 100 m (J. Schweikert, CAS): June 4, 2003, 13, June 22, 2003, 13, July 15, 2003, 13, July 31, 2003, 1♀, Aug. 23, 2003, 1♀, Apr. 13, 2004, 13, Apr. 25, 2004, 13, May 2, 2004, 13, July 18, 2004, 13, May 29, 2005, 13, May 13, 2006, 13; Tiburon, Dec. 21, 1988 (L. Cheng, T. Briggs, CAS), 1♀, mid-April 2002 (K. Passaretti, UCR), 23; Woodacre, Apr. 21, 1981, in house (P. Signet, UCB), 13. Mendocino Co.: Ukiah, June 5, 2000, in house (E. Nicolait, CAS), 13. Sonoma Co.: Petaluma, May 16, 2003, in bathroom (H. Stavropoulos, UCR), 13.</p> <p>DISTRIBUTION: Coastal California, from the San Francisco Bay area north (map 3).</p> </div>	https://treatment.plazi.org/id/038F87AEFFA3FFD8FCBCFC0DA0D55EC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
038F87AEFF9EFFD9FC91FE02A61D5C3D.text	038F87AEFF9EFFD9FC91FE02A61D5C3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Titiotus costa Platnick & Ubick 2008	<div><p>Titiotus costa, new species</p> <p>Figures 82–86; map 3</p> <p>TYPES: Male holotype and female allotype taken in pitfall traps at Canyon, Contra Costa Co., California (Aug. 25, 1981; D. Denning), deposited in AMNH.</p> <p>ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.</p> <p>DIAGNOSIS: Males resemble those of T. humboldt and T. marin but can be distinguished by the distally expanded middle prong of the tibial apophysis (fig. 84), females by the rounded epigynal septum, less protuberant posterolateral epigynal margins, and more massive posterior ducts (figs. 85, 86).</p> <p>MALE: Total length 11. Coloration as in T. flavescens. Leg spination: femora: I, II r1-2-1; III p0-2-1, r1-1-1; IV r1-2-1; tibiae: I d1-0-1, p1-0-0, v4-4-4, r0-1-1; II d1-0-1, v4-4-4, r0-1-1; III r2-0-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2. Embolus prolaterally expanded near tip, distally arched; median apophysis with only tiny subdistal projection; most ventral prong of tibial apophysis triangular, middle prong greatly expanded distally incised, dorsal prong distally narrowed (figs. 82–84).</p> <p>FEMALE: Total length 11. Coloration as in T. flavescens. Leg spination: femora: II p0-2-2, r1-2-1; III r1-1-2; IV p1-2-1; tibiae: I p0-0-0, v4-4-4; II p1-0-0, v4-4-4, r1-0-0; III r1-0-1; metatarsi III p1-2-2. Epigynal septum widest at about half its length, posterolateral epigynal margins only slightly protuberant; posterior ducts massive, triangular (figs. 85, 86).</p> <p>OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Alameda Co.: Berkeley Hills, May 19, 1976, in house (C. Prince, UCB), 13. Contra Costa Co.: Canyon, June 21, 1979, redwood grove (D. Denning, AMNH), 13, July 1–10, 1981 (D. Denning, AMNH), 13, Aug. 25, 1981, pitfalls (D. Denning, AMNH), 13; Moraga, Sept. 14, 1973 (D. Denning, AMNH), 1♀; Tilden Regional Park, pitfalls (J. Fraser, CAS): June 20, 1980, 73, 2♀, July 4, 1980, 33, July 16, 1980, 103, 1♀, Aug. 1, 1980, 53, 3♀, Aug. 15, 1980, 23, 4♀, Aug. 29, 1980, 6♀, Sept. 12, 1980, 4♀, Sept. 26, 1980, 1♀; Univ. of California Richmond Field Station, Richmond, July 11, 1961, ground, on canvas (P. Craig, CDU), 1♀. Napa Co.: N side, Howell Mountain, 2 mi NNE Angwin, all elev. 1300 ft (H. Leech, CAS): June 9, 1975, 13, May 26, 1977, 13, June 19, 1977, 23, 1♀, Aug. 16, 1977, 1♀, July 1, 1978, 13, June 27, 1980, 1♀, Aug. 18, 1980, 1♀; Napa City, May 27, 2000, in linen closet of vacation home, riparian (K. Stockwell, UCR), 13; St. Helena, May 22, 2004, in wine cellar (J. Martel, UCR), 13, 1♀. Sonoma Co.: Boyes Springs, Sept. 19, 1993, in house (J. Dorard, S Meehl, CDU), 1♀; Sonoma, Sept. 12, 2002, in garage (UCR), 1♀; Sonoma Mountain, just E Cotati, June 6, 1973 (F. DeOme, CAS), 13.</p> <p>DISTRIBUTION: Eastern San Francisco Bay region, generally to the east of T. marin (map 3).</p> </div>	https://treatment.plazi.org/id/038F87AEFF9EFFD9FC91FE02A61D5C3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Platnick, Norman I.;Ubick, Darrell	Platnick, Norman I., Ubick, Darrell (2008): A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae). American Museum Novitates 3608 (1): 1-36, DOI: 10.1206/0003-0082(2008)3608[1:AROTEC]2.0.CO;2, URL: http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2008)3608%5B1%3AAROTEC%5D2.0.CO%3B2
