taxonID	type	description	language	source
038E8029FF85FF96FF78D1930A59A76A.taxon	type_taxon	Type species: Phenice bivittata Coquerel, 1859 (Figs 2 – 16) Amended Diagnosis. Moderate to large planthoppers. Head weakly projected, vertex trapazoidal with anterior transverse carina, lateral margins bearing pits; face with frons quadrate, bearing pits relatively broad, lateral margins approximately parallel-sided, median carina absent. Antennae short. Genae bearing large subantennal foliate process. Wings held upright (perpendicular) to body in vivo (Fig. 2). Costal margin of tegmen with sensory pits basally. Forewing C 1 cell elongated, with longitudinal fold (giving appearance of an unattached, spurious vein). Lateral margin of pronotum with carinae (between tegula and head) Hind wing with stridulatory plate at apex of jugal lobe. Pygofer with bilaterally asymmetrical articulated medioventral process, angled dorsad from base. Endosoma bearing 5 processes with varying serration on margins.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF85FF96FF78D1930A59A76A.taxon	description	Amended Description. Body color usually pale, variably marked; wings transparent with extensive dark markings. Length (including wings) ~ 6 - 9 mm. Head. Head from dorsal view narrower or equal to prothorax width. From lateral view, head usually conoid, weakly projected anterodorsal (vertex posteriorly declinate or nearly level). Vertex trapezoidal in dorsal view (approximately 1.3 - 3.5 x longer at midlength than wide at posterior margin, narrowing anteriorly); with margins carinate, anterior margin of vertex (in dorsal view) bearing transverse carina (anterior margin concave between lateral carinae); posterior margin elevated, shallowly concave in dorsal view; lateral margins keeled, bearing two rows of sensory pits (a prominent row of large pits and an irregular second row of small pits) along lateral margins, disc of vertex sunken (relative to lateral margins); median carina usually present, often obscure. Frons quadrate (or nearly so), distinctly longer than wide, lateral margins subparallel (narrowed in F. analarara sp. nov.), disc of frons with row of large sensorial pits along lateral margins, starting just ventrad of vertex, ending before frontoclypeal suture, smaller pits interspersed among larger pits; disc of frons concave, median carina absent. Compound eye (in lateral view) ovoid, deeply emarginated at antenna; dorsal margin with or without horn-like callus; lateral ocelli present, inconspicuous, median ocellus absent. Gena bearing foliate process extending from beneath antenna, curved dorsocaudad, often in shape of a wide bowl. Antennae short, scape very short and inconspicuous, pedicel globular, bearing many sensory plaques, with or without with subapical cuticular eminence and with or without peg-like sensory structure in form of double setae of varying length, located on a cuticular prominence (pedestal); plate organs with basiconic-like projections of two arrangements: lateral ones arranged horizontally, usually fork-shape and vertical in central part of plate organ; plate organs surrounded by a ring of elevated spines partially directed inwards. Frontoclypeal suture almost straight. Clypeus elongated, at frontoclypeal suture as wide as frons; postclypeus with lateral and median carina; anteclypeus with only median carina. Rostrum exceeding hind coxae, apical segment as long as wide. Thorax. Pronotum relatively short (subequal to vertex in midlength), in lateral view anteriorly declinate, mostly raised above level of vertex; from dorsal view anterior margin following contours of head, posterior margin concave; tricarinate with median carina distinct, lateral carinae tracing anterior margin; postocular carina present; paradiscal region proximately quadrate, usually reaching level of ventral margin of antenna, ventral apex rounded. Mesonotum in lateral view convex, raised above level of pronotum, ascending from base and ending at the base of scutellum; disc of scutellum flattened; in dorsal view slightly longer in midline (including scutellum) than wide in lateral angles; lateral angles of the mesonotum placed caudad of midlength (including scutellum); disc of mesonotum tricarinate, median carina reaching scutellum; lateral carinae not connected with median, converging posteriorly, shorter and partly curved not touching posterior margin and not surpassing of the level of lateral angles of mesonotum; scutellum wide with lateral ridges, not continue to the tip of scutellum; tip of scutellum blunt. Hind tibiae slender, longer than femur, without lateral spines; metatibiotarsal formula 5: 5 – 6: 5 – 6. Forewings held upright (perpendicular) to body in vivo (Fig. 2). Forewing distinctly elongated, narrowly spatulate widest near 2 / 3 length, costal and claval margin subparallel, costal margin with sensory pits basally; claval margin about fusion of valval veins concave; posterior margin concave in apical region of MP 1 + 2 field; costal margin with sensory pits at basal part. Basal cell very narrow and elongated; ScP + R + MP arising from leaving basal cell as short common stem and forking distinctly before fusion of claval vein Pcu and A 1. Forking sequence of longitudinal veins ScP + R → CuA → MP; all first forks placed before tip of clavus. Branching pattern RA 2 - branched, RP 3 - branched, MP 5 or 6 - branched (MP 3 + 4 always single), CuA 1 2 - branched; branches of CuA 2 anastomosed to form closed C 5 (procubital) cell with vein CuA 1. Longitidunal vein MP 3 + 4 fused at short distance with CuA 1 a or connected by transverse veinlet. Forewing C 1 cell extremely elongated, with longitudinal fold (giving appearance of an unattached, spurious vein); C 3 cell long without sclerification, C 5 shortest also without sclerification. Cell length sequence C 1 → C 3 → C 5. Transverse veinlet icu present and additional false vein sclerification between CuP and icu present. Clavus closed; ending before midlength of tegmen; composite vein Pcu + A 1 reaching wing margin distinctly before CuP; claval veins Pcu and A 1 joined together before middle of clavus, clavus always without any transverse veinlets but false veins (sclerification) present in some species. Hind wing distinctly narrow and shorter than forewing. Longitudinal vein ScRA with single terminal, RP with single terminal, CuA with 3 branches: CuA 1 2 - branched, CuA 2 single; CuP single; A 1 and A 2 single reaching posterior margin; 2 transverse veinlets r-m (curved) and m-cu (straight) present. Jugal lobe distinctly separated from rest of hind wing with distinct stridulitrum, situated at apex of jugal lobe. Two pairs of large glands on dorsolateral surface of tergits 7 and 8 of both sexes present (Figs 10 A – D). Male terminalia. Pygofer in lateral view narrow, irregular in shape, inclined anteriorly, caudal margin concave (often with dorsolateral angle projected), anterior margin sinuate; in ventral view, medioventral process articulated, elongate and asymmetrical. Gonostyli slightly asymmetrical. in lateral view spatulate, dorsal margin bearing a variably developed proximal dorsal lobe and a hooked process near midlength; in ventral view, gonostyli often medially arched, bearing a variably developed medial process proximally. Aedeagus with shaft weakly upcurved, simple bearing complex endosoma bearing multiple elongated (often flattened, serrated and mostly retrorse) processes. Anal tube from lateral view slender (especially distad of paraproct) and elongated, apex often downcurved; from dorsal view broad (apex sometimes bilobed). Paraproct small, often inconspicuous from lateral view.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF85FF96FF78D1930A59A76A.taxon	discussion	Remarks. Fescennia is recognizable from other Phenicini by the articulated medioventral process. Other genera, such as Paraphenice Muir, Phenice Westwood and Metaphenice Emeljanov possess strongly asymmetrical processes, but they are contiguous with the pygofer (not articulated). Based on wing shape and aedeagal structure, Paraphenice is the most similar genus (from within the Phenicini). Paraphenice are usually pale taxa with wings held tectiform in repose, the vertex is longer than wide and the medioventral process is fixed and symmetrical (Locker et al. 2009, Bahder et al. 2024 b). The frons of Paraphenice tends to be narrower than Fescennia (e. g., Locker et al. 2009, figs. 6 C, 7 C; Bahder et al. 2024 b, fig. 4 C). Interestingly, a genus currently in the tribe Cenchreini (Bourgoin 2025), Aethocauda Williams, is similar to Fescennia as it also possesses an asymmetrical, articulated medioventral process and has a similar structure to the aedeagus with serrated “ fins ” and a similar wing structure. In Williams (1976), it is stated that “ It runs to couplet 15 (Paraphenice Muir, Fescennia Stål) in Fennah’s (1952) key of Cenchreini ”. It appears Aethocauda is incorrectly placed in the Derbinae and belongs in Phenicini of the Otiocerinae. Given the morphological similarities, Aethocauda may be a synonym of Fescennia. Aethocauda is here transferred to the Phenicini. It appears this genus was missed in Emeljanov (1995). Holzinger et al. (2008) reported two species attributed to Fescennia from the Seychelles that were subsequently (Locker et al. 2009) moved to Paraphenice, in the original designation by Muir (1925).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	description	(Figs 2 – 16)	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	materials_examined	Type locality. Mayotte.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	diagnosis	Diagnosis. Rust colored, head in lateral view conical with vertex posteriorly declinate, in dorsal view vertex broad, narrowed anteriorly, frons broadly quadrate and parallel-sided. Antennal pedicel apically invaginated, with dorsal peg and subapical lateral projection on ventroapical margin. Lateral margins of frons parallel, callus absent on dorsum of eye. Forewings with three large, fuscous bands. Medioventral process of pygofer broadly spatulate, apex bifurcated. Aedeagus with multiple broad, fin-like processes, serrated on margin.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	description	Description. Color. Ground color orange, darker on thorax, paler on abdomen, legs testaceous, head white towards anterior margin, yellow near posterior margin, lateral margins of vertex and frons fuscous, fuscous patches on posterior of vertex, clypeus white with yellow wash on anterior margin, sides yellow-orange, apical segment of stylet black, thorax with carinae paler, outlined fuscous, wings with fuscous mottling, abdomen with fuscous mottling (Fig. 3). Structure. Head (Figs 4 – 7). In dorsal view, head just wider than pronotum. Vertex trapezoidal (about 3.5 x wider posteriorly than long at midline); median carina present; lateral margins keeled (bearing rows of pits, one row large and evenly spaced, second row incomplete with small pits, plus few irregularly placed pits), weakly arcuate; anterior margin with transverse carina, distally concave medially, posterior margin convex; disc of vertex medially flat (sunken relative to keeled lateral margins); in lateral view, head anteriodorsally projected for length less than greatest eye width, profile generally rounded, slightly angulate at fastigium, extending slightly above and beyond eyes. In frontal view, lateral margins of frons subparallel, constricting slightly at frontoclypeal suture, row of large sensorial pits along lateral margins, from just ventrad of vertex to frontoclypeal suture, smaller pits interspersed among larger pits. Compound eye, ovoid, deeply emarginated at antenna, dorsal margin without horn-like callus; eye bearing peg-like sensory structures; cuticular eminence at postero-apical margin (Fig. 4). Lateral ocelli obscure near anteroventral margin or eye. Antennal pedicel elongate-rounded, with ventro-apical projection; flagellum directed caudolaterally (hidden from frontal view); peg-like structures present (Fig. 7). Thorax. Pronotum in lateral view declinate anteriorly, raised above level of vertex; in dorsal view narrow, midlength at midline about equal to vertex; anterior margin convex (following contour of posterior head margin), concave on posterior margin; tricarinate, disc concave with ~ 4 nodules on each side of midline, postocular carina distinct. Mesonotum from lateral view raised above level of pronotum, convex; in dorsal view, slightly wider than long at midpoint, tricarinate, mid carina complete to scutellum, lateral carinae serpentine, arising at anterolateral margin, curved posteriorly, obsolete in posterior 1 / 3 (Figs 4 – 6). Hind leg with apical tarsomere about same length (or slightly longer) than middle tarsus, basitarsus about length of telotarsus; metatibio-tarsal formula 5 - 6 - 5 (Fig. 8). Forewing (Fig. 9): branching pattern MP 1 + 2 5 - branched; CuA 1 a fused in short distance with branch MP 3 + 4; clavus without false veins (sclerification). Hind wing (Figs 10, 11) Male terminalia (Figs 12 – 15). Pygofer in lateral view narrow, strongly sinuate, narrowest near midlength, dorsally with caudal margin expanded to form triangular projection on posterior margin, expanded in ventral 1 / 3 to width to approximately equal width as dorsum, medioventral process asymmetrical, articulated at base, angled dorsally, in lateral view, narrowest basally, expanding from articulation point, bifurcated at apex, posterior bifurcation slender, curved toward anterior face, anterior bifurcation wide at base, subtriangular, gently curving anteriorly (Figs 12, 13). Gonostyli in lateral view broadly spatulate, rounded at apex, bearing distinct lateral ridge near dorsal margin for entire length; dorsal margin with hooked process near midlength, hooked posteriorly; proximally veering a small, angulate projection with thick setae (Figs. 12 – 13); in ventral view, broad basally, bearing curved median process proximally with distinct basal ridge, angled mediodorsad; gonostyli constricting distad of processes, the expanded near midlength, apex rounded, large, subtriangular projection arising on inner margin at apex (Figs 12, 13). Aedeagal shaft tubular, simple, weakly upcurved, endosoma complex, asymmetrical, bearing four large processes; first process (E 1) arising subapically on left lateral side, bifurcated proximally into two projections, posterior bifurcation (E 1 a) spinose, directed dorsad, anterior bifurcation (A 2 b) flattened, sinuate on dorsal and ventral margins, retrorse, finely serrulatedat apex, reaching just beyond shaft midlength (Figs 14 – 15); second process (E 2) arising at apex on dorsal margin, retrorse, nearly reaching aedeagal base, elongated and broadly flattened, serrated on dorsal margin near apex, apex cupped (appearing scoop-like) (Figs 14 – 15); third process (E 3) arising on right lateral side of E 1, retrorse, broadly flattened and spatulate, reaching to aedeagal midpoint (Figs 14 – 15); fourth process (E 4) arising at apex on ventral margin (Figs 14 – 15), bifurcated proximally into a pair of flattened, spatulate processes, right lateral process (E 4 a) serrated along apical margins, pinnaform in lateral view, spatulate in dorsal and ventral view, reaching to near midlength of shaft, left lateral bifurcation (E 4 b) narrower and shorter than E 4 a, spatulate, apex with serrated margin in distal 1 / 3 (Figs 14, 15). Anal tube in lateral view marrowly triangular, short, not reaching apex of gonostyli, ventral margin linear, dorsal margin irregularly sinuate (Figs 12, 13). Plant Associations. Pandanus sp. (Pandanaceae).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	distribution	Distribution. Eastern Madagascar; Analalava Forest Reserve, Isle St. Marie / Nosy Boraha. Also reported Mayotte, Comoros Islands, and Madagascar (Coquerel 1859, Stål 1866, Jacobi 1917)	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	materials_examined	Material Examined. Madagascar, Alaotra-Mangoro Region, Analalava Forest Reserve, 27. I. 2023, 24 males, 5 females (FLREC), 14 males, 12 females (MIIZ); Nosy Boraha (as. St. Marie), Voeltzkow [collector] (6 males, ASU) (Fig. 16).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF8DFF8AFF78D5A80A46A626.taxon	discussion	Remarks. This species was described from Mayotte, subsequently reported from Nosy Boraha (as Sainte- Marie), Comoros Islands, and Madagascar (Coquerel 1859, Stål 1866, Jacobi 1917, Metcalf 1945). The Comoros Island record from Jacobi (1917) may be an artifact from Jacobi attributing Mayotte as part of the Comoros; however, if the asserted distribution of the species is as widespread as it appears, F. bivittata is likely to be correctly attributed to the islands. The Coquerel collection was most likely deposited at the Société entomologique de France, which was mostly integrated into the collection of the Muséum National d'Histoire Naturelle in Paris, but the type of this species was not found. Muir (1925: 472) stated that this type was in the Signoret collection based on Stål (1866: 199), which is located in part in the Naturhistorisches Museum Wien (Natural History Museum, Vienna), but the specimens have not been located. Fescennia bivittata is characterized by a combination of the shape of the head, wing pattern and coloration of the mesonotum. The distinct banding on the wings, the angled head and nearly parallel margins of the frons, and rust colored mesonotum allow for this species to be distinguished from others. In addition, the structure of the medioventral process of the pygofer and aedeagus separates this species from congeners.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	description	(Figs 17 – 22)	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	materials_examined	Type Locality. Madagascar, Alaotra-Mangoro Region, Analalava Forest Reserve.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	diagnosis	Diagnosis. Overall body coloration orange. In lateral view, head rounded, vertex nearly level. In dorsal view, lateral margins of frons relatively short and narrowed ventrally. Pedicel of antenna, not apically invaginated (dorsal peg and subapical lateral projection absent). Forewings with fuscous areas diffuse (without three large bands). Medioventral process of pygofer slender, bifurcated, serrate on dorsal margins. Aedeagus with slender, serrate fin-like processes.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	description	Description. Color. Uniform orange, darker dorsally (Fig. 17), head paler, orange wash along posterior margin, clypeus pale, darker on lateral margins, pronotum orange, paler anteriorly and carinae. Tegulae white (fuscous dorsad) (Fig. 18). Forewings translucent with fuscous mottling (Fig. 19). Structure. Head. Vertex trapezoidal transversely (Fig. 18); about 3.4 x wider posteriorly than length at midline; anterior margin (between eyes) weakly convex, lateral margins arcuate, subparallel, posterior margin concave, two rows of sensorial pits on lateral margins (Fig. 18) (outer pits large, inner pits small and irregularly distributed) (Fig. 18), in lateral view, head angled at fastigum, profile rounded from fastigium to frontoclypeal suture, subantennal process extended from beneath antenna, arched dorsad approximate to hind margin of eye. In frontal view, frons relatively short (midlength about 1.4 x greatest width), with lateral margins narrowed from fastigium, bearing row of sensorial pits on lateral margins (Fig. 18). Eye without dorsal callus; peg-like sensory structures absent; posteroapically without cuticular eminence (Fig. 18). Antennal pedicel columnar with apical flagellum; projection and peg-like structures absent. Thorax. Pronotum in lateral view raised above level of vertex, anteriorly declined; in dorsal view, convex anteriorly (between eyes), deeply concave posteriorly, narrowest at midpoint, postocular carina fully developed (Figs 15 B, 16 A). Mesonotum in dorsal view approximately 1.5 x longer than wide at midpoint, tricarinate, median carina extending from anterior to scutellum, lateral carinae serpentine, arising at anteriolateral margin, curved posteriorly, obsolete in posterior 1 / 3 (Fig. 18). Metatibio-tarsal formula of hind leg 5 - 5 - 5. Forewing (Fig. 19): branching pattern MP 1 + 2 4 - branched; CuA 1 a connected with branch MP 3 + 4 by transverse veinlet; clavus without false veins (sclerification). Male terminalia (Figs. 20 – 22). Pygofer in lateral view strongly sinuate at anterior and posterior margins, narrowest medially, dorsolateral margin expanded into a large rounded lobe (Fig. 20); medioventral process angled dorsally, in lateral view, narrowest basally, expanding distally, bifurcated at apex, posterior process slender, sinuate, serrated on anterior margin, pointed at apex, anterior process slender, approximately half the length of posterior bifurcation, truncate at apex, serrated on anterior margin (Fig. 20). Gonostyli in lateral view irregularly spatulate irregularly sinuate on ventral margin, apex angulate, dorsal margin with pair of projections near midlength, proximal projection a rounded lobe, skewed cephalad, posterior projection a sclerotized hook, angled cephalad (Fig. 20); in ventral view, inner and outer margins irregularly sinuate, medial margin with large hooked process (sclerotized distally) near midlength and bearing a slender tooth on caudal margin; subapical medial margin expanded (creating a deep concavity between medial tooth and subapical region). Medial margin bearing large denticles (gonostyli asymmetrical apically) (Fig. 20). Aedeagal shaft irregular, margins sinuate, endosoma complex, asymmetrical with five retrorse processes; first process (E 1) arising subapically on dorsal margin, left of midline, curved over midline to right lateral side, angled slightly ventrad, dorsal margin serrated near midlength; second process (E 2) arising subapically on left lateral side, directed dorsad and cephalad, dorsal margin serrated in distal half, ventral margin with a few larger teeth; third process (E 3) arising subapically on dorsal margin, elongated, nearly reaching aedeagal base, dorsal and left lateral margin serrated in distal half; fourth process (E 4) arising subapically on ventral margin, right of midline, angled ventrad and cephalad, curving mesad to midpoint, right lateral margin serrated; fifth process (E 5) arising subapically on ventral margin near E 4, strongly bifurcated, inner bifurcation (E 5 a) short, slender, outer bifurcation (E 5 b) elongated, reaching adeagal base, lateral margins at apex serrated (Figs 21, 22). Anal tube in lateral view slender, narrowly triangular, nearly reaching apex of gonostyli, ventral margin sublinear, slightly convex, dorsal margin irregularly sinuate, apex curved ventrad (Fig. 20). Plant Associations. Pandanus sp. (Pandanaceae).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	distribution	Distribution. Madagascar, Alaotra-Mangoro Region, Analalalava Forest Reserve.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	etymology	Etymology. The specific name analarara is an amalgamation of Analalava and the Latin word for rare “ rarus ” due to only finding one specimen despite extensive surveying.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF91FF8EFF78D5E40DC9A74E.taxon	materials_examined	Material Examined. Holotype ♂, Madagascar, Alaotra-Mangoro Region / Analalava Forest Reserve / 27. I. 2023, ♂ / Coll.: B. W. Bahder / Host: Pandanus sp., Fescennia analarara (FLREC).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	description	(Figs 23 – 28)	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	materials_examined	Type Locality. Madagascar, Atsinanana Region, Parc Mitsinjo.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	diagnosis	Diagnosis. Base color pale, irregular fuscous markings. Head in lateral view slightly conical, vertex weakly convex. In dorsal view, vertex broad, narrowed anteriorly. Pedicel of antenna bearing a peg-like process on dorsal margin. Eye with conspicuous horn’like callus on dorsal margin. Forewings with four fuscous bands in dorsal view. Medioventral process of pygofer slender, trifurcated and serrate. Aedeagus thick with narrow, arched with serrate projections.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	description	Description. Color. Ground color varies from tan to white, head generally white, orange wash at ventral margin of genae, clypeus anterior margin white, lateral margins orange, mesonotum orange with fuscous wash (paler medially), forewings transluscent with yellow and fuscous mottling, fuscous band on lateral margins of abdomen, tergites orange (Fig. 23). Structure. Head. Head subequal in width to pronotum. Vertex relatively broad, trapezoidal (about 2.9 x wider posteriorly than long at midline). Anterior margin nearly truncate, posterior margin concave medially (almost straight); lateral margins weakly arcuate, subparallel, bearing two rows of sensorial pits, outer pits approximately twice as large as inner row (Fig. 24). In lateral view, head profile angled at fastigum, fastigium slightly projected, gently rounded from fastigium to frontoclypeal suture; vertex weakly convex. Gena with subantennal processes concave from beneath antenna to beneath eye (removed from ventral margin of eye). In frontal view, frons with lateral margins nearly parallel-sided (weakly constricted ventrally) bearing row of sensorial pits with smaller sensorial pits inset (Fig. 24). Antennal pedicel globular with flagellum at apex, without projections, bearing a peg-like sensory process dorsally. Eye deeply emaginate at antenna, dorsal margin bearing horn-like callus; postero-apical margin without cuticular eminence (Fig. 24). Lateral oceelus just anterior to ventroapical margin of eye. Thorax. Pronotum in dorsal view about as long at midlength as vertex. convex at anterior margin (between eyes), deeply concave at posterior margin, narrowest medially, tricarinate (Fig. 24). Mesonotum in dorsal view approximately 1.5 x longer than wide at midpoint, tricarinate with median carina reaching scutellum, lateral carinae arising laterally, running dorsally, curving mesally at dorsal margin, obsolete in posterior 1 / 3 (Fig. 24). Hind leg with metatibio-tarsal formula 5 (fused at base) / 5 / 5 Forewing (Fig. 25): branching pattern MP 1 + 2 5 - branched; CuA 1 a fused in short distance with branch MP 3 + 4; clavus with false veins (fold). Male terminalia (Figs. 26 – 28). Pygofer in lateral view narrow with strongly sinuate anterior and posterior margins, narrowest medially, dorsocaudal margin expanded into rounded lobe (Fig. 26), medioventral process articulated, angled dorsad; in lateral view, relatively slender, bearing basal knob on dorsal surface, bifurcated near midlength and bearing a flattened, serrated secondary process arising on left lateral side; apical bifurcation with dorsal process arched, directed caudad, acuminate with dorsal and ventral margins serrulated; ventral process expanded proximally, then narrowed to acuminate, finely serrulated apex (Fig. 26). Gonostyli in lateral view broad, spatulate, ventral margin smoothly convex, apex rounded, dorsal margin irregularly sinuate, bearing low, founded dorsal process proximally, and distal hook-like process (apically sclerotized) (Fig. 26); in ventral view, gonostyli weakly asymmetrical, medial margins concave bearing a large elongated sclerotized process distal medial margin bearing low, rounded lobes (Fig. 26). Aedeagus with shaft simple, smoothly sinuate, nearly straight, expanded distally to appear club-like; endosoma asymmetrical, bearing four retrorse processes (each of these bifurcated); first (E 1) arising subapically on dorsal margin, bifurcated near base, inner bifurcation (E 1 a) short, slender, curved; outer bifurcation (E 1 b) 4 x as long as E 1 a, sinuate on inner margin, curved mesad, distal half serrated (Figs 27, 28); second process (E 2) arising subapically on left lateral side, bifurcated proximally with dorsal process curved gently ventrad, slender, nearly reaching aedeagal base, ventral process a broad ventral lobe (Fig. 27 – 28); third process arising subapically on right lateral side, angled ventrad, ventral margin serrated at apex (Figs 27, 28), fourth process (E 4) arising subapically on ventral margin, curving to left lateral side, angled cephalad, bifurcated with dorsal branch (E 4 a) irregularly sinuate, subapically expanded into serrated lobe (visible from dorsal or ventral view); ventral branch half length of dorsal. serrated in distal half (Figs 27, 28). Anal tube, in lateral view, narrowly quadrate with distal portion narrowed beyond paraproct, apexs downcurved, nearly reaching apex of gonostyli (Fig. 26). Plant Associations. Pandanus sp. (Pandanaceae).	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	distribution	Distribution. Eastern Madagascar, Atsinanana Region, Parc Mitsinjo.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	etymology	Etymology. The specific name is derived from the Latin word ‘ cornigera ’ meaning ‘ horn-bearing’ (referring to the horn-like callus above the eye). The name is given the feminine termination (- a) to match the gender of the genus.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
038E8029FF95FF82FF78D47C0B32A38B.taxon	materials_examined	Material Examined. Holotype ♂, Madagascar, Atsinanana Region / Parc Mitsinjo / 30. I. 2023, ♂ / Coll.: B. W. Bahder / Host: Pandanus sp., Fescennia cornigera (FLREC). Paratypes (n = 2) same as holotype.	en	Stroiński, Adam, Łukasik, Piotr, Bartlett, Charles R., Pilet, Fabian, Hasinjaka, Rasolondalao Harin’Hala, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Bahder, Brian W. (2025): New planthoppers in the genus Fescennia (Hemiptera: Derbidae) from screw palms (Pandanus spp.) in eastern Madagascar, with redescription of F. bivittata and an updated key to the genus. Zootaxa 5729 (1): 75-104, DOI: 10.11646/zootaxa.5729.1.3, URL: https://doi.org/10.11646/zootaxa.5729.1.3
