identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038E87ACFFBB042DFF26FAB201B9ABF0.text	038E87ACFFBB042DFF26FAB201B9ABF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asceua Thorell 1887	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Asceua Thorell, 1887</p>
            <p> Type species.  Asceua elegans Thorell, 1887 , by monotypy. </p>
            <p>Diagnosis. For genus description and diagnosis, see Jocqué (1991).</p>
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	https://treatment.plazi.org/id/038E87ACFFBB042DFF26FAB201B9ABF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFBB0424FF26F99A00E4AD08.text	038E87ACFFBB0424FF26F99A00E4AD08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asceua cingulata (Simon 1905)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Asceua cingulata (Simon, 1905)</p>
            <p>Figs 1–6, 17</p>
            <p> Suffucia cingulata Simon, 1905: 174 (♀).— Gravely, 1921: 409, fig. 2d (♁ ♀). </p>
            <p> Type material.  Lectotype (here designated) ♀ and paralectotypes 2 ♀♀ from INDIA: Puducherry (formerly Pondicherry), 11°56’N 79°48’E, 5 m a.s.l., date unknown, leg. M.M. Maindron, repository MNHN, Paris (AR9978), examined based on photographs.</p>
            <p>
                 Other material examined.  INDIA: Andaman Islands: Ross Island, 11°40’N 92°45’E, 41 m a.s.l., 29 March 1911, leg. Paiva: 2 ♀♀ (NZC-ZSI 2718 /H2). West Bengal: Hooghly, Serampore, 22°45’N 88°20’E, 9 m a.s.l., July 1919, leg. Drake: 1 ♀ (NZC-ZSI 2719 /H2). Tamil Nadu: Chennai (formerly Madras), 13°04’N 80°16’E, 4 m a.s.l., 12 March, leg. Ramuni Menon: 1 ♀ (only prosoma) (NZC-ZSI 2720 /H2).  Odisha (formerly Orissa): Ganjam,  
                <a title="Search Plazi for locations around (long 76.20617/lat 11.26838)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.20617&amp;materialsCitation.latitude=11.26838">Chilika Lake</a>
                 (=  
                <a title="Search Plazi for locations around (long 76.20617/lat 11.26838)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.20617&amp;materialsCitation.latitude=11.26838">Chilka Lake</a>
                 ),  
                <a title="Search Plazi for locations around (long 76.20617/lat 11.26838)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.20617&amp;materialsCitation.latitude=11.26838">Barkuda Island</a>
                 (formerly part of Madras Presidency), 19°33’N 85°08’E, 13 m a.s.l., 3-19 August 1919, leg. F.H. Gravely: 2 ♁♁, 4 ♀♀ (NZC-ZSI 2721 /H2).  Kerala: Malappuram, Nilambur, Canolly’s Plot, 11°16’06.17’’N 76°12’22.21’’E, 20 m a.s.l., 16 May 2013, on the ground, by hand, leg. M.S. Pradeep: 3 ♀♀, 1 subadult ♁, 1 juvenile (ADSH192) ;   Ernakulam, Karimpani,  
                <a title="Search Plazi for locations around (long 76.666664/lat 10.133333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.666664&amp;materialsCitation.latitude=10.133333">Bhoothathankettu Forest Reserve</a>
                 , 10°08’N 76°40’E, 24 m a.s.l., 8 September 2013, on the ground, by hand, leg. M.S. Pradeep: 1 ♀ (ADSH193)  ;   Ernakulam,  
                <a title="Search Plazi for locations around (long 76.55016/lat 10.198873)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.55016&amp;materialsCitation.latitude=10.198873">Illithodu</a>
                 , 10°11’55.94’’N 76°33’00.57’’E, 24 m a.s.l., 25 February 2014, on the ground, by hand, leg. M.S. Pradeep: 1 ♁, 2 ♀♀, 1 juvenile (ADSH194)  . 
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            <p> Diagnosis. Males of  A. cingulata are closely related to those of  A. thrippalurense sp. nov. by the shape of retrolateral, ventral and prolateral tibial apophyses, cymbium with basomedial conical process extending between ventral and retrolateral apophyses and embolus with similar course of running, but can be distinguished by the smaller cymbial retrolateral fold with less modified basoretrolateral extremity (vs. highly modified in  A. thrippalurense sp. nov. ), anterolateral margin of cymbial fold less prominent (vs. prominent in  A. thrippalurense sp. nov. ), retrolateral process of conductor directed at 3-o’ clock ventrally and basally acute (vs. 2-o’ clock and basally rounded in  A. thrippalurense sp. nov. ), median apophysis with long proximal part (vs. short in  A. thrippalurense sp. nov. ), and embolus with distal loop (vs. absent in  A. thrippalurense sp. nov. ) (compare Figs 4A–C, 5A–D with Figs 9A–C, 10A–D). Females are closely related to the females of  Asceua similis Song &amp; Kim, 1997 in the general shape and orientation of copulatory ducts, but can be separated from the latter by epigyne with anterior plate (vs. absent in  A. similis ), and spherical spermathecae (vs. irregular in  A. similis ) (compare Figs 4D–F, 5E–F with Song &amp; Kim 1997: figs 14–15). </p>
            <p>Redescription. Male in alcohol (ADSH194) (Figs 2A–J). Body length 1.57. Carapace 0.87 long, 0.70 wide. Opisthosoma 0.74 long, 0.62 wide. Carapace, eye region, clypeus, chilum, chelicerae, endites, labium, sternum yellowish brown; leg segments creamy-white to yellowish brown; palp segments yellowish brown to brown; dorsal scutum brownish, opisthosoma shiny, greyish black with two pairs of large dull yellow patches (Fig. 2D), lateral and rear end black with creamy-white patches, streaks and dots (Figs 2C–D), venter creamy-white with yellow anterior part (Fig. 2B); spinnerets creamy-white (Fig. 2H); carapace, clypeus, chelicerae, margins of sternum, leg segments with black mottling. Carapace glossy; cephalic part without long black setae. Fovea short, longitudinal, straight, dark brown. Clypeus high (Fig. 2E). Chilum inverted triangular, unipartite (Fig. 2E). Cheliceral promargin with two tiny teeth, retromargin without tooth visible (Fig. 2F). Sternum glossy, shield-shaped with posterior end truncated, rebordered, provided with scattered black setae, with coxal and intercoxal extensions (Fig. 2G). Opisthosoma oval in dorsal view, dorsal scutum complete, without long black setae (Fig. 2D); rear and lateral opisthosoma with corrugations. Colulus and anal tubercle prominent (Figs 2C, H). Femur I without retrolateral striae; metatarsi II–III with distal preening brush (Fig. 2I); all metatarsi and tarsi without scopula (Fig. 2J); all tarsi with reduced claw tuft (Figs 2I–J). Eye diameters and interdistances: ALE 0.06, AME 0.08, PLE 0.07, PME 0.05; AME–ALE 0.03, AME– AME 0.05, AME–PME 0.05, ALE–PLE 0.04, PME–PLE 0.08, PME–PME 0.07. Clypeus height at AMEs 0.29, at ALEs 0.24. Chelicerae 0.25 long. Sternum 0.45 long, 0.44 wide. Measurements of palp and legs: palp 1.02 [0.35, 0.15, 0.10, 0.42], I 2.54 [0.66, 0.23, 0.61, 0.62, 0.42], II 2.10 [0.59, 0.23, 0.45, 0.50, 0.33], III 2.03 [0.58, 0.20, 0.41, 0.54, 0.30], IV 2.79 [0.72, 0.23, 0.62, 0.83, 0.39]. Leg formula: 4123. Spination of palp: femur do 2, patella spineless, tibia spineless, tarsus/cymbium spineless; legs: femora I–IV do 1; patellae I–IV spineless; tibiae I–IV spineless; metatarsi I–III spineless, IV plv 1 rlv 1; tarsi I–IV spineless. Palp (Figs 4A–C, 5A–D). Tibia with short retrolateral, broad ventral and narrow, finger-like prolateral apophyses (Figs 4A, C, 5A, C–D); RTA having triangular distal part (Fig. 5C); RTA and VTA enclose large concavity in which fits basomedial projection of cymbium (Figs 4C, 5C). Cymbium modified, with prominent cymbial retrolateral fold having less prominent anterolateral margin and less modified posteroretrolateral extremity (Figs 4B–C, 5B–D), with basomedial conical process extending between RTA and VTA (Fig. 5C). Tegulum broad, with posteromedian extension (Figs 4B, 5B). Median apophysis with broad proximal and long, slender, prolaterally oriented distal parts (Figs 4A–C, 5A–C), lying adjacent to retrolateral process of conductor (Figs 4B, 5B). Conductor broad, sclerotized, with retrolateral and mediolateral, blunt processes (Figs 4B, 5B). Embolus long, filiform, basal to tegulum, with short base, at first running anteriad along prolateral side of cymbium, then turning retrolaterad, then to posterolaterad forming distal loop, with blunt tip directed at 1-o’ clock ventrally (Figs 4A–B, 5A–B).</p>
            <p>Female in alcohol (ADSH194) (Figs 3A–I). Body length 2.00. Carapace 0.96 long, 0.68 wide. Opisthosoma 1.04 long, 0.88 wide. Habitus and details as in male except for the following: opisthosoma without scutum and corrugations (Fig. 3A); dorsum with three pairs of large patches, numerous tiny spots, and a single median patch at the rear end in front of anal tubercle, all creamy-white (Fig. 3A); laterals without stripes or dots (Fig. 3C); venter uniformly creamy-white (Fig. 3B). Palp segments yellowish brown to brown. Eye diameters and interdistances: ALE 0.06. AME 0.07. PLE 0.04. PME 0.05; AME–ALE 0.04. AME–AME 0.03. AME–PME 0.07. ALE–PLE 0.04. PME–PLE 0.08. PME–PME 0.07. Clypeus height at AMEs 0.25, at ALEs 0.21. Chelicerae 0.33 long. Sternum 0.45 long, 0.43 wide. Measurements of palp and legs: palp 0.85 [0.27, 0.16, 0.16, 0.26], I 2.28 [0.64, 0.21, 0.54, 0.53, 0.36], II 1.94 [0.55, 0.22, 0.40, 0.48, 0.29], III 2.00 [0.54, 0.22, 0.40, 0.53, 0.31], IV 2.70 [0.69, 0.22, 0.61, 0.79, 0.39]. Leg formula: 4132. Spination of palp: tibia pld 1; tarsus pl 2 pld 1 v 2; legs: metatarsi II–IV plv 1 rlv 1. Genitalia (Figs 4D–F, 5E–F). Epigyne with convex posterior margin, with an anteromedial, sclerotized, transversely oval plate (Figs 4D–E, 5E). Copulatory openings situated anteriorly behind epigynal plate, lying close to each other (Figs 4E, 5E). Copulatory ducts long, slender, with multiple windings (Figs 4F, 5F). Spermathecae small, globular, lying adjacent to each other, situated posteriorly (Figs 4F, 5F). Fertilization ducts long, narrow, diverging (Figs 4F, 5F).</p>
            <p>Variation. For checking variation in measurements, only the freshly collected specimens were considered. Female (n=6): body length 1.80–2.08. One of the female specimens collected from Malappuram presents a variation in the abdominal pattern as shown in Fig. 3J, and its epigyne also shows variation in the shape of the epigynal plate as shown in Fig. 4G.</p>
            <p>Remarks. All the specimens deposited in the NZC-ZSI, Kolkata are severely dried off due to bad preservation (Figs 6A–D).</p>
            <p>Distribution. India (Andaman Islands (Rose Island), Kerala (new record), Odisha, Puducherry, Tamil Nadu, West Bengal) (Gravely 1921; present data) (Fig. 17).</p>
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	https://treatment.plazi.org/id/038E87ACFFBB0424FF26F99A00E4AD08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFB20421FF26FC090159AA4C.text	038E87ACFFB20421FF26FC090159AA4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Asceua thrippalurense Sankaran 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Asceua thrippalurense sp. nov.</p>
            <p>Figs 7–10, 17</p>
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                 Type material.   Holotype ♁ (ADSH196) from INDIA: Kerala: Palakkad, Thrippalur,  
                <a title="Search Plazi for locations around (long 76.56469/lat 10.6379385)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.56469&amp;materialsCitation.latitude=10.6379385">Pullodu</a>
                 , 10°38’16.58’’N 76°33’52.87’’E, 70 m a.s.l., 24 January 2016, on the ground, by hand, leg. M.S. Pradeep.  Paratype: 2 ♀♀ (ADSH197), same data as for the holotype . 
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            <p>Etymology. The specific epithet is an adjective and referring to Thrippalur, where the type locality of the new species is located.</p>
            <p> Diagnosis. Males are similar to  A. cingulata , but can be distinguished from it as stated above (compare Figs 9A–C, 10A–D with Figs 4A–C, 5A–D). Females are closely related to the females of  Asceua wallacei Bosmans &amp; Hillyard, 1990 as both share spherical spermathecae and copulatory ducts with prominent windings near copulatory openings, but can be separated from the latter species by epigyne with anteromedian pore (vs. absent in  A. wallacei ), and without anteromedian process (vs. present in  A. wallacei ) (compare Figs 9D–F, 10E–F with Bosmans &amp; Hillyard 1990: figs 54–55). Females of  A. thrippalurense sp. nov. can easily be distinguished from the females of  A. cingulata by the presence of longitudinal striae retrolaterally on femur I (Fig. 8H) which is absent in  A. cingulata . </p>
            <p>Description. Male in alcohol (holotype, ADSH196) (Figs 7A–J). Body length 2.06. Carapace 1.09 long, 0.88 wide. Opisthosoma 0.97 long, 0.73 wide. Carapace, eye region, clypeus dark brown; chilum, chelicerae, endites, labium, sternum brown; leg and palp segments yellowish brown to creamy-white; dorsal and lateral opisthosoma black with creamy-white streaks on laterals, dorsal scutum shiny, brownish black (Fig. 7A), venter creamy-white with yellowish brown and greyish patches; spinnerets creamy-white with black shades (Fig. 7G). Carapace finely rugose; cephalic part provided with scattered few long black setae. Fovea short, longitudinal, straight, dark. Clypeus high (Fig. 7D). Chilum inverted triangular, unipartite (Fig. 7D). Cheliceral promargin with two tiny teeth, retromargin without tooth visible (Fig. 7E). Sternum rebordered, rugose, shield-shaped with posterior end truncated, provided with scattered greyish black setae, with coxal and intercoxal extensions (Fig. 7F). Opisthosoma oval, scutum covering 2/3 rd of dorsum area (Fig. 7A), covered with long black setae; rear and lateral opisthosoma with corrugations. Colulus and anal tubercle prominent (Figs 7A, G). Femur I retrolaterally provided with longitudinal striae almost along its entire length (Fig. 7H); metatarsi II–III with distal preening brush (Fig. 7I); all metatarsi and tarsi without scopula (Fig. 7J); all tarsi with reduced claw tuft (Figs 7I–J). Eye diameters and interdistances: ALE 0.08, AME 0.10, PLE 0.07, PME 0.05; AME–ALE 0.02, AME–AME 0.03, AME–PME 0.07, ALE–PLE 0.03, PME–PLE 0.08, PME–PME 0.09. Clypeus height at AMEs 0.32, at ALEs 0.30. Chelicerae 0.28 long. Sternum 0.54 long, 0.52 wide. Measurements of palp and legs: palp 1.36 [0.42, 0.17, 0.13, 0.64], I 3.22 [0.87, 0.27, 0.79, 0.82, 0.47], II 2.55 [0.73, 0.28, 0.54, 0.64, 0.36], III 2.46 [0.68, 0.26, 0.52, 0.67, 0.33], IV 3.54 [0.93, 0.28, 0.81, 1.08, 0.44]. Leg formula: 4123. Spination of palp: femur rld 1, patella do 1, tibia spineless, tarsus/cymbium spineless; legs: femora I–IV do 1; patellae I–IV spineless; tibiae I–IV spineless; metatarsus I spineless, II–IV plv 1 rlv 1; tarsi I–IV spineless. Palp (Figs 9A–C, 10A–D). Tibia with short retrolateral, broad ventral and narrow, finger-like prolateral apophyses (Figs 9A, C, 10A–D); RTA with prolaterally oriented distal part (Figs 9C, 10C); RTA and VTA enclose large concavity in which fits basomedial projection of cymbium (Figs 9C, 10C). Cymbium with large retrolateral fold having prominent anterolateral margin and highly modified basoretrolateral extremity (Figs 9B–C, 10B–C), with basomedial conical process extending between RTA and VTA (Figs 9C, 10C). Tegulum broad, with basomedian extension (Figs 9B, 10B). Median apophysis with short proximal and long, slender, prolaterally oriented distal parts (Figs 9B–C, 10A– C), lying adjacent to retrolateral process of conductor (Figs 9B, 10B). Conductor broad, sclerotized, with retrolateral and mediolateral, blunt processes (Figs 9B, 10B); retrolateral process anteriorly oriented (Figs 9B, 10B). Embolus long, filiform, with short embolar base originated basally to tegulum, at first running anteriad along prolateral side of cymbium, then turning retrolaterad, then to posterolaterad, then to anteroretrolaterad without forming distal loop, with blunt tip directed at 1-o’ clock ventrally (Figs 9A–B, 10A–B).</p>
            <p>Female in alcohol (paratype, ADSH197) (Figs 8A–J). Body length 2.46. Carapace 1.04 long, 0.88 wide. Opisthosoma 1.42 long, 1.00 wide. Habitus and details like male except for the following: leg and palp segments yellowish brown to light brown. Opisthosoma without scutum and corrugations (Fig. 8A); dorsum with three pairs of creamy-white patches, anterior C-shaped, median V-shaped and posterior dot-like (Fig. 8A); laterals black and creamy-white without streaks; venter uniformly creamy-white with black at the rear end. Eye diameters and interdistances: ALE 0.07. AME 0.08. PLE 0.04. PME 0.05; AME–ALE 0.02. AME–AME 0.04. AME–PME 0.09. ALE–PLE 0.05. PME–PLE 0.10. PME–PME 0.09. Clypeus height at AMEs 0.33, at ALEs 0.28. Chelicerae 0.42 long. Sternum 0.58 long, 0.55 wide. Measurements of palp and legs: palp 1.04 [0.34, 0.20, 0.21, 0.29], I 2.97 [0.82, 0.30, 0.67, 0.72, 0.46], II 2.44 [0.66, 0.28, 0.52, 0.62, 0.36], III 2.51 [0.69, 0.28, 0.52, 0.67, 0.35], IV 3.42 [0.91, 0.30, 0.78, 1.01, 0.42]. Leg formula: 4132. Spination of palp: tibia pld 1; tarsus pl 1 pld 1 v 2; legs: metatarsi I–III spineless. Genitalia (Figs 9D–F, 10E–F). Epigyne with W-shaped posterior margin, with a central pore leading to copulatory openings (Figs 9D–E, 10E). Copulatory openings contiguous, situated inside the central pore. Copulatory ducts long, slender, with multiple windings (Figs 9F, 10F). Spermathecae large, globular, lying adjacent to each other, situated posteriorly (Figs 9F, 10F). Fertilization ducts long, narrow, diverging (Figs 9F, 10F).</p>
            <p>Distribution. Known only from the type locality (Fig. 17).</p>
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	https://treatment.plazi.org/id/038E87ACFFB20421FF26FC090159AA4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFB70421FF26FB470108A844.text	038E87ACFFB70421FF26FB470108A844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capheris Simon 1893	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Capheris Simon, 1893</p>
            <p> Type species.  Capheris crassimana (Simon, 1877) , by subsequent designation. </p>
            <p> Diagnosis.  Capheris species are very similar to those of  Cydrela in habitus and genitalic morphology, but can be distinguished from those of  Cydrela by profile of carapace without deep concavity at level of fovea (vs. present in  Cydrela ), the ALEs are situated in front of the AMEs, touching each other or almost separated by their radius (vs. distinctly separated by more than their radius in  Cydrela ), PER straight or slightly curved in dorsal view (vs. curved in  Cydrela ), sternum triangular or shield-shaped, usually wider than long (vs. usually slightly longer than wide in  Cydrela ), sternum anterolaterally with deep indentations accommodating the endites (vs. almost straight or with small indentations in  Cydrela ), sternum with precoxal sclerites (vs. absent in  Cydrela ), and very stout palp as large or larger than leg femora (vs. normal and not enlarged in  Cydrela ) (Jocqué 1991, 2009; Henrard 2019, PhD Thesis). For a description of the genus, see Jocqué (1991, 2009). </p>
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	https://treatment.plazi.org/id/038E87ACFFB70421FF26FB470108A844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFB70423FF26F94F006FAD94.text	038E87ACFFB70423FF26F94F006FAD94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capheris escheri Reimoser 1934	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Capheris escheri Reimoser, 1934 comb. rest. </p>
            <p>Figs 11, 18</p>
            <p> Capheris escheri Reimoser, 1934: 467 , fig. 1 (♀). </p>
            <p> Cydrela escheri — Jocqué, 2009: 123 (transfer from  Capheris ). </p>
            <p>
                 Type material.   Holotype ♀ from INDIA: Tamil Nadu: Dindigul,  
                <a title="Search Plazi for locations around (long 77.36667/lat 10.266666)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.36667&amp;materialsCitation.latitude=10.266666">Kodaikanal</a>
                 , from small shola near kukkal/kookal, 10°16’N 77°22’E, 1972 m a.s.l., leg. J. Carl &amp; K. Escher, 1926-1927, repository MHNG, Geneva (ARTO-26290), examined based on photographs. 
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            <p> Diagnosis. Females of  C. escheri are very similar to the females of  Capheris oncka Lawrence, 1927 as both share semi-circular epigynal plate and short basomedian septum, but can be separated from the latter species by copulatory openings with straight anterior rim (vs. semi-circular in  C. oncka ) (compare Fig. 11G with Jocqué 2009: fig. 173). </p>
            <p>Supplementary description. Female (holotype) (Figs 11A–F) (partly after Reimoser (1934)). Body length 5.90. Carapace 3.00 long, 1.50 wide. Opisthosoma 2.90 long, 1.70 wide. Carapace yellowish brown, slightly domed, without deep concavity at level of fovea (Fig. 11D); endites, sternum yellowish red; legs reddish yellow with darker femora; dorsum of opisthosoma black with small yellow dots and dashes (Fig. 11F), with five pairs of large yellow patches in varying sizes and shapes (Fig. 11F), laterals with paired pale yellow bands of reeds and spots, venter pale yellow with rows of black bands and spots. ALE almost touching, separated from each other by less than their radius (Figs 11B–C); PER straight in dorsal view (Fig. 11B). Sternum finely wrinkled, shield-shaped, sparsely covered with long bristles, with anterolateral indentations accommodating posterior part of endites, with precoxal sclerites (Fig. 11E). Tarsi II–IV ventrolaterally with two rows of spinules. Epigyne (Fig. 11G). Epigynal plate sclerotized, semi-circular, with narrow basomedian septum. Copulatory openings narrow, slit-like, with straight, sclerotized anterior margins.</p>
            <p>Male. Unknown.</p>
            <p>Distribution. India (Tamil Nadu) (Reimoser 1934) (Fig. 18).</p>
            <p> Justification of the transfer. This species was originally placed under  Capheris (Reimoser 1934) . Jocqué (2009) erroneously transferred it to  Cydrela , without proper justification. An examination of the holotype of this species showed that it has diagnostic features of  Capheris species including carapace without deep concavity at level of fovea, ALEs situated in front of AMEs, ALEs almost separated by their radius, straight PER, sternum with anterolateral indentations and precoxal sclerites, and very stout palps (Figs. 11A–E). Based on these observations, the transfer of  C. escheri is fully justified. </p>
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	https://treatment.plazi.org/id/038E87ACFFB70423FF26F94F006FAD94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFB50423FF26FC7F06FEA85B.text	038E87ACFFB50423FF26FC7F06FEA85B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capheris nitidiceps Simon 1905	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Capheris nitidiceps Simon, 1905 comb, rest., nomen dubium </p>
            <p>Figs 12, 18</p>
            <p> Capheris nitidiceps Simon, 1905: 171 (subadult ♀). </p>
            <p> Cydrela nitidiceps — Jocqué, 2009: 123 (transfer from  Capheris ). </p>
            <p>
                 Type material.   Holotype subadult ♀ from INDIA: Tamil Nadu: Nilgiris,  
                <a title="Search Plazi for locations around (long 76.8/lat 11.333333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.8&amp;materialsCitation.latitude=11.333333">Coonoor</a>
                 , 11°20’N 76°48’E, 1591 m a.s.l., date unknown, leg. M. M. Maindron, repository MNHN, Paris (AR-16434), examined based on photographs. 
            </p>
            <p> Remarks. This species has to be considered a nomen dubium due to the immature stage of the female holotype. The type locality of this species lies hardly 267 kms away from the type locality of  C. escheri that let suggests a possible synonymy of the latter species with it; however, confirmation requires the examination of adult specimens of  C. nitidiceps from the type locality. Until adult specimens are found, it is treated as nomen dubium. </p>
            <p>Distribution. India (Tamil Nadu) (Simon 1905) (Fig. 18).</p>
            <p> Justification of the transfer. This species was originally placed under  Capheris (Simon 1905) . Jocqué (2009) erroneously transferred it to  Cydrela , without proper justification. An examination of the holotype of this species showed that it has diagnostic features of  Capheris species including ALEs almost touching, situated in front of AMEs, straight PER, sternum shied-shaped, anterolaterally with indentations accommodating endites, and very stout palps (Figs 12A–C). Based on these observations, the transfer of  C. nitidiceps is fully justified. </p>
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	https://treatment.plazi.org/id/038E87ACFFB50423FF26FC7F06FEA85B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFB5043FFF26F9320068AA90.text	038E87ACFFB5043FFF26F9320068AA90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capheris stillata Simon 1905	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Capheris stillata Simon, 1905 comb. rest. </p>
            <p>Figs 13–14, 18</p>
            <p> Capheris stillata Simon, 1905: 171 , fig. 3 (♁). </p>
            <p> Cydrela stillata — Jocqué, 2009: 123 (transfer from  Capheris ). </p>
            <p>
                 Type material.   Syntype ♁ from INDIA: Puducherry: Mahé /  
                <a title="Search Plazi for locations around (long 75.53333/lat 11.7)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=75.53333&amp;materialsCitation.latitude=11.7">Mayyazhi</a>
                 , 11°42’N 75°32’E, 2 m a.s.l., date unknown, leg. M. M. Maindron, repository MNHN, Paris (registration number unknown, Simon’s number 22495), not examined  . 
            </p>
            <p>
                 Other material examined.   INDIA: Kerala: Wayanad,  
                <a title="Search Plazi for locations around (long 76.083336/lat 11.8)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.083336&amp;materialsCitation.latitude=11.8">Kuruva Island</a>
                 , 11°48’N 76°05’E, 760 m a.s.l., 2014, from bark, by hand, collector unknown: 1 ♁ (ADSH210)  . 
            </p>
            <p> Remarks. The type specimen was not available for the present study as it is currently not found in the collection of MNHN and may be misplaced elsewhere (Elise-Anne Leguin, pers. comm.). I have found a male  Capheris specimen in the old  Arachnida collection of ADSH. The somatic features of this male specimen match with the description of  C. stillata and the shape of its palp’s RTA looks also similar to that species (compare Simon 1905: fig. 3 (Fig. 14A) with Figs 13H, J, 14D). Moreover, the collecting locality of this male specimen is lying less than 93 kms away from the type locality of  C. stillata . So, I tentatively consider this male specimen as  C. stillata , which is described hereafter. </p>
            <p> Diagnosis. Males of  C. stillata are most similar to the males of  Capheris approximata (Karsch, 1878) as both share long RTA with apical bifurcation and broad conductor, but can be separated from the latter species by RTA with a basal process (vs. absent in  C. approximata ), conductor with distal curvature (vs. median curvature in  C. approximata ) and mediolaterally originating embolus (vs. basolateral in  C. approximata ) (compare Figs 13H–J, 14C–D with Jocqué 2009: figs 155–156). </p>
            <p>Redescription. Male in alcohol (Figs 13A–G). Body length 6.15. Carapace 3.62 long, 2.43 wide. Opisthosoma 2.53 long, 1.82 wide. Carapace, eye region, clypeus, chelicerae, labium, endites, sternum reddish brown; leg and palp segments brown with green shades; dorsal and lateral opisthosoma black with chalk-white broad patches, venter brown with chalk-white spots, black patches and striae; spinnerets brown. Fovea short, longitudinal, straight, dark. PER straight in dorsal view (Fig. 13B). Clypeus high (Fig. 13D). Chelicerae without teeth (Fig. 13F). Sternum anterolaterally with two large indentations and laterally provided with three pairs of precoxal sclerites (II, III &amp; IV) (Fig. 13G). Tarsi II–IV provided with ventral spinules of varying numbers. Opisthosoma oval, hirsute. Eye diameters and interdistances: ALE 0.09, AME 0.12, PLE 0.11, PME 0.10; AME–ALE 0.14, AME–AME almost contiguous, ALE–ALE 0.04, ALE–PME 0.10, PME–PLE 0.16, PME–PME 0.09. Clypeus height at AMEs 0.47, at ALEs 0.74. Chelicerae 0.81 long. Sternum 1.59 long, 1.32 wide. Measurements of palp and legs: Palp 4.15 [1.67, 0.97, 0.38, 1.13], I (right) 8.04 [2.27, 0.86, 1.90, 1.78, 1.23], II 6.60 [1.88, 0.97, 1.23, 1.58, 0.94], III 6.08 [1.83, 0.92, 0.93, 1.57, 0.83], IV 8.54 [2.36, 1.00, 1.57, 2.39, 1.22]. Leg formula: 4123. Spination of palp: no spines, instead with short but thick macrosetae; legs: femur I do 2 rl 1, II–IV do 2; patellae I–II spineless, III pl 1 pld 1 rl 1, IV pl 1 pld 1 do 1 rl 1; tibia I plv 2 rlv 1, II pld 1 plv 2 rlv 2, III pld 3 plv 3 do 2 rl 2 rld 1 rlv 3, IV pld 3 plv 3 do 2 rl 2 rld 1 rlv 3; metatarsus I plv 2 rlv 3, II pl 1 pld 1 plv 1 rlv 3 vt 1, III pl 2 pld 2 plv 3 rl 1 rld 3 rlv 3 vt 1, IV pl 1 pld 3 plv 5 do 1 rl 2 rld 3 rlv 3; tarsi I–IV spineless. Palp (Figs 13H–J, 14B–D). Femur strongly developed, larger than leg femora (Fig. 13H). Tibia with RTA large, flat, slightly bifid at its extremity as seen ventrally (Figs 13H–J, 14C–D), dorsal prong slightly longer than ventral one, with a basolateral triangular process (Figs 13I–J, 14C–D). Cymbium with numerous short, thick macrosetae: basodorsally with cluster of eight, prolateroapically two and prolaterally nine (Figs 13I–J, 14B–D), without cymbial fold. Subtegulum membranous (Figs 14B–C). Tegulum simple, with slight posterolateral swelling (Fig. 14C). Embolus narrow, originating mediolaterally to tegulum, partially covered by conductor (Figs 13I, 14C). Conductor broad, membranous, longitudinally folded, distoprolaterally originating, with distoretrolaterad curvature, directed at 2-o’ clock ventrally (Figs 13I–J, 14C–D).</p>
            <p>Female. Unknown.</p>
            <p>Distribution. India (Kerala (new record), Puducherry) (Simon 1905; present data) (Fig. 18).</p>
            <p> Justification of the transfer. This species was originally placed under  Capheris (Simon 1905) . Jocqué (2009) erroneously transferred it to  Cydrela , without proper justification. This species also has diagnostic features of  Capheris species including carapace without deep concavity at level of fovea, ALEs situated in front of AMEs, ALEs almost separated by their radius, straight PER, sternum shied-shaped, with precoxal sclerites, anterolaterally with deep indentations accommodating endites, and very stout palps (Figs 13B–E, G–H). Based on these observations, the transfer of  C. stillata is fully justified. </p>
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	https://treatment.plazi.org/id/038E87ACFFB5043FFF26F9320068AA90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFA90438FF26FAB20667AB25.text	038E87ACFFA90438FF26FAB20667AB25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laminion jatashankar (Talwar, Majagi, Bodkhe & Kamble 2018) Sankaran 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laminion jatashankar (Talwar, Majagi, Bodkhe &amp; Kamble, 2018) comb. nov.</p>
            <p>Figs 15, 19</p>
            <p> Euryeidon jatashankarae Talwar et al., 2018: 72 , figs 5.1–7, 6.1–8 (♀) (remarks: in figures, partly named jatashankara (lapsus )). </p>
            <p>
                 Type material.   Holotype ♀ from INDIA: Madhya Pradesh: Hoshangabad, Pachmarhi (not Panchmarhi)  
                <a title="Search Plazi for locations around (long 77.3127/lat 20.922138)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.3127&amp;materialsCitation.latitude=20.922138">Wildlife Sanctuary</a>
                 , 22°31’30.00’’N 78°25’40.90’’E, September 2015; leg. S. Kamble; repository SRL JDPSM (no registration number), not examined.  Three paratypes (male(s) or female(s) unspecified): one with same data as holotype; two from Maharashtra: Amravati, J. D. Patil Sangludkar Mahavidyalya College campus, 20°55’19.70’’N 77°18’45.70’’E, August 2015; collector unknown; repository SRL JDPSM (no registration number), not examined. (Talwar et al. (2018: figs 5.6, 6.3, 5, 7) presented the colour images of the habitus and genitalia, which are diagnostic and were used for comparative purposes).
            </p>
            <p> Diagnosis. Females of  L. jatashankar comb. nov. are similar to the females of  Laminion birenifer (Gravely, 1921) , the type species of the genus in the general appearance of the epigyne, but can be separated from the latter species by smoothly constricted lateral margins of median, sclerotized plate of epigyne (vs. strongly constricted in  L. birenifer ), and slightly concave anterior margin of median, sclerotized plate (vs. anterior margin with a conical protrusion in  L. birenifer ) (compare Fig. 15A with Sankaran et al. 2020: fig. 3D). </p>
            <p>Description. For description of the female, see Talwar et al. (2018).</p>
            <p>Male. Unknown.</p>
            <p> Justification of the transfer. Talwar et al. (2018) described  E. jatashankarae based on four specimens collected from Madhya Pradesh and Maharashtra. They erroneously placed this species under  Euryeidon as it lacks the transverse band connecting the spermathecae, which is the main diagnostic feature of the female  Euryeidon species (Dankittipakul &amp; Jocqué 2004: figs 16, 27, 29, 34, 40, 42). Instead, it fits with  Laminion based on the following features: domed prosoma in lateral view, high clypeus and epigyne with median and lateral plates (compare Talwar et al. 2018: figs 5.4, 6.3, 5, herein Fig. 15A with Sankaran et al. 2020: figs 1E–F, 3D, 6B). Moreover, the internal genitalia of this species have similarities with that of  L. debasrae (Biswas &amp; Biswas, 1992) (compare Talwar et al. 2018: fig. 6.7, herein Fig. 15B with Sankaran et al. 2020: fig. 6C). Based on these observations, the transfer of  E. jatashankarae is fully justified. </p>
            <p> Nomenclatural note. Since the specific epithet is based on the name of a natural cave and a holistic place of Lord Shiva (jatashankar) located in the type locality (as it was considered a noun in apposition), it should therefore be amended as  jatashankar instead and rather not as  jatashankarae or jatashankara (lapsus) as mentioned in the original description (Talwar et al. 2018). </p>
            <p>Distribution. India (Madhya Pradesh, Maharashtra) (Talwar et al. 2018) (Fig. 19).</p>
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	https://treatment.plazi.org/id/038E87ACFFA90438FF26FAB20667AB25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
038E87ACFFAE043BFF26FAEF0794AC50.text	038E87ACFFAE043BFF26FAEF0794AC50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laminion katepagai (Talwar, Majagi, Bodkhe & Kamble 2018) Sankaran 2023	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Laminion katepagai (Talwar, Majagi, Bodkhe &amp; Kamble, 2018) comb. nov.</p>
            <p>Figs 16, 19</p>
            <p> Euryeidon katepagai Talwar et al., 2018: 69 , figs 1.1–10, 2.1–6, 3.1–7, 4.8–11 (♁ ♀) (remarks: in figures, partly named katepagae (lapsus )). </p>
            <p>
                 Type material.   Holotype (male or female unspecified) from INDIA: Karnataka: Kalaburagi (= Kalburgi),  
                <a title="Search Plazi for locations around (long 77.873055/lat 17.310278)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.873055&amp;materialsCitation.latitude=17.310278">Chincholi Wildlife Sanctuary</a>
                 (formerly  
                <a title="Search Plazi for locations around (long 77.873055/lat 17.310278)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.873055&amp;materialsCitation.latitude=17.310278">Konchavaram Forest</a>
                 ), 17°18’37’’N 77°52’23’’E, collector unknown, possibly  
                <a title="Search Plazi for locations around (long 77.873055/lat 17.310278)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.873055&amp;materialsCitation.latitude=17.310278">Supriya Talwar</a>
                 ; August 2016; repository SRL JDPSM (no registration number), not examined.  Two paratypes (male(s) or female(s) unspecified), with same data as holotype except July 2017, not examined. (Talwar et al. (2018: figs 1.4, 6, 2.3, 5, 4.8–9, 11) presented the colour images of the habitus and genitalia, which are diagnostic and were used for comparative purposes).
            </p>
            <p> Diagnosis. Male of  L. katepagai comb. nov. is similar to the males of  Laminion gujaratense (Tikader &amp; Patel, 1975) in the general appearance of palp, but can be distinguished from the latter species by RTA with retrolaterally oriented tip (vs. prolaterally oriented in  L. gujaratense ), wide embolic base (vs. narrow in  L. gujaratense ), first cymbial process with narrow apex (vs. broad in  L. gujaratense ), and beak-like retrolateral branch of median apophysis (vs. widely triangular in  L. gujaratense ) (compare Figs 16A–B with Sankaran et al. 2020: figs 8D–F). Female is most similar to  L. jatashankar comb. nov. in the general appearance of the epigyne, but can be separated from the latter species by the convex anterior margin of median, sclerotized plate (vs. slightly concave in  L. jatashankar comb. nov. ) (compare Fig. 15A with Fig. 16C). </p>
            <p>Description. For description of the male and female, see Talwar et al. (2018).</p>
            <p> Justification of the transfer. Talwar et al. (2018) described  E. katepagai based on four specimens collected from Karnataka. This species is also misplaced in  Euryeidon as it lacks the prolateral extension of cymbium and the transverse band that connects the spermathecae. It seems that the authors misinterpreted the shadow of the anterior epigynal margin as the transverse band connecting the spermathecae (Talwar et al. 2018: fig. 2.6, herein Figs 16C–D). This species is a member of  Laminion based on the following features: domed prosoma in lateral view, high clypeus, patella with a dorsal apophysis, cymbium with large retrolateral flange and processes, long, filiform embolus, and epigyne with median and lateral plates (compare Talwar et al. 2018: figs 2.3, 4.8, 9, 11, herein Figs 16A–C with Sankaran et al. 2020: figs 1E–F, 3D, 6B). Based on these observations, the transfer  E. katepagai is fully justified. </p>
            <p>Distribution. India (Karnataka) (Talwar et al. 2018) (Fig. 19).</p>
            <p> Remarks. Previously,  Laminion is known to occur in the Indian States of Andhra Pradesh, Gujarat, Odisha and West Bengal (Sankaran et al. 2020). The transfer of the two Indian  Euryeidon species extended its distribution to Karnataka, Madhya Pradesh and Maharashtra, indicating a wide distribution range of this genus in India (Fig. 19). </p>
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	https://treatment.plazi.org/id/038E87ACFFAE043BFF26FAEF0794AC50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sankaran, Pradeep M.	Sankaran, Pradeep M. (2023): Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocqué, 2004 (Araneae: Zodariidae). Zootaxa 5296 (3): 381-405, DOI: 10.11646/zootaxa.5296.3.4, URL: http://dx.doi.org/10.11646/zootaxa.5296.3.4
