taxonID	type	description	language	source
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	materials_examined	Type: — CHINA. Sichuan: Tongolo (= Dongeluo), près Ta-tsien-lou (now Kangding county), 5 August 1893, J. A. Soulié 391 (lectotype P 00145766!, designated here; isolectotypes K 000694194!, P 00145767!, P 00145768!, P 00145769!). Figs. 4, 5.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	materials_examined	Type: — CHINA. Yunnan: Ninglang county, Cuiyu town, Shuicaoba village, on slopes in ravine, alt. 3100 m, 10 July 1981, Y. F. Han et al. 81 - 1070 (holotype PE 00477951!; isotypes PE 00477950!, PE 00477952!). Fig. 1.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	description	Description: — Perennial herbs, glabrous throughout. Roots fibrous. Stem 40 – 120 cm tall, striate, distally branched, glabrous. Leaves 2 – 4 - ternate; leaf blade pentagonal, 7 – 30 cm long, 7 – 30 cm broad; leaflets ovate, obovate, obliquely ovate or broadly ovate, 0.8 – 4 cm long, 0.8 – 4 cm broad, base cuneate, rounded or subcordate, apex obtuse or acute, 3 - lobate; lobes entire or 2 ‒ 4 - dentate or 2 ‒ 4 - crenate, obtuse or acute; nerves obscure adaxially, prominent abaxially; petioles 0.5 – 5 cm long, with stipules at base; stipules obliquely oblong, membranous, margin torn. Inflorescence a panicle, leafy, raceme-like; branches narrow. Pedicels 1 ‒ 3 (5) mm long. Flowers bisexual, erect. Sepals elliptic, narrowly oblong or narrowly obovate, 2 ‒ 2.5 mm long, 1.3 ‒ 1.8 mm broad, margin membranous, 1 - nerved, greenish or greenish white. Stamens 5 ‒ 16, 2.5 ‒ 4 mm long; filaments 2 ‒ 3.5 mm long, filiform, apex slightly dilatate, greenish or greenish white; anthers 0.5 ‒ 1 mm long, ovate or elliptic, apex mucronate, pale yellow. Carpels 4 – 8, subsessile; ovaries obliquely compressed-ovoid; styles ca. 1.5 mm long, apex dorsally uncinate or circinate; stigmas inconspicuous. Achenes 3.5 ‒ 5 mm long, subsessile; body obliquely compressed-ovoid, inconspicuously ribbed when dry; style persistent, dorsally uncinate or circinate at apex.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	biology_ecology	Phenology: — Flowering from June to August; fruiting from July to September.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	distribution	Distribution and habitat: — Thalictrum atriplex is widely distributed in southwestern Qinghai, western Sichuan, northwestern Yunnan, and southeastern Xizang, China (Fig. 13). It grows in bushes or sparse forests, along forest margins or on grassy slopes at elevations of 2500 ‒ 4270 m above sea level.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	discussion	Typification: — In the protologue of Thalictrum atriplex, Finet & Gagnepain (1903) cited a single collection, i. e. J. A. Soulié 391. We have been able to trace five sheets of this collection, with one at K (barcode K 000694194) and another four at P (barcodes P 00145766, P 00145767, P 00145768, P 00145769). These sheets are all well preserved and match the original description, and thus are syntypes, as the author did not specify which one was used to prepare the original description. In conformity with ICN Art. 9.10 and Art. 9.11 (Turland et al. 2018) and as discussed by McNeill (2014), a lectotypification is proposed for T. atriplex. In the above, the P sheet (barcode P 00145766) that bears a dissection drawing has been designated as the lectotype, and the other four sheets cited as isolectotypes.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	materials_examined	Additional specimens examined: — CHINA. Qinghai: Yushu, T. N. Ho et al. 2594 (BM, barcode unavailable, E 00064269, HNWP, herb. no. 0287328, PE 00447985). Sichuan: Baiyu, D. E. Boufford et al. 36969 (PE 01459307); Barkam, C. Y. Chang & H. F. Chow 22644 (KUN 0690567, NAS 00186888, PE 00447961, SZ 00091928), C. Y. Chang & H. F. Chow 22837 (KUN 0690576, NAS 00186890, PE 00447959, SZ 00091927, WUK, herb. no. 252334), C. Y. Chang & H. F. Chow 22987 (KUN 0690529, NAS 00186887, PE 00447962, SZ 00091945, WUK, herb. no. 242216), L. Xie 200109020 (PE 01569323), L. Xie 200109021 (PE 01569324), L. Xie 200209008 (PE 01569384); Batang, K. Y. Lang, L. Q. Li & Y. Fei 2307 (PE 01108726, PE 01108727), K. Y. Lang, L. Q. Li & Y. Fei 2328 (KUN 0691916, PE 01108735, PE 01108734); Daocheng, Anonymous 515 (SM 704604444), D. E. Boufford et al. 28108 (E 00293940, PE 01578928), Sichuan Econ. Plant Exped. 171 (CDBI 0025918, CDBI 0025920, CDBI 0025921), Sichuan Econ. Plant Exped. 198 (CDBI 0025932), Sichuan Veg. Exped. 2060 (CDBI 0025923, CDBI 0025924, KUN 0689148, PE 00427859); Dawu, Anonymous 140 (SM 704604437), Anonymous 141 (SM 704604438), Anonymous 5558 (CDBI 0025926, CDBI 0025930, CDBI 0025931, PE 00447942), D. E. Boufford et al. 27697 (E 00320420, PE 01458750), K. L. Chu 7337 (NAS 00186891, NAS 00186892, NAS 00186894), P. C. Kuo & W. Y. Wang 20923 (HNWP, herb. no. 65655, KUN 0692064, NAS, herb. no. 504538, WUK, herb. no. 344291), Sichuan Med. Res. Exped. 15327 (CDBI 0025912, NAS 00186889), Z. P. Wang et al. 931 (KUN 0690578, WUK, herb. nos. 315057, 316613), T. S. Ying et al. 4646 (PE 00447967, SZ 00091939, SZ 00091940, SZ 00091941); Dege, Y. W. Tsui 5063 (PE 00447957); Jinchuan, No. 8 Forest Manag. Exped. 2336 (PE 00447966); Jiulong, D. E. Boufford et al. 33109 (KUN 0769895, KUN 0769896, PE 01459309), D. E. Boufford et al. 33271 (KUN 0770335, KUN 0770336, PE 01459308), T. T. Yu 6787 (KUN 0689149, KUN 0689150), Y. P. Zeng & Q. L. Huang 384 (IBSC, barcode unavailable); Kangding, Anonymous 4823 (CDBI 0025914, CDBI 0025915, CDBI 0025933, PE 00447949), X. J. He & Q. S. Zhao 180308 (SZ, herb. no. 512295), X. J. He & Q. S. Zhao 183754 (SZ, herb. no. 594447), S. Jiang et al. 2060 (KUN 0689155, PE 00447955, SZ 00091942), K. C. Kuan & W. T. Wang 656 (PE 00447948, PE 00447954, PE 00447965, PE 00447968, PE 01040678), C. R. Lancaste 941 A (PE 00447946), K. Y. Lang, L. Q. Li & Y. Fei 700 (PE 01108728, PE 01108741), K. Y. Lang, L. Q. Li & Y. Fei 1076 (KUN 0689159, PE 01108738, PE 01108739), Y. L. Peng et al. Gaoxf 0511 (CDBI 0207060), J. A. Soulié 2365 (P 00145762), J. A. Soulié 3365 (P 00145764, P 00145765), Y. W. Tsui 4194 (PE 00447958), Y. P. Zeng & Q. L. Huang 365 (IBSC, barcode unavailable), Y. P. Zeng & Q. L. Huang 370 (IBSC, barcode unavailable), Y. P. Zeng & Q. L. Huang 374 (IBSC, barcode unavailable), Z. J. Zhao et al. 114956 (CDBI 0026464, SZ 00091929), Y. C. Zhu 29645 (CDBI 0025904); Lixian, W. K. Hu & D. Z. Deng 1893 (SZ 00091926); Muli, C. Y. Wu et al. 152 (KUN 0689889), S. K. Wu 2607 (KUN 0689151), Q. S. Zhao 6276 (CDBI 0025910, SZ 00091935), Q. S. Zhao et al. 6745 (CDBI 0025929, SZ 00091931), Q. S. Zhao et al. 7840 (CDBI 0025936, SZ 00091936); Xiangcheng, Anonymous 989 (CDBI 0025919), Anonymous 4361 (CDBI 0025934), D. E. Boufford et al. 30691 (KUN 0769874, PE 01459304), Qinghai-Xizang Exped. 4704 (CDBI 0025937, HITBC, herb. no. 003442, KUN 0689156, PE 01108742), Xiangcheng Exped. 344 (SM 704604439); Xinlong, Y. P. Zeng & Q. L. Huang 400 (IBSC, barcode unavailable); Yajiang, D. E. Boufford et al. 35843 (KUN 0770303, PE 01459306), C. L. Wu 40983 (NAS, herb. no. 492149), Yajiang Exped. 124 (SM 704604441); Zamtang, L. Wang, X. Q. Guo & Y. P. Zeng 3570 (IBSC, barcode unavailable), Y. H. Wu 31 - 004 (HNWP, herb. no. 0205545), Y. H. Wu 31 - 023 (HNWP, herb. no. 0205558). Xizang (Tibet): Bomi, Anonymous 1063 (PE 00558464), Y. Fei et al. 129 (KUN 0689141), Y. Fei et al. 278 (KUN 0689143), Inst. Bio. Xizang Exped. 637 (HNWP, herb. no. 37753), L. Wang, X. Q. Guo & Y. P. Zeng 3316 (IBSC, barcode unavailable); Chagyab, Qinghai-Tibetan Sci. Exped. 12282 (HNWP, herb. no. 62155, KUN 0690780, PE 00427873); Cona, C. Y. Wu et al. 75 - 1100 (HNWP, herb. no. 54108, KUN 0689163, PE 00470001); Gongbo’gyamda, Y. T. Chang & K. Y. Lang 1374 (PE 00447990), Y. T. Chang & K. Y. Lang 2754 (PE 00447989), P. C. Kuo & W. Y. Wang 21981 (HNWP, herb. no. 66731), F. Ludlow, G. Sherriff & H. H. Elliot 14131 (BM, barcode unavailable), Qinghai-Xizang Exped. 74 - 3704 (KUN 0689138, PE 00447996); Gyirong, PE-Xizang Exped. 4153 (PE, herb. no. 2446929); Jomda, Inst. Bio. Xizang Exped. 2202 (HNWP, herb. no. 38489), Qinghai-Xizang Sci. Exped. 12415 (HNWP, herb. no. 61331, KUN 0689160, PE 00427870), L. Wang, X. Q. Guo & Y. P. Zeng 3431 (IBSC, barcode unavailable); Lhunze, Qinghai-Xizang Exped. Suppl. Group 750421 (HNWP, herb. no. 96821, KUN 0689162, PE 00447999); Mainling, F. Ludlow, G. Sherriff & G. Taylor 5680 (BM, barcode unavailable), Z. C. Ni et al. 2870 (PE 00447986), Qinghai-Xizang Exped. Suppl. Group 750911 (HNWP, herb. no. 51537, KUN 0689140, PE 00447997), Xizang Trad. Med. Exped. 3806 (HNWP, herb. no. 32862, PE 00447994); Markam, W. Q. Fei 22 (IBSC, barcode unavailable); Nangxian, S. Bowes Lyon 11024 (E 00080172), F. Ludlow & G. Sherriff 2690 (BM, barcode unavailable), L. Wang, X. Q. Guo & Y. P. Zeng 3209 (IBSC, barcode unavailable); Nyingchi, J. Luo & S. L. Wang Luojian-XZ- 09261 (HNWP, herb. no. 0288254, PE 01898340), Z. C. Ni et al. 1771 (PE 00447988, XZ 0001682), Xizang Inst. Bio. Plant Res. Exped. 3453 (XZ 0001679); Precise locality unknown, S. X. Jia 341 (PE 00447947, PE 00447950), P. C. Tsoong 5521 (PE 00447945); Qamdo, L. Wang, X. Q. Guo & Y. P. Zeng 3407 (IBSC, barcode unavailable), C. Y. Wu 4694 (KUN 0689145); Zayu, R. F. Huang 556 (HNWP, herb. no. 85417), X. H. Jin et al. DLJ-ET 1989 (PE 02006357), C. W. Wang 65517 (IBSC 0090116, KUN 0689153, NAS 00186897, PE 00447964, WUK, herb. no. 35700), C. W. Wang 65599 (IBSC 0090115), Y. P. Zeng & Q. L. Huang 550 (IBSC, barcode unavailable); Zhanang, PE-Xizang Exped. 5263 (PE, herb. no. 2553942). Yunnan: Deqen, D. M. Anderson et al. 170 (KUN 1280720), J. W. Gregory & C. J. Gregory s. n. (BM, barcode unavailable), Hengduan Exped. Veg. Group 4648 (PE 01108732), IBCAS Hengduan Exped. 3459 (PE 01108745), Qinghai-Xizang Exped. 2318 (CDBI 0025938, HITBC, herb. no. 003451, KUN 0689136, PE 01108743), H. Sun & Z. G. Qian 709 (KUN 0689173), L. Xie DQ 1 (PE 00471477), L. Xie DQ 2 (PE 00471475), L. Xie & J. F. Mao 114 (PE 01569442), L. Xie & J. F. Mao 115 (PE 01569441), L. Xie & J. F. Mao 148 (PE 01569439), J. S. Yang 8531 (KUN 0689178), J. S. Yang 8541 (KUN 0690328), J. S. Yang 8686 (KUN 0689177), T. T. Yu 9515 (KUN 0689130, PE 00447983), T. T. Yu 10185 (BM, barcode unavailable, KUN 0689132, PE 00447972), T. T. Yu 10511 (BM, barcode unavailable, KUN 0690768, PE 00447982), T. T. Yu 10568 (BM, barcode unavailable, KUN 0689133, PE 00447973), Y. P. Zeng & Y. F. Luo 525 (IBSC, barcode unavailable); Gongshan, T. T. Yu 6787 (PE 00447974); Lijiang, Anonymous 38 (IBSC 0090788, KUN 0689183), Anonymous 4104 (KUN 1265294), K. M. Feng 21352 (KUN 0689170, NAS, barcode unavailable, PE 00447977, SZ 00091943), K. M. Feng 22486 (HNWP, herb. no. 46167, KUN 0689181, PE 00447980), C. Schneider 2023 (K, barcode unavailable), C. W. Wang 70849 (KUN 0690795, NAS 00186896, PE 00447978, WUK, herb. no. 66448); Shangri-la, Anonymous 63 - 3336 (KUN 0689186), K. M. Feng 1723 (KUN 0689167), K. M. Feng 23452 (KUN 0689180, PE 00447979), H. H. Kong & L. Li YQE 3023 (IBSC, barcode unavailable), C. Y. Wu 4278 (KUN 0689174), L. Xie ZD 9 (PE 00471478), L. Xie & J. F. Mao 71 (PE 01569398), L. Xie & J. F. Mao 72 (PE 01569438), L. Xie & J. F. Mao 73 (PE 01569437), L. Xie & J. F. Mao 91 (PE 01569479), L. Xie & J. F. Mao 97 (PE 01569435), J. S. Yang 7649 (KUN 0690891), J. S. Yang 8002 (KUN 0689179), J. S. Yang 84 - 551 (KUN 1265273), Q. E. Yang & H. Z. Kong 98 - 255 (PE 00471474), Q. E. Yang & Q. Yuan 1821 (PE 02032593), T. T. Yu 12346 (KUN 0689131, PE 00447971), Y. P. Zeng & Y. F. Luo 503 (IBSC, barcode unavailable), Zhongdian Exped. 683 (KUN 0689189); Weixi, C. W. Wang 63686 (PE 00447981), Q. E. Yang & Q. Yuan 2052 (PE, herb. no. 2430710).	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
0381BD2FFFD9FFACFF27FAC851E0A094.taxon	discussion	Notes: — The specimen J. A. Soulié s. n. (MPU 023569), although collected from Kangding in western Sichuan, the type locality of Thalictrum atriplex, does not belong to original material of this species. In the protologue of T. atriplex, Finet & Gagnepain (1903) did not refer to this specimen, citing only J. A. Soulié 391. It was wrongly annotated as an isosyntype of T. atriplex on the determination slip. It actually belongs to a new species which we will describe elsewhere. Of the collections cited under Thalictrum atriplex by Wang (2018), the collection information of some were wrongly given and some were probably misidentified. The collection Z. M. Zhang & H. F. Chow 22837 from Barkam in western Sichuan should be corrected as C. Y. Chang & H. F. Chow 22837. The collection Hengduan Exped. Veg. Group 8648 from Deqen in northwestern Yunnan should be corrected as Hengduan Exped. Veg. Group 4648. The collections T. S. Ying & D. Y. Hong 650211 from Bomi in southeastern Xizang, Y. W. Tsui 4248 from Garze and C. S. Liu 695, H. L. Tsiang 36027 and K. C. Kuan & W. T. Wang 69 all from Kangding in western Sichuan (four sheets shown in Figure 14) should not belong to T. atriplex because the leaflets are smaller (0.5 – 1.5 cm long and broad vs. 0.8 – 4 cm long and broad) (Figs. 2 G, 7 F, 8 F, 14) and the pedicles are longer, over 5 mm long (vs. 1 ‒ 3 (5) mm long) (Figs. 2 N, 3 B, 7 L, 8 M, 14 B, D). In general appearance they resemble T. uncatum but differ by having 6 ‒ 14 - denticulate (vs. 3 - lobate) leaflets (Figs. 11 C, D, 12 C, D, 14), shorter styles ca. 1 mm (vs. ca. 2 mm) long (Figs. 11 I, 12 H, 14 A, C), and achenes subsessile, obliquely compressed-ovoid (vs. shortly stipitate with the stipes 1 ‒ 2 mm long, compressed-lunate) (Figs. 11 K, 12 J, 14 B, D). Further studies are needed to determine the identity of these collections.	en	Zeng, You-Pai, Yuan, Qiong, Yang, Qin-Er (2022): Thalictrum spiristylum (Ranunculaceae), described from northwestern Yunnan, China, is merged with T. atriplex. Phytotaxa 538 (1): 1-20, DOI: 10.11646/phytotaxa.538.1.1
