taxonID	type	description	language	source
03838780FFF0FFC97DBEFF22FE64C22E.taxon	materials_examined	Type material (8 exs). Holotype: CHINA: ♂: ‘ China: Hunan, Shaoyang City, Chengbu Miao autonomous county, Leng-shui-ping Village, Baiyun Cave, 26 ° 20 ’ 52.38 ’’ N, 110 ° 19 ’ 25.70 ’’ E, 468 m, 10. xi. 2017, Gu-Chun Zhou leg. [湖南冷水坪村白云洞] ’ (SNUC). Paratypes: CHINA: 2 ♂♂, 3 ♀♀, same collecting data as the holotype; 2 ♀♀: also from Leng-shui-ping Village and same date, except ‘ Biyun Cave, 26 ° 20 ’ 33.84 ’’ N, 110 ° 19 ’ 45.39 ’’ E, 473 m [湖南冷水坪村碧云洞] ’ (SNUC).	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF0FFC97DBEFF22FE64C22E.taxon	diagnosis	Diagnosis. Body length 2.28 – 2.33 mm. Vertex with a short mediobasal carina, lateral postantennal carinae present; clypeus short, less than one-third of total head length. Pronotum with a shallow median longitudinal impression and transverse antebasal sulci, with dorsal carinae and antebasal protuberances. Elytra with shallow, indistinct discal striae. Tergite IV with two mediobasal and four basolateral foveae. Sternite IV (second visible sternite) with two mediobasal and six basolateral foveae. Male with antennomeres IX – X, protarsomere II, apices of the mesofemur, mesotibia, and metatibia modified. Aedeagus with a large and transverse basal capsule.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF0FFC97DBEFF22FE64C22E.taxon	description	Description. Male (Fig. 1 A). Body length (combined length of head, pronotum, elytra, and abdomen) 2.28 – 2.33 mm. Body uniformly reddish-brown, mouthparts, tibiae, and tarsi paler. Head (Fig. 2 A) about as long as wide, length from clypeal anterior margin to head base 0.46 mm, width across eyes 0.43 – 0.44 mm, finely punctate; vertex with short median carina extending from near posterior margin of head toward level slightly past mid-length of eyes; pair of lateral postantennal carinae extending from posterior margins of antennal tubercles toward postocular margins; clypeus relatively short, slightly more than one-fifth of total head length. Each eye composed of about 35 facets. Antennal club formed by three apical enlarged antennomeres (Fig. 2 B), antennomeres IX and X enlarged and concave at mesal surfaces, XI asymmetrically narrowed at apical one-third. Pronotum (Fig. 2 A) slightly narrower than head, longer than wide, length along midline 0.49 mm, maximum width 0.41 – 0.42 mm; disc coarsely punctate, with wide and shallow median longitudinal impression and transverse antebasal sulcus, with dorsal carinae and antebasal tubercles. Elytra slightly longer than wide, length along suture 0.73 – 0.75 mm, maximum width 0.70 – 0.71 mm, with shallow discal striae at basal third. Metaventrite lacking modification. Protarsomere II expanded at ventral margin near middle (Fig. 2 C); apex of mesofemur roundly projected at ventral margin (Fig. 2 D); apex of mesotibia with large, acute spine (Fig. 2 E); metatrochanter distinctly projected ventrally (Fig. 2 E); metatibia strongly curved, with long pubescence at middle and dilated at apical third (Fig. 2 G). Length of dorsally visible part of abdomen along midline 0.58 – 0.65 mm, maximum width of abdomen 0.62 mm. Tergite IV with complete and oblique inner marginal carinae; basal impression subdivided into three parts by one pair of mediobasal and two pairs of basolateral foveae, tergites V – VII each with one pair of basolateral foveae. Sternite IV (second visible sternite) with one pair of mediobasal and three pairs of basolateral foveae, sternites V – VII each with one pair of basolateral foveae. Length of aedeagus (Fig. 2 H – J) 0.31 mm, median lobe with stout and slightly transverse basal bulb and large foramen; apex of elongate median lobe deeply narrowed; dorsal lobe broad, apex truncate. Female. General appearance similar to male. Antennae and legs lacking modifications. Eyes smaller, each composed of about 8 facets. Elytra relatively shorter. Measurements: body length (combined length of head, pronotum, elytra, and abdomen) 2.73 – 2.80 mm, length of head from clypeal anterior margin toward head base 0.48 mm, width across eyes 0.45 mm, length of pronotum along midline 0.49 mm, maximum width of pronotum 0.42 mm, length of elytra along suture 0.70 mm, maximum width of elytra 0.68 – 0.69 mm, length of dorsally visible part of abdomen along midline 0.63 – 0.66 mm, maximum width of abdomen 0.64 – 0.65 mm. Comparative notes. Using the key in Yin et al. (2016), the new species could be keyed out at couplet 4 together with A. grossepunctatus Yin, Jiang & Li. These two species share a similar external morphology of the head and pronotum, a similar form of the aedeagal basal capsule, and a close distribution (both in southern Hunan). However, the unique structures of antennomeres IX – X, protarsomere II, apex of the mesofemur, metatrochanter, and metatibia in the male of A. curvitibialis, and the configuration of the aedeagal median and dorsal lobes readily separate the new species from A. grossepunctatus, as well as from all other congeners of the genus.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF0FFC97DBEFF22FE64C22E.taxon	distribution	Distribution. China: Hunan.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF0FFC97DBEFF22FE64C22E.taxon	etymology	Etymology. The new specific epithet refers to the curved metatibiae of the male of the new species, which are a species characteristic.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF2FFCF7DBEFB58FCE0C73C.taxon	materials_examined	Type material (2 exs). Holotype: CHINA: ♂: ‘ China: Hunan, Shaoyang City, Chengbu Miao autonomous county, Leng-shui-ping Village, Biyun Cave, 26 ° 20 ’ 33.84 ’’ N, 110 ° 19 ’ 45.39 ’’ E, 473 m, 10. xi. 2017, Gu-Chun Zhou leg. [湖 南冷水坪村碧云洞] ’ (SNUC). Paratype: CHINA: 1 ♀, also from Leng-shui-ping Village and same date, except ‘ Baiyun Cave, 26 ° 20 ’ 52.38 ’’ N, 110 ° 19 ’ 25.70 ’’ E, 468 m [湖南冷水坪村白云洞] ’ (SNUC).	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF2FFCF7DBEFB58FCE0C73C.taxon	diagnosis	Diagnosis. Male with simple vertex; antennomeres IX – XI modified; each eye composed of about 45 facets; pronotum with small lateral spines; metaventrite raised above metacoxae to form high projections, with a row of dense setae along each ridge; aedeagus with apical portion of the median lobe split to two parts, dorsal part shorter than ventral part. Female with each eye composed of about 15 facets, and may be identified only when associated with a male.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF2FFCF7DBEFB58FCE0C73C.taxon	description	Description. Male. Body (Fig. 3 A) reddish brown, mouth parts, antennae and tarsi paler. Body length (combined length of head, pronotum, elytra, and abdomen) 2.80 mm. Length of head from clypeal anterior margin to head base 0.58 mm, width across eyes 0.58 mm; median longitudinal carina extending from base to level of midline of eyes; each eye composed of about 45 facets; antennomeres IX – XI modified (Fig. 3 B). Pronotum about as long as wide, length along midline 0.62 mm, maximum width 0.61 mm; with small discal and lateral spines. Elytra about as long as wide, length along suture 0.91 mm, maximum width 0.92 mm; discal striae extending to apical 4 / 5 of elytral length. Metaventrite (Fig. 3 C – D) strongly raised above metacoxae to form high projections, with rows of dense setae along projections; metacoxa and metatrochanter simple. Abdomen narrower than elytra, length of dorsally visible part along midline 0.69 mm, maximum width 0.79 mm. Aedeagus (Figs. 3 E – F) asymmetric, length 0.55 mm. Female. General appearance similar to male. Eyes smaller, each composed of about 15 facets. Antennae and metaventrite simple. Measurements: body length (combined length of head, pronotum, elytra, and abdomen) 2.73 mm, length of head from clypeal anterior margin toward head base 0.55 mm, width across eyes 0.56 mm, length of pronotum along midline 0.61 mm, maximum width of pronotum 0.60 mm, length of elytra along suture 0.83 mm, maximum width of elytra 0.88 mm, length of dorsally visible part of abdomen along midline 0.74 mm, maximum width of abdomen 0.82 mm. Comparative notes. Thirteen species from China and Thailand have been included in the Tribasodites bedosae species-group (Yin et al. 2015), which exclusively contains cavernicolous species. Using the key provided in Yin et al. (2015), T. biyun could be keyed out at couplet 10 together with T. bama Yin, Nomura & Li. These two species share the spinose pronotal lateral margins, simple metacoxa and metatrochanter, and short rows of setae present only on the posterior half of the metaventrite. Tribasodites biyun is the first species of the T. bedosae group from Hunan Province, and its males can be readily separated from those of T. bama by the simple antennomere VIII, lateral projection of antennomere IX short and not curved, antennomere X less expanded and lacking a large excavation at the base, and the much longer dorsal branch of the apical fork of the aedeagal median lobe. Among the Chinese species, the simple antennomere VIII of T. biyun is shared only with T. abnormalis Yin, Nomura & Li from Guizhou, but males of the latter species possess unmodified antennae, a modified vertex, an unforked apex of the aedeagal median lobe, and thus can be readily separated from T. biyun.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF2FFCF7DBEFB58FCE0C73C.taxon	distribution	Distribution. China: Hunan.	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
03838780FFF2FFCF7DBEFB58FCE0C73C.taxon	etymology	Etymology. The new specific epithet is taken directly from the name of the cave where the holotype was collected, i. e., Biyun Cave [literally: Bluish Cloud Cave].	en	Yin, Zi-Wei, Zhou, Gu-Chun (2018): Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae) from southern China. Zootaxa 4457 (4): 589-594, DOI: 10.11646/zootaxa.4457.4.9
