occurrenceID	taxonID	catalogNumber	collectionCode	institutionCode	typeStatus	verbatimLabel	sex	individualCount	eventDate	recordedBy	recordNumber	decimalLatitude	decimalLongitude	minimumElevationInMeters	maximumElevationInMeters	minimumDepthInMeters	maximumDepthInMeters	country	stateProvince	municipality	locality	references	associatedOccurrences	associatedReferences	associatedSequences	basisOfRecord
03828798D377646CFDAFF9CDAE27FC50.mc.3B433CD3D371646CFF13FD4AAFB5FC8E	03828798D377646CFDAFF9CDAE27FC50.taxon			FMNH	paratype	MADAGASCAR • 1 ♀; “ envir. de Moramanga ” [around Moramanga]; “ (Q. 1000 c. a) ” [1000 m a. s. l.]; 21 Jul. 1970; N. Sanfilippo leg.; FMNH	female	1	1970-07-21	N. Sanfilippo				1000				Madagascar			MADAGASCAR	https://treatment.plazi.org/id/03828798D377646CFDAFF9CDAE27FC50#3B433CD3D371646CFF13FD4AAFB5FC8E				MaterialCitation
03828798D371646DFDE4FB95ACC5FCD5.mc.3B433CD3D371646CFF13F8EDAE9CF835	03828798D371646DFDE4FB95ACC5FCD5.taxon			FMNH		MADAGASCAR • 5 specs; “ envir. de Perinet riv. Farimbany ” [around Perinet, Farimbany river]; “ (Q 800) ” [800 m a. s. l.]; 18 Jul. 1970; N. Sanfilippo leg.; FMNH		1	1970-07-18	N. Sanfilippo				800				Madagascar		Farimbany river	Perinet	https://treatment.plazi.org/id/03828798D371646DFDE4FB95ACC5FCD5#3B433CD3D371646CFF13F8EDAE9CF835				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFF13FAE4A9D4FA66	03828798D3726460FDC3FE68AFD3F80A.taxon				holotype	MADAGASCAR • 1 ♂; Antananarivo former province, Analamanga region, 10 km E of Anjozorobe, Anjozorobe-Angavo protected area, marsh next to the stream in Saha forest; 18.4128 ° S, 47.9439 ° E; 1320 m a. s. l.; 23 Nov. 2014; J. Bergsten, R. Bukontaite, S. Holmgren, J. H. Randriamihaja and T. Ranarilalatiana leg.; collecting event MAD 14 - 78; forest marsh; NHRS-JLKB 000025033.		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFF13FAE4A9D4FA66				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFF13F99AA9D2F9F9	03828798D3726460FDC3FE68AFD3F80A.taxon				holotype	MADAGASCAR • 1 spec.; same collecting data as for holotype; NHRS-JLKB 000025034		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFF13F99AA9D2F9F9				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFABEF99AA997F9DE	03828798D3726460FDC3FE68AFD3F80A.taxon				holotype	27 specs (preserved in ethanol); same collecting data as for holotype; NHRS-JLKB 000025027		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFABEF99AA997F9DE				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFB7CF9F7AFA1F933	03828798D3726460FDC3FE68AFD3F80A.taxon			FMNH	holotype	5 specs; same collecting data as for holotype; FMNH		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFB7CF9F7AFA1F933				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFD39F9D0A822F933	03828798D3726460FDC3FE68AFD3F80A.taxon			BMNH	holotype	5 specs; same collecting data as for holotype; BMNH		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFD39F9D0A822F933				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFAB4F9D0AE01F910	03828798D3726460FDC3FE68AFD3F80A.taxon			PBZT, MBC	holotype	5 specs; same collecting data as for holotype; PBZT / MBC		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFAB4F9D0AE01F910				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFCECF92DAC2CF975	03828798D3726460FDC3FE68AFD3F80A.taxon			DEUA	holotype	5 specs; same collecting data as for holotype; DEUA		1	2014-11-23	J. Bergsten & R. Bukontaite & S. Holmgren & J. H. Randriamihaja & T. Ranarilalatiana		-18.4128	47.9439	1320				Madagascar	Antananarivo	Anjozorobe-Angavo	Saha forest	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFCECF92DAC2CF975				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFE88F909A844F8AE	03828798D3726460FDC3FE68AFD3F80A.taxon				paratype	1 ♂; Antananarivo former province, Analamanga region, 10 km E of Anjozorobe, Anjozorobe Angavo protected area, Antsahabe, circuit bamboo; 18.41266 ° S, 47.9438 ° E; 1341 m a. s. l.; 5 Apr. 2016; T. Ranarilalatiana leg.; collecting event MAD 16 - 40; marsh and stream pool; NHRS-JLKB 000065489	male	1	2016-04-05	T. Ranarilalatiana		-18.41266	47.9438	1341				Madagascar	Antananarivo	Anjozorobe Angavo	Antsahabe	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFE88F909A844F8AE				MaterialCitation
03828798D3726460FDC3FE68AFD3F80A.mc.3B433CD3D372646FFF13F8A0AEC3F883	03828798D3726460FDC3FE68AFD3F80A.taxon				paratype	1 ♀; same collecting data as for preceding; NHRS-JLKB 000065490	female	1	2016-04-05	T. Ranarilalatiana		-18.41266	47.9438	1341				Madagascar	Antananarivo	Anjozorobe Angavo	Antsahabe	https://treatment.plazi.org/id/03828798D3726460FDC3FE68AFD3F80A#3B433CD3D372646FFF13F8A0AEC3F883				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FF13FB91AE0AFB0B	03828798D37F6464FD98FEB5A856FDBC.taxon	NHRS-JLKB000025032		NHRS-JLKB		MADAGASCAR • 1 ♂; Fianarantsoa former province, Amoron’i Mania region, 3 km south of Ambalamanakana next to RN 7, Ankazomivady forest; 20.7722 ° S, 47.1809 ° E; 1700 m a. s. l.; 1 Nov. 2014; J. Bergsten, T. Ranarilalatiana and S. Holmgren leg.; collecting event MAD 14 - 02; hygropetric rocks and marsh with vegetation; NHRS-JLKB 000025032	male	1	2014-11-01	J. Bergsten & T. Ranarilalatiana & S. Holmgren		-20.7722	47.1809	1700				Madagascar	Fianarantsoa former province		Amoron'i Mania region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FF13FB91AE0AFB0B				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FCE6FB28AFFEFB68	03828798D37F6464FD98FEB5A856FDBC.taxon	NHRS-JLKB000025024		NHRS-JLKB		5 specs (preserved in ethanol); same collection data as for preceding; NHRS-JLKB 000025024		5	2014-11-01	J. Bergsten & T. Ranarilalatiana & S. Holmgren		-20.7722	47.1809	1700				Madagascar	Fianarantsoa former province		Amoron'i Mania region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FCE6FB28AFFEFB68				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FD58FB05A847FB68	03828798D37F6464FD98FEB5A856FDBC.taxon			FMNH, PBZT, MBC		1 spec.; same collection data as for preceding; FMNH		2	2014-11-01	J. Bergsten & T. Ranarilalatiana & S. Holmgren		-20.7722	47.1809	1700				Madagascar	Fianarantsoa former province		Amoron'i Mania region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FD58FB05A847FB68				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FF13FB61AE46FB4D	03828798D37F6464FD98FEB5A856FDBC.taxon			FMNH, PBZT, MBC		1 spec.; same collection data as for preceding; PBZT / MBC		2	2014-11-01	J. Bergsten & T. Ranarilalatiana & S. Holmgren		-20.7722	47.1809	1700				Madagascar	Fianarantsoa former province		Amoron'i Mania region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FF13FB61AE46FB4D				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FCD0FB61ACA6FA38	03828798D37F6464FD98FEB5A856FDBC.taxon	NHRS- JLKB000025025		NHRS		1 spec. (preserved in ethanol); Fianarantsoa former province, Matsiara Ambony region, Ranomafana NP, 500 m from the road at Vohiparara village; 21.2397 ° S, 47.3774 ° E; 1110 m a. s. l.; 3 Nov. 2014; J. Bergsten, T. Ranarilalatiana and S. Holmgren leg.; collecting event MAD 14 - 10; small forest lake with floating margins of vegetation; NHRSJLKB 000025025		1	2014-11-03	J. Bergsten & T. Ranarilalatiana & S. Holmgren		-21.2397	47.3774	1110					Fianarantsoa former province		Matsiara Ambony region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FCD0FB61ACA6FA38				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FE35FAD5AF11FA56	03828798D37F6464FD98FEB5A856FDBC.taxon	NHRS-JLKB000025026		NHRS-JLKB		1 spec.; Toamasina former province, Alaotra Mangoro region, 6 km W of Moramanga, Betsabora river by RN 2 near Antsampanana village; 18.9247 ° S, 48.1828 ° E; 900 m a. s. l.; 24 Nov. 2014; J. Bergsten, J. H. Randriamihaja and T. Ranarilalatiana leg.; collecting event MAD 14 - 81; river with side pools; NHRS-JLKB 000025026		1	2014-11-24	J. Bergsten & J. H. Randriamihaja & T. Ranarilalatiana		-18.9247	48.1828	900					Toamasina former province		Alaotra Mangoro region	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FE35FAD5AF11FA56				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FDE2FA6FAFEAF989	03828798D37F6464FD98FEB5A856FDBC.taxon			CMM		2 specs; Fianarantsoa former province, ca 9 km SW Ivato, Bemanta (inselberg); 20 ° 40 ′ 00 ″ S, 47 ° 07 ′ 41.5 ″ E; 1531 m a. s. l.; 18 May 2016; Ramahandrison and Manuel leg.; wide and calm portion of small river; CMM		2	2016-05-18			-20.666666	47.128193	1531					Fianarantsoa former province		Ramahandrison and Manuel	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FDE2FA6FAFEAF989				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FD49F9A5AE59F927	03828798D37F6464FD98FEB5A856FDBC.taxon			CMM		16 specs; Fianarantsoa former province, along road N 7 between Ambositra and Fianarantsoa, ca 3.5 km S of Ambalamanakana; 20 ° 46 ′ 33 ″ S, 47 ° 10 ′ 53 ″ E; 1702 m a. s. l.; 19 May 2016; Ramahandrison and Manuel leg.; large pond with Nymphaeaceae and surrounding peat bog with Sphagnum and Drosera; CMM		16	2016-05-19	Ramahandrison & Manuel		-20.775833	47.18139	1702					Fianarantsoa former province		Fianarantsoa former province	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FD49F9A5AE59F927				MaterialCitation
03828798D37F6464FD98FEB5A856FDBC.mc.3B433CD3D37F6462FC38F93FAE20F8BD	03828798D37F6464FD98FEB5A856FDBC.taxon			CMM		13 specs; Fianarantsoa former province, ca 8 km WNW of Ranomafana, ca 1 km SW of Vohiparara; 21 ° 14 ′ 51 ″ S, 47 ° 22 ′ 34.5 ″ E; 1134 m a. s. l.; 29 Dec. 2017; Ramahandrison and Manuel leg.; small marsh, very shallow, with small Poaceae and Cyperaceae, along slowly flowing stream; CMM.		13	2017-12-29			-21.2475	47.376247	1134					Fianarantsoa former province		Ramahandrison and Manuel	https://treatment.plazi.org/id/03828798D37F6464FD98FEB5A856FDBC#3B433CD3D37F6462FC38F93FAE20F8BD				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FF13FC32AC89FC4B	03828798D3796465FDE2FDF8A910F86A.taxon			MNHN	holotype	MADAGASCAR • 1 ♂; Ankaratra, Tsiafajavona; 2500 m a. s. l.; Jun. 1947; J. Millot leg.; MNHN. [Note that Bistr ̂ m (1985) during his revision of the group, was unable to locate this holotype that we have since rediscovered.]	male	1	1947-06	J. Millot				2500				Madagascar		Tsiafajavona	Ankaratra	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FF13FC32AC89FC4B				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FF13FB87A856FB20	03828798D3796465FDE2FDF8A910F86A.taxon					MADAGASCAR • 1 spec.; Antananarivo former province, Vakinankaratra region, Manjakatompo Stn Forestière, Anosiarivo; 19.34489 ° S, 047.30414 ° E; 2073 m a. s. l.; 24 Jan. 2012; T. Ranarilalatiana and J. H. Randriamihaja leg.; collecting event MAD-MJK 12 - 13; lake near source; NHRS-JLKB 000025031		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FF13FB87A856FB20				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FF7DFB19A840FB06	03828798D3796465FDE2FDF8A910F86A.taxon					20 specs (preserved in ethanol); same collection data as for preceding; NHRS-JLKB 000024967		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FF7DFB19A840FB06				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FF13FB7BAE1AFB5A	03828798D3796465FDE2FDF8A910F86A.taxon			FMNH		5 specs; same collection data as for preceding; FMNH		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FF13FB7BAE1AFB5A				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FCFAFB7BAC6FFABF	03828798D3796465FDE2FDF8A910F86A.taxon			PBZT, MBC		5 specs; same collection data as for preceding; PBZT / MBC		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FCFAFB7BAC6FFABF				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FEC9FB57AEC9FABF	03828798D3796465FDE2FDF8A910F86A.taxon			BMNH		5 specs; same collection data as for preceding; BMNH		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FEC9FB57AEC9FABF				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FBACFB57AC9BFA9C	03828798D3796465FDE2FDF8A910F86A.taxon			DEUA		5 specs; same collection data as for preceding; DEUA		1	2012-01-24	T. Ranarilalatiana & J. H. Randriamihaja		-19.34489	47.30414	2073				Madagascar	Antananarivo	Manjakatompo	Anosiarivo	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FBACFB57AC9BFA9C				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FE7DFAB0AF01FA2A	03828798D3796465FDE2FDF8A910F86A.taxon					18 specs (preserved in ethanol); Fianarantsoa former province, Haute Matsiatra region, Andringitra NP, lake north east of camp 3; 22.17373 ° S, 46.9016 ° E; 2026 m a. s. l.; 2 Dec. 2013; J. H. Randriamihaja and T. Ranarilalatiana leg.; collecting event MAD 13 - 33; small lake in open area; NHRS-JLKB 000025023		1	2013-12-02	J. H. Randriamihaja & T. Ranarilalatiana		-22.17373	46.9016	2026				Madagascar	Fianarantsoa	Haute Matsiatra region	Andringitra	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FE7DFAB0AF01FA2A				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FD95FACBA9DCFA2A	03828798D3796465FDE2FDF8A910F86A.taxon			PBZT, MBC		5 specs; same collection data as for preceding; PBZT / MBC		1	2013-12-02	J. H. Randriamihaja & T. Ranarilalatiana		-22.17373	46.9016	2026				Madagascar	Fianarantsoa	Haute Matsiatra region	Andringitra	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FD95FACBA9DCFA2A				MaterialCitation
03828798D3796465FDE2FDF8A910F86A.mc.3B433CD3D3786465FAB6FACAA90EFA41	03828798D3796465FDE2FDF8A910F86A.taxon			CMM		4 specs; Fianarantsoa former province, Ankaratra massif, ca 2.7 km WNW of Manjakatompo; 19 ° 21 ′ 57 ″ S, 47 ° 18 ′ 22.5 ″ E; 1814 m a. s. l.; 19 Dec. 2017; Ramahandrison and Manuel leg.; large grassy temporary pond, beneath slope with pine trees, with Conchostraca crustaceans; CMM		1	2017-12-19	Ramahandrison & Manuel & Conchostraca		-19.365833	47.306248	1814				Madagascar	Fianarantsoa		Ankaratra	https://treatment.plazi.org/id/03828798D3796465FDE2FDF8A910F86A#3B433CD3D3786465FAB6FACAA90EFA41				MaterialCitation
03828798D37B6467FDFCFEB5A953FE2C.mc.3B433CD3D37B6466FF13FBB0AE9BFBF0	03828798D37B6467FDFCFEB5A953FE2C.taxon			NHRS, CAS, NMW, PBZT, MBC, BMNH, DEUA		478 specimens (NHRS, CAS, NMW, PBZT / MBC, BMNH, DEUA) from provinces Antsiranana, Mahajanga, Toliara and Fianarantsoa (see Supplementary File 1).		478											Antsiranana		Antsiranana	https://treatment.plazi.org/id/03828798D37B6467FDFCFEB5A953FE2C#3B433CD3D37B6466FF13FBB0AE9BFBF0				MaterialCitation
