taxonID	type	description	language	source
03F99722FFEFFFA9FF25266DFDFCFCED.taxon	diagnosis	Diagnosis. Medium-sized Eisenia worms with long clitellum extending beyond segment 35. Tubercles beginning behind segment 30, ending on or behind 35. Included species. Eisenia kontschani sp. nov.; Eisenia malekae sp. nov.; Eisenia malevici Perel, 1962; Eisenia muganiensis (Michaelsen, 1910); Eisenia patriciae Szederjesi, Pavlíček, Coşkun & Csuzdi, 2014; Eisenia transcaucasica (Perel, 1967) (Table 1).	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEFFFAAFF252252FC6BFD62.taxon	materials_examined	Holotype. HNHM / 16953. Grazed Mediterranean garrigue near Hasankeyf, Batman Province, Turkey, 37 o 42 ' N, 41 o 24 ' E. Leg. T. Pavlíček & P. Cardet, 23.04.2009. Paratypes. HNHM / 16954, 5 ex., locality and date same as that of the Holotype.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEFFFAAFF252252FC6BFD62.taxon	etymology	Derivatio nominis. The new species is named in honour of our college, the renowned mite taxonomist Dr. Jenő Kontschán for his continuous help in our earthworm researches.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEFFFAAFF252252FC6BFD62.taxon	diagnosis	Diagnosis. Length 56 – 79 mm, diameter 3.5 – 4 mm, setae closely paired. Colour red-violet dorsally. First dorsal pore in 4 / 5. Clitellum on 26 – 37, 1 / 2 38, tubercles on 32 – 36. Male pore on 15, surrounded by glandular crescents. Nephridial pores not visible. Four pairs of vesicles in 9 – 12; spermathecae two pairs in 9 / 10 – 10 / 11, open near d. Calciferous glands in 11 – 13. Hearts in segments 7 – 11. Nephridial bladders simple, sausage-shaped. Longitudinal musculature pinnate. External characters. Holotype 78 mm long and 4 mm wide. Number of segments 169. Paratypes 56 – 79 mm long and 3.5 – 4 mm wide. Number of segments 117 – 173. Colour red-violet dorsally. Prostomium tanylobous. First dorsal pore at intersegmental furrow 4 / 5. Setae closely paired. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 18.6: 1.3: 10: 1: 38.6 (Fig. 2 A). Male pores on segment 15, surrounded by glandular crescents protruding into the neighbouring segments. Nephridial pores not visible. Clitellum on segments 26 – 37, 1 / 2 38. Tubercula pubertatis on segments 32 – 36. Glandular tumescence on segments 10 – 12, 14, 16, 21 – 23, 32 – 37 ab (Fig. 2 B). Internal characters. Septa 6 / 7 – 8 / 9 strongly thickened, 9 / 10 – 10 / 11 less so. Testes and funnels paired in segments 10 – 11, covered with peri-oesophageal testis sac. Four pairs of seminal vesicles in 9 – 12. Spermathecae two pairs in 9 / 10, 10 / 11 with external openings near setal line d. Calciferous glands in segment 11 – 13 without well demarcated diverticula. Paired hearts appear in segments 7 – 11, with a pair of small extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in segments 15 – 16, and gizzard in segments 17 – 18. Typhlosolis T-shaped, large. Longitudinal musculature is of pinnate type.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEFFFAAFF252252FC6BFD62.taxon	discussion	Remarks. The new species is close to E. malevici Perel, 1962 but differs from it by the somewhat longer clitellum, the tanylobic head and the number of seminal vesicles (Table 1).	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFECFFABFF2520D6FC43F836.taxon	materials_examined	Holotype. HNHM / 16956. Moist oak forest at Mah Parviz in Kohgiluyeh and Boyer-Ahmad Province, Iran, 30 ° 37 ' 14 '' N, 51 ° 39 ' 0 '' E. Leg. R. Latif, 29.03.2010. Paratypes. HNHM / 16957, 1 ex. ZUTC 1134, 3 ex., locality and date same as that of the holotype.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFECFFABFF2520D6FC43F836.taxon	etymology	Derivatio nominis. The new species is named in honour of Dr. Masoumeh Malek (University of Tehran) who initiated the current earthworm research in Iran.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFECFFABFF2520D6FC43F836.taxon	diagnosis	Diagnosis. Length 94 – 107 mm, diameter 4 mm, setae closely paired. Colour alive dark red-violet, conserved pale. First dorsal pore in 3 / 4. Clitellum on 27, 28 – 41, 42, tubercles on 37, 38 – 41. Male pore on 15, surrounded by glandular crescents. Nephridial pores irregularly alternated between setal line b and above d. Two pairs of vesicles in 11, 12; spermathecae two pairs in 9 / 10 – 10 / 11, open near d. Calciferous glands in 11 – 12 with small diverticulum in 12. Hearts in segments 7 – 11. Nephridial bladders simple, sausage-shaped. Longitudinal musculature pinnate. External characters. Holotype 94 mm long and 4 mm wide. Number of segments 219. Paratypes 100 – 104 mm long and 4 mm wide. Number of segments 214 – 227. Colour dark red-violet especially dorsally in alive specimens, bleached out to pale in preserved specimens. Prostomium epilobous 1 / 2 opened. First dorsal pore at intersegmental furrow 3 / 4. Setae closely paired. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 11: 1.5: 8.5: 1: 34 (Fig. 3 A). Male pores on segment 15, surrounded by glandular crescents, protruding into the neighbouring segments. Nephridial pores irregularly alternated between setal line b and above d. Clitellum on segments 27, 28 – 41, 42. Tubercula pubertatis on segments 37, 38 – 41. Glandular tumescence on segments 11, 12, 16 – 18, 24, 26, 29, 32, 35 – 42 ab (Fig. 3 B). Internal characters. Septa 5 / 6 – 14 / 15 thickened, 6 / 7 – 10 / 11 strongly thickened. Testes and funnels paired in segments 10 – 11, covered with perioesophageal testis sac in 11. Two pairs of seminal vesicles in 11, 12. Spermathecae two pairs in 9 / 10, 10 / 11 with external openings near setal line d. Calciferous glands in segment 11 – 12 with small diverticulum in 12. Paired hearts appear in segments 7 – 11, with a pair of small extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in segments 15 – 16, and gizzard in segments 17 – 18. Typhlosolis large, anchor-shaped. Longitudinal musculature is of pinnate type.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFECFFABFF2520D6FC43F836.taxon	discussion	Remarks. E. malekae sp. nov. is similar to E. muganiensis in the position of the clitellum however differs from it in the number of seminal vesicles and the more backward stretching tubercles (Table 1). brackets Head tanylobous epilobous epilobous epilobous epilobous epilobous numbers in Musculature pinnate pinnate pinnate? pinnate fasciculated column glands) glands " Calciferous) (- – 11 13 11 –) 12 12 (11 11 (12), – 11 - 12 ()) 12 - (1 – (10 2 / - 12 Calciferous " Vesicles 12 – 9, 11 12, 12 11 12 9 – 12 9 – 9 – 12 the In M. d d d> cd d cd species group. Spermatheca / 11 10, 10 / 9, 10 / 10 9 / 11 10, 10 / 11 9 / / 10 9 10 11 /, 10 10 9 / 11, / 10, 11 9 10 / / diverticula) 39 muganiensis missing Tubercles – 32 36 37,38 – 41 36 31,32 – 35 36) 38 (– (– 34 38 – 35 31 Eisenia denotes 40 " 2 the in) " - (; 38 42 / 39,1 36 species of diverticula Clitellum 37, 2 / 26 – 1 27,28 – 41, 29 36 – – 41 28 26,27 1 – / 2 26,1 – 26 / 2 characters calciferous 1910,) 2014,. al et), 1967 Distinguishing. 1 the of position the .. kontschani sp nov. malekae nov sp. malevici, Perel 1962 muganiensis Michaelsen (Szederjesi patriciae (transcaucasica Perel TABLE refer to Species Eisenia Eisenia Eisenia Eisenia Eisenia Eisenia	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFACFF2526E6FE1AFBB8.taxon	materials_examined	Material examined. HNHM / 12708, 1 ex. Paratype. Kemerovskoy Obl. Prokopyevska. Leg. U. B. Byzov, 1960. Diagnosis. Length 95 – 110 mm, diameter 6 – 7 mm, setae closely paired. Colour dark red-violet dorsally. First dorsal pore in 4 / 5. Clitellum on 29 – 36, tubercles on (31) 32 – 36. Male pore on 15, surrounded by glandular crescents. Two pairs of vesicles in 11 – 12 and two pairs of spermathecae in 9 / 10 – 10 / 11 open above setal line d, around 2 / 3 distance to M. Calciferous glands in 11 – 12 with large diverticula in 11. Nephridial bladders simple, sausage-shaped. Longitudinal musculature pinnate.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFACFF2526E6FE1AFBB8.taxon	distribution	Distribution. Salair Ridge, Kemerovo Region, Russia.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFACFF2526E6FE1AFBB8.taxon	discussion	Remarks. The diagnosis agrees completely with the original description except for the calciferous glands, recorded here also in 12.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFACFF2520FEFCFFF993.taxon	diagnosis	Diagnosis. Length 53 – 68 mm, diameter 4 – 6 mm, setae closely paired. Colour dark reddish-brown dorsally. First dorsal pore in 4 / 5. Clitellum on 28 – 41, tubercles on 35, 36 – 38, 39. Male pore on 15, surrounded by glandular crescents. Four pairs of vesicles in 9 – 12; spermathecae two pairs in 9 / 10 – 10 / 11 cd. Calciferous glands in 11 – 12 without distinct diverticula. Nephridial bladders simple, sausage-shaped.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFACFF2520FEFCFFF993.taxon	distribution	Distribution. Mughan Plain (Michaelsen 1910) and the Talysh Mts. (Perel & Kvavadze 1979) Azerbaijan. Remarks. This diagnosis is from the original description.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFADFF2522ECFCFFFF33.taxon	diagnosis	Diagnosis. Length 55 – 72 mm, diameter 4 – 4.5 mm, setae closely paired. Colour red-violet. First dorsal pore in 3 / 4. Clitellum on 1 / 2 26, 27 – 39, 1 / 2 40, tubercles on 34 – 38. Male pore on 15, surrounded by glandular crescents. Nephridial pores irregularly alternate between b and above d. Four pairs of vesicles in 9 – 12. Spermathecae two pairs in 9 / 10, 10 / 11, open near d. Calciferous glands in 12 without distinct diverticula. Nephridial bladders simple, sausage-shaped. Longitudinal musculature pinnate.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFADFF2522ECFCFFFF33.taxon	distribution	Distribution. Diyarbakır Province, Turkey.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEAFFADFF2522ECFCFFFF33.taxon	discussion	Remarks. This diagnosis is from the original description.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEBFFADFF252568FC99FC74.taxon	diagnosis	Diagnosis. Length 49 – 62 mm, diameter 4 – 5 mm, setae closely paired. Colour pale. First dorsal pore in 4 / 5. Clitellum on 1 / 2 26, 26 – 36, tubercles on 31 – 35. Male pore on 15, surrounded by glandular crescents. Four pairs of vesicles in 9 – 12. Spermathecae two pairs in 9 / 10, 10 / 11, open near cd. Calciferous glands in 11, 12 without distinct diverticula. Nephridial bladders simple, sausage-shaped. Longitudinal musculature fasciculated.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEBFFADFF252568FC99FC74.taxon	distribution	Distribution. Western Caucasus around Krasnaya Polyana and in the Caucasian Biosphere Reserve, Russia (Kvavadze 1985).	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
03F99722FFEBFFADFF252568FC99FC74.taxon	discussion	Remarks. This diagnosis is from the original description. Kvavadze (1985) described E. transcaucasica breviclitelata from a locality near the Laura River, Caucasian Biosphere Reserve, Russia. However, this species has much shorter clitellum (25 – 33) and tubercles (29 – 33). The specimens possess large calciferous diverticula in 10 – 12, proving that they are not related to E. transcaucasica. Unfortunately, there is no data available on the shape of the nephridial bladders. However, absent pigmentation and large calciferous diverticula in 10 could indicate that breviclitelata can even belong to the genus Allolobophora.	en	Szederjesi, Tímea, Pavlíček, Tomás, Latif, Robabeh, Csuzdi, Csaba (2014): Review of the Eisenia muganiensis (Michaelsen, 1910) species group with description of two new species (Oligochaeta: Lumbricidae). Zootaxa 3884 (3): 282-288, DOI: 10.11646/zootaxa.3884.3.7
