taxonID	type	description	language	source
03F887989F63A0663AA5098F7F6B44FC.taxon	diagnosis	Diagnosis amended. Body small, less than 1 mm in length. Pd 4 and Pd 5 in both sexes fused. Pd 5 with rounded triangular corners and without sensilla. In female and male genital somites in both sexes without sensilla; genital double-somite in female 1.8 – 2.0 times as long as wide. Rostral spines well developed, bifid. Antennule in female 25 - segmented; number and position of seta / spine on left antennule characteristic for Speodiaptominae (Arabic numerals = evident segments; S — seta; sp — spine): 1 (S), 2 (3 S [4 S *]), 3 (S), 4 (S), 5 (S), 6 (S), 7 (S), 8 (S + sp [1 S *]), 9 (2 S), 10 (S), 11 (S), 12 (S + sp [2 S *]), 13 (S), 14 (S), 15 (S), 16 (S), 17 (S), 18 (S), 19 (S), 20 (S), 21 (S), 22 (2 S), 23 (2 S), 24 (2 S), 25 (5 S [4 S ”]) (* – in Microdiaptomus; ” — in Microdiaptomus and Nannodiaptomus). Right antennule in male without unarticulated armature on segment 21; none of segments especially broadened; small spine-like seta on segments 8, 10 – 13. P 1 – P 4 with 2 - segmented Exp, 1 - segmented Endp. P 1 – P 4 Exp- 1 without inner seta and no spines on outer margin; Endp with 3 setae terminally. Left basis of P 5 in female with reduced Endp; Endp lack on right side; Exp- 2 without lateral spine, inner attenuation of Exp- 2 long and robust; Exp- 3 completely absent. Right P 5 in male with evident hyaline lobe on inner border of basis; without Endp. Left P 5 as long as right P 5; with setulose pad on Exp; Endp 1 - segmented.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
03F887989F63A06F3AA50C2D7DC245E9.taxon	materials_examined	Type locality. River in Hang Phong Nha Cave (17 ° 34 ' 59.0 '' N, 106 ° 16 ' 55.0 '' E), Quang Binh province, Central Vietnam. Neotype designation. As the type material was lost, a neotype is designated. Neotype (total length 625 µm; IEBR-COP 3418): male, collected on 0 9 August 2014; dissected and mounted in glycerol; deposited in the Institute of Ecology and Biological Resources (IEBR), Hanoi, Vietnam. Additional material. 10 males, 10 females, prepared for SEM examination; one adult male (IEBR-COP 3419), one adult female (IEBR-COP 3420), both dissected and mounted in glycerol; 8 males, 13 females (IEBR-COP- AED 08.14.013) preserved in 70 % ethanol; deposited in the IEBR. 15 males, 15 females preserved in 70 % ethanol; deposited in the National Institute of Biology (NIB), Ljubljana, Slovenia. All specimens collected at the same locality and date as neotype. One male (IEBR-COP 3421), one female (IEBR-COP 3422), both dissected and mounted in glycerol; 30 males, 30 females (IEBR-COP-AED 04.14.021) preserved in 70 % ethanol; all deposited in the IEBR. Material collected on 19 April 2014 from two pools inside Hang Va Cave (17 ° 29 ' 22.3 " N, 106 ° 17 ' 04.8 " E), Quang Binh province, Central Vietnam.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
03F887989F63A06F3AA50C2D7DC245E9.taxon	description	Description of female. Total length (without caudal setae) 595 – 670 µm, mean 640 µm (N = 25). Preserved specimens colourless and eyeless. Body (Fig. 2 A) slender with maximum width at middle of Pd 1. Mean prosome length: 445 µm; mean urosome length: 194 µm; prosome / urosome ratio 2.3 (N = 10). Rostrum represented by pair of large finger-like filaments, with pointed tips (Figs. 5 C, D; 6 L). Pd 4 and Pd 5 fused, dorsal and lateral spinules lacking. Lateral wings of Pd 5 moderately developed, nearly symmetrical, with rounded posterolateral margins, smooth, without sensilla / spines (Figs. 2 C, D; 6 H). Urosome (Fig. 2 A) of 3 free somites (ratio of Ur 1: Ur 2: Ur 3: CR = 0.53: 0.12: 0.16: 0.19). Ur 1 (genital doublesomite) (Figs. 2 C – E, 6 H) about 2 - times as long as wide, slightly asymmetrical in dorsal view; without sensilla. Genital field with pair of gonopores located mid-ventrally, gonopores completely covered by large genital operculum; genital area large, sac-like (Fig. 2 E); spermatophore insertion point on left posterolateral corner of somite. Ur 2 small, cylindrical, about 0.7 times as long as wide. Ur 3 (anal somite) slightly shorter than caudal rami, with concave lateral margins. Anal operculum small, semicircular, posterior margin smooth. Caudal rami (Fig. 2 A) symmetrical, 1.8 – 2.1 times as long as wide (mean 1.9, N = 10). Inner and outer margins smooth; with 5 plumose setae (setae II – VI) and articulated smooth dorsal seta (VII), inserted on inner distal corner; seta I lacking. Length of setae II – VI increases from outer to inner; seta VI 1.6 times as long as seta II; seta VII longest. Antennule (Fig. 3 A) 25 - segmented, reaching beyond posterior margins of caudal rami. Seta on segment 1 short, not reaching distal margin of segment 2. Largest setae on segments 3, 7, 9, 14, 21 – 25. Armature of segments as follows (Arabic numerals = evident segments; a — aesthetasc; S — seta; sp — spine): 1 (S + a), 2 (3 S + a), 3 (S + a), 4 (S), 5 (S + a), 6 (S), 7 (S + a), 8 (S + sp), 9 (2 S + a), 10 (S), 11 (S), 12 (S + sp + a), 13 (S), 14 (S + a), 15 (S), 16 (S), 17 (S), 18 (S), 19 (S + a), 20 (S), 21 (S), 22 (S + S), 23 (S + S), 24 (S + S), 25 (4 S + a). Antenna (Fig. 3 B) with Exp as long as Endp; coxa small, with 1 seta; basis robust with 2 setae on inner distal corner. Endp 2 - segmented; proximal segment elongated, 3.7 times as long as wide, with 2 setae at 1 / 2 length of inner margin; distal segment with 2 lobes. Outer lobe with 6 long setae terminally and 1 short sub-terminal seta; inner lobe with 1 short seta, 2 medium-sized and 5 long setae. Exp 7 - segmented; segment 1 with 1 seta, segment 2 with 3 setae, 1 seta on each of segments 3 – 6, terminal segment with 1 lateral and 3 apical setae. Mandible (Fig. 3 C) with 7 small teeth on gnathobase; ventral tooth medium sized. Basis with 4 sub-equal setae, proximal one plumose. Endp 2 - segmented, proximal segment with 4 setae, distal segment with 8 setae. Exp 4 - segmented with 1, 1, 1, 3 setae; ancestral segments 4 and 5 fused. Maxillule (Fig. 3 D) with 9 apical spiniform and 4 plumose setae on posterior surface of praecoxal arthrite. Coxal epipodite with 7 long setae and 2 short spiniform setae. Basal exite with 1 small spine-like seta. Coxal and first basal endites with 4 setae each. Basis and Endp fused, with 5 setae laterally and 8 setae apically. Exp with 6 long setae. Maxilla (Fig. 3 E) with 2 praecoxal and 2 coxal lobes, with 5, 3, 3, 3 setae, respectively. Basal lobe well developed, with 3 long setae. Endp 2 - segmented, proximal one with 2 setae, distal one with 3 setae. Maxilliped (Fig. 3 F) well developed. Syncoxa with 4 endites with 1, 2, 3, 4 unequal setae. Distal endite of syncoxa with well developed protuberance projecting over basis. Basis elongated, with 3 plumose setae increasing in length distally, inserted near 1 / 2 length of inner margin. Endp 6 - segmented, segment 1 partially incorporated into the basis. Segment 1 with 2, segment 2 with 3 sub-equal setae; segments 3 – 5 with 2, 2, 1 + 1, segment 6 with 4 setae, respectively. P 1 – P 4 (Figs. 2 F – I) with 2 - segmented Exp and 1 - segmented Endp; coxa and basis without seta. P 1 – P 4 Endp- 1 and Exp- 2 with setules along inner margin. Armament of swimming legs as follows (Roman numerals — spines; Arabic numerals — seta; outer-apical-inner position of element): P 5 (Figs. 5 E, 6 I, K). Coxa sub-cylindrical, about 1.4 times as long as wide; posterior surface with small protuberance bearing short spine at 2 / 3 of coxa length. Bases asymmetrical; left basis triangular, slightly larger than right one; right one’s inner margin convex. Left Endp small, triangular, with rounded tip, no armature; right Endp lack. Exp 2 - segmented on both sides; Exp- 1 about 2.1 times as long as wide, cylindrical; Exp- 2 short, prolonged into robust terminal claw-like apophysis, with row of small spinules along both margins; lateral spine lack. Description of male. Total length (without caudal setae) 525 – 640 µm, mean 615 µm (N = 25). Mean prosome length: 438 µm; mean urosome length: 193 µm; prosome / urosome ratio 2.3 (N = 10). Preserved specimens colourless and eyeless. Body slender, with typical diaptomid shape (Fig. 4 A). Rostrum represented by pair of finger-like filaments with pointed tips. Pd 4 and Pd 5 fused. Pd 5 slightly asymmetrical (Figs. 4 A, C), with rounded posterolateral margins, lack dorsal ornamentation or sensilla. Urosome (Fig. 4 A) with 5 free somites; somites ratio 1 – 5 + CR as follows: 0.22: 0.18: 0.13: 0.13: 0.14: 0.20. Ur 1 (= genital somite) asymmetrical; middle part of left lateral margin slightly protuberant, expanded; sensilla lack on both margins of somite. Dorsal and ventral surface of Ur 2 – 5 un-ornamented; posterior margins smooth. Caudal rami symmetrical, 1.7 – 2.0 times as long as wide (mean 1.8, N = 10), with inner and outer margins smooth. Setae II – VI plumose; dorsal seta (VII) articulated, smooth, inserted on inner distal corner of caudal ramus. Length of caudal setae II – VI increases from outermost to innermost; seta VI 1.2 times as long as outer seta (II). Dorsal seta (VII) 1.3 times as long as seta VI. Antennule shorter than in female, reaching distal part of caudal rami. Right antennule with 22 evident segments (Figs. 5 A, B), with largest setae on segments 3, 7, 9, 14, 19, 20, 21; none of segments especially broadened. Segments 8, 10 – 13 each with short spine-like seta. Segments 17 - 19 with 1 large, knife-like spine each. One small aesthetasc on segments 1, 2, 3, 5, 7, 9, 12, 14, 15, 16, 19, 22. Armature segments as follows (Arabic numerals = evident segments; a — aesthetasc, S — seta, sp — spine): 1 (S + a), 2 (3 S + a), 3 (S + a), 4 (S), 5 (S + a), 6 (S), 7 (S + a), 8 (S + sp), 9 (2 S + a), 10 (S + sp), 11 (S + sp), 12 (S + sp + a), 13 (S + sp), 14 (2 S + a), 15 (S + a), 16 (2 S + a), 17 (S + sp), 18 (sp), 19 (S + sp + a), 20 (2 S + 2 S), 21 (S + S), 22 (4 S + a). Setation of left antennule as in female. Antennae, mouthparts and P 1 – P 4 as in female. P 5 asymmetrical, coxae on posterior side each with 1 spine at 2 / 3 of length of segment, unequal in size (Fig. 4 F). Right P 5 (Figs. 4 F; 6 A – C, G): basis about 1.4 times as long as wide, with large, semicircular and flattened hyaline lobe along full length of inner margin. Exp- 1 short and trapezoidal, 1.1 times as long as wide, with narrow hyaline lamella along inner margin; distal margin with small rounded lobe. Exp- 2 oval, about 1.4 times as long as wide, outer margin convex, outer lateral spine curved, inserted at 2 / 3 length of segment, extending beyond distal margin of Exp- 2. Exp- 2 claw slender, almost semicircular, 2.2 times as long as Exp- 2; inner margin with a short row of tiny spinules. Endp lack. Left P 5 (Figs. 4 E – G; 6 A, C – F) reaching posterior margins of right P 5 Exp- 2. Basis cylindrical, elongated, 1.8 times as long as wide, without lateral seta. Exp- 1 about 2.2 times as long as wide, with row of setules on inner margin. Exp- 2 rounded, inner margin with field of short setules. Apical process slightly shorter than segment bearing it, with 3 – 4 denticular protrusions along inner margin. Subterminal lateral seta stout, spiniform, about twice as long as apical process, with 3 denticular protrusions on inner 2 / 3 margin length. Endp 1 - segmented, elongated, with width / length ratio of 2.3, laterally curved, about half as long as basis; with group of short setules apically.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
03F887989F6AA0753AA50EDE78734511.taxon	materials_examined	Type locality. A small siphon lake inside Hang Toi Cave (coordinates of the entrance to the cave: 17 ° 34 ' 26.4 '' N, 106 ° 15 ' 15.0 '' E), about 900 m from the entrance; Phong Nha-Ke Bang National Park, Quang Binh province, Central Vietnam. Type material. Holotype (female; total length 655 µm; IEBR-COP 3423; collected on 21 April 2014, by D. L. Tran); dissected and mounted in glycerol. Allotype (male; total length 640 µm; IEBR-COP 3424; collected on 21 April 2014 by D. L. Tran); dissected and mounted on slide in glycerol. Paratypes: 3 males (IEBR-COP 3425 – 3427), 3 females (IEBR-COP 3428 – 3430), dissected and mounted in glycerol; collection details the same as the holotype and allotype. The holotype, allotype and 6 paratypes deposited in the Institute of Ecology and Biological Resources (IEBR), Hanoi, Vietnam; 15 males and 15 females undissected, preserved in 70 % ethanol, collection details the same as the holotype; deposited at the National Institute of Biology (NIB), Ljubljana, Slovenia; 10 males and 10 females prepared for SEM examinations. Collection details the same as the holotype. Additional material. 40 males, 30 females (IEBR-COP-AED 04.14.028), preserved in 70 % ethanol, deposited at the IEBR, Vietnam. Collection details the same as the holotype.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
03F887989F6AA0753AA50EDE78734511.taxon	description	Description of female. Total length (without caudal setae) 610 – 680 µm, mean 655 µm (N = 25). Mean prosome length: 476 µm; mean urosome length: 196 µm; prosome / urosome ratio 2.4 (N = 10). Preserved specimens colourless and eyeless. Body (Fig. 7 A) slender, with a typical diaptomid shape. Rostrum represented by pair of soft, finger-like filaments, pointed backward (Figs. 7 C, D; 11 A). Pd 4 and Pd 5 fused, with rounded posterolateral margins, without sensilla / spines (Figs. 7 A; 11 B). Pd 5 asymmetrical, with slightly larger left wing; posterolateral margins rounded, with no dorsal ornamentation; sensilla lack on both sides (Figs. 7 A; 11 B). Urosome of 3 free somites (ratio of Ur 1: Ur 2: Ur 3: CR = 0.48: 0.12: 0.19: 0.21). Ur 1 (genital double-somite) (Figs. 7 A, E; 11 B) about 1.8 times as long as wide, slightly asymmetrical in dorsal view; right proximal margin slightly swollen on proximal part; without lateral sensilla. Genital field with pair of gonopores, divided by median longitudinal septum; gonopores covered by large genital operculum; genital area large (Fig. 7 E); spermatophore inserted on posterolateral margin of genital field. Ur 2 small, cylindrical, about 0.7 times as long as wide. Ur 3 (anal segment) as long as wide and as long as caudal rami; deep cleft posteriorly, lateral margins almost straight. Anal operculum small, semicircular, posterior margin smooth. Caudal rami (Figs. 7 A, E) symmetrical, 1.8 – 2.1 times as long as wide (mean 1.9, N = 10). Inner and outer margins smooth; with 5 plumose setae (setae II – VI) and smooth, articulated dorsal seta (VII), inserted on inner distal corner; seta I lack. Seta II shortest, other similar in length. Seta VII 1.3 times as long as setae III – VI. Antennule 25 - segmented (Fig. 8 A), reaching beyond posterior margins of caudal rami. Details of armament the same as in N. phongnhaensis, including position of the longest setae. Antenna, mandible, maxillule, maxilla and maxilliped (Figs. 8 B, C – D, E, F, G, respectively) similar to N. phongnhaensis. P 1 – P 4 (Figs. 7 F – I) armature the same as in N. phongnhaensis. P 5 (Figs. 10 G; 11 I, K). Coxa cylindrical, about 1.3 times as long as wide, posterior surface with small protuberance bearing short spiniform structure at 3 / 4 of segment length. Bases triangular, asymmetrical; left one larger than right one; without seta on lateral margins. Left Endp very small, triangular, with rounded tip, no armature; right Endp lack. Exp 2 - segmented on both sides; Exp- 1 cylindrical, elongated, about 2.2 times as long as wide; Exp- 2 short, prolonged into terminal claw-like apophysis, with row of tiny spinules on both margins, lateral spine lack. Description of male. Total length (without caudal setae) 605 – 655 µm, mean 629 µm (N = 25). Mean prosome length: 463 µm; mean urosome length: 193 µm; prosome / urosome ratio 2.4 (N = 10). Preserved specimens colourless and eyeless. Body slender, with typical diaptomid shape (Fig. 9 A). Rostral field as in female. Pd 4 and Pd 5 fused, with suture limited to lateral margins only. Pd 5 slightly asymmetrical, with rounded posterior margins, right lobe larger; without sensilla (Figs. 9 C, 11 C). Urosome of 5 free somites, simmilar to N. phongnhaensis with urosomites ratio 1 – 5 + CR as follows: 0.16: 0.13: 0.13: 0.15: 0.21: 0.22. Caudal rami symmetrical, 1.8 – 1.9 times as long as wide (mean 1.8, N = 10). Inner and outer margins smooth; setae II – VI plumose; dorsal seta (VII) articulated, smooth, inserted on inner distal corner of caudal ramus. Seta II shortest, about 0.8 as long as seta VI; dorsal seta (VII) 1.4 times as long as seta VI. Antennulae (Figs. 9 D, E) relatively shorter than in female, reaching middle of caudal rami. Armature of antennules as in N. phongnhaensis. Structure and armature of antennae, mouthparts and P 1 – P 4 as in female. P 5 asymmetrical, coxae each with 1 spine at 3 / 4 of segment length, right one larger. Right P 5 (Figs. 10 A, B, F; 11 D, E) basis sub-rectangular, about 1.2 times as long as wide; with large, triangular hyaline lobe with broadly rounded tip on distal inner corner; no seta on lateral margin. Exp- 1 trapezoidal, as long as wide, distal outer corner with small rounded process. Exp- 2 elongate, slightly curved inward, about 2.2 times as long as wide; outer lateral spine slightly curved, inserted at distal 3 / 4 of segment length, extending well beyond distal margin of Exp- 2. Exp- 2 claw sickle-like, slender, about 1.8 times as long as Exp- 2; inner margin with row of tiny spines. Endp lack. Left P 5 (Figs. 10 C – E; 11 D – H) long, reaching middle part of right Exp- 2. Basis about 1.3 times as long as wide; outer margin straight, without lateral seta; inner margin slightly convex. Exp- 1 1.9 times as long as wide, curved, with row of short setules on inner margin. Exp- 2 short, rectangular, about 0.7 times as long as wide; inner margin with field of short setules; apical process digitiform, elongated, as long as segment bearing it, with small serrated membrane on inner margin and tip divided with two small chitinous processes. Subterminal seta stout, spiniform, 2 times as long as apical process; proximal part 2 short serrated membrane on inner margin. Endp 1 - segmented, short, triangular, 1.4 times as long as wide; tip with group of short setules.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
03F887989F6AA0753AA50EDE78734511.taxon	etymology	Etymology. The new species is named after Assoc. Prof. Ho Thanh Hai from the Institute of Ecology and Biological Resources (IEBR), in recognition of his contribution to the knowledge of Vietnamese copepods.	en	Tran, Duc Luong, Brancelj, Anton (2017): Amended diagnosis of the genus Nannodiaptomus (Copepoda, Calanoida), based on redescription of N. phongnhaensis and description of a new species from caves in central Vietnam. Zootaxa 4221 (4): 457-476, DOI: 10.11646/zootaxa.4221.4.3
