taxonID	type	description	language	source
03FB87CBFF90FFBCFF28FB2FFB41F904.taxon	materials_examined	Type species. Hemerobius priscus Weyenbergh, 1869 a, by original designation.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF90FFBCFF28FB2FFB41F904.taxon	diagnosis	Diagnosis. Forewing elongate, broad proximally [strongly narrowed proximally in Liminympha Ren et Engel, 2007]; all distal veinlets of ScP once or twice forked [all veinlets of ScP simple in Mesonymphes Carpenter, 1929]; MP with four-five pectinate branches, each connected by one crossvein [crossveins absent between branches of MP in Liminympha]; CuP space broad, more than twice as wide as intracubital space [narrow, nearly as wide as intracubital space in Liminympha]. Hind wing MP with four-five pectinate branches [eight branches in Sialium Westwood, 1854]; at least proximal branches of CuA connected by crossveins [not connected in Mesonymphes]; CuP short, not pectinately branched. Legs (at least hind) with tibial spurs. Composition. Two species from the Jurassic of Eurasia: Nymphites bimaculatus sp. nov. from the Middle Jurassic of China, and Nymphites priscus (Weyenbergh, 1869) from the Late Jurassic of Germany.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF90FFBAFF28F8E4FE62F8A8.taxon	diagnosis	Diagnosis. Distinguished from Nymphites bimaculatus sp. nov. and from Nymphites sp. A by the absence of crossveins between distal branches of CuA and branches of MP.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF90FFBAFF28F8E4FE62F8A8.taxon	description	Description. Thorax fragmentarily preserved; details not discernible. Hind wing approximately 29 mm long as preserved (estimated complete length 31 mm); 8.0 mm wide as preserved (estimated complete width 8.5 mm). Margins poorly-preserved; trichosors not visible. Costal space narrow. Subcostal veinlets simple in proximal 2 / 3, some distal veinlets apparently shallowly forked. ScP, RA preserved as very closely approaching veins; subcostal space not visible. Preserved veinlets of ScP + RA dichotomously branched. RP smooth, only slightly zigzagged; originating near wing base. Preserved crossveins between RA, RP before fusion of ScP + RA widely spaced, rather regularly arranged. RP with 11 or 12 branches (poorly preserved); at least RP 1, RP 2 deeply forked. Crossveins in radial space relatively scarce, irregularly spaced proximally; rare or absent distally. Origin of M not preserved. MA dichotomously branched relatively close to wing margin. MP pectinately branched, with four branches, two deeply forked; one preserved crossvein evident between them. CuA long, pectinately branched with 11 long branches, at least one deeply forked; four-six crossveins between proximal branches forming gradate series. CuP incompletely preserved, short, apparently deeply forked. Anal veins not preserved. Crossveins between RP 1 and MA, MA and MP, MP and CuA rather scarce, irregularly spaced.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF90FFBAFF28F8E4FE62F8A8.taxon	materials_examined	Type material. Holotype No. 15453 (part; formerly No. 10336), No. 15454 (counterpart), deposited in TM. A poorly-preserved specimen with two incomplete hind wings. Type locality and horizon. Solnhofen, Germany; Late Jurassic, early Tithonian (Solnhofen Formation).	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF90FFBAFF28F8E4FE62F8A8.taxon	discussion	Discussion. Handlirsch (1906 – 1908) thought that these are forewings, apparently following Weyenbergh (1869 a) and Haase (1890).	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	diagnosis	Diagnosis. May be distinguished from Nymphites priscus by crossveins present between all branches of CuA, MP; from Nymphites sp. A by a more obtuse angle between Cu and its branches.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	description	Description. Holotype CNU-NEU-NN 2011116 PC. Thorax, abdomen fragmentarily preserved; details not visible. Metafemora relatively stout. Metatibiae long, slender with remnants of terminal spurs (Fig. 4 A); covered with dense, relatively short hairs. Tarsus covered with fine, short hairs; four proximal tarsomeres distally with row of strong dark bristles. Claws moderately large; not strongly curved; basally not strongly dilated. Forewing: left wing 37.5 mm long as preserved (estimated complete length 38 mm), 11 mm wide; right wing 33 mm as preserved, 12.2 mm wide. One trichosor between tips of each two veins along apical, outer margins of wings; three-four trichosors between tips of each two veins along hind margin in proximal half of wing (Fig. 5). Costal space narrow, strongly dilated after fusion of ScP, RA. Proximal subcostal veinlets mostly simple, few forked; all distal subcostal veinlets once or twice forked; no crossveins between them. ScP, RA stout for entire length. Subcostal space narrow, slightly dilated distally; no distinct crossveins detected (but see Remarks). Veinlets of ScP + RA dichotomously branched; no crossveins between them. RP smooth, not zigzagged; originating near wing base. Ten irregularly spaced crossveins between RA, RP before fusion of ScP + RA; not detected after fusion. RP with 14 (left wing), 13 (right wing) branches; all dichotomously branched (RP 1, RP 2 more deeply than others). Crossveins in radial space rather numerous, irregularly spaced in proximal two thirds; rare in distal portion, most form outer gradate series. M incurved; forked far proximad origin of RP 1. One preserved crossvein connecting RP, M before fork. MA, MP parallel. MA branched (probably dichotomously) relatively close to wing margin (incompletely preserved). MP zigzagged distally, pectinately branched; with four branches, two deeply forked (left wing), five shallowly forked branches (right wing); four crossveins between them. CuA long, pectinately branched with four long branches, two-three deeply forked; four crossveins between these. CuP long, parallel to posterior margin; pectinately forked: left wing, with six long branches, two proximal-most deeply forked; right wing, nine branches, four of them deeply forked; no crossveins between these. All preserved branches of CuA once or twice shallowly forked. CuP space twice wider than intracubital space. AA 3 + 4 long, pectinately branched, with two long branches, very shallowly forked (left wing); dichotomous, with anterior branch very shallowly forked, posterior branch deeply forked (right wing). AP 1 + 2 deeply forked; anterior branch forked, posterior at most shallowly forked (incompletely preserved). Crossveins between RP 1 and MA, MA and MP, MP and CuA, CuA and CuP rather numerous, irregularly spaced. Two crossveins between CuP, AA 3 + 4; one crossvein between AA 3 + 4, AP 1 + 2. Wing patterns consisted of two spots near hind wing margin, more or less rounded at MA, elongate at outer gradate series in radial space. Hind wing: left wing about 36 mm long as preserved (estimated complete length 37 mm), 10 mm wide; right wing 31.5 mm as preserved, 10.8 mm wide. Trichosors preserved between each vein pair terminating along margin, one medially, three-four basally (Fig. 5). Costal space narrower than in forewing. Subcostal veinlets mostly simple in proximal 2 / 3; once or twice shallowly forked distally. ScP, RA stout for entire length. Subcostal space narrow; no crossveins detected. Preserved veinlets of ScP + RA dichotomously branched; no crossveins connecting them detected. RP smooth, not zigzagged; originating near wing base. Eleven irregularly arranged crossveins between RA, RP before fusion of ScP + RA detected; not detected after fusion. RP with 11 branches; RP 1 to RP 3 deeply forked. Crossveins in radial space rather numerous, irregularly spaced in proximal proximally; rare or absent distally. Origin of M not preserved. MA dichotomously branched relatively close to wing margin. MP pectinate branched, with five branches, two-three deeply forked; three-four crossveins between them. CuA long, pectinately branched with 12 long branches, three-four deeply forked; 11 – 12 crossveins between them forming gradate series. All preserved branches of MP, CuA once or twice shallowly forked. CuP incompletely preserved, short, deeply forked. Anal veins not preserved. Crossveins between RP 1 and MA, MA and MP, MP and CuA rather numerous, irregularly spaced. Two-three crossveins between CuP and AA 3 + 4. Paratype CNU-NEU-NN 2011117 PC. Forewing 38.4 mm long, 11.5 mm wide as preserved (estimated complete length about 40 – 41 mm, width about 12 mm). Trichosors preserved in distal part of anterior wing margin. Costal space relatively narrow; strongly dilated after fusion of ScP, RA. Subcostal veinlets not preserved proximally; mainly forked medially; all once or twice shallowly forked distally; no crossveins connecting them. ScP, RA stout for entire length. Subcostal space moderately broad; slightly dilated distally; no crossveins detected. ScP + RA entering margin well after wing apex; its veinlets dichotomously branched; no crossveins connecting these. RP smooth not zigzagged; originating near wing base. Ten crossveins between RA, RP before fusion of ScP + RA widely spaced; not detected after fusion. RP with 11 branches; all dichotomously branched (RP 1, RP 3, RP 4 more deeply than others). Crossveins in radial space rather numerous, irregularly spaced in proximal two thirds; rare in distal portion, most form outer gradate series. M slightly incurved; forked far proximad origin of RP 1. One preserved crossvein connecting RP, M before fork. MA, MP parallel. MA branched (probably dichotomously) relatively close to wing margin. MP pectinately branched, with four branches; three crossveins between them. CuA long, pectinately branched with five long branches; four crossveins between them. CuP long, parallel to posterior margin; pectinate branched with six long branches; no crossveins between these. Branches of CuA, CuP incompletely preserved; some rather deeply forked. CuP space more than twice wider than intracubital space. Several irregularly spaced crossveins between CuA, CuP. AA 3 + 4 long, pectinately branched, with three branches; preserved branches shallowly forked; proximal branch deeply forked. AA 3 + 4, CuP closely approach basally. AP 1 + 2 deeply forked. Crossveins between RP 1 and MA, MA and MP, MP and CuA, CuA and CuP rather numerous, irregularly spaced. One crossvein between CuP and AA 3 + 4, AA 3 + 4 and AP 1 + 2, AP 1 + 2 and AP 3 + 4. Wing color pattern: two spots near hind wing margin, at MA, at outer gradate series in radial space. Hind wing about 37 mm long, 10.4 mm wide. Trichosors preserved in distal part of anterior wing margin. Costal space narrower than in forewing. Subcostal veinlets simple proximally, medially; once or twice shallowly forked distally. ScP, RA stout for entire length. Subcostal space moderately broad; slightly dilated distally; no crossveins detected. ScP + RA entering margin well after wing apex; its veinlets dichotomously branched; no crossveins connecting these. RP smooth, not zigzagged; originating near wing base. Nine crossveins between RA, RP before fusion of ScP + RA detected, widely spaced; not detected after fusion. RP with 12 branches; RP 1 to RP 4 deeply forked. Crossveins in radial space rather numerous, irregularly spaced in proximal proximally; rare distally, most of them form outer gradate series. Origin of M not preserved. MA branched relatively close to wing margin. MP pectinately branched, with four branches, two deeply forked; two crossveins between these. CuA long, pectinately branched, with eight long branches; all but one deeply forked; seven crossveins between them forming gradate series. CuP incompletely preserved, short, probably deeply forked. Anal veins not preserved. Crossveins between RP 1 and MA, MA and MP, MP and CuA rather numerous, irregularly spaced. One crossvein between CuP and AA 3 + 4, AA 3 + 4 and AP 1 + 2, AP 1 + 2 and AP 3 + 4.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	materials_examined	Material. Holotype CNU-NEU-NN 2011116 PC (part, counterpart), four well preserved wings and part of a poorly preserved thorax and abdomen. Paratype CNU-NEU-NN 2011117 PC (part, counterpart), a forewing and hind wing, overlapping. Both specimens are deposited in CNUB.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	etymology	Etymology. The specific epithet is formed from the Latin bi, two, and maculatus, spotted, in reference to the presence of two small spots on the forewing.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	materials_examined	Type locality and horizon. Daohugou Village, Wuhua Township, Ningcheng County, Inner Mongolia, China. Middle Jurassic, Bathonian / Callovian; Jiulongshan Formation.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF97FFB4FF28FF51FA35FC27.taxon	discussion	Remarks. The new species is assigned to Nymphites based on the strong similarity of its hind wing venation to that of the type species N. priscus, e. g., irregularly arranged crossveins in the radial space; the four pectinate branches of MP; the presence of a series of gradate crossveins between branches of CuA; short CuP. Nevertheless, this assignment is preliminary because of the incompleteness of the holotype. The holotype and paratypes of Nymphites bimaculatus sp. nov. have similar venation in general, and identical forewing patterning (two small spots). Their venation differs in details, but these differences do not extend beyond those of the holotype left and right wings. Although subcostal crossveins in this species are not confidently detected, we assume that this is likely an artefact of preservation, and that a few fine crossveins (not distinctly visible) and features of thyridiate crossveins (ScP with small scarce projections posteriorly) are probably present. We explain the clear difference in width noted between the right (wider) and left (narrower) wings as a result of post-mortem plastic distortion, extension of the matrix approximately along the plane of the length of the left wings. Therefore, we estimate that the actual forewing length of the holotype is probably 34 – 35 mm (not 37 mm).	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF9EFFB2FF28FF51FE21FCB4.taxon	description	Description. Thorax, abdomen fragmentary preserved; details not visible. Meso-, metafemora long, rather swollen. Mesotibiae relatively stout, short. Metatibiae long, slender with short terminal spurs (Fig. 4 B). All preserved segments covered with dense, relatively short hairs. Hind wing 25.8 mm long, 10.2 mm wide as preserved. Trichosors not preserved. Costal space rather narrow. Subcostal veinlets mostly simple in proximal 2 / 3; once or twice shallowly forked distally. ScP, RA stout for entire length. Subcostal space moderately broad; no crossveins detected. RP smooth, not zigzagged; originating near wing base. Eleven irregularly arranged crossveins between RA, RP before fusion of ScP + RA; not detected after fusion. RP with eight preserved branches (estimated 11 – 12 complete). Crossveins in radial space rather rare, irregularly spaced in proximal proximally. MA, MP fragmentarily preserved, branching not preserved. Intermedial crossveins long, somewhat oblique. CuA long, pectinate branched with nine preserved long branches, some deeply forked; branches inclined to CuA at rather acute angle; all preserved branches connect by one crossveins forming gradate series. CuP fragmentarily preserved, short. Anal veins not preserved.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF9EFFB2FF28FF51FE21FCB4.taxon	materials_examined	Material. CNU-NEU-NN 2011118 PC (part, counterpart), deposited in CNUB. Fragmentarily preserved body with two incomplete hind wings.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
03FB87CBFF9EFFB2FF28FF51FE21FCB4.taxon	discussion	Remarks. This species is clearly not conspecific with Nymphites bimaculatus sp. nov. It differs from the latter by the more acute angle between Cu and its branches, more scarce crossveins, and probably wider hind wing. However, as this specimen is very incompletely preserved, we refrain from naming the species pending more complete specimens.	en	Shi, Chaofan, Makarkin, Vladimir N., Yang, Qiang, Archibald, S. Bruce, Ren, Dong (2013): New species of Nymphites Haase (Neuroptera: Nymphidae) from the Middle Jurassic of China, with a redescription of the type species of the genus. Zootaxa 3700 (3): 393-410, DOI: 10.11646/zootaxa.3700.3.4
