taxonID	type	description	language	source
03FD357F970AFFE377927061A028FA20.taxon	description	Habitat: emergent or collected from many trees including maple, American elm, ash, oak, and eastern redbud; collected on but not reared from gymnospermous plant genera. Collection Method: emergence.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F970AFFE377927061A028FA20.taxon	biology_ecology	Biology: infests heartwood, collected from stumps, polyphagous. Present Study: not significantly as- sociated with any substrate or forest type. References: Packard 1890; Blatchley 1910; Nelson 1975; Downie and Arnett 1996; MacRae 2006; Nelson et al. 2008; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9709FFE0779276C2A01CFB60.taxon	description	Range: eastern North America, south to Florida, west to Texas and Kansas. Habitat: recorded from 19 + trees, shrubs, and vines including maple, hickory, oak, walnut, ash, cherry, and mulberry. Collection Method: emergence. Biology: flight period March to August. Present Study: significantly higher abundance in FWD 1 and secondary forest. References: Linsley and Chemsak 1984, 1997; Downie and Arnett 1996; Peck and Thomas 1998; Lingafelter 2007; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9709FFE0779270A1A0E6F980.taxon	biology_ecology	Biology: flight period June to August, feeds under bark and enters wood to pupate. Present Study: significantly higher abundance in primary forest. References: Linsley and Chemsak 1984, 1997; Downie and Arnett 1996; Turnbow and Thomas 2002; Lingafelter 2007; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9709FFE0779273C2A62AF8A0.taxon	biology_ecology	Biology: flight period April to September. Present Study: significantly higher abundance in primary forest. References: Packard 1890; Linsley and Chemsak 1995, 1997; Downie and Arnett 1996; Peck and Thomas 1998; Lingafelter 2007.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9708FFE177927642A038FBE0.taxon	description	Range: northern and eastern North America, British Columbia east to Maine, south to Texas. Habitat: under bark of maple, beech, and spruce. Collection Method: emergence, searching under bark of dead logs. Biology: collected from numerous fungi, larvae collected in July under bark. Present Study: significantly higher abundance in primary forest. References: Blatchley 1910; Lawrence and Stephan 1975; Lawrence 1982 a; Downie and Arnett 1996; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9708FFE1779275A2A5BCFE20.taxon	description	Range: eastern North America, south to Florida, west to Ontario. Habitat: recorded from birch, hornbeam, Ostrya virginiana (Mill.) K. Koch, and pine. Collection Method: emergence. Biology: flight period May to August, attracted to flowers including goatsbeard, hydrangea, grape, false Solomon’s seal. Present Study: significantly higher abundance in FWD 2. References: Linsley and Chemsak 1976, 1997; Downie and Arnett 1996; Peck and Thomas 1998; Turnbow and Thomas 2002; Lingafelter 2007; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9708FFEE779273A1A0C7FE80.taxon	description	Octotemnus laevis Casey (Fig. 22) Range: widespread throughout northern North America from Alaska to Nova Scotia, south into California, Kansas, and Alabama. Habitat: recorded from 14 + fungi, especially associated with Coriolus spp. Collection Method: emergence. Biology: breeds in polypore fungi. Present Study: significantly higher abundance in FWD 1 and primary forest. References: Blatchley 1910; Lawrence 1971, 1973, 1982 b; Downie and Arnett 1996; Thayer and Lawrence 2002; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9706FFEF77927442A741FDE0.taxon	discussion	Comments. Cophes Champion needs to be revised and redefined. The last treatment (Sleeper 1955) does not cover all species in North America. Range: eastern North America, south to Florida and west to Texas. Habitat: emergent from hickory limbs dead two years and Cassia sp., collected from maple, woods trash, hollow tree. Collection Method: at lights, sifting, emergence. Biology: unknown other than host plants. Present Study: significantly higher abundance in FWD 1 and primary forest. References: Blatchley and Leng 1916 (as Cryptorhynchus fallax LeConte); Downie and Arnett 1996; Peck and Thomas 1998; Anderson 2002; Ciegler 2010.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9706FFEF779276E2A7E0FB60.taxon	description	Range: eastern North America, south to Florida and west to Texas. Habitat: reported from 27 + tree species including hardwoods and softwoods (see Atkinson et al. 1990 for a complete list). Collection Method: Lindgren funnel trap with ethanol, MV and UV lights, emergence. Biology: introduced species from Asia, first collected in North America in 1988. Present Study: not significantly associated with any substrate or forest type. References: Atkinson et al. 1990; Downie and Arnett 1996; Peck and Thomas 1998; Vandenberg et al. 2000; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9706FFEF779271A2A596FA80.taxon	description	Collection Method: emergence, Frontalin-baited trap. Biology: introduced species from Asia, reported to attack cut or injured trees 1.5 cm diameter to large logs. Present Study: significantly higher abundance in secondary forest. References: Wood 1982 (and references therein); Peck and Thomas 1998; Rabaglia 2003.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9706FFEF779270C2A0B5F8E0.taxon	description	Xyloterinus politus (Say) (Fig. 33) Range: eastern North America, south to Florida, west to Minnesota. Habitat: numerous hardwoods including maple, hickory, beech, oak, elm, occasionally pine. Collection Method: emergence, traps.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9706FFEF779270C2A0B5F8E0.taxon	biology_ecology	Biology: monogamous. Present Study: significantly higher abundance in CWD 1 and primary forest.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9705FFEC77927742A11FFD20.taxon	description	Collection Method: collected at lights, emergence, beaten from vegetation at the margins of low woods.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9705FFEC77927742A11FFD20.taxon	biology_ecology	Biology: poorly known. Present Study: significantly higher abundance in CWD 2 and primary forest.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9705FFEC77927662A778FC60.taxon	description	Collection Method: beating foliage, emergence. Biology: poorly known. Present Study: not significantly associated with any substrate or forest type. References: LeConte 1853 (as Elater luteolus LeConte); Dietrich 1945; Downie and Arnett 1996.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9705FFEC77927362A7FFF8E3.taxon	description	Range: Sevier Co., Tennessee, and Macon Co., North Carolina. Habitat: rotten wood, very rarely in leaf litter. Collection Method: dung trap, sifting / Berlese litter and CWD 5, emergence chamber. Biology: adults have unique suctorial mouthparts, possibly feed on slime molds. Ferro et al. (2012 a): significantly higher abundance in CWD 5, primary forest, and spring; previously known from five specimens (Vit 1995 and C. E. Carlton collection). Present Study: significantly higher abundance in FWD 2, CWD 3 – 4, and primary forest. References: Vit 1995.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9704FFED77927062A51EF963.taxon	description	Bacanius tantillus LeConte (Fig. 43) Range: probably throughout eastern United States, reported from Pennsylvania, New York, Indiana, Tennessee, and Louisiana. Habitat: reported from leaf litter and decaying wood. Collection Method: emergence, sifting / Berlese. Biology: reported from dry organic debris in tree cavities and sawdust piles, feeds mainly on fungal spores. Present Study: not significantly associated with any substrate or forest type. References: Horn 1873; Casey 1893 (key to most species); Blatchley 1910; Wenzel 1960; Downie and Arnett 1996; Peck and Thomas 1998; Kovarik and Caterino 2002. LEIODIDAE Leiodinae	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9704FFED77927462A05AFE40.taxon	biology_ecology	Biology: overwinter as larvae. Present Study: significantly higher abundance in primary forest.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9704FFED77927482A5CAFB40.taxon	description	Melasis pectinicornis Melsheimer (Fig. 41) Range: northeastern North America, south to Florida, west to Louisiana. Habitat: reported from maple, blackgum, oak, elm, birch, and beech. Collection Method: emergence. Biology: under bark of fallen maple and blackgum, emergent from dead birch and beech. Present Study: significantly higher abundance in CWD 2 and secondary forest. References: Dury 1888; Blatchley 1910; Kirk 1922; Hopping 1926; Peterson 1960 (illustrated larva); Downie and Arnett 1996; Peck and Thomas 1998; Muona 2000. Microrhagus subsinuatus LeConte (Fig. 42) Range: northeastern North America, south to North Carolina, west to Wisconsin. Habitat: emergence from dead decayed beech, collected from alder, swept from milkweed. Collection Method: emergence, flight intercept trap, on vegetation. Biology: reported to be active on dead beech in daytime in June. Present Study: not significantly associated with any substrate or forest type. References: Dury 1888; Knull 1946 (as Dirhagus imperfectus (LeConte )); Downie and Arnett 1996; Muona 2000; Majka 2007 b; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9703FFEA779277E1A603FC00.taxon	description	Hypulus simulator Newman (Fig. 46) Range: northeastern North America south to Mississippi. Habitat: poorly known. Collection Method: emergence. Biology: poorly known. Present Study: not significantly associated with any substrate or forest type. References: Packard 1890; Downie and Arnett 1996 (as Mystaxus simulator Newman); Pollock 2002 a; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9702FFEB77927742A0D1FD00.taxon	description	Range: northeastern Canada and United States, south to Tennessee, west to Michigan. Habitat: reared from beech. Collection Method: emergence. Biology: unknown other than host plant. Present Study: significantly higher abundance in FWD 2 and secondary forest. References: Blatchley 1910; Champlain and Knull 1922; White 1962, 1976, 1982; Downie and Arnett 1996.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9702FFEB77927122A739FA80.taxon	description	Range: eastern North America, south to Florida, west to North Dakota. Habitat: larvae subcortical on the upper side of dead logs. Collection Method: at lights, beating foliage, emergence. Biology: reared larvae required wood inoculated by fungi to fully develop but could survive on sterilized wood, larval predators include millipedes, Elateridae larvae, and fly larvae, larvae are attached by the para- site Zelia vertebrata (Say) (Diptera: Tachinidae). Present Study: significantly higher abundance in FWD 2, CWD 2, and primary forest. References: Payne 1931; Young 1975, 1983, 2002; Downie and Arnett 1996; Peck and Thomas 1998; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9702FFEB779273E2A766F880.taxon	description	Range: Pennsylvania south to Florida and west to Missouri. Habitat: Collected from American chestnut and white oak. Collection Method: emergence chamber. Biology: under bark of moist rotting wood of logs or stumps, brachypterous. Present Study: not significantly associated with any substrate or forest type. References: Bell 1970; Bell and Bell 1985; Bousquet and Larochelle 1993; Downie and Arnett 1996; Peck and Thomas 1998; Ciegler 2000.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9701FFE8779277F8A1CCFC09.taxon	discussion	Comments. These specimens could not be reliably identified to genus. Aleocharinae are the largest subfamily of Staphylinidae with 21 tribes, 183 genera, and 1385 described species known from North America and are badly in need of a comprehensive revision. See Newton et al. (2001) and references therein, for further information about this subfamily. Range: throughout North America. Habitat: ubiquitous in terrestrial habitats. Collection Method: sifting / Berlese organic material, pitfall traps, bait traps, UV light, emergence chamber. Biology: virtually every mode of life (many very specialized) is known in this subfamily: free living, parasitic, herbivore, carnivore, fungivore, flier, walker, runner, swimmer, social, solitary, etc., but life history is almost unknown at the species level. Ferro et al. (2012 a): indifferent to substrate, forest type, and season. Present Study: significantly higher abun- dance FWD 1, CWD 2, and primary forest. References: Downie and Arnett 1996; Newton et al. 2001.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9701FFE977927174A7E8FEC0.taxon	discussion	Comments. These specimens could only be reliably identified to Athetini, which is a very large difficult tribe. Seevers’ (1978) characterization of the tribe and genera is inadequate. Currently 64 genera are recognized within the tribe in North America (Newton et al. 2001) but a complete revision is needed. Gusarov (2002 a – e, 2003 a – e, 2004 a – b) has greatly contributed to our knowledge of many genera and Elven et al. (2010) provided the first molecular phylogeny of the tribe, but more work needs to be done. Range: throughout North America. Habitat: ubiquitous; decaying plants and animals, dung, bird and mammal nests, riparian areas, ant nests, under bark and logs. Collection Method: sifting / Berlese organic material, emergence. Biology: unknown; predators. Ferro et al. (2012 a): significantly higher abundance in leaf litter and spring, indifferent to forest type. Present Study: not significantly associ- ated with any substrate or forest type. References: Seevers 1978; Downie and Arnett 1996; Newton et al. 2001; Gusarov 2002 a – e, 2003 a – e, 2004 a – b; Elven et al. 2010. Atheta spp. (Fig. 58) Comments. These specimens could only be reliably identified to the genus Atheta Thomson. Gusarov (2003 e) discussed the varying definitions of the genus and reviewed the types of the known species. Newton et al. (2001) report 176 species known in North America (as Xenota Mulsant and Rey). Range: throughout North America. Habitat: ubiquitous; decaying plants and animals, dung, bird and mam- mal nests, riparian areas, ant nests, under bark and logs. Collection Method: sifting / Berlese organic material, flight intercept trap, emergence. Biology: unknown, predators. Present Study: significantly higher abundance in CWD 2 and primary forest. References: Blatchley 1910; Downie and Arnett 1996 (as Xenota sp.); Peck and Thomas 1998; Newton et al. 2001 (as Xenota sp.); Gusarov 2003 e; Majka et al. 2011. Leptusa carolinensis Pace (Fig. 59) Range: northeastern North America south to North Carolina, west to Quebec. Habitat: occurs in sugar maple, red spruce, black spruce, and young, mature, and old growth red spruce / hemlock forests. Collection Method: pitfall traps, flight intercept traps, emergence. Biology: found under bark of beech, under maple log, in bracket fungi, associated with woody debris. Present Study: not significantly associated with any substrate or forest type. References: Pace 1989; Klimaszewski et al. 2004; Gouix and Klimaszewski 2007; Webster et al. 2009; Park et al. 2010.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9701FFE877927462A51CFE00.taxon	description	Habitat: damp, shaded sites in association with dead wood, hanging dead leaves, moss, under lichens. Collection Method: beating dead branches, emergence. Biology: adults feed on bark of unthrifty alder and maple twigs, larvae feed on inner brown bark (see Howden and Howden (1981) notes on life history). Present Study: significantly higher abundance in FWD 1 and primary forest. References: Blatchley 1910; Blair 1932; Howden and Howden 1981; Downie and Arnett 1996; Pollock 2002 b; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9700FFE977927082A6DCF880.taxon	description	Range: Kentucky, Mississippi, New York, Pennsylvania, Tennessee. Habitat: lives in nests of Lasius spp. ants (Hymenoptera: Formicidae); nests have been found in beech logs in advance stages of decay. Collection Method: sifting Lasius spp. ant nests, rarely sifting / Berlese forest litter. Biology: obligate myrmecophile on Lasius spp. ants; adults feed on fluids obtained from their adult and immature hosts; possibly feed on dead immature ants; see Park (1932 a) and Akre and Hill (1973) for interesting behavioral observations of the genus. Ferro et al. (2012 a): significantly higher abundance in secondary forest, indifferent to substrate or season, probably heavily influenced by their host. Present Study: significantly higher abundance in CWD 3 – 4 and secondary forest. References: Wickham 1901; Blatchley 1910; Park 1932 a (with notes on life history), 1935, 1964; Akre and Hill 1973; Hill et al. 1976; Downie and Arnett 1996; Newton et al. 2001.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F9700FFE9779271C2A5BCFA40.taxon	description	Habitat: under bark, in dead wood, forest litter. Collection Method: sifting litter, debris, and dead wood (with Berlese funnel), emergence chamber. Biology: unknown. Ferro et al. (2012 a): signifi- cantly higher abundance in CWD 5, secondary forest, and spring. Present Study: significantly higher abundance in FWD 2, CWD 3 – 4, and primary forest. References: Horn 1871 (as Glyptoma costale Erichson, figure and key to common species in North America); Blatchley 1910; Notman 1920; Irmler 1985; Downie and Arnett 1996 (figure is not T. costalis); Peck and Thomas 1998; Ferro and Gimmel 2011; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F973FFFD777927201A092FE60.taxon	description	Range: mostly midwest, northeast, and southeastern United States. Habitat: forest floor litter, moss, tree holes, rotting logs, and other moist habitats. Collection Method: sifting / Berlese litter, pitfalls, flight intercept traps, UV lights, looking under stones. Biology: adults and immatures feed on oribatid mites. Ferro et al. (2012 a): only Euconnus (Napochus) sp. was found in significantly higher abundance in leaf litter and secondary forest. Present Study: not significantly associated with any substrate or forest type. References: Blatchley 1910; Downie and Arnett 1996 (usefulness of keys uncertain); Peck and Thomas 1998; O’Keefe 2001; Grebennikov and Newton 2009; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F973EFFD777927761A030FD20.taxon	description	Habitat: found on old trees, fermenting sap under bark of oak, decaying wood of old fallen trees. Collection Method: at lights, malt trap, sifting / Berlese, emergence. Biology: poorly known. Present Study: significantly higher abundance in primary forest. References: Horn 1884 (as Philonthus apicalis (Say )); Blatchley 1910 (as Hesperus apicalis (Say )); Moore 1958; Frank 1983; Smetana 1995; Downie and Arnett 1996 (as H. apicalis); Peck and Thomas 1998; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F973EFFD777927662A5CAFC00.taxon	description	Range: eastern North America from Maine south to Mississippi, west to Texas and Iowa. Habitat: in leaf litter, wood trash, on fungus, underside of log. Collection Method: sifting / Berlese, emergence.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F973EFFD777927662A5CAFC00.taxon	biology_ecology	Biology: recorded from undetermined Agaricales. Present Study: significantly higher abundance in CWD 3 – 4 and primary forest. References: Campbell 1976; Newton 1984; Downie and Arnett 1996; Majka et al. 2011.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
03FD357F973DFFD477927422A7DBFDC0.taxon	discussion	Comments. The entire family, including this genus, is in need of revision. Range: reported from Massachusetts, New York, North Carolina, and Tennessee. Habitat: unknown. Collection Method: emergence. Biology: unknown, adults of the family are thought to be generalist pollen and mold feeders. Present Study: significantly higher abundance in FWD 1, CWD 1, and secondary forest. References: Blanchard 1917; Downie and Arnett 1996; Johnson 2002 a.	en	Ferro, Michael L., Gimmel, Matthew L., Harms, Kyle E., Carlton, Christopher E. (2012): Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA. Insecta Mundi 2012 (260): 1-8, DOI: 10.5281/zenodo.5175284
