taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03FD8227FFF5E30BFF78FD45FEEE8291.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF5E30BFF78FD45FEEE8291.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF4E30DFF78FE52FD3D85D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232530/files/figure.png	https://doi.org/10.5281/zenodo.11232530	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF4E30DFF78FE52FD3D85D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232517/files/figure.png	https://doi.org/10.5281/zenodo.11232517	FIGURE 7. Section 2 Jbel Issoumour	FIGURE 7. Section 2 Jbel Issoumour	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF4E30DFF78FE52FD3D85D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232515/files/figure.png	https://doi.org/10.5281/zenodo.11232515	FIGURE 6. Section 1 Jbel Ou-Driss	FIGURE 6. Section 1 Jbel Ou-Driss	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF4E30DFF78FE52FD3D85D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232532/files/figure.png	https://doi.org/10.5281/zenodo.11232532	PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF2E30EFF78FA87FBBC8058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232547/files/figure.png	https://doi.org/10.5281/zenodo.11232547	PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF2E30EFF78FA87FBBC8058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232517/files/figure.png	https://doi.org/10.5281/zenodo.11232517	FIGURE 7. Section 2 Jbel Issoumour	FIGURE 7. Section 2 Jbel Issoumour	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF2E30EFF78FA87FBBC8058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232549/files/figure.png	https://doi.org/10.5281/zenodo.11232549	PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF2E30EFF78FA87FBBC8058.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232553/files/figure.png	https://doi.org/10.5281/zenodo.11232553	PLATE 7. A–M Harpes lentigo n. sp., from a coral horizon 5 m (12.2 m down slope) below the “Morocconites Couche”, Tazoulait Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 4). A–G dorsal exoskeleton, thorax disarticulated and partially rolled, holotype, NHMUK It 29236.A, dorsal, B, anterior, C, posterior and D, lateral view. E, lateral view of eye lobe (uncoated), F, laterodorsal view of eye lobe (uncoated) and G, dorsal view of brim perforations. H–M Harpes lentigo n. sp., plate of three disarticulated dorsal exoskeletons, paratypes, NHMUK It 29237 (1 to 3) (Fig. 2B, Map 8, site 5). H, dorsal, I, anterodorsal and J, lateral view. K, dorsal view of brim perforations, L, lateral view of pleurae and M, anterior view of rim. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 7. A–M Harpes lentigo n. sp., from a coral horizon 5 m (12.2 m down slope) below the “Morocconites Couche”, Tazoulait Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 4). A–G dorsal exoskeleton, thorax disarticulated and partially rolled, holotype, NHMUK It 29236.A, dorsal, B, anterior, C, posterior and D, lateral view. E, lateral view of eye lobe (uncoated), F, laterodorsal view of eye lobe (uncoated) and G, dorsal view of brim perforations. H–M Harpes lentigo n. sp., plate of three disarticulated dorsal exoskeletons, paratypes, NHMUK It 29237 (1 to 3) (Fig. 2B, Map 8, site 5). H, dorsal, I, anterodorsal and J, lateral view. K, dorsal view of brim perforations, L, lateral view of pleurae and M, anterior view of rim. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF1E310FF78FF0CFBE68568.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232553/files/figure.png	https://doi.org/10.5281/zenodo.11232553	PLATE 7. A–M Harpes lentigo n. sp., from a coral horizon 5 m (12.2 m down slope) below the “Morocconites Couche”, Tazoulait Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 4). A–G dorsal exoskeleton, thorax disarticulated and partially rolled, holotype, NHMUK It 29236.A, dorsal, B, anterior, C, posterior and D, lateral view. E, lateral view of eye lobe (uncoated), F, laterodorsal view of eye lobe (uncoated) and G, dorsal view of brim perforations. H–M Harpes lentigo n. sp., plate of three disarticulated dorsal exoskeletons, paratypes, NHMUK It 29237 (1 to 3) (Fig. 2B, Map 8, site 5). H, dorsal, I, anterodorsal and J, lateral view. K, dorsal view of brim perforations, L, lateral view of pleurae and M, anterior view of rim. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 7. A–M Harpes lentigo n. sp., from a coral horizon 5 m (12.2 m down slope) below the “Morocconites Couche”, Tazoulait Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 4). A–G dorsal exoskeleton, thorax disarticulated and partially rolled, holotype, NHMUK It 29236.A, dorsal, B, anterior, C, posterior and D, lateral view. E, lateral view of eye lobe (uncoated), F, laterodorsal view of eye lobe (uncoated) and G, dorsal view of brim perforations. H–M Harpes lentigo n. sp., plate of three disarticulated dorsal exoskeletons, paratypes, NHMUK It 29237 (1 to 3) (Fig. 2B, Map 8, site 5). H, dorsal, I, anterodorsal and J, lateral view. K, dorsal view of brim perforations, L, lateral view of pleurae and M, anterior view of rim. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF1E310FF78FF0CFBE68568.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232555/files/figure.png	https://doi.org/10.5281/zenodo.11232555	PLATE 8. A–L Harpes cf. lentigo, El Otfal Formation, Eifelian, Jbel Issoumour. A–H from an horizon halfway between “Lobopyge Couche” and “Phylonix Couche”, dorsal exoskeleton of cephalon, NHMUK It 29239 (Fig. 2B, Map 7, site 14). A, dorsal, B, anterior and C, lateral view. D, anterior view of perforations on brim and genal roll, E, dorsal view of perforations on brim, F, lateral view of eye lobe (uncoated), G, lateral view of genal area (uncoated) and H, lateral view of the internal rim. I–L from the “Lobopyge Couche”, dorsal exoskeleton of cephalon, NHMUK It 29238 (Fig. 2B, Map 7, site 7). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 8. A–L Harpes cf. lentigo, El Otfal Formation, Eifelian, Jbel Issoumour. A–H from an horizon halfway between “Lobopyge Couche” and “Phylonix Couche”, dorsal exoskeleton of cephalon, NHMUK It 29239 (Fig. 2B, Map 7, site 14). A, dorsal, B, anterior and C, lateral view. D, anterior view of perforations on brim and genal roll, E, dorsal view of perforations on brim, F, lateral view of eye lobe (uncoated), G, lateral view of genal area (uncoated) and H, lateral view of the internal rim. I–L from the “Lobopyge Couche”, dorsal exoskeleton of cephalon, NHMUK It 29238 (Fig. 2B, Map 7, site 7). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFF1E310FF78FF0CFBE68568.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232530/files/figure.png	https://doi.org/10.5281/zenodo.11232530	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEFE311FF78FA17FC07849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232532/files/figure.png	https://doi.org/10.5281/zenodo.11232532	PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 2. A–D Harpes cf. boudibensis, from horizon 7, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 11). An enrolled dorsal exoskeleton NHMUK It 29225. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–M Harpes oudris, from horizon 5, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4). E–H, enrolled dorsal exoskeleton, paratype, NHMUK It 29240 (site 21). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations. I–M Harpes oudris, enrolled dorsal exoskeleton, holotype, NHMUK It 29241 (site 22). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations and M, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEFE311FF78FA17FC07849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232515/files/figure.png	https://doi.org/10.5281/zenodo.11232515	FIGURE 6. Section 1 Jbel Ou-Driss	FIGURE 6. Section 1 Jbel Ou-Driss	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEFE311FF78FA17FC07849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232598/files/figure.png	https://doi.org/10.5281/zenodo.11232598	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEFE311FF78FA17FC07849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232549/files/figure.png	https://doi.org/10.5281/zenodo.11232549	PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 6. A–H Harpes cf. oudris, dorsal exoskeleton with ventral surfaces exposed on anterior half of cephalon, NHMUK It 29242; from El Otfal Formation, Eifelian, horizon 4, Section 1 (Fig. 6), Jbel Ou-Driss (Fig. 2B, Map 4, site 10). A, dorsal, B, anterior and C, lateral view. D, lateral view of glabella (uncoated) and muscle insertion scars, E, ventral view of anterior half of cephalon, F, ventral view of hypostome in place, G, ventral view of anterior of rim, H, dorsal view of brim perforations. I–N Harpes cf. lahceni, dorsal exoskeleton of partial cephalon only, NHMUK It 29235, from “Erbenochile Couche”, Timrhanrhart Formation, Emsian, 23.8 km SW of Foum Zguid (Fig. 2A, Map 2, site 6). I, dorsal, J, anterior and K, lateral view. L, dorsal view of brim perforations and M and N, dorsal and lateral view of eye lobe respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEFE311FF78FA17FC07849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232530/files/figure.png	https://doi.org/10.5281/zenodo.11232530	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 1. A–Q Harpes boudibensis n. sp., El Otfal Formation, Eifelian. A–D, articulated dorsal exoskeleton, paratype, NHMUK It 29226, type locality horizon 1 (site 1). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J, dorsal exoskeleton, pygidium enrolled, paratype, NHMUK It 29228, type locality, horizon 2 (site 5). E, anterior, F, dorsal and G, lateral view. H, ventral view of anterior part of cephalon, I, hypostome and J, dorsal view of brim perforations. K–Q, enrolled articulated dorsal exoskeleton, holotype, NHMUK It 29227, horizon 1, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 4). K, dorsal, L, anterior, M, lateral view, N, anterior dorsal view of genal roll and brim perforations, O, dorsal view of brim perforations, P, lateral dorsal view of eye lobe and muscle impression areas at S0, S1, S2 and S3, and Q, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEEE312FF78FC43FD9283D0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232561/files/figure.png	https://doi.org/10.5281/zenodo.11232561	PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 9. A–I Harpes prescheri n. sp., from an horizon 100 m upslope of “Couche Rouge”, close to the Pyschopyge horizon, Er Remlia Formation, Emsian, Jbel Issoumour (Fig. 2B, Map 7). A–E dorsal exoskeleton of cephalon and disarticulated thorax, holotype, NHMUK It 29244 (site 9). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of eye lobe (uncoated). F–I dorsal exoskeleton of cephalon, paratype, NHMUK It 29243 (site 8);, F dorsal, G, anterior and H, lateral view. I, dorsal view of brim perforations. J–N Harpes cf. prescheri, dorsal exoskeleton of cephalon, NHMUK It 29245, from “Lobopyge Couche”, El Otfal Formation, Emsian/Eifelian boundary, Jbel Issoumour (Fig. 2B, Map 7, site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEDE313FF78FC8FFD7F80E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232545/files/figure.png	https://doi.org/10.5281/zenodo.11232545	PLATE 4. A–D Harpes supercilium n. sp., partial dorsal exoskeleton, paratype, NHMUK It 29248, horizon 10, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 14). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Harpes sphenocephalus n. sp., dorsal exoskeleton and partial thorax enrolled, holotype, NHMUK It 29246, horizon 1, Section 1 (Fig. 6), Er Remlia Formation, lower Emsian, Jbel Ou-Driss (Fig. 2B, Map 4, site 15). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, anterior view of genal roll, J, anterior-dorsal view of perforations across brim, exsag. K, lateral view of eye lenses. L–O Harpes cf. sphenocephalus, partial dorsal exoskeleton, NHMUK It 29247, “Cherurid Couche”, Er Remlia Formation, lower Emsian Jbel Issoumour (Fig. 2B, Map 7, site 18). L, dorsal, M, anterior and N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 4. A–D Harpes supercilium n. sp., partial dorsal exoskeleton, paratype, NHMUK It 29248, horizon 10, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 14). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Harpes sphenocephalus n. sp., dorsal exoskeleton and partial thorax enrolled, holotype, NHMUK It 29246, horizon 1, Section 1 (Fig. 6), Er Remlia Formation, lower Emsian, Jbel Ou-Driss (Fig. 2B, Map 4, site 15). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, anterior view of genal roll, J, anterior-dorsal view of perforations across brim, exsag. K, lateral view of eye lenses. L–O Harpes cf. sphenocephalus, partial dorsal exoskeleton, NHMUK It 29247, “Cherurid Couche”, Er Remlia Formation, lower Emsian Jbel Issoumour (Fig. 2B, Map 7, site 18). L, dorsal, M, anterior and N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEDE313FF78FC8FFD7F80E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232515/files/figure.png	https://doi.org/10.5281/zenodo.11232515	FIGURE 6. Section 1 Jbel Ou-Driss	FIGURE 6. Section 1 Jbel Ou-Driss	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232539/files/figure.png	https://doi.org/10.5281/zenodo.11232539	PLATE 3. A–K Harpes supercilium n. sp., from horizon 5, Section 2 (Fig. 7), El Otfal Formation, Eifelian, Jbel Issoumour (Fig. 2B, Map 7). A–F, articulated dorsal exoskeleton, paratype, NHMUK It 29251 (site 3). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of glabella muscle insertion scars and F, lateral view of eye lobe. G–K, dorsal exoskeleton thorax enrolled, holotype, NHMUK It 29252 (site 10). G, dorsal view, H, anterior view, I, lateral view, J, dorsal view of brim perforations, K, lateral view of eye lobe. L–P Harpes supercilium n. sp., from horizon 8, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 24), dorsal exoskeleton of cephalon, paratype, NHMUK It 29250. L, dorsal, M, anterior and N, lateral view. O, lateral view of muscle insertion scars on glabella, P, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 3. A–K Harpes supercilium n. sp., from horizon 5, Section 2 (Fig. 7), El Otfal Formation, Eifelian, Jbel Issoumour (Fig. 2B, Map 7). A–F, articulated dorsal exoskeleton, paratype, NHMUK It 29251 (site 3). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations, E, lateral view of glabella muscle insertion scars and F, lateral view of eye lobe. G–K, dorsal exoskeleton thorax enrolled, holotype, NHMUK It 29252 (site 10). G, dorsal view, H, anterior view, I, lateral view, J, dorsal view of brim perforations, K, lateral view of eye lobe. L–P Harpes supercilium n. sp., from horizon 8, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 24), dorsal exoskeleton of cephalon, paratype, NHMUK It 29250. L, dorsal, M, anterior and N, lateral view. O, lateral view of muscle insertion scars on glabella, P, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232545/files/figure.png	https://doi.org/10.5281/zenodo.11232545	PLATE 4. A–D Harpes supercilium n. sp., partial dorsal exoskeleton, paratype, NHMUK It 29248, horizon 10, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 14). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Harpes sphenocephalus n. sp., dorsal exoskeleton and partial thorax enrolled, holotype, NHMUK It 29246, horizon 1, Section 1 (Fig. 6), Er Remlia Formation, lower Emsian, Jbel Ou-Driss (Fig. 2B, Map 4, site 15). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, anterior view of genal roll, J, anterior-dorsal view of perforations across brim, exsag. K, lateral view of eye lenses. L–O Harpes cf. sphenocephalus, partial dorsal exoskeleton, NHMUK It 29247, “Cherurid Couche”, Er Remlia Formation, lower Emsian Jbel Issoumour (Fig. 2B, Map 7, site 18). L, dorsal, M, anterior and N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 4. A–D Harpes supercilium n. sp., partial dorsal exoskeleton, paratype, NHMUK It 29248, horizon 10, Section 1 (Fig. 6), El Otfal Formation, Eifelian, Jbel Ou-Driss (Fig. 2B, Map 4, site 14). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Harpes sphenocephalus n. sp., dorsal exoskeleton and partial thorax enrolled, holotype, NHMUK It 29246, horizon 1, Section 1 (Fig. 6), Er Remlia Formation, lower Emsian, Jbel Ou-Driss (Fig. 2B, Map 4, site 15). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, anterior view of genal roll, J, anterior-dorsal view of perforations across brim, exsag. K, lateral view of eye lenses. L–O Harpes cf. sphenocephalus, partial dorsal exoskeleton, NHMUK It 29247, “Cherurid Couche”, Er Remlia Formation, lower Emsian Jbel Issoumour (Fig. 2B, Map 7, site 18). L, dorsal, M, anterior and N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232517/files/figure.png	https://doi.org/10.5281/zenodo.11232517	FIGURE 7. Section 2 Jbel Issoumour	FIGURE 7. Section 2 Jbel Issoumour	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232515/files/figure.png	https://doi.org/10.5281/zenodo.11232515	FIGURE 6. Section 1 Jbel Ou-Driss	FIGURE 6. Section 1 Jbel Ou-Driss	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFEBE315FF78F8EBFBD68087.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE9E317FF78F8EBFEB8854C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232563/files/figure.png	https://doi.org/10.5281/zenodo.11232563	PLATE 10. A–D Lioharpes venulosa, from the Natural History Museum London collection, NHMUK In 4180. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Lioharpes galea n. sp., from coral in an upper Pragian horizon, Ihandar formation, Jbel Otfal (Fig. 2B, Map 9, site 10). E–H partial dorsal exoskeleton, holotype, NHMUK It 29268, E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforation. I–K dorsal exoskeleton, paratype, NHMUK It 29269, from type locality (site 11), I, dorsal, J, anterior and K, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 10. A–D Lioharpes venulosa, from the Natural History Museum London collection, NHMUK In 4180. A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Lioharpes galea n. sp., from coral in an upper Pragian horizon, Ihandar formation, Jbel Otfal (Fig. 2B, Map 9, site 10). E–H partial dorsal exoskeleton, holotype, NHMUK It 29268, E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforation. I–K dorsal exoskeleton, paratype, NHMUK It 29269, from type locality (site 11), I, dorsal, J, anterior and K, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE9E317FF78F8EBFEB8854C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232697/files/figure.png	https://doi.org/10.5281/zenodo.11232697	PLATE 54. A–D Lioharpes venulosus (Hawle & Corda, 1847); type locality Upper Koněprusy Limestones, (Pragian) Koněprusy, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L12545. A dorsal view, B lateral view, C anterior view and D dorsal view of brim perforations. E–H Bohemoharpes buphthalmus (Bouček, 1935). Type locality the horizon with Ananaspis fecunda (Ludlow stage), Dlouhá hora near Beroun, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L17145. E, dorsal view, F lateral view, G anterior view and H dorsal view of brim perforations. I–L Bohemoharpes naumanni (Barrande 1852); type horizon and zone Cyrtograptus lundgreni zone (Upper Wenlock) Lodenice near Beroun, Czech Republic; dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague, lectotype (subsequent designation, Prantl and Přibyl, 1954) L6127, IT 48. I dorsal view, J anterior view, K lateral view and L dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl,1954 pl. iv fig. 1 and Ebach and McNamara, 2002 Fig 2A. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 54. A–D Lioharpes venulosus (Hawle & Corda, 1847); type locality Upper Koněprusy Limestones, (Pragian) Koněprusy, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L12545. A dorsal view, B lateral view, C anterior view and D dorsal view of brim perforations. E–H Bohemoharpes buphthalmus (Bouček, 1935). Type locality the horizon with Ananaspis fecunda (Ludlow stage), Dlouhá hora near Beroun, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L17145. E, dorsal view, F lateral view, G anterior view and H dorsal view of brim perforations. I–L Bohemoharpes naumanni (Barrande 1852); type horizon and zone Cyrtograptus lundgreni zone (Upper Wenlock) Lodenice near Beroun, Czech Republic; dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague, lectotype (subsequent designation, Prantl and Přibyl, 1954) L6127, IT 48. I dorsal view, J anterior view, K lateral view and L dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl,1954 pl. iv fig. 1 and Ebach and McNamara, 2002 Fig 2A. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE8E318FF78FAFFFCB58580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232567/files/figure.png	https://doi.org/10.5281/zenodo.11232567	PLATE 11. A–M Lioharpes morocconensis n. sp. A–E dorsal exoskeleton of cephalon only, paratype, NHMUK It 29272, from a lower Emsian horizon, Boutiskaouine Formation, Jbel el Otfal (Fig. 2B, Map 9, site17). A, dorsal, B, anterior and C, lateral view. D, lateral view of eye lobe and E, dorsal view of brim perforation. F–J dorsal exoskeleton of cephalon only, holotype, NHMUK It 29271 (1), from type locality and horizon (Fig. 2B, Map 9, site 6). F, dorsal, G, lateral and H, anterior view. I, dorsal view of glabella and J dorsal view of brim perforation. K–L dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29271 (2)), on same plate as holotype, K, dorsal view and L, lateral view of eye lobe. M, dorsal view of NHMUK It 29271 (1 and 2) plate. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 11. A–M Lioharpes morocconensis n. sp. A–E dorsal exoskeleton of cephalon only, paratype, NHMUK It 29272, from a lower Emsian horizon, Boutiskaouine Formation, Jbel el Otfal (Fig. 2B, Map 9, site17). A, dorsal, B, anterior and C, lateral view. D, lateral view of eye lobe and E, dorsal view of brim perforation. F–J dorsal exoskeleton of cephalon only, holotype, NHMUK It 29271 (1), from type locality and horizon (Fig. 2B, Map 9, site 6). F, dorsal, G, lateral and H, anterior view. I, dorsal view of glabella and J dorsal view of brim perforation. K–L dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29271 (2)), on same plate as holotype, K, dorsal view and L, lateral view of eye lobe. M, dorsal view of NHMUK It 29271 (1 and 2) plate. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE8E318FF78FAFFFCB58580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232571/files/figure.png	https://doi.org/10.5281/zenodo.11232571	PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE7E319FF78FB3EFCD0849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232640/files/figure.png	https://doi.org/10.5281/zenodo.11232640	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE7E319FF78FB3EFCD0849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232579/files/figure.png	https://doi.org/10.5281/zenodo.11232579	PLATE 15. A-N Lioharpes saredra n. sp. A–C partial dorsal exoskeleton of cephalon from “Odontochile Couche” Boutiskaouine Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 7), paratype, NHMUK it 29276. A, dorsal and B, lateral view. C, dorsal view of brim perforations. D–J disarticulated dorsal exoskeleton, from the Destombina Horizon, Tazoulait Formation, upper Emsian,Jbel Shebras (Fig. 2B, Map 12, site 1), paratype, NHMUK It 29278. D, dorsal, E, lateral and F, anterior view. Dorsal view of: G, genal area showing internal mould, H, brim perforations, I, pygidium and J external rim. K–N dorsal exoskeleton of cephalon only, from coral at the top of the Pragian, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 1), paratype, NHMUK It 29277. K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 15. A-N Lioharpes saredra n. sp. A–C partial dorsal exoskeleton of cephalon from “Odontochile Couche” Boutiskaouine Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 7), paratype, NHMUK it 29276. A, dorsal and B, lateral view. C, dorsal view of brim perforations. D–J disarticulated dorsal exoskeleton, from the Destombina Horizon, Tazoulait Formation, upper Emsian,Jbel Shebras (Fig. 2B, Map 12, site 1), paratype, NHMUK It 29278. D, dorsal, E, lateral and F, anterior view. Dorsal view of: G, genal area showing internal mould, H, brim perforations, I, pygidium and J external rim. K–N dorsal exoskeleton of cephalon only, from coral at the top of the Pragian, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 1), paratype, NHMUK It 29277. K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE7E319FF78FB3EFCD0849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232674/files/figure.png	https://doi.org/10.5281/zenodo.11232674	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE7E319FF78FB3EFCD0849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232672/files/figure.png	https://doi.org/10.5281/zenodo.11232672	PLATE 43. A–B Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 13, site 11). A, anterior and B, lateral view. C– P Pinnuloharpes aff. haustrum, from base of upper Emsian, Hamar Laghdad. C–L dorsal exoskeleton with thorax enrolled, NHMUK It 29347 (1) (Fig. 2B, Map 11, site 7). C, dorsal, D, lateral and E, anterior view. F, dorsal view of brim perforations, G, anterior view of external rim. H, I and J dorsal view of sculpture on dorsal surface of genal area, brim perforations and occipital node, respectively. K, anterior view of perforations on the genal roll and L dorsal view of eye lobe. M–P ventral exoskeleton of cephalon only, NHMUK It 29347 (2), from same site as NHMUK It 29347 (1). M, ventral and N, lateral view. O and P, ventral view of hypostome and external rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 43. A–B Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 13, site 11). A, anterior and B, lateral view. C– P Pinnuloharpes aff. haustrum, from base of upper Emsian, Hamar Laghdad. C–L dorsal exoskeleton with thorax enrolled, NHMUK It 29347 (1) (Fig. 2B, Map 11, site 7). C, dorsal, D, lateral and E, anterior view. F, dorsal view of brim perforations, G, anterior view of external rim. H, I and J dorsal view of sculpture on dorsal surface of genal area, brim perforations and occipital node, respectively. K, anterior view of perforations on the genal roll and L dorsal view of eye lobe. M–P ventral exoskeleton of cephalon only, NHMUK It 29347 (2), from same site as NHMUK It 29347 (1). M, ventral and N, lateral view. O and P, ventral view of hypostome and external rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE6E31AFF78FC42FC9B81E8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232571/files/figure.png	https://doi.org/10.5281/zenodo.11232571	PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 12. A–F Lioharpes cf. morocconensis, from a lower Emsian horizon, Boutiskaouine Formation, 28.3 km SSW of Rissani (Fig. 2B, Map 14, site 10), dorsal exoskeleton of partial cephalon, NHMUK It 29273. A, dorsal, B, anterior and C and D, lateral view. E, lateral view of eye lobe and F, dorsal view of brim perforation. G–M Lioharpes ammari n. sp., from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 9), dorsal exoskeleton with pygidium enrolled, holotype, NHMUK It 29263. G, dorsal, H, anterior and I, lateral view. J and K, dorsal and lateral views, respectively, of eye lobe. L and M, dorsal view of brim perforations and exterior rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE6E31AFF78FC42FC9B81E8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232573/files/figure.png	https://doi.org/10.5281/zenodo.11232573	PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE6E31AFF78FC42FC9B81E8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232515/files/figure.png	https://doi.org/10.5281/zenodo.11232515	FIGURE 6. Section 1 Jbel Ou-Driss	FIGURE 6. Section 1 Jbel Ou-Driss	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE6E31AFF78FC42FC9B81E8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232695/files/figure.png	https://doi.org/10.5281/zenodo.11232695	PLATE 53. A–F Lioharpes ammari n. sp. (?), a small holaspid or late meraspid from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 25), dorsal exoskeleton of partial cephalon, NHMUK It 29266. A, dorsal view, B anterior view and C and D lateral views. E, dorsal view of brim perforations and F, laterodorsal view of eye lobe. G–L Eoharpes beningensis (Barrande 1872), from Dobrotivá beds, Llandeilo, Lower Ordovician Svatá Dobrovtivá central Bohemia. G–H partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173331. G dorsal view and H, lateral view. I–L partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173316. I, dorsal view and L, lateral view. J, dorsal view of brim perforations and K, lateral view of eye lobe and caeca on genal area. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 53. A–F Lioharpes ammari n. sp. (?), a small holaspid or late meraspid from an upper Emsian horizon, Tazoulait Formation, North flank of Jbel Ou-Driss (Fig. 2B, Map 4, site 25), dorsal exoskeleton of partial cephalon, NHMUK It 29266. A, dorsal view, B anterior view and C and D lateral views. E, dorsal view of brim perforations and F, laterodorsal view of eye lobe. G–L Eoharpes beningensis (Barrande 1872), from Dobrotivá beds, Llandeilo, Lower Ordovician Svatá Dobrovtivá central Bohemia. G–H partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173331. G dorsal view and H, lateral view. I–L partial dorsal exoskeleton, Museum of Comparative Zoology Harvard University, IP 173316. I, dorsal view and L, lateral view. J, dorsal view of brim perforations and K, lateral view of eye lobe and caeca on genal area. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232577/files/figure.png	https://doi.org/10.5281/zenodo.11232577	PLATE 14. A–N Lioharpes saredra n. sp., from “Odontochile Couche”, Boutiskaouine Formation, Emsian, Jbel Saredrar (Fig. 2B, Map 5). A–I dorsal exoskeleton, thorax disarticulated and enrolled, holotype, NHMUK It 29274 (site 1).A, dorsal, B, anterior, C, lateral and H, posterior view. D, posteroventral view of prolongation, E, dorsal surface of genal area. F, dorsal view of brim perforation, G, lateral view of eye lobe, I, dorsal lateral view of pleurae. J–N dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29275 (site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 14. A–N Lioharpes saredra n. sp., from “Odontochile Couche”, Boutiskaouine Formation, Emsian, Jbel Saredrar (Fig. 2B, Map 5). A–I dorsal exoskeleton, thorax disarticulated and enrolled, holotype, NHMUK It 29274 (site 1).A, dorsal, B, anterior, C, lateral and H, posterior view. D, posteroventral view of prolongation, E, dorsal surface of genal area. F, dorsal view of brim perforation, G, lateral view of eye lobe, I, dorsal lateral view of pleurae. J–N dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29275 (site 2). J, dorsal, K, anterior and L, lateral view. M, lateral view of glabella (uncoated) and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232579/files/figure.png	https://doi.org/10.5281/zenodo.11232579	PLATE 15. A-N Lioharpes saredra n. sp. A–C partial dorsal exoskeleton of cephalon from “Odontochile Couche” Boutiskaouine Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 7), paratype, NHMUK it 29276. A, dorsal and B, lateral view. C, dorsal view of brim perforations. D–J disarticulated dorsal exoskeleton, from the Destombina Horizon, Tazoulait Formation, upper Emsian,Jbel Shebras (Fig. 2B, Map 12, site 1), paratype, NHMUK It 29278. D, dorsal, E, lateral and F, anterior view. Dorsal view of: G, genal area showing internal mould, H, brim perforations, I, pygidium and J external rim. K–N dorsal exoskeleton of cephalon only, from coral at the top of the Pragian, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 1), paratype, NHMUK It 29277. K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 15. A-N Lioharpes saredra n. sp. A–C partial dorsal exoskeleton of cephalon from “Odontochile Couche” Boutiskaouine Formation, Emsian, Jbel Oufatene (Fig. 2B, Map 8, site 7), paratype, NHMUK it 29276. A, dorsal and B, lateral view. C, dorsal view of brim perforations. D–J disarticulated dorsal exoskeleton, from the Destombina Horizon, Tazoulait Formation, upper Emsian,Jbel Shebras (Fig. 2B, Map 12, site 1), paratype, NHMUK It 29278. D, dorsal, E, lateral and F, anterior view. Dorsal view of: G, genal area showing internal mould, H, brim perforations, I, pygidium and J external rim. K–N dorsal exoskeleton of cephalon only, from coral at the top of the Pragian, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 1), paratype, NHMUK It 29277. K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232703/files/figure.png	https://doi.org/10.5281/zenodo.11232703	PLATE 57.A–F Fritchaspis montagnei (Hawle and Corda, 1847) type horizon and locality Acanthopyge Limestones (Eifelian), Koněprusy area, Czech Republic, five dorsal exoskeleton of partial cephala, Museum of Natural History, Prague, syntype Nos L 2996 (one of four syntypes) and L 2990. A–C, L 2996, a partial cephalon, A dorsal view, B anterior view and C lateral view. L 2990, D–F a plate with four partial cephala, D dorsal view, E lateral view of largest cephalon and F dorsal vie of brim perforations. G–J Declivoharpes praecedens by (Prantl and Přibyl 1954,) type horizon and locality Radotín Limestones (Lochkovian), praharadotín (Černá rokle gorge) dorsal exoskeleton of partial cephalon, Museum of Natural History, Prague holotype No. L19103. G dorsal view, H anterior view, I lateral view and J dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl. 8 fig.2). Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 57.A–F Fritchaspis montagnei (Hawle and Corda, 1847) type horizon and locality Acanthopyge Limestones (Eifelian), Koněprusy area, Czech Republic, five dorsal exoskeleton of partial cephala, Museum of Natural History, Prague, syntype Nos L 2996 (one of four syntypes) and L 2990. A–C, L 2996, a partial cephalon, A dorsal view, B anterior view and C lateral view. L 2990, D–F a plate with four partial cephala, D dorsal view, E lateral view of largest cephalon and F dorsal vie of brim perforations. G–J Declivoharpes praecedens by (Prantl and Přibyl 1954,) type horizon and locality Radotín Limestones (Lochkovian), praharadotín (Černá rokle gorge) dorsal exoskeleton of partial cephalon, Museum of Natural History, Prague holotype No. L19103. G dorsal view, H anterior view, I lateral view and J dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl. 8 fig.2). Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE5E31CFF78FEA3FDBA81E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232701/files/figure.png	https://doi.org/10.5281/zenodo.11232701	PLATE 56. A–F Bohemoharpes vittatus (Barrande 1852); type horizon and locality with Prionopeltis archiaci, Kopanina Formation (Ludlow) Praha-Lochkov, dorsal exoskeletons of 2 partial cephala, Natural History Museum Prague Nos L 12549 and L 12565. A–C, L 12549, A dorsal view, B lateral view and C anterior view. D–F, No. L 12549, D dorsal view of cephalon, E brim perforations and F genal area including eye lobe and genal ridge. G–K Fritchaspis perneri, Prantl & Přibyl,1954 type horizon and locality Třebotov Limestones, Praha-Holyně Czech Republic, three dorsal exoskeletons of three partial cephala, Natural History Museum Prague No. L6140 and Czech Geological Survey Nos JV15912 and JV15897. G–I, No. L6140 holotype, G dorsal view of partial cephalon, H anterior of glabella with axial tubercle and I brim perforations. J, No. JV15912 dorsal view of genicranium and K, No. JV 15897 dorsal view of partial cephalon. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 56. A–F Bohemoharpes vittatus (Barrande 1852); type horizon and locality with Prionopeltis archiaci, Kopanina Formation (Ludlow) Praha-Lochkov, dorsal exoskeletons of 2 partial cephala, Natural History Museum Prague Nos L 12549 and L 12565. A–C, L 12549, A dorsal view, B lateral view and C anterior view. D–F, No. L 12549, D dorsal view of cephalon, E brim perforations and F genal area including eye lobe and genal ridge. G–K Fritchaspis perneri, Prantl & Přibyl,1954 type horizon and locality Třebotov Limestones, Praha-Holyně Czech Republic, three dorsal exoskeletons of three partial cephala, Natural History Museum Prague No. L6140 and Czech Geological Survey Nos JV15912 and JV15897. G–I, No. L6140 holotype, G dorsal view of partial cephalon, H anterior of glabella with axial tubercle and I brim perforations. J, No. JV15912 dorsal view of genicranium and K, No. JV 15897 dorsal view of partial cephalon. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE3E31EFF78FE9EFD8F8273.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232585/files/figure.png	https://doi.org/10.5281/zenodo.11232585	PLATE 18. A–F Fritchaspis edgecombei n. sp., dorsal exoskeleton of cephalon only, NHMUK It 29224, from “Cherurid Couche”, Er Remlia Formation, lower Emsian, Jbel Issoumour (Fig. 2B, Map 7, site 19). A, dorsal, B, antero-dorsal, C, anterior and D, lateral view. E, dorsal view of brim perforation and F, lateral view of eye lobe. G–N Maghroharpes hammii n. sp., dorsal exoskeleton, thorax enrolled, holotype, NHMUK It 29292, from an horizon below the “Dicranurus Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, map 8, site 21). G, dorsal, H, anterior, J, lateral and I, posterior view. L, dorsal view of brim perforation, N, lateral view of glabella and muscle insertion scars and M, dorsal view of cuticle on glabella, genal area, ala, occipital ring and posterior border. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 18. A–F Fritchaspis edgecombei n. sp., dorsal exoskeleton of cephalon only, NHMUK It 29224, from “Cherurid Couche”, Er Remlia Formation, lower Emsian, Jbel Issoumour (Fig. 2B, Map 7, site 19). A, dorsal, B, antero-dorsal, C, anterior and D, lateral view. E, dorsal view of brim perforation and F, lateral view of eye lobe. G–N Maghroharpes hammii n. sp., dorsal exoskeleton, thorax enrolled, holotype, NHMUK It 29292, from an horizon below the “Dicranurus Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, map 8, site 21). G, dorsal, H, anterior, J, lateral and I, posterior view. L, dorsal view of brim perforation, N, lateral view of glabella and muscle insertion scars and M, dorsal view of cuticle on glabella, genal area, ala, occipital ring and posterior border. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE3E31EFF78FE9EFD8F8273.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232581/files/figure.png	https://doi.org/10.5281/zenodo.11232581	PLATE 16. A–L Kielania tumula n. sp., from a lower Pragian horizon, Ihandar Formation, Iminwasief, 4.6 km W of Jbel Ou- Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, thorax disarticulated and enrolled, holotype, NHMUK It 29259 (site 1). A, dorsal, B, lateral, C, anterior and E, posterior view. D, dorsal view of brim perforations. F–L dorsal exoskeleton, thorax disarticulated and enrolled, paratype, NHMUK It 29260 (site 2). F, dorsal, G, lateral, H, anterior and J, posterior view. I, dorsal view of brim perforations, K, lateral view of glabella (uncoated) and L, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 16. A–L Kielania tumula n. sp., from a lower Pragian horizon, Ihandar Formation, Iminwasief, 4.6 km W of Jbel Ou- Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, thorax disarticulated and enrolled, holotype, NHMUK It 29259 (site 1). A, dorsal, B, lateral, C, anterior and E, posterior view. D, dorsal view of brim perforations. F–L dorsal exoskeleton, thorax disarticulated and enrolled, paratype, NHMUK It 29260 (site 2). F, dorsal, G, lateral, H, anterior and J, posterior view. I, dorsal view of brim perforations, K, lateral view of glabella (uncoated) and L, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232654/files/figure.png	https://doi.org/10.5281/zenodo.11232654	PLATE 36. A–K Harpes igaouii n. sp., Ihandar Formation, Pragian-Emsian boundary. A–H dorsal exoskeleton of cephalon only, holotype, NHMUK It 29360, from coral NW of Jbel Ou-Driss (Fig. 2B, Map 4, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal anterior view of perforations on genal roll and brim. E, dorsal view of glabellar tubercles, F, dorsal view of brim perforations, G, lateral view of glabellar muscle insertion scars (uncoated) and H, lateral view of eye lobe (uncoated). I–K dorsal exoskeleton of cephalon and partial thorax, paratype, NHMUK It 29361, from Ceratocephala horizon, North Jbel Ou-Driss (Fig. 2B, Map 4, site 8). I, dorsal anterior lateral view (uncoated). J, ventral view of hypostome and K, lateral view of eye lobe (uncoated). L–O Harpes aff. igaouii, dorsal exoskeleton of cephalon only, NHMUK It 29362, from Fezna, “Struveaspis Couche”, El Otfal Formation, Eifelian.(Fig. 2B, Map 6, site 1). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. P–R Kielania aff. tumula, Ihandar Formation, upper Pragian (Fig. 2B, Map 10, Site 5) dorsal exoskeleton of a partial cephalon, NHMUK It 29262. P, dorsal, Q, anterior and R, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 36. A–K Harpes igaouii n. sp., Ihandar Formation, Pragian-Emsian boundary. A–H dorsal exoskeleton of cephalon only, holotype, NHMUK It 29360, from coral NW of Jbel Ou-Driss (Fig. 2B, Map 4, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal anterior view of perforations on genal roll and brim. E, dorsal view of glabellar tubercles, F, dorsal view of brim perforations, G, lateral view of glabellar muscle insertion scars (uncoated) and H, lateral view of eye lobe (uncoated). I–K dorsal exoskeleton of cephalon and partial thorax, paratype, NHMUK It 29361, from Ceratocephala horizon, North Jbel Ou-Driss (Fig. 2B, Map 4, site 8). I, dorsal anterior lateral view (uncoated). J, ventral view of hypostome and K, lateral view of eye lobe (uncoated). L–O Harpes aff. igaouii, dorsal exoskeleton of cephalon only, NHMUK It 29362, from Fezna, “Struveaspis Couche”, El Otfal Formation, Eifelian.(Fig. 2B, Map 6, site 1). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. P–R Kielania aff. tumula, Ihandar Formation, upper Pragian (Fig. 2B, Map 10, Site 5) dorsal exoskeleton of a partial cephalon, NHMUK It 29262. P, dorsal, Q, anterior and R, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232705/files/figure.png	https://doi.org/10.5281/zenodo.11232705	PLATE 58.A –F Kielania waageni (Novák MS) Prantl and Přibyl, 1954 type horizon and locality Prokop Limestones (Pragian), Praha-Malá Chuchle, Czech Republic, dorsal exoskeleton of a partial cephalon Museum of Natural History, Prague, holotype No. L6141.A dorsal view, B lateral view, C anterior view, D anterolateral view, E anterodorsal view and F lateral view of girder and perforations on genal roll and brim. This specimen was illustrated by Prantl and Přibyl (1954, pl. 9 fig.2). G–J Helioharpes dorbignyanus (Barrande 1846) type horizon and locality Dvorce Limestones (Pragian), Praha-Brank (“Za Branickou skalou” locality), Czech Republic, dorsal exoskeleton of a partial cephalon Natural History Museum Prague, No. L12563; G dorsal view, H lateral view, I anterior view and J dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 58.A –F Kielania waageni (Novák MS) Prantl and Přibyl, 1954 type horizon and locality Prokop Limestones (Pragian), Praha-Malá Chuchle, Czech Republic, dorsal exoskeleton of a partial cephalon Museum of Natural History, Prague, holotype No. L6141.A dorsal view, B lateral view, C anterior view, D anterolateral view, E anterodorsal view and F lateral view of girder and perforations on genal roll and brim. This specimen was illustrated by Prantl and Přibyl (1954, pl. 9 fig.2). G–J Helioharpes dorbignyanus (Barrande 1846) type horizon and locality Dvorce Limestones (Pragian), Praha-Brank (“Za Branickou skalou” locality), Czech Republic, dorsal exoskeleton of a partial cephalon Natural History Museum Prague, No. L12563; G dorsal view, H lateral view, I anterior view and J dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232583/files/figure.png	https://doi.org/10.5281/zenodo.11232583	PLATE 17. A–K and M–P Helioharpes aougili n. sp., from an Eifelian horizon, El Otfal Formation, Tamgoute Bulgan, 11.6 km SWt of El Jorf (Fig. 2B, Map 6). A–K dorsal exoskeleton, pygidium missing, holotype, NHMUK It 29254 (site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim proximal to girder across the anterior of genicranium, E, anterior view of perforations on brim and genal roll, F, dorsal view of brim perforations. G and H, lateral and dorsal view of eye lobe, respectively. I, dorsal view of cuticle on glabella and genal area. J and K, lateral and dorsal view of pleurae, respectively. M–P dorsal exoskeleton of cephalon only, paratype, NHMUK It 29255 (site 17). M, dorsal, N, lateral and O, anterior view. P, dorsal view of brim perforation. L, H. perradiatus, dorsal view of brim proximal to girder across the anterior of genicranium (reprint), Richter & Richter, 1943, abb 2. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 17. A–K and M–P Helioharpes aougili n. sp., from an Eifelian horizon, El Otfal Formation, Tamgoute Bulgan, 11.6 km SWt of El Jorf (Fig. 2B, Map 6). A–K dorsal exoskeleton, pygidium missing, holotype, NHMUK It 29254 (site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim proximal to girder across the anterior of genicranium, E, anterior view of perforations on brim and genal roll, F, dorsal view of brim perforations. G and H, lateral and dorsal view of eye lobe, respectively. I, dorsal view of cuticle on glabella and genal area. J and K, lateral and dorsal view of pleurae, respectively. M–P dorsal exoskeleton of cephalon only, paratype, NHMUK It 29255 (site 17). M, dorsal, N, lateral and O, anterior view. P, dorsal view of brim perforation. L, H. perradiatus, dorsal view of brim proximal to girder across the anterior of genicranium (reprint), Richter & Richter, 1943, abb 2. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFE1E360FF78FD2FFC9684D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9FE363FF78FB97FEFB84B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232583/files/figure.png	https://doi.org/10.5281/zenodo.11232583	PLATE 17. A–K and M–P Helioharpes aougili n. sp., from an Eifelian horizon, El Otfal Formation, Tamgoute Bulgan, 11.6 km SWt of El Jorf (Fig. 2B, Map 6). A–K dorsal exoskeleton, pygidium missing, holotype, NHMUK It 29254 (site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim proximal to girder across the anterior of genicranium, E, anterior view of perforations on brim and genal roll, F, dorsal view of brim perforations. G and H, lateral and dorsal view of eye lobe, respectively. I, dorsal view of cuticle on glabella and genal area. J and K, lateral and dorsal view of pleurae, respectively. M–P dorsal exoskeleton of cephalon only, paratype, NHMUK It 29255 (site 17). M, dorsal, N, lateral and O, anterior view. P, dorsal view of brim perforation. L, H. perradiatus, dorsal view of brim proximal to girder across the anterior of genicranium (reprint), Richter & Richter, 1943, abb 2. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 17. A–K and M–P Helioharpes aougili n. sp., from an Eifelian horizon, El Otfal Formation, Tamgoute Bulgan, 11.6 km SWt of El Jorf (Fig. 2B, Map 6). A–K dorsal exoskeleton, pygidium missing, holotype, NHMUK It 29254 (site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim proximal to girder across the anterior of genicranium, E, anterior view of perforations on brim and genal roll, F, dorsal view of brim perforations. G and H, lateral and dorsal view of eye lobe, respectively. I, dorsal view of cuticle on glabella and genal area. J and K, lateral and dorsal view of pleurae, respectively. M–P dorsal exoskeleton of cephalon only, paratype, NHMUK It 29255 (site 17). M, dorsal, N, lateral and O, anterior view. P, dorsal view of brim perforation. L, H. perradiatus, dorsal view of brim proximal to girder across the anterior of genicranium (reprint), Richter & Richter, 1943, abb 2. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9FE363FF78FB97FEFB84B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232707/files/figure.png	https://doi.org/10.5281/zenodo.11232707	PLATE 59. A–D Helioharpes dvorcensis (Prantl and Přibyl, 1954), type horizon and locality Dvorce Limestones (Pragian), Praha-Podoli (formerly Dvorce), Czech Republic, dorsal exoskeleton of a partial cephalon from the Natural History Museum Prague No. L19105 holotype. A dorsal view, B lateral view, C anterior,view and D dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl.8 fig. 5). E –G Declivoharpes novaki (Prantl and Přibyl 1954). Type horizon Prokop Limestone (Pragian), St. Prokop quarry, Praha- Hlubocepy, Czech Republic, dorsal exoskeleton of a partial cephalon from the Natural History Museum Prague holotype No. L6120, E dorsal view of exoskeleton, F anterior view and G lateral view. This specimen was illustrated by Prantl and Přibyl (1954, pl.9 fig. 1). H–L Stoloharpes convexus. (Hawle and Corda 1847) Suchomasty Limestones (Eifelian), Měňany, Czech Republic, dorsal exoskeletons of two partial cephalon from the Natural History Museum Prague Nos L 6138 and L 6139. H, L 6139, dorsal view of exoskeleton. I–L, L 6138, J dorsal view of exoskeleton, K lateral view, L anterior view and I dorsal view of brim perforations. L 6138 and L 6139 were illustrated by Prantl and Přibyl (1954, pl. 9 figs 3 and 4 respectively). Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 59. A–D Helioharpes dvorcensis (Prantl and Přibyl, 1954), type horizon and locality Dvorce Limestones (Pragian), Praha-Podoli (formerly Dvorce), Czech Republic, dorsal exoskeleton of a partial cephalon from the Natural History Museum Prague No. L19105 holotype. A dorsal view, B lateral view, C anterior,view and D dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl.8 fig. 5). E –G Declivoharpes novaki (Prantl and Přibyl 1954). Type horizon Prokop Limestone (Pragian), St. Prokop quarry, Praha- Hlubocepy, Czech Republic, dorsal exoskeleton of a partial cephalon from the Natural History Museum Prague holotype No. L6120, E dorsal view of exoskeleton, F anterior view and G lateral view. This specimen was illustrated by Prantl and Přibyl (1954, pl.9 fig. 1). H–L Stoloharpes convexus. (Hawle and Corda 1847) Suchomasty Limestones (Eifelian), Měňany, Czech Republic, dorsal exoskeletons of two partial cephalon from the Natural History Museum Prague Nos L 6138 and L 6139. H, L 6139, dorsal view of exoskeleton. I–L, L 6138, J dorsal view of exoskeleton, K lateral view, L anterior view and I dorsal view of brim perforations. L 6138 and L 6139 were illustrated by Prantl and Přibyl (1954, pl. 9 figs 3 and 4 respectively). Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9FE363FF78FB97FEFB84B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232703/files/figure.png	https://doi.org/10.5281/zenodo.11232703	PLATE 57.A–F Fritchaspis montagnei (Hawle and Corda, 1847) type horizon and locality Acanthopyge Limestones (Eifelian), Koněprusy area, Czech Republic, five dorsal exoskeleton of partial cephala, Museum of Natural History, Prague, syntype Nos L 2996 (one of four syntypes) and L 2990. A–C, L 2996, a partial cephalon, A dorsal view, B anterior view and C lateral view. L 2990, D–F a plate with four partial cephala, D dorsal view, E lateral view of largest cephalon and F dorsal vie of brim perforations. G–J Declivoharpes praecedens by (Prantl and Přibyl 1954,) type horizon and locality Radotín Limestones (Lochkovian), praharadotín (Černá rokle gorge) dorsal exoskeleton of partial cephalon, Museum of Natural History, Prague holotype No. L19103. G dorsal view, H anterior view, I lateral view and J dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl. 8 fig.2). Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 57.A–F Fritchaspis montagnei (Hawle and Corda, 1847) type horizon and locality Acanthopyge Limestones (Eifelian), Koněprusy area, Czech Republic, five dorsal exoskeleton of partial cephala, Museum of Natural History, Prague, syntype Nos L 2996 (one of four syntypes) and L 2990. A–C, L 2996, a partial cephalon, A dorsal view, B anterior view and C lateral view. L 2990, D–F a plate with four partial cephala, D dorsal view, E lateral view of largest cephalon and F dorsal vie of brim perforations. G–J Declivoharpes praecedens by (Prantl and Přibyl 1954,) type horizon and locality Radotín Limestones (Lochkovian), praharadotín (Černá rokle gorge) dorsal exoskeleton of partial cephalon, Museum of Natural History, Prague holotype No. L19103. G dorsal view, H anterior view, I lateral view and J dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl (1954, pl. 8 fig.2). Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9FE363FF78FB97FEFB84B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9FE363FF78FB97FEFB84B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9CE365FF78FC6EFD348344.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232642/files/figure.png	https://doi.org/10.5281/zenodo.11232642	PLATE 33. A–K Eskoharpes sicarius n. sp., from the ammonoid mining horizon, Achguig Formation Group, lower Frasnian, Bouia, 7 km SE of Jorf (Fig., 2B, Map 6, site 5). A–C paratype, NHMUK It 29256 (6), dorsal exoskeleton of cephalon. A, dorsal view, B, anterior view and C, lateral view. D–F holotype, NHMUK It 29256 (1), dorsal exoskeleton partial cephalon and thorax, and a complete pygidium. D, dorsal view of whole exoskeleton, E, dorsal view of thorax and pygidium and F, dorsal posterior view of pygidium. G assemblage of partial dorsal exoskeletons of material shown in Figs 43 and 44. H paratype, NHMUK It 29256 (8), partial dorsal exoskeleton of cephalon and thorax, dorsal view. I–K paratype, NHMUK It 29256 (3), dorsal exoskeleton of lower lamella. I, dorsal, J, lateral and K, anterior view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 33. A–K Eskoharpes sicarius n. sp., from the ammonoid mining horizon, Achguig Formation Group, lower Frasnian, Bouia, 7 km SE of Jorf (Fig., 2B, Map 6, site 5). A–C paratype, NHMUK It 29256 (6), dorsal exoskeleton of cephalon. A, dorsal view, B, anterior view and C, lateral view. D–F holotype, NHMUK It 29256 (1), dorsal exoskeleton partial cephalon and thorax, and a complete pygidium. D, dorsal view of whole exoskeleton, E, dorsal view of thorax and pygidium and F, dorsal posterior view of pygidium. G assemblage of partial dorsal exoskeletons of material shown in Figs 43 and 44. H paratype, NHMUK It 29256 (8), partial dorsal exoskeleton of cephalon and thorax, dorsal view. I–K paratype, NHMUK It 29256 (3), dorsal exoskeleton of lower lamella. I, dorsal, J, lateral and K, anterior view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9CE365FF78FC6EFD348344.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232646/files/figure.png	https://doi.org/10.5281/zenodo.11232646	PLATE 34. A–I Eskoharpes sicarius n. sp., from Type horizon, lower Frasnian, Bouia, 7 km SE of Jorf (Fig. 2B, Map 6, site 5). A–E paratype, NHMUK it 29256 (2), dorsal exoskeleton of partial cephalon only. A, dorsal view, B, lateral view, C, anterior view, D, dorsal view of eye lobe and eye ridge and E, anterior dorsal view of brim perforations. F–G paratype, NHMUK it 29256 (5), dorsal exoskeleton partial cephalon. F, dorsal, G, lateral view of posterior of prolongation including genal spine. H paratype, NHMUK it 29256 (4), dorsal exoskeleton of partial cephalon and disarticulated thorax, dorsal view. I paratype, NHMUK it 29256 (1&7), dorsal exoskeleton of partial concealed cephalon, dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 34. A–I Eskoharpes sicarius n. sp., from Type horizon, lower Frasnian, Bouia, 7 km SE of Jorf (Fig. 2B, Map 6, site 5). A–E paratype, NHMUK it 29256 (2), dorsal exoskeleton of partial cephalon only. A, dorsal view, B, lateral view, C, anterior view, D, dorsal view of eye lobe and eye ridge and E, anterior dorsal view of brim perforations. F–G paratype, NHMUK it 29256 (5), dorsal exoskeleton partial cephalon. F, dorsal, G, lateral view of posterior of prolongation including genal spine. H paratype, NHMUK it 29256 (4), dorsal exoskeleton of partial cephalon and disarticulated thorax, dorsal view. I paratype, NHMUK it 29256 (1&7), dorsal exoskeleton of partial concealed cephalon, dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9CE365FF78FC6EFD348344.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9CE365FF78FC6EFD348344.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9AE367FF78FCFBFDF6849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232585/files/figure.png	https://doi.org/10.5281/zenodo.11232585	PLATE 18. A–F Fritchaspis edgecombei n. sp., dorsal exoskeleton of cephalon only, NHMUK It 29224, from “Cherurid Couche”, Er Remlia Formation, lower Emsian, Jbel Issoumour (Fig. 2B, Map 7, site 19). A, dorsal, B, antero-dorsal, C, anterior and D, lateral view. E, dorsal view of brim perforation and F, lateral view of eye lobe. G–N Maghroharpes hammii n. sp., dorsal exoskeleton, thorax enrolled, holotype, NHMUK It 29292, from an horizon below the “Dicranurus Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, map 8, site 21). G, dorsal, H, anterior, J, lateral and I, posterior view. L, dorsal view of brim perforation, N, lateral view of glabella and muscle insertion scars and M, dorsal view of cuticle on glabella, genal area, ala, occipital ring and posterior border. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 18. A–F Fritchaspis edgecombei n. sp., dorsal exoskeleton of cephalon only, NHMUK It 29224, from “Cherurid Couche”, Er Remlia Formation, lower Emsian, Jbel Issoumour (Fig. 2B, Map 7, site 19). A, dorsal, B, antero-dorsal, C, anterior and D, lateral view. E, dorsal view of brim perforation and F, lateral view of eye lobe. G–N Maghroharpes hammii n. sp., dorsal exoskeleton, thorax enrolled, holotype, NHMUK It 29292, from an horizon below the “Dicranurus Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, map 8, site 21). G, dorsal, H, anterior, J, lateral and I, posterior view. L, dorsal view of brim perforation, N, lateral view of glabella and muscle insertion scars and M, dorsal view of cuticle on glabella, genal area, ala, occipital ring and posterior border. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9AE367FF78FCFBFDF6849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232590/files/figure.png	https://doi.org/10.5281/zenodo.11232590	PLATE 19. A–O Maghroharpes hammii n. sp. A–F dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29296, from main trilobite ‘couche’ Ihandar Formation, Pragian, southern flank of Jbel el Mrakib (Fig. 2B, Map 9, site 16). A, dorsal, B, anterior, C, lateral and D, posterior view. E, dorsal view of brim perforations and F, lateral view of muscle insertion scars on glabella (uncoated). G–L dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29294, from a Ceratonurus horizon, Ihandar Formation, Pragian, Jbel Remlia (Fig. 2B, Map 9, site 12). G, dorsal, H, anterior and I, lateral view. J, lateral view of eye lobe (uncoated), K, dorsal view of brim perforations, dorsal view of tubercles on axial rings. M–O dorsal exoskeleton with pygidium missing, paratype, NHMUK It 29295, from main trilobite ‘couche’, Ihandar Formation, Pragian, southern flank of Jbel el Mrakib (Fig. 2B, Map 9, site 15). M, dorsal, N, anterior and O, lateral view. P, dorsal view of tubercles on rim. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 19. A–O Maghroharpes hammii n. sp. A–F dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29296, from main trilobite ‘couche’ Ihandar Formation, Pragian, southern flank of Jbel el Mrakib (Fig. 2B, Map 9, site 16). A, dorsal, B, anterior, C, lateral and D, posterior view. E, dorsal view of brim perforations and F, lateral view of muscle insertion scars on glabella (uncoated). G–L dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29294, from a Ceratonurus horizon, Ihandar Formation, Pragian, Jbel Remlia (Fig. 2B, Map 9, site 12). G, dorsal, H, anterior and I, lateral view. J, lateral view of eye lobe (uncoated), K, dorsal view of brim perforations, dorsal view of tubercles on axial rings. M–O dorsal exoskeleton with pygidium missing, paratype, NHMUK It 29295, from main trilobite ‘couche’, Ihandar Formation, Pragian, southern flank of Jbel el Mrakib (Fig. 2B, Map 9, site 15). M, dorsal, N, anterior and O, lateral view. P, dorsal view of tubercles on rim. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9AE367FF78FCFBFDF6849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232592/files/figure.png	https://doi.org/10.5281/zenodo.11232592	PLATE 20. A–O Maghroharpes hammii n. sp., from Ihandar Formation, Pragian. A–D dorsal exoskeleton, cephalon and disarticulated thorax, paratype, NHMUK It 29290, from Jbel Oufatene (Fig. 2B, Map 8, site 8). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations and F, lateral view of muscle insertion scars on glabella (uncoated). E–I dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29300, from North Zireg (Fig. 2B, Map 13, site 1). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations, I, lateral view of eye lobe. J–M dorsal exoskeleton of cephalon, paratype, NHMUK It 29291, from Jbel Oufatene (Fig. 2B, Map 8, site 9). J, dorsal, K, anterior and L, lateral view. M, dorsal view of brim perforations. N–P upper lamella only, paratype, NHMUK It 29293, from an horizon between the “Dicranurus Couche” and the “Kolihapeltis Couche”, Talhadjate, Jbel Oufatene (Fig. 2B, Map 8, site 22). N, dorsal, O, anterior and P, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 20. A–O Maghroharpes hammii n. sp., from Ihandar Formation, Pragian. A–D dorsal exoskeleton, cephalon and disarticulated thorax, paratype, NHMUK It 29290, from Jbel Oufatene (Fig. 2B, Map 8, site 8). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations and F, lateral view of muscle insertion scars on glabella (uncoated). E–I dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29300, from North Zireg (Fig. 2B, Map 13, site 1). E, dorsal, F, lateral and G, anterior view. H, dorsal view of brim perforations, I, lateral view of eye lobe. J–M dorsal exoskeleton of cephalon, paratype, NHMUK It 29291, from Jbel Oufatene (Fig. 2B, Map 8, site 9). J, dorsal, K, anterior and L, lateral view. M, dorsal view of brim perforations. N–P upper lamella only, paratype, NHMUK It 29293, from an horizon between the “Dicranurus Couche” and the “Kolihapeltis Couche”, Talhadjate, Jbel Oufatene (Fig. 2B, Map 8, site 22). N, dorsal, O, anterior and P, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9AE367FF78FCFBFDF6849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232622/files/figure.png	https://doi.org/10.5281/zenodo.11232622	PLATE 27. A–D Maghroharpes ihmadii n. sp., dorsal exoskeleton of cephalon and disarticulated thorax, paratype, NHMUK It 29303, from type horizon and locality, Ihandar Formation, Pragian. Jbel Ou-Driss (Fig. 2B, Map 4, site 18). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Maghroharpes cf. ihamadii, dorsal exoskeleton pygidium enrolled, NHMUK It 29304, from “Kolihapeltis Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, Map 8, site 14). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe. J and K, dorsal view of brim perforations and pleurae, respectively. L–O Magroharpes hammii, dorsal exoskeleton, thorax and disarticulated pygidium, NHMUK It 29297, from Ihandar Formation, Iminwasief (Fig. 2B, Map 4, site 4). L, dorsaI, M, anterior, N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 27. A–D Maghroharpes ihmadii n. sp., dorsal exoskeleton of cephalon and disarticulated thorax, paratype, NHMUK It 29303, from type horizon and locality, Ihandar Formation, Pragian. Jbel Ou-Driss (Fig. 2B, Map 4, site 18). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Maghroharpes cf. ihamadii, dorsal exoskeleton pygidium enrolled, NHMUK It 29304, from “Kolihapeltis Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, Map 8, site 14). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe. J and K, dorsal view of brim perforations and pleurae, respectively. L–O Magroharpes hammii, dorsal exoskeleton, thorax and disarticulated pygidium, NHMUK It 29297, from Ihandar Formation, Iminwasief (Fig. 2B, Map 4, site 4). L, dorsaI, M, anterior, N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF9AE367FF78FCFBFDF6849C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232697/files/figure.png	https://doi.org/10.5281/zenodo.11232697	PLATE 54. A–D Lioharpes venulosus (Hawle & Corda, 1847); type locality Upper Koněprusy Limestones, (Pragian) Koněprusy, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L12545. A dorsal view, B lateral view, C anterior view and D dorsal view of brim perforations. E–H Bohemoharpes buphthalmus (Bouček, 1935). Type locality the horizon with Ananaspis fecunda (Ludlow stage), Dlouhá hora near Beroun, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L17145. E, dorsal view, F lateral view, G anterior view and H dorsal view of brim perforations. I–L Bohemoharpes naumanni (Barrande 1852); type horizon and zone Cyrtograptus lundgreni zone (Upper Wenlock) Lodenice near Beroun, Czech Republic; dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague, lectotype (subsequent designation, Prantl and Přibyl, 1954) L6127, IT 48. I dorsal view, J anterior view, K lateral view and L dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl,1954 pl. iv fig. 1 and Ebach and McNamara, 2002 Fig 2A. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 54. A–D Lioharpes venulosus (Hawle & Corda, 1847); type locality Upper Koněprusy Limestones, (Pragian) Koněprusy, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L12545. A dorsal view, B lateral view, C anterior view and D dorsal view of brim perforations. E–H Bohemoharpes buphthalmus (Bouček, 1935). Type locality the horizon with Ananaspis fecunda (Ludlow stage), Dlouhá hora near Beroun, Czech Republic, a dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague L17145. E, dorsal view, F lateral view, G anterior view and H dorsal view of brim perforations. I–L Bohemoharpes naumanni (Barrande 1852); type horizon and zone Cyrtograptus lundgreni zone (Upper Wenlock) Lodenice near Beroun, Czech Republic; dorsal exoskeleton of a partial cephalon, Museum of Natural History, Prague, lectotype (subsequent designation, Prantl and Přibyl, 1954) L6127, IT 48. I dorsal view, J anterior view, K lateral view and L dorsal view of brim perforations. This specimen was illustrated by Prantl and Přibyl,1954 pl. iv fig. 1 and Ebach and McNamara, 2002 Fig 2A. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF98E368FF78FC42FB8D8159.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232628/files/figure.png	https://doi.org/10.5281/zenodo.11232628	PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF98E368FF78FC42FB8D8159.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232632/files/figure.png	https://doi.org/10.5281/zenodo.11232632	PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 30. A–E Maghroharpes azmamarensis n. sp., from type locality and horizon, Ihandar Formation, Pragian, dorsal exoskeleton of cephalon, paratype, NHMUK It 29282 (Fig., 2B, Map 8, site 19). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. 1 azmamarensis, from top of Pragian, Ihandar Formation, ElAtchane Jbel Oufatene, dorsal exoskeleton of partial cephalon, NHMUK It 29285 (Fig. 2B, Map 8, site 17). F, dorsal, G, lateral and H, anterior view. I, lateral and J, dorsal view of eye lobe, respectively. K, dorsal view of brim perforations. L–O Maghroharpes cf. 2 azmamarensis, from an upper Pragian horizon, Ihandar Formation, Maharch, Jbel Mrakib, dorsal exoskeleton of partial cephalon, NHMUK It 29286 (Fig. 2B, Map 9, site 14). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF98E368FF78FC42FB8D8159.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232664/files/figure.png	https://doi.org/10.5281/zenodo.11232664	PLATE 39. A–M Pinnuloharpes apteros n. sp., El Otfal Formation, upper Eifelian. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29356 from Ksar Hannabou (Fig. 2B, Map 6, site 8). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–K dorsal exoskeleton with thorax and pygidium disarticulated, paratype, NHMUK It 29359, from type locality and horizon (Fig. 2B, Map 6, site 19). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, lateral view of eye lobe, J, anterior lateral view of rim and marginal band, and K, dorsal view of pleurae close to pygidium. L–M dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29357, from Ksar Hannabou (Fig. 2B, Map 6, site 15). L, dorsal view and M, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 39. A–M Pinnuloharpes apteros n. sp., El Otfal Formation, upper Eifelian. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29356 from Ksar Hannabou (Fig. 2B, Map 6, site 8). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–K dorsal exoskeleton with thorax and pygidium disarticulated, paratype, NHMUK It 29359, from type locality and horizon (Fig. 2B, Map 6, site 19). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, lateral view of eye lobe, J, anterior lateral view of rim and marginal band, and K, dorsal view of pleurae close to pygidium. L–M dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29357, from Ksar Hannabou (Fig. 2B, Map 6, site 15). L, dorsal view and M, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF98E368FF78FC42FB8D8159.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232666/files/figure.png	https://doi.org/10.5281/zenodo.11232666	PLATE 40.A–N Pinnuloharpes hannabouensis n. sp., from an horizon 5 m down slope of “Radiaspis Couche”, upper Eifelian/ Givetian boundary, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29334 (Fig. 2B, Map 6, site 12). A, dorsal and B, lateral view. C, lateral view of eye lobe (uncoated) and D, dorsal view of brim perforations. E–I dorsal exoskeleton cephalon and thorax only, holotype, NHMUK It 29333 (Fig. 2B, Map 6, site 11). E, dorsal, F, anterior and G, lateral view. H, lateral view of eye lobes, I, dorsal view of brim perforations. J–N dorsal exoskeleton pygidium incomplete, paratype, NHMUK It 29335 (Fig. 2B, Map 6, site 13). J, dorsal, K, anterior and L, lateral view. M, lateral view of eye lobe and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 40.A–N Pinnuloharpes hannabouensis n. sp., from an horizon 5 m down slope of “Radiaspis Couche”, upper Eifelian/ Givetian boundary, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29334 (Fig. 2B, Map 6, site 12). A, dorsal and B, lateral view. C, lateral view of eye lobe (uncoated) and D, dorsal view of brim perforations. E–I dorsal exoskeleton cephalon and thorax only, holotype, NHMUK It 29333 (Fig. 2B, Map 6, site 11). E, dorsal, F, anterior and G, lateral view. H, lateral view of eye lobes, I, dorsal view of brim perforations. J–N dorsal exoskeleton pygidium incomplete, paratype, NHMUK It 29335 (Fig. 2B, Map 6, site 13). J, dorsal, K, anterior and L, lateral view. M, lateral view of eye lobe and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF97E36AFF78FE05FB82842C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232616/files/figure.png	https://doi.org/10.5281/zenodo.11232616	PLATE 26. A–Q Maghroharpes ihmadii n. sp., Ihandar Formation, Pragian, Jebl Ou-Driss. A–G dorsal exoskeleton of cephalon, holotype, NHMUK It 29301 (Fig. 2B, Map 4, site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. Eye lobe in lateral view E and G, and in dorsal view, F. H–Q dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29302 (Fig. 2B, Map 4, site 17). H, dorsaI, I, lateral, J, anterior and L, posterior-dorsal view. K and Q, dorsal view of brim perforations and pleurae, respectively, M, anterior dorsal view, O and P, lateral views of muscle insertion scars on glabella. N, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 26. A–Q Maghroharpes ihmadii n. sp., Ihandar Formation, Pragian, Jebl Ou-Driss. A–G dorsal exoskeleton of cephalon, holotype, NHMUK It 29301 (Fig. 2B, Map 4, site 16). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. Eye lobe in lateral view E and G, and in dorsal view, F. H–Q dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29302 (Fig. 2B, Map 4, site 17). H, dorsaI, I, lateral, J, anterior and L, posterior-dorsal view. K and Q, dorsal view of brim perforations and pleurae, respectively, M, anterior dorsal view, O and P, lateral views of muscle insertion scars on glabella. N, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF97E36AFF78FE05FB82842C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232622/files/figure.png	https://doi.org/10.5281/zenodo.11232622	PLATE 27. A–D Maghroharpes ihmadii n. sp., dorsal exoskeleton of cephalon and disarticulated thorax, paratype, NHMUK It 29303, from type horizon and locality, Ihandar Formation, Pragian. Jbel Ou-Driss (Fig. 2B, Map 4, site 18). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Maghroharpes cf. ihamadii, dorsal exoskeleton pygidium enrolled, NHMUK It 29304, from “Kolihapeltis Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, Map 8, site 14). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe. J and K, dorsal view of brim perforations and pleurae, respectively. L–O Magroharpes hammii, dorsal exoskeleton, thorax and disarticulated pygidium, NHMUK It 29297, from Ihandar Formation, Iminwasief (Fig. 2B, Map 4, site 4). L, dorsaI, M, anterior, N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 27. A–D Maghroharpes ihmadii n. sp., dorsal exoskeleton of cephalon and disarticulated thorax, paratype, NHMUK It 29303, from type horizon and locality, Ihandar Formation, Pragian. Jbel Ou-Driss (Fig. 2B, Map 4, site 18). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–K Maghroharpes cf. ihamadii, dorsal exoskeleton pygidium enrolled, NHMUK It 29304, from “Kolihapeltis Couche”, Ihandar Formation, Pragian, Jbel Oufatene (Fig. 2B, Map 8, site 14). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe. J and K, dorsal view of brim perforations and pleurae, respectively. L–O Magroharpes hammii, dorsal exoskeleton, thorax and disarticulated pygidium, NHMUK It 29297, from Ihandar Formation, Iminwasief (Fig. 2B, Map 4, site 4). L, dorsaI, M, anterior, N, lateral view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF97E36AFF78FE05FB82842C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232636/files/figure.png	https://doi.org/10.5281/zenodo.11232636	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF95E36BFF78FBD2FD0E823A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232606/files/figure.png	https://doi.org/10.5281/zenodo.11232606	PLATE 24. A– L Maghroharpes zguidensis n. sp., Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–E dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29325 (Fig. 2A, Map 2, site1). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. F– I dorsal exoskeleton of cephalon, paratype, NHMUK It 29326, from type locality (Fig. 2A, Map 2, site 2). F, dorsal, G, lateral and H, anterior view. I, dorsal view of brim perforations. J–L dorsal exoskeleton of cephalon, paratype, NHMUK It 29327, from type locality (Fig. 2A, Map 2, site 3). J, dorsal, K, anterior and L, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 24. A– L Maghroharpes zguidensis n. sp., Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–E dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29325 (Fig. 2A, Map 2, site1). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. F– I dorsal exoskeleton of cephalon, paratype, NHMUK It 29326, from type locality (Fig. 2A, Map 2, site 2). F, dorsal, G, lateral and H, anterior view. I, dorsal view of brim perforations. J–L dorsal exoskeleton of cephalon, paratype, NHMUK It 29327, from type locality (Fig. 2A, Map 2, site 3). J, dorsal, K, anterior and L, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF95E36BFF78FBD2FD0E823A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232646/files/figure.png	https://doi.org/10.5281/zenodo.11232646	PLATE 34. A–I Eskoharpes sicarius n. sp., from Type horizon, lower Frasnian, Bouia, 7 km SE of Jorf (Fig. 2B, Map 6, site 5). A–E paratype, NHMUK it 29256 (2), dorsal exoskeleton of partial cephalon only. A, dorsal view, B, lateral view, C, anterior view, D, dorsal view of eye lobe and eye ridge and E, anterior dorsal view of brim perforations. F–G paratype, NHMUK it 29256 (5), dorsal exoskeleton partial cephalon. F, dorsal, G, lateral view of posterior of prolongation including genal spine. H paratype, NHMUK it 29256 (4), dorsal exoskeleton of partial cephalon and disarticulated thorax, dorsal view. I paratype, NHMUK it 29256 (1&7), dorsal exoskeleton of partial concealed cephalon, dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 34. A–I Eskoharpes sicarius n. sp., from Type horizon, lower Frasnian, Bouia, 7 km SE of Jorf (Fig. 2B, Map 6, site 5). A–E paratype, NHMUK it 29256 (2), dorsal exoskeleton of partial cephalon only. A, dorsal view, B, lateral view, C, anterior view, D, dorsal view of eye lobe and eye ridge and E, anterior dorsal view of brim perforations. F–G paratype, NHMUK it 29256 (5), dorsal exoskeleton partial cephalon. F, dorsal, G, lateral view of posterior of prolongation including genal spine. H paratype, NHMUK it 29256 (4), dorsal exoskeleton of partial cephalon and disarticulated thorax, dorsal view. I paratype, NHMUK it 29256 (1&7), dorsal exoskeleton of partial concealed cephalon, dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF95E36BFF78FBD2FD0E823A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232636/files/figure.png	https://doi.org/10.5281/zenodo.11232636	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF95E36BFF78FBD2FD0E823A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232602/files/figure.png	https://doi.org/10.5281/zenodo.11232602	PLATE 23. A–D Maghroharpes aff. zguidensis, dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29287, from Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–P Maghroharpes minutipunctatus n. sp. E–J holotype, NHMUK It 29312, from “Dicranurus Couche”, Ihandar Formation, Pragian, ElAtchane, Jbel Oufatene (Fig. 2B, Map 8, site 11), dorsal exoskeleton. E, dorsal, F, anterior and G, lateral view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations and J, lateral view of eye lobe. K–P paratype, NHMUK It 29313, from type locality (Fig. 2B, Map 8, site 12), partial dorsal exoskeleton with pygidium enrolled. K, dorsal and L, lateral view. M, N, O, and P, dorsal view of pleurae, brim perforations, lower lamella and hypostome, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 23. A–D Maghroharpes aff. zguidensis, dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29287, from Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–P Maghroharpes minutipunctatus n. sp. E–J holotype, NHMUK It 29312, from “Dicranurus Couche”, Ihandar Formation, Pragian, ElAtchane, Jbel Oufatene (Fig. 2B, Map 8, site 11), dorsal exoskeleton. E, dorsal, F, anterior and G, lateral view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations and J, lateral view of eye lobe. K–P paratype, NHMUK It 29313, from type locality (Fig. 2B, Map 8, site 12), partial dorsal exoskeleton with pygidium enrolled. K, dorsal and L, lateral view. M, N, O, and P, dorsal view of pleurae, brim perforations, lower lamella and hypostome, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF94E36CFF78FDE1FE188057.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232640/files/figure.png	https://doi.org/10.5281/zenodo.11232640	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF93E36EFF78FF0BFB3387A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232612/files/figure.png	https://doi.org/10.5281/zenodo.11232612	PLATE 25 A–G Maghroharpes cf. 2 laatchanensis, Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–D dorsal exoskeleton of cephalon only, NHMUK It 29310 (i), from Foum Zguid (Fig. 2A, Map 1, site 4). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–G dorsal exoskeleton of cephalon and disarticulated thorax, NHMUK It 29310 (ii), on the same plate as NHMUK It 29310 (i). E, dorsal, F, anterior and G, lateral view. H Maghroharpes oufatenensis n. sp., from coral at top of Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 2), 9 cephala only, paratypes, NHMUK It 29318 (1 to 9), dorsal view of cephala. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 25 A–G Maghroharpes cf. 2 laatchanensis, Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–D dorsal exoskeleton of cephalon only, NHMUK It 29310 (i), from Foum Zguid (Fig. 2A, Map 1, site 4). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–G dorsal exoskeleton of cephalon and disarticulated thorax, NHMUK It 29310 (ii), on the same plate as NHMUK It 29310 (i). E, dorsal, F, anterior and G, lateral view. H Maghroharpes oufatenensis n. sp., from coral at top of Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 2), 9 cephala only, paratypes, NHMUK It 29318 (1 to 9), dorsal view of cephala. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF93E36EFF78FF0BFB3387A4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232624/files/figure.png	https://doi.org/10.5281/zenodo.11232624	PLATE 28. A–O Maghroharpes oufatenensis n. sp., A–E dorsal exoskeleton of cephalon, holotype, NHMUK It 29320 (1), from lower Pragian horizon, Hamar Laghdad (Fig. 2B, Map 11, site 8). A, dorsal and B, lateral view. C, lateral view and D, (uncoated) dorsal-lateral view of eye lobe, respectively. E, dorsal view of brim perforations. F–I dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29316, Ihandar Formation, Jbel Oufatene (Fig. 2B, Map 8, site 23). F, dorsal, G, lateral and H, anterior view. I, dorsal view of brim perforations. J–O dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29319, Ihandar Formation, Jbel Oufatene (Fig. 2B, Map 8, site 24). J, dorsal, K, anterior and L, lateral view. M, N and O, eye lobe in lateral, anterior lateral and posterior lateral view, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 28. A–O Maghroharpes oufatenensis n. sp., A–E dorsal exoskeleton of cephalon, holotype, NHMUK It 29320 (1), from lower Pragian horizon, Hamar Laghdad (Fig. 2B, Map 11, site 8). A, dorsal and B, lateral view. C, lateral view and D, (uncoated) dorsal-lateral view of eye lobe, respectively. E, dorsal view of brim perforations. F–I dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29316, Ihandar Formation, Jbel Oufatene (Fig. 2B, Map 8, site 23). F, dorsal, G, lateral and H, anterior view. I, dorsal view of brim perforations. J–O dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29319, Ihandar Formation, Jbel Oufatene (Fig. 2B, Map 8, site 24). J, dorsal, K, anterior and L, lateral view. M, N and O, eye lobe in lateral, anterior lateral and posterior lateral view, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF91E36FFF78F95AFDE38448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232602/files/figure.png	https://doi.org/10.5281/zenodo.11232602	PLATE 23. A–D Maghroharpes aff. zguidensis, dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29287, from Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–P Maghroharpes minutipunctatus n. sp. E–J holotype, NHMUK It 29312, from “Dicranurus Couche”, Ihandar Formation, Pragian, ElAtchane, Jbel Oufatene (Fig. 2B, Map 8, site 11), dorsal exoskeleton. E, dorsal, F, anterior and G, lateral view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations and J, lateral view of eye lobe. K–P paratype, NHMUK It 29313, from type locality (Fig. 2B, Map 8, site 12), partial dorsal exoskeleton with pygidium enrolled. K, dorsal and L, lateral view. M, N, O, and P, dorsal view of pleurae, brim perforations, lower lamella and hypostome, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 23. A–D Maghroharpes aff. zguidensis, dorsal exoskeleton of partial cephalon, holotype, NHMUK It 29287, from Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–P Maghroharpes minutipunctatus n. sp. E–J holotype, NHMUK It 29312, from “Dicranurus Couche”, Ihandar Formation, Pragian, ElAtchane, Jbel Oufatene (Fig. 2B, Map 8, site 11), dorsal exoskeleton. E, dorsal, F, anterior and G, lateral view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations and J, lateral view of eye lobe. K–P paratype, NHMUK It 29313, from type locality (Fig. 2B, Map 8, site 12), partial dorsal exoskeleton with pygidium enrolled. K, dorsal and L, lateral view. M, N, O, and P, dorsal view of pleurae, brim perforations, lower lamella and hypostome, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF91E36FFF78F95AFDE38448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232598/files/figure.png	https://doi.org/10.5281/zenodo.11232598	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF90E370FF78FBF6FCB1815C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232594/files/figure.png	https://doi.org/10.5281/zenodo.11232594	PLATE 21. A–O Maghroharpes laatchanensis n. sp., from Ihandar Formation, Pragian. A–G dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29308, from “Kolihapeltis Couche”,Oued el Atchane Jbel Oufatene (Fig. 2B, Map 8, site 15). A, dorsal, B, anterior, C, posterior and D, lateral view. E, dorsal view of brim perforations and F, lateral view of axial rings and pleurae. H–K dorsal exoskeleton with pygidium missing, paratype, NHMUK It 29307 from type locality (Fig. 2B, Map 8, site 13). H, dorsal, I, anterior and J, lateral view. K, dorsal view of brim perforations. L–O dorsal exoskeleton with pygidium disarticulated, paratype, NHMUK It 29306, from type locality (Fig. 2B, Map 8, site 3). L, dorsal, M, anterior, N, lateral and O, posterior view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 21. A–O Maghroharpes laatchanensis n. sp., from Ihandar Formation, Pragian. A–G dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29308, from “Kolihapeltis Couche”,Oued el Atchane Jbel Oufatene (Fig. 2B, Map 8, site 15). A, dorsal, B, anterior, C, posterior and D, lateral view. E, dorsal view of brim perforations and F, lateral view of axial rings and pleurae. H–K dorsal exoskeleton with pygidium missing, paratype, NHMUK It 29307 from type locality (Fig. 2B, Map 8, site 13). H, dorsal, I, anterior and J, lateral view. K, dorsal view of brim perforations. L–O dorsal exoskeleton with pygidium disarticulated, paratype, NHMUK It 29306, from type locality (Fig. 2B, Map 8, site 3). L, dorsal, M, anterior, N, lateral and O, posterior view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF90E370FF78FBF6FCB1815C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232598/files/figure.png	https://doi.org/10.5281/zenodo.11232598	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 22. A–E Maghroharpes cf. 1 laatchanensis, Ihandar Formation, Pragian, Jbel el Otfal (Fig. 2B, Map 9, site 8), dorsal exoskeleton of partial cephalon, NHMUK It 29311. A, dorsal, B, anterior and C, lateral view. D and E, dorsal view of brim perforations and occipital node and tubercles, respectively. F–M Maghroharpes aff.1 minutipunctus, Boutiskaoine Formation? lower Emsian. F–J dorsal exoskeleton of partial cephalon, NHMUK It 29322, Jbel Ou-Driss (North) (Fig. 2B, Map 4, site 7). F, dorsal, G, anterior and H, lateral view. I, lateral and J, dorsal view of eye lobe. K–M dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29323, Boutiskaouine Formation, lower Emsian, Ahware (Fig. 2B, Map 9, site 13). K, L and M, dorsal view of cephalon, brim perforations and eye lobe, respectively. N–Q Maghroharpes aff. 2 minutipunctus, Ihandar Formation Jbel Ou-Driss (West) (Fig. 2B, Map 4, site 20), dorsal exoskeleton of partial cephalon, NHMUK It 29324. N, dorsal, O, lateral and P, anterior view. Q, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF90E370FF78FBF6FCB1815C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232612/files/figure.png	https://doi.org/10.5281/zenodo.11232612	PLATE 25 A–G Maghroharpes cf. 2 laatchanensis, Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–D dorsal exoskeleton of cephalon only, NHMUK It 29310 (i), from Foum Zguid (Fig. 2A, Map 1, site 4). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–G dorsal exoskeleton of cephalon and disarticulated thorax, NHMUK It 29310 (ii), on the same plate as NHMUK It 29310 (i). E, dorsal, F, anterior and G, lateral view. H Maghroharpes oufatenensis n. sp., from coral at top of Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 2), 9 cephala only, paratypes, NHMUK It 29318 (1 to 9), dorsal view of cephala. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 25 A–G Maghroharpes cf. 2 laatchanensis, Merzâ Akhsai Formation?, Pragian, Foum Zguid. A–D dorsal exoskeleton of cephalon only, NHMUK It 29310 (i), from Foum Zguid (Fig. 2A, Map 1, site 4). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–G dorsal exoskeleton of cephalon and disarticulated thorax, NHMUK It 29310 (ii), on the same plate as NHMUK It 29310 (i). E, dorsal, F, anterior and G, lateral view. H Maghroharpes oufatenensis n. sp., from coral at top of Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 2), 9 cephala only, paratypes, NHMUK It 29318 (1 to 9), dorsal view of cephala. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232628/files/figure.png	https://doi.org/10.5281/zenodo.11232628	PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 29. A–D Maghroharpes forteyi n. sp., from top of Pragian, Ihandar Formation, El Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29288 (Fig. 2B, Map 8, site 16). A, dorsal B, anterior and C, lateral view. D, dorsal view of brim perforations. E–J,O Maghroharpes azmamarensis n. sp., from top of Pragian, Ihandar Formation, Oued el Atchane Jbel Oufatene, dorsal exoskeleton of cephalon, holotype, NHMUK It 29281 (Fig. 2B, Map 8, site 18). E, dorsal, F, anterior and G, lateral view. H, dorsal and I, lateral view of eye lobe, respectively. J, dorsal view of brim perforations. O, from type locality partial dorsal exoskeleton of lower Lamella, paratype NHMUK It 29283 (Fig. 2B, Map 8, site 19)). K–N Maghroharpes forteyi, from top of Pragian, Ihandar Formation, bou Lachrhal, dorsal exoskeleton of partial cephalon only, paratype, NHMUK It 29289 (Fig. 2B, Map 9, Site 7). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232636/files/figure.png	https://doi.org/10.5281/zenodo.11232636	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 31. A–E Maghroharpes aff. ihmadii, dorsal exoskeleton of partial cephalon, NHMUK It 29305, from an upper Pragian horizon, Ihandar Formation, bou Lachrhal (Fig. 2B, Map 9, site 4). A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–K Maghroharpes cf. forteyi, dorsal exoskeleton of partial cephalon, NHMUK It 29284, from top of the Pragian, Ihandar Formation, Oued el Atchane, Jbel Oufatene (Fig. 2B, Map 8, site 20). F, dorsal, G, anterior and H, lateral view. I, lateral view of eye lobe, J, dorsal view of brim perforations and K, pits on axial node. L–N Maghroharpes cf. zguidensis, dorsal exoskeleton of cephalon, NHMUK It 29328, from MerzâAkhsai Formation?, Pragian, Douar Laayoune, Tata (Fig. 2B, Map 1, site 1). L, dorsal and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232511/files/figure.png	https://doi.org/10.5281/zenodo.11232511	FIGURE 3. Souss–Massa, Tiinghir, West and North Ma’der regions	FIGURE 3. Souss–Massa, Tiinghir, West and North Ma’der regions	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232513/files/figure.png	https://doi.org/10.5281/zenodo.11232513	FIGURE 5. East Ma’der, Tafilalt and Taouz	FIGURE 5. East Ma’der, Tafilalt and Taouz	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8FE372FF78FE02FDB68398.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8DE373FF78FD46FD6F8224.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232648/files/figure.png	https://doi.org/10.5281/zenodo.11232648	PLATE 35. A–P Pinnuloharpes segaouii n. sp., El Otfal Formation, upper Eifelian. A–G dorsal exoskeleton partially enrolled, holotype, NHMUK It 29351, from Taghroute near Timerzit (Fig. 2B, Map 10, site 4). A, dorsal, B, posterior, C, anterior and D, lateral view. E and F, dorsal view of pleurae and brim perforations, respectively. G, lateral view of eye lobe (uncoated). H–K dorsal exoskeleton enrolled, paratype, NHMUK It 29352, the lower “Harpes Couche” (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 11). H, dorsal, I, anterior and J, lateral view. K, dorsal view of brim perforations. L–P dorsal exoskeleton of an incomplete cephalon, paratype, NHMUK It.29350 from the main “Harpes” horizon, 2 m upslope of the “Acanthopyge” horizon, El Otfal Formation? Eifelian/Givetian boundary, Timerzit (Fig. 2B, Map 10, site 1). L, dorsal, M, lateral and N, anterior view. O and P, dorsal view of and brim perforations and impression of ventral surface cuticle on genal area, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 35. A–P Pinnuloharpes segaouii n. sp., El Otfal Formation, upper Eifelian. A–G dorsal exoskeleton partially enrolled, holotype, NHMUK It 29351, from Taghroute near Timerzit (Fig. 2B, Map 10, site 4). A, dorsal, B, posterior, C, anterior and D, lateral view. E and F, dorsal view of pleurae and brim perforations, respectively. G, lateral view of eye lobe (uncoated). H–K dorsal exoskeleton enrolled, paratype, NHMUK It 29352, the lower “Harpes Couche” (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 11). H, dorsal, I, anterior and J, lateral view. K, dorsal view of brim perforations. L–P dorsal exoskeleton of an incomplete cephalon, paratype, NHMUK It.29350 from the main “Harpes” horizon, 2 m upslope of the “Acanthopyge” horizon, El Otfal Formation? Eifelian/Givetian boundary, Timerzit (Fig. 2B, Map 10, site 1). L, dorsal, M, lateral and N, anterior view. O and P, dorsal view of and brim perforations and impression of ventral surface cuticle on genal area, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8DE373FF78FD46FD6F8224.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232517/files/figure.png	https://doi.org/10.5281/zenodo.11232517	FIGURE 7. Section 2 Jbel Issoumour	FIGURE 7. Section 2 Jbel Issoumour	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8CE375FF78FDE2FE3E86F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232660/files/figure.png	https://doi.org/10.5281/zenodo.11232660	PLATE 38.A–E Pinnuloharpes aff. torquis, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 20), NHMUK It 29329. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F–I Pinnuloharpes aff. apteros, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, “Harpes Couche”, Ksar Hannabou (Fig. 2B, Map 6, site 6), NHMUK It 29379. F, dorsal, G, lateral and H, anterior view. I, lateral view of eye lobe. J–O Pinnuloharpes apteros n. sp.,dorsal exoskeleton, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 18), holotype, NHMUK It 29358. J, dorsal, K, anterior and L, lateral view. M, lateral view of pleurae, N, dorsal view of brim perforations and O, posterior view of pygidium. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 38.A–E Pinnuloharpes aff. torquis, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 20), NHMUK It 29329. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F–I Pinnuloharpes aff. apteros, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, “Harpes Couche”, Ksar Hannabou (Fig. 2B, Map 6, site 6), NHMUK It 29379. F, dorsal, G, lateral and H, anterior view. I, lateral view of eye lobe. J–O Pinnuloharpes apteros n. sp.,dorsal exoskeleton, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 18), holotype, NHMUK It 29358. J, dorsal, K, anterior and L, lateral view. M, lateral view of pleurae, N, dorsal view of brim perforations and O, posterior view of pygidium. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8CE375FF78FDE2FE3E86F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232664/files/figure.png	https://doi.org/10.5281/zenodo.11232664	PLATE 39. A–M Pinnuloharpes apteros n. sp., El Otfal Formation, upper Eifelian. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29356 from Ksar Hannabou (Fig. 2B, Map 6, site 8). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–K dorsal exoskeleton with thorax and pygidium disarticulated, paratype, NHMUK It 29359, from type locality and horizon (Fig. 2B, Map 6, site 19). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, lateral view of eye lobe, J, anterior lateral view of rim and marginal band, and K, dorsal view of pleurae close to pygidium. L–M dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29357, from Ksar Hannabou (Fig. 2B, Map 6, site 15). L, dorsal view and M, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 39. A–M Pinnuloharpes apteros n. sp., El Otfal Formation, upper Eifelian. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29356 from Ksar Hannabou (Fig. 2B, Map 6, site 8). A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations. E–K dorsal exoskeleton with thorax and pygidium disarticulated, paratype, NHMUK It 29359, from type locality and horizon (Fig. 2B, Map 6, site 19). E, dorsal, F, anterior and G, lateral view. H, dorsal view of brim perforations, I, lateral view of eye lobe, J, anterior lateral view of rim and marginal band, and K, dorsal view of pleurae close to pygidium. L–M dorsal exoskeleton of a partial cephalon, paratype, NHMUK It 29357, from Ksar Hannabou (Fig. 2B, Map 6, site 15). L, dorsal view and M, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF8AE376FF78F9A6FD6C85BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232668/files/figure.png	https://doi.org/10.5281/zenodo.11232668	PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF89E377FF78FB6FFDFE81CC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232674/files/figure.png	https://doi.org/10.5281/zenodo.11232674	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF88E379FF78FF72FB5E866C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232666/files/figure.png	https://doi.org/10.5281/zenodo.11232666	PLATE 40.A–N Pinnuloharpes hannabouensis n. sp., from an horizon 5 m down slope of “Radiaspis Couche”, upper Eifelian/ Givetian boundary, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29334 (Fig. 2B, Map 6, site 12). A, dorsal and B, lateral view. C, lateral view of eye lobe (uncoated) and D, dorsal view of brim perforations. E–I dorsal exoskeleton cephalon and thorax only, holotype, NHMUK It 29333 (Fig. 2B, Map 6, site 11). E, dorsal, F, anterior and G, lateral view. H, lateral view of eye lobes, I, dorsal view of brim perforations. J–N dorsal exoskeleton pygidium incomplete, paratype, NHMUK It 29335 (Fig. 2B, Map 6, site 13). J, dorsal, K, anterior and L, lateral view. M, lateral view of eye lobe and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 40.A–N Pinnuloharpes hannabouensis n. sp., from an horizon 5 m down slope of “Radiaspis Couche”, upper Eifelian/ Givetian boundary, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29334 (Fig. 2B, Map 6, site 12). A, dorsal and B, lateral view. C, lateral view of eye lobe (uncoated) and D, dorsal view of brim perforations. E–I dorsal exoskeleton cephalon and thorax only, holotype, NHMUK It 29333 (Fig. 2B, Map 6, site 11). E, dorsal, F, anterior and G, lateral view. H, lateral view of eye lobes, I, dorsal view of brim perforations. J–N dorsal exoskeleton pygidium incomplete, paratype, NHMUK It 29335 (Fig. 2B, Map 6, site 13). J, dorsal, K, anterior and L, lateral view. M, lateral view of eye lobe and N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF86E37AFF78F912FF5C827C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232670/files/figure.png	https://doi.org/10.5281/zenodo.11232670	PLATE 42. A–M Pinnuloharpes haustrum n. sp., upper Emsian, Ksar Lbour. A–F dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29341 (Fig. 2B, Map 3, site 3). A, dorsal, B, lateral and C, anterior view. D, and E, dorsal view of brim perforations and genal area sculpture respectively. G–M dorsal exoskeleton pygidium enrolled, paratype, NHMUK It 29342 (Fig. 2B, Map 3, site 4). G, dorsal, H, lateral and I, anterior view. J, lateral view of eye lobe (uncoated), K, anterior view of genal roll perforations and L, dorsal view of brim perforations. M, anterior view of marginal band. N–O Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 11, site 13). N, dorsal view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 42. A–M Pinnuloharpes haustrum n. sp., upper Emsian, Ksar Lbour. A–F dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29341 (Fig. 2B, Map 3, site 3). A, dorsal, B, lateral and C, anterior view. D, and E, dorsal view of brim perforations and genal area sculpture respectively. G–M dorsal exoskeleton pygidium enrolled, paratype, NHMUK It 29342 (Fig. 2B, Map 3, site 4). G, dorsal, H, lateral and I, anterior view. J, lateral view of eye lobe (uncoated), K, anterior view of genal roll perforations and L, dorsal view of brim perforations. M, anterior view of marginal band. N–O Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 11, site 13). N, dorsal view. O, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF86E37AFF78F912FF5C827C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232672/files/figure.png	https://doi.org/10.5281/zenodo.11232672	PLATE 43. A–B Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 13, site 11). A, anterior and B, lateral view. C– P Pinnuloharpes aff. haustrum, from base of upper Emsian, Hamar Laghdad. C–L dorsal exoskeleton with thorax enrolled, NHMUK It 29347 (1) (Fig. 2B, Map 11, site 7). C, dorsal, D, lateral and E, anterior view. F, dorsal view of brim perforations, G, anterior view of external rim. H, I and J dorsal view of sculpture on dorsal surface of genal area, brim perforations and occipital node, respectively. K, anterior view of perforations on the genal roll and L dorsal view of eye lobe. M–P ventral exoskeleton of cephalon only, NHMUK It 29347 (2), from same site as NHMUK It 29347 (1). M, ventral and N, lateral view. O and P, ventral view of hypostome and external rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 43. A–B Pinnuloharpes haustrum n. sp., upper Emsian, Hamar Laghdad, dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29345 (Fig. 2B, Map 13, site 11). A, anterior and B, lateral view. C– P Pinnuloharpes aff. haustrum, from base of upper Emsian, Hamar Laghdad. C–L dorsal exoskeleton with thorax enrolled, NHMUK It 29347 (1) (Fig. 2B, Map 11, site 7). C, dorsal, D, lateral and E, anterior view. F, dorsal view of brim perforations, G, anterior view of external rim. H, I and J dorsal view of sculpture on dorsal surface of genal area, brim perforations and occipital node, respectively. K, anterior view of perforations on the genal roll and L dorsal view of eye lobe. M–P ventral exoskeleton of cephalon only, NHMUK It 29347 (2), from same site as NHMUK It 29347 (1). M, ventral and N, lateral view. O and P, ventral view of hypostome and external rim, respectively. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF86E37AFF78F912FF5C827C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232674/files/figure.png	https://doi.org/10.5281/zenodo.11232674	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 44. A–M Pinnuloharpes hamarlaghdadensis (Cronier et al. 2018), from Hamar Laghdad, upper Emsian. A–F dorsal exoskeleton with pygidium enrolled, NHMUK It 29337 (Fig. 2B, Map 11, site 1) A, dorsal, B, anterior and C, lateral view. D, dorsal view of brim perforations, E, dorsal view of occipital node and F, lateral view of eye lobe. G–J dorsal exoskeleton of cephalon and 2 pleural segments of thorax, NHMUK It 29338 (Fig. 2B, Map 11, site 2) G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–M dorsal exoskeleton with thorax disarticulated and pygidium absent, NHMUK It 29339 (Fig. 2B, Map 11, site 3) K, dorsal and I, anterior view. L, dorsal view of brim perforations and M, lateral view of distal ends of pleurae. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF86E37AFF78F912FF5C827C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232640/files/figure.png	https://doi.org/10.5281/zenodo.11232640	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 32. A–D Maghroharpes terridus n. sp., Ihandar Formation, Pragian, Maharch, dorsal exoskeleton, thorax and pygidium disarticulated, holotype, NHMUK It 29321 (Fig. 2B, Map 9, site 18). A, dorsal, B, lateral and C, posterior view. D, dorsal view of brim perforations. E–Q Lioharpes scopulum n. sp., Ihandar Formation, bou Lachrhal. E–L dorsal exoskeleton of cephalon only, holotype, NHMUK It 29279 (Fig. 2B, Map 9, site 14). E, Dorsal, F, lateral and G, anterior view. H, lateral view of glabella (uncoated), I, dorsal view of brim perforations. J, and K, dorsal and lateral view of eye lobe respectively. L, dorsal view of occipital node. M–Q dorsal exoskeletons of partial cephala only, from type locality and horizon, paratypes, NHMUK It 29280 (1 to 3) (Fig. 2B, Map 9, site 3). M–N NHMUK It 29280 (1). M, dorsal view and N, dorsal view of eye lobe. O–P NHMUK It 29280 (2). O, dorsal view and P, dorsal view of brim perforations. Q, NHMUK It 29280 (3), dorsal view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF85E37BFF78FD2FFB0180C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232654/files/figure.png	https://doi.org/10.5281/zenodo.11232654	PLATE 36. A–K Harpes igaouii n. sp., Ihandar Formation, Pragian-Emsian boundary. A–H dorsal exoskeleton of cephalon only, holotype, NHMUK It 29360, from coral NW of Jbel Ou-Driss (Fig. 2B, Map 4, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal anterior view of perforations on genal roll and brim. E, dorsal view of glabellar tubercles, F, dorsal view of brim perforations, G, lateral view of glabellar muscle insertion scars (uncoated) and H, lateral view of eye lobe (uncoated). I–K dorsal exoskeleton of cephalon and partial thorax, paratype, NHMUK It 29361, from Ceratocephala horizon, North Jbel Ou-Driss (Fig. 2B, Map 4, site 8). I, dorsal anterior lateral view (uncoated). J, ventral view of hypostome and K, lateral view of eye lobe (uncoated). L–O Harpes aff. igaouii, dorsal exoskeleton of cephalon only, NHMUK It 29362, from Fezna, “Struveaspis Couche”, El Otfal Formation, Eifelian.(Fig. 2B, Map 6, site 1). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. P–R Kielania aff. tumula, Ihandar Formation, upper Pragian (Fig. 2B, Map 10, Site 5) dorsal exoskeleton of a partial cephalon, NHMUK It 29262. P, dorsal, Q, anterior and R, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 36. A–K Harpes igaouii n. sp., Ihandar Formation, Pragian-Emsian boundary. A–H dorsal exoskeleton of cephalon only, holotype, NHMUK It 29360, from coral NW of Jbel Ou-Driss (Fig. 2B, Map 4, site 5). A, dorsal, B, anterior and C, lateral view. D, dorsal anterior view of perforations on genal roll and brim. E, dorsal view of glabellar tubercles, F, dorsal view of brim perforations, G, lateral view of glabellar muscle insertion scars (uncoated) and H, lateral view of eye lobe (uncoated). I–K dorsal exoskeleton of cephalon and partial thorax, paratype, NHMUK It 29361, from Ceratocephala horizon, North Jbel Ou-Driss (Fig. 2B, Map 4, site 8). I, dorsal anterior lateral view (uncoated). J, ventral view of hypostome and K, lateral view of eye lobe (uncoated). L–O Harpes aff. igaouii, dorsal exoskeleton of cephalon only, NHMUK It 29362, from Fezna, “Struveaspis Couche”, El Otfal Formation, Eifelian.(Fig. 2B, Map 6, site 1). L, dorsal, M, lateral and N, anterior view. O, dorsal view of brim perforations. P–R Kielania aff. tumula, Ihandar Formation, upper Pragian (Fig. 2B, Map 10, Site 5) dorsal exoskeleton of a partial cephalon, NHMUK It 29262. P, dorsal, Q, anterior and R, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF85E37BFF78FD2FFB0180C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232573/files/figure.png	https://doi.org/10.5281/zenodo.11232573	PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 13. A–J Lioharpes ammari n. sp., from horizon 2, Section 1 (Fig. 6), Tazoulait Formation, upper Emisian, South flank of Jbel Ou-Driss (Fig. 2B, Map 4). A–E dorsal exoskeleton, pygidium missing, paratype, NHMUK It 29264 (site 12).A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe and E, dorsal view of brim perforations. F–H dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29265 (1) (site 13). F, dorsal, G, lateral and H, anterior view. I–J ventral exoskeleton of a partial upper lamella, paratype, NHMUK It 29265 (2) (site 13). I, ventral view and J, lateral view of ventral surface. K–O Pinnuloharpes igaouii n. sp., from a horizon 20 metres down slope from type horizon of Lioharpes ammari n. sp. Boutiskaouine Formation?, lower Emsian North flank of Jbel Ou-Driss (Fig. 2B Map 4 site 6). Dorsal exoskeleton of cephalon only, paratype NHMUK it 29267. K dorsal, L anterior and M lateral view. N lateral view of eye lobe and O dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF85E37BFF78FD2FFB0180C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF83E37DFF78F8EBFAFE8568.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232676/files/figure.png	https://doi.org/10.5281/zenodo.11232676	PLATE 45. A–L Pinnuloharpes merzanensis n. sp., upper Emsian, Hamar Laghdad. A–H dorsal exoskeleton partly enrolled, holotype, NHMUK It 29349 from 7.5 m below P. hamarlaghdadensis horizon (Fig. 2B, Map 11, site 6). A, dorsal, B, anterior, C, lateral, D, posteroventral and E, posterolateral view. F, dorsal view of brim perforation, G, dorsal view of pygidium and H, dorsal view of pitting on genal area. I–L dorsal exoskeleton of a partial cephalon only, paratype, NHMUK It 29348 from 2.5 m above type horizon (Fig. 2B, Map 11, site 5). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 45. A–L Pinnuloharpes merzanensis n. sp., upper Emsian, Hamar Laghdad. A–H dorsal exoskeleton partly enrolled, holotype, NHMUK It 29349 from 7.5 m below P. hamarlaghdadensis horizon (Fig. 2B, Map 11, site 6). A, dorsal, B, anterior, C, lateral, D, posteroventral and E, posterolateral view. F, dorsal view of brim perforation, G, dorsal view of pygidium and H, dorsal view of pitting on genal area. I–L dorsal exoskeleton of a partial cephalon only, paratype, NHMUK It 29348 from 2.5 m above type horizon (Fig. 2B, Map 11, site 5). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF83E37DFF78F8EBFAFE8568.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232547/files/figure.png	https://doi.org/10.5281/zenodo.11232547	PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 5. A–J Harpes lahceni, dorsal exoskeleton partially enrolled, holotype, NHMUK It 29232, from El Otfal Formation, Eifelian, Horizon 3, Section 2 (Fig. 7), Jbel Issoumour (Fig. 2B, Map 7, site 15). A, dorsal, B, lateraland C, anterior view. D, posterior view, showing dorsal surface partially enrolled thorax and pygidium, and ventral surface of prolongations, E, lateral view of eye lobe, F, ventral view of prolongations, G, dorsal lateral view of eye lobe, H, dorsal view of brim perforations, I, dorsal view of dorsal surface of rim and J, dorsal view of pygidium. K–L Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29234, from El Otfal Formation, Eifelian, “Harpes Couche” above Acanthopyge horizon, Timerzit (Fig. 2B, Map 10). K, dorsal view and L, lateral view. M–N Harpes lahceni, dorsal exoskeleton partially enrolled, paratype, NHMUK It 29233, from El Otfal Formation, “Phylonix Couche”, Jbel Oufatene (Fig. 2B, Map 8). M, dorsal and N, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF82E37EFF78FA16FCCF8033.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232656/files/figure.png	https://doi.org/10.5281/zenodo.11232656	PLATE 37. A–O Pinnuloharpes torquis n. sp., from “Harpes Couche”, El Otfal Formation, upper Eifelian, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29353 (Fig. 2B, Map 6, site 7). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforation. E–J 2 dorsal exoskeletons of cephala only (one cephalon largely covering the other), holotype, NHMUK It 29354 (1) and paratype NHMUK It 29354 (2) (Fig. 2B, Map 6, site 9). NHMUK It 29354 (1) E, dorsal view, F, lateral and G, anterior view. H, anterior dorsal view of perforations on genal roll and brim. I and J, dorsal view of impression of ventral surface of cuticle in genal area and brim perforations, respectively. NHMUK It 29354 (2) E, dorsal view. K–O dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29355 (1) (Fig. 2B, Map 6, site 10). K, dorsal and L, lateral view. M, dorsal view of brim perforations, N and O, lateral view of eye lobes (uncoated). Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 37. A–O Pinnuloharpes torquis n. sp., from “Harpes Couche”, El Otfal Formation, upper Eifelian, Ksar Hannabou. A–D dorsal exoskeleton of cephalon only, paratype, NHMUK It 29353 (Fig. 2B, Map 6, site 7). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforation. E–J 2 dorsal exoskeletons of cephala only (one cephalon largely covering the other), holotype, NHMUK It 29354 (1) and paratype NHMUK It 29354 (2) (Fig. 2B, Map 6, site 9). NHMUK It 29354 (1) E, dorsal view, F, lateral and G, anterior view. H, anterior dorsal view of perforations on genal roll and brim. I and J, dorsal view of impression of ventral surface of cuticle in genal area and brim perforations, respectively. NHMUK It 29354 (2) E, dorsal view. K–O dorsal exoskeleton of partial cephalon, paratype, NHMUK It 29355 (1) (Fig. 2B, Map 6, site 10). K, dorsal and L, lateral view. M, dorsal view of brim perforations, N and O, lateral view of eye lobes (uncoated). Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF82E37EFF78FA16FCCF8033.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232660/files/figure.png	https://doi.org/10.5281/zenodo.11232660	PLATE 38.A–E Pinnuloharpes aff. torquis, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 20), NHMUK It 29329. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F–I Pinnuloharpes aff. apteros, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, “Harpes Couche”, Ksar Hannabou (Fig. 2B, Map 6, site 6), NHMUK It 29379. F, dorsal, G, lateral and H, anterior view. I, lateral view of eye lobe. J–O Pinnuloharpes apteros n. sp.,dorsal exoskeleton, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 18), holotype, NHMUK It 29358. J, dorsal, K, anterior and L, lateral view. M, lateral view of pleurae, N, dorsal view of brim perforations and O, posterior view of pygidium. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 38.A–E Pinnuloharpes aff. torquis, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 20), NHMUK It 29329. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F–I Pinnuloharpes aff. apteros, dorsal exoskeleton of cephalon only, El Otfal Formation, upper Eifelian, “Harpes Couche”, Ksar Hannabou (Fig. 2B, Map 6, site 6), NHMUK It 29379. F, dorsal, G, lateral and H, anterior view. I, lateral view of eye lobe. J–O Pinnuloharpes apteros n. sp.,dorsal exoskeleton, El Otfal Formation, upper Eifelian, Tamgoute Bulgan (Fig. 2B, Map 6, site 18), holotype, NHMUK It 29358. J, dorsal, K, anterior and L, lateral view. M, lateral view of pleurae, N, dorsal view of brim perforations and O, posterior view of pygidium. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF82E37EFF78FA16FCCF8033.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232519/files/figure.png	https://doi.org/10.5281/zenodo.11232519	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	FIGURE 9. Cladogram consensus of 636 trees with a 31 node collapse and each tree L = 1968, Ci = 12 and Ri =48	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF82E37EFF78FA16FCCF8033.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232521/files/figure.png	https://doi.org/10.5281/zenodo.11232521	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	FIGURE 10a. Lioharpes and Maghroharpes from tree 1 of 60 found using seed 7	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF81E340FF78FFF4FCBB84F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232678/files/figure.png	https://doi.org/10.5281/zenodo.11232678	PLATE 46. A–O Stoloharpes rissaniensis n. sp., El Otfal formation, South Rissani, Eifelian. A–M articulated dorsal exoskeleton with thorax partly enrolled, holotype, NHMUK It 29373 (Fig. 2B, Map 14, site 4). A, dorsal, B, anterior, C, lateral and D, antero-dorsal view. E, Antero-dorsal view of perforations on genal roll and brim, F, G and H, dorsal view of pleurae thorax and pygidium, respectively. I, lateral view of thorax and J, ventral view of exterior rim. K, dorsal view of brim perforations, L, lateral view and M, anterior view of glabella muscle insertion scars (uncoated). N–O partial dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29376 (Fig. 2B, Map 14, site 7). N, dorsal view and O, anterior view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 46. A–O Stoloharpes rissaniensis n. sp., El Otfal formation, South Rissani, Eifelian. A–M articulated dorsal exoskeleton with thorax partly enrolled, holotype, NHMUK It 29373 (Fig. 2B, Map 14, site 4). A, dorsal, B, anterior, C, lateral and D, antero-dorsal view. E, Antero-dorsal view of perforations on genal roll and brim, F, G and H, dorsal view of pleurae thorax and pygidium, respectively. I, lateral view of thorax and J, ventral view of exterior rim. K, dorsal view of brim perforations, L, lateral view and M, anterior view of glabella muscle insertion scars (uncoated). N–O partial dorsal exoskeleton with pygidium enrolled, paratype, NHMUK It 29376 (Fig. 2B, Map 14, site 7). N, dorsal view and O, anterior view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF81E340FF78FFF4FCBB84F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232681/files/figure.png	https://doi.org/10.5281/zenodo.11232681	PLATE 47. A–J Stoloharpes rissaniensis n. sp., El Otfal formation, Eifelian, South Rissani. A–F dorsal exoskeleton of cephalon only, paratype, NHMUK It 29374 (Fig. 2B, Map 14, site 5). A, dorsal, B, anterior, C, lateral view. D, dorsal view of brim perforations. E, lateral view of glabella muscle insertion scars (uncoated) and F, lateral view of eye lobe (uncoated). G–J disarticulated dorsal exoskeleton, paratype, NHMUK It 29375 (Fig. 2B, Map 14, site 6). G, dorsal, H, lateral and I, anterodorsal view. J, dorsal view of brim perforation. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 47. A–J Stoloharpes rissaniensis n. sp., El Otfal formation, Eifelian, South Rissani. A–F dorsal exoskeleton of cephalon only, paratype, NHMUK It 29374 (Fig. 2B, Map 14, site 5). A, dorsal, B, anterior, C, lateral view. D, dorsal view of brim perforations. E, lateral view of glabella muscle insertion scars (uncoated) and F, lateral view of eye lobe (uncoated). G–J disarticulated dorsal exoskeleton, paratype, NHMUK It 29375 (Fig. 2B, Map 14, site 6). G, dorsal, H, lateral and I, anterodorsal view. J, dorsal view of brim perforation. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF81E340FF78FFF4FCBB84F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232683/files/figure.png	https://doi.org/10.5281/zenodo.11232683	PLATE 48. A–M Stoloharpes rissaniensis n. sp., “Basseiarges Couche”, Fezna, Eifelian. A–D dorsal exoskeleton cephalon only, paratype, NHMUK It 29371 (Fig. 2B, Map 6, site 3). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J dorsal exoskeleton cephalon and disarticulated thorax, paratype, NHMUK It 29372 (Fig. 2B, Map 6, site 22). E, dorsal, F, lateral and G, anterior view. H and I, lateral view of eye lobes. J, dorsal view of brim perforations. K–M Ighas El Otfal Formation, Eifelian (Fig. 2B, Map 14, site 8), dorsal exoskeleton of cephalon only, paratype, NHMUK It 29377. K, dorsal, L, anterior and M, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 48. A–M Stoloharpes rissaniensis n. sp., “Basseiarges Couche”, Fezna, Eifelian. A–D dorsal exoskeleton cephalon only, paratype, NHMUK It 29371 (Fig. 2B, Map 6, site 3). A, dorsal, B, lateral and C, anterior view. D, dorsal view of brim perforations. E–J dorsal exoskeleton cephalon and disarticulated thorax, paratype, NHMUK It 29372 (Fig. 2B, Map 6, site 22). E, dorsal, F, lateral and G, anterior view. H and I, lateral view of eye lobes. J, dorsal view of brim perforations. K–M Ighas El Otfal Formation, Eifelian (Fig. 2B, Map 14, site 8), dorsal exoskeleton of cephalon only, paratype, NHMUK It 29377. K, dorsal, L, anterior and M, lateral view. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FF81E340FF78FFF4FCBB84F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232687/files/figure.png	https://doi.org/10.5281/zenodo.11232687	PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFBFE341FF78FBAEFC3F82EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232687/files/figure.png	https://doi.org/10.5281/zenodo.11232687	PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 49. A–F Stoloharpes aff. rissaniensis, El Otfal formation, upper Eifelian, Ksar Hannabou (Fig. 2B, Map 6, site 14), dorsal exoskeleton, thorax disarticulated and pygidium missing, NHMUK It 29380. A, dorsal B, lateral and C, anterior view. D, dorsal and E, lateral view of eye lobe (uncoated). F, dorsal view of brim perforations. G–K Stoloharpes calceolus n. sp., “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 4), dorsal exoskeleton pygidium enrolled, holotype, NHMUK It 29363. G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations and K, lateral view of pleurae. L–P Stoloharpes aff. calceolus, “Akantharges Couche”, Eifelian, Ksar Lbor (Fig. 2B, Map 3, site 5), dorsal exoskeleton, thorax disarticulated pygidium enrolled, holotype, NHMUK It 29381. L, dorsal, M, anterior, N, lateral and O, posterior view. P, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFBEE343FF78FD92FAA28685.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232689/files/figure.png	https://doi.org/10.5281/zenodo.11232689	PLATE 50. A–F Stoloharpes capricornus n. sp., El Otfal Formation, upper Eifelian, Taouz East (Fig. 2B, Map 15, site 1), dorsal exoskeleton of cephalon only, holotype, NHMUK It 29364. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) E, lateral view of muscle insertion scars on glabella (uncoated). F, dorsal view of brim perforations. G–J Stoloharpes aff. capricornus, upper Eifelian, Hamar Laghdad (Fig. 2B, Map 11, site 14), dorsal exoskeleton of cephalon only, NHMUK It 29366. G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–O Stoloharpes capricornus n. sp., El Otfal Formation, upper Eifelian, TaouzEast (Fig.2B, Map 15, site 2), dorsal exoskeleton thorax enrolled, paratype, NHMUK It 29365. K, dorsal, L, lateral and M, posterior view. N, dorsal view of brim perforations and O, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 50. A–F Stoloharpes capricornus n. sp., El Otfal Formation, upper Eifelian, Taouz East (Fig. 2B, Map 15, site 1), dorsal exoskeleton of cephalon only, holotype, NHMUK It 29364. A, dorsal, B, lateral and C, anterior view. D, lateral view of eye lobe (uncoated) E, lateral view of muscle insertion scars on glabella (uncoated). F, dorsal view of brim perforations. G–J Stoloharpes aff. capricornus, upper Eifelian, Hamar Laghdad (Fig. 2B, Map 11, site 14), dorsal exoskeleton of cephalon only, NHMUK It 29366. G, dorsal, H, lateral and I, anterior view. J, dorsal view of brim perforations. K–O Stoloharpes capricornus n. sp., El Otfal Formation, upper Eifelian, TaouzEast (Fig.2B, Map 15, site 2), dorsal exoskeleton thorax enrolled, paratype, NHMUK It 29365. K, dorsal, L, lateral and M, posterior view. N, dorsal view of brim perforations and O, lateral view of eye lobe. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFBCE344FF78FA3AFED784B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232691/files/figure.png	https://doi.org/10.5281/zenodo.11232691	PLATE 51. A–L Helioharpes retiarius n. sp., El Otfal Formation, upper Eifelian, Ottara. A–G dorsal exoskeleton of cephalon only, holotype, NHMUK It 29368 (Fig. 2B, Map 14, site 2). A, dorsal, B, anterior and C, lateral view. Dorsal view of brim perforations: E, F and G lateral view of eye lobe. H–L dorsal exoskeleton of cephalon only, paratype, NHMUK it 29369 (Fig. 2B, Map 14, site 3). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 51. A–L Helioharpes retiarius n. sp., El Otfal Formation, upper Eifelian, Ottara. A–G dorsal exoskeleton of cephalon only, holotype, NHMUK It 29368 (Fig. 2B, Map 14, site 2). A, dorsal, B, anterior and C, lateral view. Dorsal view of brim perforations: E, F and G lateral view of eye lobe. H–L dorsal exoskeleton of cephalon only, paratype, NHMUK it 29369 (Fig. 2B, Map 14, site 3). I, dorsal, J, lateral and K, anterior view. L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
03FD8227FFBCE344FF78FA3AFED784B8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/11232693/files/figure.png	https://doi.org/10.5281/zenodo.11232693	PLATE 52. A–L Helioharpes retiarius n. sp., El Otfal Formation, upper Eifelian, Ottara. A–F dorsal exoskeleton of a partial cephalon only, paratype, NHMUK It 29367 (Fig. 2B, Map 14, site 1). A, dorsal, B, anterior and C, lateral view. D, antero-dorsal view of genal roll and brim perforations. Dorsal view of E, brim perforations and F, perforations on genal area. G–L dorsal exoskeleton of cephalon only, paratype, NHMUK It 29370 (Fig. 2B, Map 14, site 11). G, dorsal, H, anterior and I, lateral view. J and K, lateral view of eye lobes and L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	PLATE 52. A–L Helioharpes retiarius n. sp., El Otfal Formation, upper Eifelian, Ottara. A–F dorsal exoskeleton of a partial cephalon only, paratype, NHMUK It 29367 (Fig. 2B, Map 14, site 1). A, dorsal, B, anterior and C, lateral view. D, antero-dorsal view of genal roll and brim perforations. Dorsal view of E, brim perforations and F, perforations on genal area. G–L dorsal exoskeleton of cephalon only, paratype, NHMUK It 29370 (Fig. 2B, Map 14, site 11). G, dorsal, H, anterior and I, lateral view. J and K, lateral view of eye lobes and L, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated.	2024-05-15	Johnson, Robert G.		Zenodo	biologists	Johnson, Robert G.			
