taxonID	type	description	language	source
03FD87C8FFDAC3226055CDC6FBA4F968.taxon	diagnosis	Diagnosis. Deutonymph relatively small. Idiosomal setae smooth, setae r 4, Z 3, and Z 5 distinctly longer than remaining dorsal setae. Chelicerae with 5 teeth on fixed digit, 3 teeth on movable digit. Gnathotectum with smooth anterior free margin. Tarsus I with a distinct acrotarsus.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFDAC3226055CDC6FBA4F968.taxon	description	Description. Idiosomal length 639 (51) and width 421 (33) (N = 6). Gnathosoma. Chelicerae well developed, fixed digit with 5 teeth, movable digit with 3 distinct teeth (Fig. 1 B). Dorsal seta stout and smooth, dorsal lyrifissure present, poorly developed, lateral lyrifissure i not observed (only two specimens had the chelicerae in a lateral position, and in both the relevant part of the chelicerae was unclear); pilus dentilis small and thin. Gnathotectum with smooth anterior margin extending to a prominent, median prong with a semi blunt point (Fig. 1 A). Hypostomal setae smooth; setae hyp 1 and hyp 3 much longer than hyp 2. Subcapitular setae (cs) with a few small barbs. Corniculi smooth, hornlike; inserted slightly dorsal. Deutosternum with 7 – 9 files of teeth; teeth vary in sharpness, relative size, and number per row (from 4 – 11), and are irregularly distributed within rows. Palps with five distinct segments, tibia and tarsus not fused. Setation standard for free living Dermanyssina (Evans, 1963 b), with 2 setae on the trochanter, 5 on the femur, 6 on the genu, and 14 on the tibia; sensilla on tarsus not counted. Palp pretarsus with three tines, one of which very small. Idiosoma. Dorsum almost completely covered by two shields of subequal size; shields covered with a reticulate pattern (Fig. 3 A). Off shield cuticle weakly striate (see Fig. 5). All dorsal setae smooth, setiform. Setae r 4, Z 3, and Z 5 relatively long, median and marginal opisthosomal setae shorter; setae j 1 and z 1 short; setae Z 3 about three times the length of setae j 6; setae Z 5 distinctly longer than Z 3. One pair of lateral propodosomal setae, designated px, associated with the peritrematal shield. Dorsal setal complement nearly complete for jJ, zZ, and sS series (only s 1 lacking). Most R and Rv setae (designations tentative) inserted in soft cuticle. Complement of dorsal lyrifissures and glands similar to that listed for Zerconidae (Johnston & Moraza, 1991), but lacking lyrifissures idz 4, idS 2, and, possibly idR 3, and opisthosomal glands gdJ 2, gdJ 3, gdJ 4, and gdZ 1. The pairs of lyrifissures and glands designated in Zerconidae as ids 4 and gds 5 (Johnston & Moraza, 1991) are most probably homologous to the pair designated as ids 5 and gds 6 in Parasitidae (Al­Atawi et al., 2002). Glands gds 6 relatively large. Venter (Fig. 3 B). Tritosternum with well developed laciniae. Large sternal shield with faint reticulation, including insertions of setae st 1 – st 4. Setae st 5 inserted in soft cuticle, flanking posterior end of sternal shield. Relative size of sternal setae decreases gradually from st 1 to st 5. Endopodal shields distinct, exopodal shields weakly developed, not fused to peritrematal shields. Stigmata at level of middle of coxal acetabula IV; peritremes extending anteriorly until the middle of coxal acetabula I. Peritrematal shields not very extensive. Metapodal shields small, without setae, glands or lyrifissures. Anal shield nearly round; para­anal setae inserted at the level of center of the anal valves; unpaired postanal seta slightly longer than para­anal setae. Cribrum transverse, linear, poorly developed. Opisthogastral setae relatively short, only Sv 5 slightly longer. All ventral setae smooth and setiform. All 3 sternal lyrifissures well developed, peritrematal shield with lyrifissures ip 1 and ip 2, and gland gp. Opisthogastral glands gv 2 close to coxae IV, multiple; glands gv 3 at margin of anal shield; sternal glands gv 1 not observed. Opisthogastral lyrifissures as in Zerconidae (Johnston & Moraza, 1991), but lyrifissure pair ivo 3 may be absent (the identity of the lyrifissure between setae Rv 3 and Rv 4 is unclear; it may be homologous to ivo 3 or idR 3). Legs. Tarsus I with distinctly delineated acrotarsus, and partial basitarsal fissure. Legs II thickened. Leg setation: Coxae: 2 – 2 – 2 – 1; Trochanters: 6 – 5 – 5 – 5; Femora I: 2 3 / 1 2 / 3 2; II: 2 3 / 1 2 / 2 1; III – IV: 1 2 / 1 2 / 0 0. Genua I: 2 3 / 2 3 / 1 2; II: 2 3 / 1 2 / 1 2; III: 2 2 / 1 2 / 1 1; IV: 2 2 / 1 3 / 1 1. Tibiae I: 2 3 / 2 3 / 2 2; II: 2 2 / 1 2 / 1 2; III: 2 1 / 1 2 / 1 1; IV: 2 1 / 1 3 / 1 2. Tarsi II – IV 3 3 / 2 1 / 1 3 / 2 3 / 3; setae ad 1, pd 1 small. thin and somewhat ribbon like, occasionally not observed. Ventral setae of tarsus and tibia II smooth, setiform, only slightly thicker than dorsal setae. All legs with a well developed pretarsus including two claws.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFDAC3226055CDC6FBA4F968.taxon	materials_examined	Material examined (all deutonymphs). CANADA: British Columbia, Hope, 11.5 mi SW of, 49 ° 23 ' N, 121 ° 26 ' W, ex Neurotrichus gibbsii (Talpidae) (GSJ 1746), 8 Jul 1973, coll. Jones, G. S., OSAL 013769 (Holotype). Paratypes: Same locality and date, ex Neurotrichus gibbsii (GSJ 1741), coll. Jones, G. S., OSAL 013768; Jct BC 16 & " Nantleg " Rd, ex Microtus sp. (Muridae) (GSJ 1244), 2 Aug 1972, coll. Jones, G. S., OSAL 013766. U. S. A: Colorado, Grand Co., Jones Pass, 1 mi S of, 39 ° 46 ' 25 " N, 105 ° 53 ' 19 " W, 3475 m, ex Microtus longicaudus (DMA 2021), coll. Armstrong, D. M., OSAL 013795; Oregon, Benton Co., Mary's Peak, 4 mi from jct. sr 344 & forest rd 1244, 44 ° 30 ' 16 " N, 123 ° 33 ' 00 " W, ex Sorex bendirii (Soricidae) (GSJ 1035), 14 Jul 1972, coll. Jones, G. S., OSAL 013763; Mary's Peak, 32 km SW Philomath, ex Sorex pacificus (GSJ 1047), 14 Jul 1972, coll. Jones, G. S., OSAL 013764; Mary's Peak, station 1, 32 km SW Philomath, ex Arborimus albipes (Muridae) (GSJ 1645), 28 Jun 1973, coll. Jones, G. S., OSAL 013767; Benton Co., 44 ° 30 ' N, 123 ° 25 ' W, ex Sorex pacificus (CM 1034), 24 Mar 1968, coll. Maser, C. O., OSAL 013772 – 73; ex Sorex trowbridgii (CM 1042), 21 Mar 1968, coll. Maser, C. O., OSAL 013771; Clackamas Co., Estacada Quadrangle, 4517 ' 23 " N, 122 ° 19 ' 57 " W, ex Mustela erminea (Mustelidae), 31 Jul 1969, coll. Maser, C. O., OSAL 013787 – 90; Coos Co., Bandon, SE of, 43 ° 07 ' 09 " N, 124 ° 24 ' 26 " W, ex Sorex pacificus (CM 2962), 27 Dec 1971, coll. Maser, C. O., OSAL 013780; Bandon, 4 mi NNE of, ex Peromyscus maniculatus (Muridae) (CM 2511), 25 Feb 1971, coll. Maser, C. O., OSAL 013776; Bandon, E of, ex Peromyscus maniculatus (CM 3267), 16 Jul 1972, coll. Maser, C. O., OSAL 013804; Coos Co., 43 ° 29 ' N, 124 ° 00 ' W, ex Sorex trowbridgii (CM 2495), 21 Jan 1971, coll. Maser, C. O., OSAL 013759; ex Arborimus albipes (RML 59592), 3 Jun 1970, coll. Maser, C. O., OSAL 013756; Curry Co., Brookings, 16 km E of, ex Scapanus orarius (Talpidae) (CM 3144), 21 Apr 1970, coll. Maser, C. O., OSAL 013803; Curry Co., 42 ° 30 ' N, 124 ° 13 ' W, CM 2462, ex Neurotrichus gibbsii (CM 2462), 4 Dec 1970, coll. Maser, C. O., OSAL 013785 – 86; Douglas Co., Reedsport, SW of, 43 ° 42 ' 09 " N, 124 ° 05 ' 44 " W, 3 m, ex Arborimus albipes (RML 62993), 22 Mar 1972, coll. Maser, C. O., OSAL 013760; Douglas Co., 43 ° 20 ' N, 123 ° 10 ' W, ex Sorex pacificus (CM 2091), 5 Feb 1970, coll. Maser, C. O., OSAL 013775; Lane Co., H. J. Andrews Exp. Forest, 43 ° 55 ' N, 122 ° 50 ' W, ex Neurotrichus gibbsii (CM 6115 A), 14 Aug 1972, coll. Maser, C. O., OSAL 013793 – 94; ex Sorex pacificus, (CM 3387), 12 Aug 1972, coll. Maser, C. O., OSAL 013784; ex Peromyscus maniculatus, 5 Apr 1972, coll. RAN, OSAL 013791; Lincoln Co., Cascade Head Exp. Forest, 45 ° 02 ' 03 " N, 123 ° 55 ' 20 " W, ex Sorex bendirii (CM 3305), 13 Aug 1972, coll. Maser, C. O., OSAL 013781; ex Sorex bendirii (CM 3310), 15 Aug 1972, coll. Maser, C. O., OSAL 013782; ex Sorex pacificus (CM 2600), 27 Apr 1971, coll. Maser, C. O., OSAL 013778; ex Arborimus albipes (RML 59829), 4 Aug 1972, coll. Maser, C. O., OSAL 013757; ex Arborimus albipes (RML 59835), 7 Aug 1972, coll. Maser, C. O., OSAL 013758; ex Arborimus albipes (RML 63008), 12 Aug 1972, coll. Maser, C. O., OSAL 013761; ex Arborimus albipes (RML 63010), 19 Aug 1972, coll. Maser, C. O., OSAL 013762; ex Microtus oregoni (CM 2573), 14 Apr 1971, coll. Maser, C. O., OSAL 013777; ex Zapus trinotatus (Dipodidae) (CM 2743), 8 Aug 1971, coll. Maser, C. O., OSAL 013779; Lincoln Co., 44 ° 40 ' N, 123 ° 50 ' W, ex Sorex bendirii (CM 3329), 18 Aug 1972, coll. Maser, C. O., OSAL 013783; ex Sorex pacificus, 16 Aug 1972, coll. Maser, C. O., OSAL 013792; Linn Co., 44 ° 30 ' N, 122 ° 35 ' E, ex Peromyscus maniculatus (CM 1949), 2 Nov 1969, coll. Maser, C. O., OSAL 013774; Washington, Clallam Co., Olympic Nat. Pk., forest rd. 304, 48 ° 16 ' 00 " N, 124 ° 40 ' 30 " W, ex Peromyscus sp. (GSJ 1090), 18 Jul 1972, coll. Jones, G. S., OSAL 013765; Snohomish Co., Gold Bar, 48 ° 03 ' N, 121 ° 43 ' W, ex Neurotrichus gibbsii (JOW 6953), 24 Jun 1971, coll. Whitaker, J. O., Jr., OSAL 013770. Deposition of types. Holotype deutonymph, OSAL 013769, in OSAL. Paratypes in collection of OSAL, CNC, JOW, NMNH, UMMZ, and ZIN.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFDAC3226055CDC6FBA4F968.taxon	etymology	Etymology. The species name refers to the western North American distribution of this species.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFDAC3226055CDC6FBA4F968.taxon	discussion	Remarks. Euryparasitus tori, E. longicheta, E. goncharovi, and E. occidentalis differ from all other described Euryparasitus deutonymphs by the complete absence of barbed setae on the idiosoma, the presence of a gnathotectum with a completely smooth margin, the presence of an acrotarsus on legs I, and the presence of 5 teeth on the movable digit, and 3 teeth on the fixed digit of the chelicerae. However, E. longicheta has much longer setae than all others (Bondartchuk & Buyakova, 1978), and E. goncharovi has a more or less pentagonal shaped anal shield with dorsal setae that are all subequal in length (Bondartchuk & Buyakova, 1976). Euryparasitus occidentalis and E. tori share elongate setae Z 5 and an almost round anal shield (especially distinct in E. tori). However, setae Z 3 in E. tori are similar in length to setae Z 2 and S 3 (Davydova, 1970), while they are distinctly longer in E. occidentalis. Setae Z 5 in E. occidentalis are almost half as long as the opisthosomal shield, in E. tori those setae are at most 1 / 3 the length of that shield. The presence of an acrotarsus on legs I is an unusual character for Dermanyssina, and is more commonly associated with deutonymphs of Parasitidae. Continued assignment of the above group of species to Euryparasitus, rather than some genus of Parasitidae is based in part on the morphology of the cribrum (a thin row lining most of the posterior margin of the anal shield in all Euryparasitus, a triangular field in parasitid deutonymphs), and the multiplication of ventral glands gv 2 (single in Parasitidae). There is also evidence from the adults. While only the deutonymphs of E. occidentalis, E. longicheta, and E. tori are known, E. goncharovi is known from both nymphal instars and the adults. Its female has the sternal and metasternal shields fused, with seta st 4 on the sternal shield, and a subrectangular genital shield, characteristics of, respectively, Rhodacaroidea, and Euryparasitidae sensu Anthony (1980). The male shows a free spermatotodactyl (fused in Parasitidae) with the massive elongate shape typical for Euryparasitus and Cyrtolaelaps. Based on this evidence we continue to assign these species to Euryparasitus. This assignment implies that the occurence of an acrotarsus on legs I of the deutonymph is homoplasious, with separate origin or retention in Parasitidae and some Euryparasitus. Lee (1970) noted a ventral setal complement in adult E. emarginatus of 3 Jv, 3 Zv, 3 Sv, and 3 Rv setae. E. occidentalis deutonymphs carry one less Rv seta.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD7C3206055C83CFE3BFBC0.taxon	description	Description. Idiosomal length 655 (36) and width 414 (15) (N = 6). Gnathosoma. Cheliceral dentition, gnathotectal shape, and palp segmentation and setation as in E. occidentalis. Hypostomal setae relatively longer, lengths of setae hyp 2 far surpassing the insertion of setae cs (Fig. 2 B). Idiosoma. Dorsal shields subequal in size, covering most of dorsum (Fig. 3 A). Podonotal shield length 325 (16), width 355 (16), opisthonotal shield length 301 (19), width 356 (40). All setae smooth; most setae long, although setae j 1, z 1, and some marginal setae are short; setae r 4, Z 3, and Z 5 longer than most other setae, but less obviously so than in E. occidentalis. Distribution of setae, lyrifissures, and glands as in E. occidentalis. Venter. Sternal shield large. Peritremes extending beyond the anterior edge of coxal acetabula I. Peritrematal, metapodal, and anal shields as in E. occidentalis. Setae slightly longer than in E. occidentalis, but much less so than on the dorsum. Lyrifissures ivp, situated posterior to the anus not observed. This may be an artifact of inadequate material. Legs. Leg morphology and setation pattern as in E. occidentalis.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD7C3206055C83CFE3BFBC0.taxon	materials_examined	Material examined (all deutonymphs). U. S. A.: Oregon, Union Co., Starkey Exp. Forest, 45 ° 14 ' N, 118 ° 33 ' W, ex Sorex vagrans (CM 7004), 19 Oct 1976, coll. Maser, C. O., OSAL 013796 – 800; ex Clethrionomys gapper i (Muridae) (CM 6994) 26 Oct 1976, coll. Maser, C. O., OSAL 013801. Deposition of specimens. Specimens deposited in JOW and OSAL.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD7C3206055C83CFE3BFBC0.taxon	discussion	Remarks. Euryparasitus longicheta was described from Western Siberia associated with Eutamias sibericus (Sciuridae) and Cricetulus barabensis (Muridae) (Bondartchuk & Buyakova, 1978).	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	diagnosis	Diagnosis. Deutonymph of medium size. Some idiosomal and all hypostomal setae barbed. Setae S 3 absent from opisthosomal shield. Chelicerae with 4 teeth on the fixed, 3 teeth on the movable digit. Gnathotectum with a serrate anterior, free, margin. Acrotarsus on legs I absent.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	description	Description. Idiosomal length 797 (57), width 536 (51) (N = 6). Gnathosoma. Chelicerae with 4 large teeth on the fixed digit, and 3 on the movable digit (Fig. 1 D). Lyrifissures id and i present, well developed; dorsal seta vestigial or absent; pilus dentilis much larger than in E. occidentalis. Gnathotectum developed into a single median point; with serrate anterior margin (Fig. 1 C). All hypostomal setae barbed, tip of setae hyp 2 surpassing the insertion points of setae cs; setae hyp 2 subequal in length to setae hyp 1 (Fig. 2 D). Deutosternum with 14 – 16 rows of irregular shaped and arranged teeth. Palps as in E. occidentalis, but third tine of palpal pretarsus distinct. Idiosoma. Podonotal shield distinctly larger than opisthonotal shield (Fig. 5 A). Most setae inserted on dorsal shields smooth, but setae s 2, r 2, r 4, Z 5, and all marginal opisthosomal setae barbed. Setal complement on shields largely as in E. occidentalis, but opisthonotal seta S 3 absent. The occasional presence of an undesignated seta off the shield, near the usual position of seta S 3, suggests that S 3 might still be present, but inserted off the shield (see Fig. 5 A). Number of marginal setae in soft cuticle increased by presence of five additional setae, tentatively designated as setae R 3, R 4, R 5, Rv 2, and Rv 5. Shield setae j 2, r 4, Z 3 and Z 5 distinctly longer than remaining dorsal setae; setae Z 3 and Z 5 subequal in length, almost twice as long as the other opisthonotal setae; setae j 1 and z 1 similar in length to other podonotal shield setae; marginal opisthosomal setae generally shorter. Gland pattern and lyrifissure complement as in E. occidentalis. Venter (Fig. 5 B). Sternal shield large. Setae st 4 inserted off shield in soft cuticle in one specimen, on sternal shield in all other specimens examined. Peritremes extending anteriorly to the middle of coxal acetabula I. Peritrematal, metapodal, and anal shields as in E. occidentalis, although anal shield not quite as rounded. Sternal setae st 1 and st 2, and opisthogastral seta Rv 5 strongly barbed, remaining sternal and opisthogastral setae with weak barbs or smooth. Opisthogastral setal complement richer than in E. occidentalis by the addition of setae Zv 4 and Sv 2. Para­anal setae positioned slightly posterior to the anus. Lyrifissure and gland pattern generally as in E. occidentalis; cluster of gv 2 glands posterior to each coxa IV more prominent; lyrifissure ivp, situated posterior to the anus in E. occidentalis, not observed (may be an artifact). Legs. Leg setation pattern similar to that in E. occidentalis. Acrotarsus I absent or poorly developed. Legs II more conspicuously thickened than in E. occidentalis. Tarsal setae av 2, pv 2, and mv, and tibial setae pv 1 of legs II strongly thickened, spinelike, with distinct barbs. Tibial seta av 1 barbed, but less thick and spinelike. Lateral and dorsal setae may be barbed, but remain setiform.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	materials_examined	Material examined (all deutonymphs). U. S. A.: Oregon, Malheur Co., Whitehorse Ranch, 10 km SE of, 43 ° 15 ' N 117 ° 40 ' W, ex Onychomys leucogaster (CM 7223), 19 Aug 1976, coll. Maser, C. O., OSAL 013968 (Holotype). Paratypes: Same collection data, OSAL 013956 – 67; Malheur Co., 10 km SSE Vale, 43 ° 15 ' N, 117 ° 40 ' W, ex Dipodomys ordii (Dipodidae) (CM 6478), coll. Maser, C. O., OSAL 013969; ex Spermophilus townsendii (Sciuridae) (CM 6471), 20 Mar 1975, coll. Maser, C. O., OSAL 013971; ex Spermophilus townsendii (CM 6496), 21 Mar 1975, coll. Maser, C. O., OSAL 013970; Malheur Co., 43 ° 15 ' N, 117 ° 40 ' W, ex Lemmiscus curtatus (Muridae) (CM 7233), 27 Aug 1976, coll. Maser, C. O., OSAL 013972; ex Lemmiscus curtatus (CM 7239), 29 Aug 1976, coll. Maser, C. O., OSAL 013973. CANADA: Alberta, Bow Island, 49 ° 52 ' N, 111 ° 22 ' W, ex Onychomys sp. (Muridae) (GSJ 1307), 11 Aug 1972, coll. Jones, G. S., OSAL 013974 – 75.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	description	Deposition of types. Holotype deutonymph, OSAL 013968, in OSAL. Paratypes in OSAL, JOW, ZIN.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	etymology	Etymology. This species is named in honor of Chris Maser, who collected many of the mites in this study.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD5C3266055CB54FBECFA20.taxon	discussion	Remarks. Relative to the ventral setal complement of adult E. emarginatus (Lee, 1970) this deutonymph features one additional Zv and one Sv seta. The additional Rv setae are considered less significant as some of these lateral setae may appear dorsal or ventral on slide mounted specimens. They may be homologous with any of the large number of marginal setae figured for E. emarginatus in dorsolateral position.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD3C33A6055C906FE50F9F0.taxon	diagnosis	Diagnosis. Very large species Some idiosomal and all hypostomal setae barbed. Chelicerae with 6 teeth on the fixed, 3 teeth on the movable digit. Gnathotectum with serrate anterior, free, margin. Acrotarsus on legs I absent.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD3C33A6055C906FE50F9F0.taxon	description	Description. Idiosomal length 1183 (132), width 773 (91) (N = 6). Gnathosoma. Chelicerae with 6 teeth on the fixed digit, 2 of which are small, and 3 large teeth on the movable digit (Fig. 1 F). Lyrifissures id and i present, well developed; dorsal seta vestigial or absent; pilus dentilis as in E. maseri. Gnathotectum developed into a single median point and with serrate anterior margin (Fig. 1 E); serrations less robust than in E. maseri. Subcapitulum (Fig. 2 D), including relative size and shape of the hypostomal setae, quite similar to that of E. maseri. Deutosternum with 13 – 15 rows of irregular teeth. Palps as in E. maseri. Idiosoma. Podonotal shield slightly larger than opisthonotal shield (Fig. 6 A). Most setae inserted on dorsal shields smooth, but setae r 4, and most marginal opisthosomal setae barbed. Setal complement on shields as in E. occidentalis. Number of marginal setae in soft cuticle as in E. maseri. Shield setae r 4, Z 3, and Z 5 distinctly longer than remaining dorsal setae; setae j 2 only slightly longer than setae z 2 and z 3; setae j 1 and z 1 similar in length to most other podonotal setae; setae Z 5 distinctly longer than setae Z 3; both at least twice as long as all other opisthonotal setae; marginal setae not shorter than opisthonotal ones. Gland pattern and lyrifissure complement as in E. occidentalis. Venter (Fig. 6 B). Sternal shield large. Stigmatal position, peritremes and peritrematal shields as in E. occidentalis; metapodal shields relatively large; anal shield distinctly wider than long (see Table 1). All sternal and most marginal opisthogastral setae barbed, remaining opisthogastral setae with weak barbs or smooth; opisthogastral setal complement identical to that in E. maseri; para­anal setae positioned slightly posterior to the anus. Lyrifissure and gland pattern as in E. maseri. Legs. Leg setation pattern similar to that in E. occidentalis. Leg morphology more similar to that in E. maseri: acrotarsus I absent and ventral setae of tarsus and tibia II thickened and barbed, although less obviously so.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD3C33A6055C906FE50F9F0.taxon	materials_examined	Material examined (unless indicated otherwise all specimens are deutonymphs). CANADA: British Columbia, Hope, 11.5 mi SW of, 49 ° 23 ' N, 121 ° 26 ' W, ex Zapus trinotatus (GSJ 1704), 6 Jul 1973, coll. Jones, G. S., OSAL 014022; Cottonwood River & BC 94 Jct, 59 ° 06 ' 03 " N, 129 ° 46 ' 46 " W, ex Zapus princeps (GSJ 1207), 30 Jul 1972, coll. Jones, G. S., OSAL 013984; Fraser River valley, 25 km W jct. BC 1 & BC 3 on BC 1, 53 ° 29 ' N, 122 ° 43 ' W, ex Microtus sp. (GSJ 1166), 23 Jul 1972, coll. Jones, G. S., OSAL 014023; New Brunswick, Mt. Carleton Park, 47 ° 23 ' 00 " N, 066 ° 53 ' 00 " W, ex Condylura cristata (Talpidae) (TF 102), 16 Jun – 20 Jul 1980, coll. French, T. W., OSAL 013992; ex Blarina brevicauda (Soricidae) (TF 644), coll. French, T. W., OSAL 013993; ex Blarina brevicauda (TF 675), coll. French, T. W., OSAL 013994; ex Sorex cinereus (TF 764), coll. French, T. W., OSAL 013999; ex Sorex fumeus (TF 118), coll. French, T. W., OSAL 013989 – 90; ex Sorex fumeus (TF 479), coll. French, T. W., OSAL 013996; ex Sorex gaspensis (TF 372), coll. French, T. W., OSAL 013991; ex Microtus chrotorrhinus (TF 601), coll. French, T. W., OSAL 014000 – 01; ex Microtus pennsylvanicus (TF 107), coll. French, T. W., OSAL 014002; ex Napaeozapus insignis (Dipodidae) (TF 765), coll. French, T. W., OSAL 013997 – 98; up the Tabusintac River, ex Blarina brevicauda (TF 761), 15 Jun 29 Jul 1980, coll. French, T. W., OSAL 013995. U. S. A.: Colorado, Routt Co., 40 ° 31 ' N, 106 ° 59 ' W, ex Tamiasciurus hudsonicus (Sciuridae) (CM 2360), 7 Sep 1970, coll. Maser, C. O., OSAL 014006; ex Tamias quadrivittatus (Sciuridae) (CM 1736), 30 Jul 1969, coll. Maser, C. O., OSAL 014008; Colorado, ex Clethrionomys gapperi (DMA 1972), coll. Armstrong, D. M., OSAL 014020; Illinois, Clark Co., Marshall, 10 km NE of, 39 ° 23 ' 29 " N, 087 ° 41 ' 37 " W, ex Microtus pinetorum (DDP 76), 20 Nov 1967, coll. Pascal, D. D., Jr., OSAL 014018; ex Microtus pinetorum (DDP 500), 5 Dec 1968, coll. Pascal, D. D., Jr., OSAL 014019; Indiana, Jasper Co., 41 ° 02 ' 29 " N, 087 ° 09 ' 05 " W, ex Sorex cinereus (REM 6323), 20 Mar 1970, coll. Whitaker, J. O., Jr., OSAL 014017; Pulaski Co., 41 ° 03 ' 05 " N, 086 ° 36 ' 11 " W, ex Tamias striatus (JOW 9225), 30 Mar 1975, coll. Whitaker, J. O., Jr., OSAL 013982; Vigo Co., Terre Haute, 39 ° 28 ' 00 " N, 087 ° 24 ' 50 " W, 153 m, ex Scalopus aquaticus (Talpidae) (JOW 7587), 17 Oct 1971, coll. Whitaker, J. O., Jr., OSAL 014015; New York, Delaware Co., Dry Brook Ridge, Catskills, 42 ° 06 ' 08 " N, 074 ° 35 ' 51 " W, ex Microtus chrotorrhinus (TF 1197), 23 Sep 1981, coll. French, T. W., OSAL 014025; ex Microtus chrotorrhinus (TF 1564), 8 Oct 1981, coll. French, T. W., OSAL 014024; Otsego Co., Milford, 42 ° 35 ' 26 " N, 074 ° 56 ' 44 " W, ex Tamias striatus (JOW 5926), 24 Jun 1970, coll. Whitaker, J. O., Jr., OSAL 013983; Schuyler Co., Arnot Forest, 42 ° 25 ' N, 076 ° 28 ' W, ex Blarina brevicauda, 14 Sep 1974, coll. Northam, M., UMMZ; Tompkins Co., jct. Warren Rd & Rte 13, 42 ° 27 ' N, 076 ° 28 ' W, ex Microtus pennsylvanicus, 6 Mar 1975, coll. OConnor, B. M., UMMZ; jct. Rte 13 & Rte 366, ex Blarina brevicauda, 24 Feb 1976, coll. OConnor, B. M., UMMZ (Female); North Carolina, Clay Co., Fire Creek, Bristol cabin, 35 ° 04 ' 05 " N, 083 ° 52 ' 06 " W, ex Ochrotomys nuttalli (Muridae) (JOW 8266), 22 Apr 1973, coll. Whitaker, J. O., Jr., OSAL 014016; Oregon, Benton Co., 4429 ' N, 123 ° 25 ' W, ex Sorex bendirii (CM 1046), 25 Mar 1968, coll. Maser, C. O., OSAL 014012; Clackamas Co., Estacada Quadrangle, 45 ° 17 ' 23 " N, 122 ° 19 ' 57 " W, ex Mustela erminea, 31 Jul 1969, coll. Maser, C. O., OSAL 013980; Coos Co., Bandon, SE of, 43 ° 07 ' 09 " N, 124 ° 24 ' 26 " W, ex Sorex pacificus (CM 2962), 27 Dec 1971, coll. Maser, C. O., OSAL 014009; Coos Co., 43 ° 29 ' N, 124 ° 00 ' W, ex Sorex vagrans (CM 1795), 26 Aug 1969, coll. Maser, C. O., OSAL 014013; Harney Co., 43 ° 10 ' N, 119 ° 00 ' W, ex Lemmiscus curtatus (CM 6981), 3 Aug 1975, coll. Maser, C. O., OSAL 013979; Lane Co., H. J. Andrews Exp. Forest, 43 ° 55 ' N, 122 ° 50 ' W, ex Sorex trowbridgii (CM 3359), 31 Aug 1972, coll. Maser, C. O., OSAL 013977; ex Tamias townsendii (CM 5606), 28 Sep 1972, coll. Maser, C. O., OSAL 014007; Lane Co., 43 ° 55 ' N, 122 ° 50 ' W, ex Sorex bendirii (CM 6100), 15 Aug 1972, coll. Maser, C. O., OSAL 014011; ex Sorex trowbridgii (CM 1104), 8 May 1968, coll. Maser, C. O., OSAL 013976; Lincoln Co., Cascade Head Exp. Forest, 45 ° 02 ' 03 " N, 123 ° 55 ' 20 " W, ex Microtus oregoni (CM 2570), 13 Apr 1971, coll. Maser, C. O., OSAL 014004; ex Microtus longicaudus (CM 2801), 24 Aug 1971, coll. Maser, C. O., OSAL 014005; ex Arborimus albipes (RML 63010), 19 Aug 1972, coll. Maser, C. O., OSAL 014010; Linn Co., 44 ° 29 ' N, 122 ° 35 ' E, ex Sorex trowbridgii (CM 4134), 26 Sep 1972, coll. Maser, C. O., OSAL 013978; Washington, Clallam Co., Olympic Nat. Pk, E border Nat. For. nr rd 349, 48 ° 12 ' N, 124 ° 23 ' W, ex Sorex trowbridgii (GSJ 1102), 19 Jul 1972, coll. Jones, G. S., OSAL 013988; Clallam Co., Olympic Nat. Pk., 48 ° 16 ' 00 " N, 124 ° 40 ' 30 " W, ex Phenacomys intermedius (Muridae) (MLJ 594349), 15 Sep 1976, coll., OSAL 013985; Pierce Co., Tacoma, 16 km SE of, between McChord & Spanaway, 47 ° 06 ' 15 " N, 122 ° 26 ' 00 " W, ex Peromyscus maniculatus (PGF 979), 18 Apr 1972, coll. Fish, P. G., OSAL 014021; Snohomish Co., Gold Bar, 48 ° 03 ' N, 121 ° 43 ' W, ex Sorex bendirii (JOW 6945), 24 Jun 1971, coll. Whitaker, J. O., Jr., OSAL 013987; ex Neurotrichus gibbsii (JOW 6953), coll. Whitaker, J. O., Jr., OSAL 013986; Wyoming, Carbon Co., Medicine Bow Natl. For., 41 ° 15 ' N, 106 ° 15 ' W, ex Zapus sp. (GSJ 2136), 12 Aug 1973, coll. Jones, G. S., OSAL 013981. No locality information, ex Scalopus aquaticus (71777), 17 Nov 1971, coll. Forsyth, D. J., OSAL 014026; ex Scalopus aquaticus (71772), 14 Nov 1971, coll. Forsyth, D. J., OSAL 014027; ex Scalopus aquaticus (DJF 82), coll. Forsyth, D. J., OSAL 014028; ex Scalopus aquaticus (DJF 83), coll. Forsyth, D. J., OSAL 014029. Deposition of specimens. Specimens deposited in collection of OSAL, CNC, JOW, NMNH, UMMZ, and ZIN.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
03FD87C8FFD3C33A6055C906FE50F9F0.taxon	discussion	Remarks. Euryparasitus calcarator was described and redescribed for the adults only (Banks, 1910; Hennessey & Farrier, 1988). The species is quite similar to E. emarginatus, but differs by the presence of 3, rather than 4, teeth on the movable digit. The association of this name with the series of deutonymphs described below is tentative. It is based on similarity in the structure of the chelicerae, and the fact that so far only one type of deutonymph of a very large Euryparasitus has been collected in eastern North America, despite extensive collecting. Within North America, this deutonymph is most similar to that of E. maseri n. sp. The two species share a serrate anterior margin of the gnathotectum, presence of a large complement of opisthosomal setae, position of the para­anal setae posterior to the anus, absence of an acrotarsus on tarsus I, and the morphology of the ventral setae of tarsus and tibia II. The two species differ in absolute size, presence / absence of setae S 3, the relatively larger size of legs II and the smaller size of the opisthonotal shield in E. maseri, and the dentition of the fixed digit of the chelicerae. This species also shows some clear intraspecific variation. Specimens from western North America (Oregon, Washington) are larger than those from eastern North America (New Brunswick, New York, Indiana), e. g. podonotal shield length 747 (63) vs. 576 (12). However, these size differences did not correlate with qualitative characters, and may be allometric changes representing an East – West cline. The available collections were insufficient to exclude that possibility, and we therefore elect to retain all specimens in a single species.	en	Hagele, Thomas, Kaufman, Bruce, Whitaker, John O., Jr, Klompen, Hans (2005): The genus Euryparasitus in North America (Mesostigmata: Euryparasitidae). Zootaxa 1036: 1-20, DOI: 10.5281/zenodo.169744
