taxonID	type	description	language	source
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	etymology	Etymology. A combination of the Latin brevis (short), and the Latin loba (lobed), referring to the short clypeal lobe.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	distribution	Distribution. Malaysia: Peninsular Malaysia (Selangor).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	diagnosis	Worker. Diagnosis. In full-face view, head distinctly longer than wide (CI: 82 - 84). Mandible broadly curved throughout. Clypeus with lateral lobe, smoothly incurved between lateral lobe and median extension; apex of median extension convex. In profile, petiole longer than or almost as long as high (LPI: 94 - 102), highest near middle, with subhorizontal dorsal face.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	description	Measurements (n = 5, holotype in parentheses). HL: 1.35 - 1.42 (1.39), HLL: 1.07 - 1.09 (1.09), HLA: 0.12 - 0.16 (0.16), HW: 1.11 - 1.20 (1.14), CML: 0.18 - 0.21 (0.21), CI: 82 - 84 (82), CLI: 12 - 15 (15), SL: 1.32 - 1.38 (1.38), SI: 112 - 121 (121), EL: 0.34 - 0.37 (0.36), OI: 32 - 34 (33), PrL: 0.84 - 0.95 (0.84), PrH: 0.59 - 0.67 (0.67), PrW: 0.87 - 0.97 (0.90), WL: 2.22 - 2.37 (2.27), PeL: 0.85 - 0.88 (0.85), PeH: 0.81 - 0.89 (0.81), PeW: 0.60 - 0.67 (0.62), LPI: 94 - 102 (94), DPI: 71 - 76 (72).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	description	Description. In full-face view, head distinctly longer than wide. Mandible broadly curved throughout, with subapical tooth near apical tooth. Eye strongly prominent, measuring one-third of head lateral margin length. Antennal scape distinctly longer than width of head, surpassing posterior margin of head by one-fourth of its length; antennomere III ca. 1.6 times as long as wide. Clypeus with lateral lobe, smoothly incurved between lateral lobe and median extension; median extension short, with convex apex. In dorsal view, pronotum shorter than or slightly longer than wide. Metanotal groove weakly impressed. In profile, propodeal dorsum almost straight. Petiole in profile longer than or almost as long as high, highest near middle, with subrectangular node; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face subhorizontal, but weakly sloping anteriad; posterior margin almost straight to slightly concave. Head distinctly areolate-rugose. Mandible smooth. Gastral segment I with scalloped depressions, which are irregular in size but generally small; distance between small depressions greater than diameter of depressions; large depressions close to each other; segments II smooth. Body black, slightly tinged with red; clypeus, mandible, antenna, legs and ventral half of petiole dark-red. Apical two or three segments of gaster red-brown. Scalloped depressions on gastral segments I bearing hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	description	Queen. Measurements (n = 2). HL: 1.32 - 1.46, HLL: 1.05 - 1.12, HLA: 0.15 - 0.16, HW: 1.14 - 1.21, CML: 0.21, CI: 83 - 87, CLI: 14 - 16, SL: 1.25 - 1.36, SI: 110 - 112, EL: 0.35 - 0.39, OI: 33 - 35, PrL: 0.88 - 0.98, PrH: 0.66 - 0.76, PrW: 0.94 - 0.98, WL: 2.27 - 2.41, PeL: 0.83 - 0.88, PeH: 0.87 - 0.95, PeW: 0.68 - 0.70, LPI: 105 - 108, DPI: 79 - 82. Mandible shorter than that of worker. Petiole in profile shorter than high, in dorsal view distinctly longer than wide.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
C7B762B3EED3ABC3BB4C42FDFFBACCD9.taxon	description	Male. Unknown.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	etymology	Etymology. From the Latin curvus, meaning curved, referring to the curved mandible.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	distribution	Distribution. Indonesia (West Sumatra).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	diagnosis	Worker. Diagnosis. In full-face view, head distinctly longer than wide (CI: 77 - 78). Mandible distinctly curved throughout. Clypeus without lateral lobe; apex of median extension strongly projecting. In profile, petiole higher than long (LPI: 115 - 117), highest just anterior to posterodorsal angle; anterior and dorsal faces of node forming continuous curve; dorsal face distinctly sloping anteriad. Gastral segment II extensively covered with scalloped depressions. Body covered with understory layer of short, dense and recumbent pubescence.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	description	Measurements (n = 3, holotype in parentheses). HL: 1.71 - 1.75 (1.75), HLL: 1.18 - 1.22 (1.22), HLA: 0.14 - 0.15 (0.15), HW: 1.32 - 1.35 (1.35), CML: 0.29 - 0.31 (0.31), CI: 77 - 78 (77), CLI: 17 - 18 (18), SL: 1.43 - 1.52 (1.52), SI: 108 - 113 (113), EL: 0.34 - 0.36 (0.34), OI: 28 - 31 (28), PrL: 1.03 - 1.06 (1.03), PrH: 0.63 - 0.64 (0.64), PrW: 0.97 - 1.00 (0.97), WL: 2.48 - 2.53 (2.48), PeL: 0.81 - 0.82 (0.82), PeH: 0.93 - 0.95 (0.95), PeW: 0.65 - 0.67 (0.65), LPI: 115 - 117 (116), DPI: 80 - 82 (80).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	description	Description. In full face view, head distinctly longer than wide. Mandible distinctly curved throughout, with subapical tooth near apical tooth. Eye prominent, measuring one-fourth to one-third of head lateral margin length. Antennal scape longer than width of head, surpassing posterior margin of head by less than one-fifth of its length; antennomere III ca. 2.2 times as long as wide. Clypeus without lateral lobe; median extension moderately long, with apex strongly projecting. In dorsal view, pronotum longer than wide. Metanotal groove distinctly impressed. In profile, propodeal dorsum broadly convex. Petiole in profile higher than long, with fan-shaped node, highest just anterior to posterodorsal angle of node; anterior and dorsal faces of node forming continuous curve; dorsal face distinctly sloping anteriad; posterior face almost straight. Head weakly areolate-rugose, longitudinally striate anteriorly; vertex weakly and transversally striate. Mandible longitudinally striate. Prosternum weakly rugose. Gastral segments I-II extensively covered with scalloped depressions that are irregular in size and close to each other; generally depressions on segment I larger than those on segment II. Body black-gray; clypeus, mandible, antenna, legs, and ventral half of petiole dark-red. Body covered with understory layer of short, dense and recumbent pubescence. Scalloped depressions on gastral segments I-II bearing hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	description	Queen. Measurements (n = 1). HL: 1.65, HLL: 1.19, HLA: 0.17, HW: 1.30, CML: 0.32, CI: 79, CLI: 20, SL: 1.41, SI: 109, EL: 0.36, OI: 31, PrL: 1.05, PrH: 0.75, PrW: 0.99, WL: 2.50, PeL: 0.76, PeH: 1.01, PeW: 0.74, LPI: 132, DPI: 97. Petiole in profile distinctly higher than long, in dorsal view almost as long as wide.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
F665A935A37578297EA1FB333C78F203.taxon	description	Male. Unknown.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	etymology	Etymology. Dedicated to Professor Fuminori Ito, collector of the material examined.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	distribution	Distribution. Malaysia: Peninsular Malaysia (Selangor and Pahang).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	diagnosis	Worker. Diagnosis. In full-face view, head slightly longer than wide (CI: 93 - 95). Mandible curved near base, becoming straight apically. Antennal scape long (SI: 99 - 110), surpassing posterior margin of head by two-fifths to one-third of its length. Clypeus with lateral lobe, with blunt angle between lateral lobe and median extension; median extension long, with broadly convex apex. In profile, petiole distinctly higher than long (LPI: 109 - 118), highest just anterior to posterodorsal angle; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face distinctly sloping anteriad.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	description	Measurements (n = 5, holotype in parentheses). HL: 1.66 - 1.73 (1.73), HLL: 1.12 - 1.18 (1.18), HLA: 0.13 - 0.15 (0.14), HW: 1.54 - 1.64 (1.64), CML: 0.29 - 0.36 (0.35), CI: 93 - 95 (95), CLI: 17 - 21 (20), SL: 1.57 - 1.80 (1.80), SI: 99 - 110 (110), EL: 0.31 - 0.35 (0.35), OI: 28 - 30 (30), PrL: 0.99 - 1.10 (1.10), PrH: 0.67 - 0.80 (0.79), PrW: 1.00 - 1.07 (1.05), WL: 2.47 - 2.72 (2.72), PeL: 0.84 - 0.95 (0.95), PeH: 0.99 - 1.04 (1.04), PeW: 0.68 - 0.74 (0.74), LPI: 109 - 118 (109), DPI: 76 - 84 (78).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	description	Description. In full face view, head slightly longer than wide. Mandible curved near base, becoming straight apically; subapical tooth obsolete. Eye prominent, measuring one-fourth to one-third of head lateral margin length. Antennal scape longer than width of head, surpassing posterior margin of head by two-fifths to one-third of its length; antennomere III ca. 2.5 times as long as wide. Clypeus with lateral lobe, with blunt angle between lateral lobe and median extension; median extension long, with broadly convex apex. In dorsal view, pronotum almost as long as wide. Metanotal groove slightly impressed. In profile, propodeal dorsum almost straight. Petiole in profile distinctly higher than long, highest just anterior to posterodorsal angle of node, with fan-shaped node; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face distinctly sloping anteriad; posterior face slightly concave dorsally and slightly convex ventrally. Head strongly areolate-rugose, longitudinally striate near clypeus; vertex weakly and transversally striate. Mandible weakly striate longitudinally. Prosternum smooth and without punctures. Gastral segment I with scalloped depressions that are irregular in size; distance between depressions distinctly greater than diameter of depressions; interspace between depressions with small punctures; segments II smooth. Body black, slightly tinged with red; clypeus, mandible, antenna, legs and ventral half of petiole dark-red. Apical two or three segments of gaster red-brown. Scalloped depressions on gastral segment I bearing hairs; small punctures without hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	description	Queen. Measurements (n = 5). HL: 1.58 - 1.68, HLL: 1.07 - 1.14, HLA: 0.10 - 0.14, HW: 1.43 - 1.59, CML: 0.29 - 0.34, CI: 89 - 94, CLI: 18 - 20, SL: 1.51 - 1.62, SI: 102 - 107, EL: 0.30 - 0.36, OI: 27 - 31, PrL: 0.98 - 1.09, PrH: 0.66 - 0.79, PrW: 1.01 - 1.09, WL: 2.40 - 2.72, PeL: 0.84 - 0.90, PeH: 1.02 - 1.07, PeW: 0.72 - 0.78, LPI: 119 - 126, DPI: 85 - 90. Mandible shorter than that of worker. Petiole in profile distinctly higher than long, in dorsal view distinctly longer than wide.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	description	Male. Measurements (n = 2). HL: 1.08 - 1.17, HW: 0.83 - 0.87, CML: 0.35 - 0.38, CI: 75 - 77, CLI: 33, EL: 0.63 - 0.64, MeL: 1.01 - 1.10, MeW: 1.05 - 1.20, WL: 2.44 - 2.59, PeL: 0.68 - 0.75, PeH: 0.81 - 0.85, PeW: 0.54, LPI: 113 - 120, DPI: 71 - 80.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
DC4EF4E5096598E0D9ED9D290F58176C.taxon	description	Description. In full-face view, head longer than wide excluding eyes, wider posteriorly; posterior margin distinctly rounded. Mandible small, without basal angle and denticles, with rounded apex. Eye occupying over half of head length. Three prominent ocelli present; maximum length of lateral ocellus shorter than minimum distance between lateral ocellus and eye. Antenna 13 - segmented; scape distinctly shorter than width of head; antennomere II shortest, roughly as long as wide; III longest; XIII shorter than III. Clypeus triangular and projecting anteriad, convex on midline; anterolateral margin straight; apex moderately acute; epistomal suture broadly rounded. Mesoscutum in profile convex, in dorsal view shorter than wide, widest posterior to mid-length; notaulus weakly notable; parapsidal line more than half of mesoscutal length; transscutal line almost straight. Anterior margin of mesoscutellar disc rounded. In profile, propodeal dorsum forming continuous curve with declivity; metapleural-propodeal sulcus present. Forewing ca. 3.0 times as long as wide, with stigma. Hindwing less than 0.8 times as long as forewing. Petiole in profile higher than long, in dorsal view longer than wide, highest posteriorly, with short peduncle, with fan-shaped node; anterior and dorsal faces forming convex line; dorsal face sloping anteriad; posterior face convex; in dorsal view, lateral margin broadly convex. With waist in profile, anterior lobe of subpetiolar process short-subtriangular, anterior margin convex, ventral apex moderately acute, posterior margin concave. With gaster in profile, prora low, long; continuously curved between prora and anteroventral edge of gastral sternite I. Constriction between gastral segments I and II distinct. Head, mesosoma, and petiole weakly rugose, with piligerous small punctures. Mandible smooth. Clypeus with oblique to longitudinal striae. Subpetiolar process finely striate. Gastral segment I with irregularly sized depressions; distance between depressions distinctly greater than diameter of depressions; interspace between depressions with small punctures; segments II-V smooth; constriction between gastral segments I and II finely scrobiculate Body black, tinged with red; clypeus, mandible, antenna, legs, subpetiolar process, and gaster red-brown. Hairs of various lengths, scattered, white-yellow; antennal scape and legs with dense short pubescence, mixed with long scattered hairs. Scalloped depressions and small punctures on gastral segment I bearing hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	etymology	Etymology. Dedicated to Dr. Taisuke Kanao, collector of the material examined.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	distribution	Distribution. Malaysia: Borneo (Sarawak and Sabah).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	diagnosis	Worker. Diagnosis. In full-face view, head distinctly longer than wide (CI: 78 - 83). Mandible strongly curved throughout. Clypeus without lateral lobe; apex of median extension strongly projecting. Petiole in profile higher than long (LPI: 112 - 122), highest just anterior to posterodorsal angle; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face sloping anteriad. Gastral segment II smooth.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	description	Measurements (n = 5, holotype in parentheses). HL: 1.66 - 1.78 (1.77), HLL: 1.18 - 1.29 (1.24), HLA: 0.16 - 0.19 (0.19), HW: 1.34 - 1.43 (1.43), CML: 0.30 - 0.34 (0.34), CI: 78 - 83 (81), CLI: 17 - 19 (19), SL: 1.50 - 1.63 (1.56), SI: 109 - 116 (109), EL: 0.34 - 0.39 (0.36), OI: 27 - 30 (29), PrL: 1.04 - 1.09 (1.09), PrH: 0.69 - 0.74 (0.69), PrW: 1.03 - 1.07 (1.07), WL: 2.57 - 2.69 (2.69), PeL: 0.83 - 0.91 (0.91), PeH: 0.96 - 1.06 (1.02), PeW: 0.70 - 0.75 (0.75), LPI: 112 - 122 (112), DPI: 78 - 87 (82).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	description	Description. In full face view, head distinctly longer than wide. Mandible strongly curved throughout, with subapical tooth near apical tooth. Eye prominent, measuring one-fourth to one-third of head lateral margin length. Antennal scape distinctly longer than width of head, surpassing posterior margin of head by one-fifth of its length; antennomere III ca. 2.2 times as long as wide. Clypeus without lateral lobe; median extension moderately long, with apex strongly projecting. In dorsal view, pronotum longer than wide. Metanotal groove distinctly impressed. In profile, propodeal dorsum broadly convex. Petiole in profile higher than long, highest near mid-length, with roughly fan-shaped node; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face sloping anteriad; posterior face almost straight. Head weakly areolate-rugose, longitudinally striate anteriorly; vertex weakly and transversally striate. Mandible weakly and longitudinally striate. Prosternum weakly rugose. Gastral segment I with scalloped depressions that are irregular in size and close to each other; segments II smooth. Body black, gaster slightly tinged with red; clypeus, mandible, antenna, legs and ventral half of petiole dark-red. Apical two or three segments of gaster red-brown. Scalloped depressions on gastral segment I bearing hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	description	Queen. Measurements (n = 5). HL: 1.59 - 1.74, HLL: 1.13 - 1.26, HLA: 0.16 - 0.18, HW: 1.30 - 1.39, CML: 0.26 - 0.33, CI: 80 - 82, CLI: 15 - 19, SL: 1.39 - 1.52, SI: 107 - 110, EL: 0.33 - 0.40, OI: 29 - 32, PrL: 0.96 - 1.03, PrH: 0.65 - 0.71, PrW: 1.00 - 1.06, WL: 2.39 - 2.56, PeL: 0.82 - 0.86, PeH: 1.03 - 1.09, PeW: 0.72 - 0.78, LPI: 123 - 126, DPI: 88 - 94. Petiole in profile distinctly higher than long, in dorsal view longer than wide.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
B0D944DA3F95D371983F685DC489BFCF.taxon	description	Male. Unknown.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	etymology	Etymology. The species epithet, malayana, refers to the fact that this species was found on Peninsular Malaysia.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	distribution	Distribution. Malaysia: Peninsular Malaysia (Trengganu).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	diagnosis	Worker. Diagnosis. In full-face view, head almost as long as wide (CI: 98 - 101). Mandible curved near base, becoming straight apically. Antennal scape distinctly longer than width of head (SI: 115 - 121), surpassing posterior margin of head by two-fifths of its length. Clypeus with lateral lobe, with blunt angle between lateral lobe and median extension; apex of median extension truncate. Petiole in profile distinctly higher than long (LPI: 112 - 116), highest just anterior to posterodorsal angle; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face distinctly inclined anteriad.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	description	Measurements (n = 3, holotype in parentheses). HL: 1.57 - 1.63 (1.63), HLL: 1.16 - 1.21 (1.18), HLA: 0.17 - 0.19 (0.17), HW: 1.59 - 1.61 (1.60), CML: 0.23 - 0.24 (0.23), CI: 98 - 101 (98), CLI: 14 - 15 (14), SL: 1.83 - 1.95 (1.91), SI: 115 - 121 (120), EL: 0.36 - 0.38 (0.37), OI: 31 (31), PrL: 1.01 - 1.09 (1.09), PrH: 0.74 - 0.89 (0.89), PrW: 1.12 - 1.14 (1.14), WL: 2.57 - 2.78 (2.78), PeL: 0.88 - 0.97 (0.97), PeH: 1.02 - 1.10 (1.09), PeW: 0.72 - 0.76 (0.76), LPI: 112 - 116 (112), DPI: 77 - 82 (78).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	description	Description. In full face view, head almost as long as wide. Mandible curved near base, becoming straight apically; subapical tooth distinct in holotype or absent in paratypes. Eye prominent, measuring one-third of head lateral margin length. Antennal scape distinctly longer than width of head, surpassing posterior margin of head by two-fifths of its length; antennomere III ca. 2.7 times as long as wide. Clypeus with lateral lobe, with blunt angle between lateral lobe and median extension; median extension short, with truncate apex. In dorsal view, pronotum shorter than wide. Metanotal groove slightly impressed. In profile, propodeal dorsum weakly convex to almost straight. Petiole in profile distinctly higher than long, highest just anterior to posterodorsal angle, with fan-shaped node; anterior face of node ventrally vertical and dorsally forming continuous curve with dorsum; dorsal face distinctly inclined anteriad; posterior face slightly concave dorsally and slightly convex ventrally. Head strongly areolate-rugose, longitudinally striate near clypeus; vertex weakly and transversally striate. Mandible distinctly striate longitudinally. Gastral segment I with scalloped depression that are irregular in size but generally large; large depressions close to each other; distance between small depressions greater than diameter of depressions; segments II smooth. Body black, gaster slightly tinged with red; clypeus, mandible, antenna, legs and ventral half of petiole dark-red. Apical two or three segments of gaster red-brown. Scalloped depressions on gastral segment I bearing hairs.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
3898E8781DB13C9E1BF9109A3889ED3A.taxon	description	Queen and male. Unknown.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
937A70D7D95B2BBDDB299888BE75D696.taxon	distribution	Distribution. Indonesia: Enggano Island.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
937A70D7D95B2BBDDB299888BE75D696.taxon	diagnosis	Worker. Diagnosis. In full-face view, head distinctly longer than wide (CI: 88 - 90). Mandible curved near base, becoming straight apically. Antennal scape slightly longer than width of head (SI: 103 - 105), surpassing posterior margin of head by one-fifth to one-fourth of its length. Clypeus with lateral lobe, smoothly incurved between lateral lobe and median extension; median extension with convex apex. In profile, petiole slightly higher than long (LPI: 103 - 108), highest just anterior to posterodorsal angle; anterior and dorsal faces forming continuous curve; dorsal face distinctly sloping anteriad.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
937A70D7D95B2BBDDB299888BE75D696.taxon	description	Measurements (n = 5, lectotype in parentheses). HL: 1.52 - 1.68 (1.68), HLL: 1.11 - 1.22 (1.22), HLA: 0.18 - 0.20 (0.18), HW: 1.34 - 1.50 (1.50), CML: 0.25 - 0.28 (0.28), CI: 88 - 90 (90), CLI: 15 - 17 (17), SL: 1.40 - 1.58 (1.58), SI: 103 - 105 (105), EL: 0.30 - 0.34 (0.34), OI: 26 - 28 (28), PrL: 0.99 - 1.09 (1.09), PrH: 0.59 - 0.75 (0.75), PrW: 0.95 - 1.02 (1.02), WL: 2.49 - 2.68 (2.68), PeL: 0.85 - 0.92 (0.92), PeH: 0.91 - 0.98 (0.98), PeW: 0.64 - 0.67 (0.67), LPI: 103 - 108 (106), DPI: 73 - 76 (73).	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
937A70D7D95B2BBDDB299888BE75D696.taxon	description	Queen and male. Unknown.	en	Arimoto, Koichi (2017): Taxonomy of the Leptogenysmodiglianii species group from southeast Asia (Hymenoptera, Formicidae, Ponerinae). ZooKeys 651: 79-106, DOI: http://dx.doi.org/10.3897/zookeys.651.10336, URL: http://dx.doi.org/10.3897/zookeys.651.10336
