taxonID	type	description	language	source
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	description	(Figs 1 A-C; 2; 3 A, B)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	materials_examined	TYPE MATERIAL. — Holotype: China, Sichuan Prov., Xin Long County, Snake Mouth Cave, No. 389, 5. VIII. 1999, leg. J. & B. Lips, ♂ (MNHN DB 21). Paratypes: same locality, 1 ♂ (MNHN DB 21); 1 ♂ (ZMUM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	etymology	ETYMOLOGY. — To emphasize the prevailing marbled, brown coloration.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by peculiar male tarsal papillation, as well as certain details of leg and gonopod structure.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	description	DESCRIPTION Length c. 32 mm, maximum width 2.8 mm. Coloration rather pale brown, largely marbled; head, antennae (except for a pallid tip), dorsum and metazona below paraterga being slightly darker brown, pattern becoming somewhat annulate. Legs usually paler. Eye patches blackish. Head width = segment 4 >> collum> segment 2> 3 <5 <6 = 20 <7; after segment 20 or 21, body very gradually tapering toward telson. Body with 30 segments (29 pleurotergites, or rings, in terms of Enghoff et al. 1993). Head densely setose, clypeolabral region slightly convex. Eye patches triangular, each composed of 15 - 17 convex ocelli. Antennae very long and slender (Figs 1 A; 2 A), reaching beyond body segment 5 dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third. Gnathochilarium without promentum (Fig. 2 B). Collum usual, obcordate in shape, with rudimentary paraterga (Fig. 1 A). Tegument smooth, shining, only prozona distinctly and densely striolate transversely. Metatergal setation 3 + 3, typical, macrochaetae short, rather thick but pointed, positioned on minute knobs; stricture between pro- and metazona shallow, inconspicuous (Fig. 1 A-C). Paraterga small dorsolateral swellings / shoulders, not set off by peritremata, rather regularly rounded in dorsal view (Fig. 1 A-C). CIX (15) = 0.58; MIX (15) = 1.26; MA (15) ≈ 160 °; PIX (15) impossible to evaluate due to insufficiently developed paraterga. Axial suture distinct, pallid, usual; segment 7 only very slightly broader than adjacent ones, conspicuous due to slightly upturned, ridge-like pleural arches, without ventral apophyses. Limbus regularly microdentate, quite distinct ventrolaterally; similar minute denticles covering entire surface of pleural ridges. Legs long and slender, slightly longer in male, c. 1.4 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes but without papillae; coxa 2 perforated by gonopore orifice. All subsequent legs conspicuously papillate over caudal face ofventral half (Fig. 2 C- E, G, H). Legs 3 - 7 distinctly incrassate, pairs 3 and 4 particularly so. Femora 3 and 4 each with a small but evident mushroom-like protuberance at midway ventrally (Fig. 2 C). Coxae 5 - 7 microgranulate ventrally, prefemora 5 - 7 each with a microgranulate bulge dorso-parabasally (Fig. 2 D). Coxa 7 and trochanter 7 with a distoventral digitiform outgrowth (Fig. 2 E, F). Legs 10 and 11 with coxal glands; coxae 10 conspicuously enlarged and setose (Fig. 2 G); each coxa 11 with a large, apically pilose, coniform funnel with a long parabasal seta on front face; prefemur 11 with a strong parabasal process ventrally (Fig. 1 H). Claws invariably long, sabre-shaped, simple. Anterior gonopods (leg-pair 8) as usual very strongly reduced, sternum with a median and a lateral lobule, coxites spike-like (Fig. 2 I). Posterior gonopods (leg-pair 9) hypertophied (Fig. 3 A, B), with two ear-shaped folds at base on front face of colpocoxites and two highly conspicuous, extremely ramified, plumose, fimbriate, bush-like structures at base on caudal face; colpocoxites prominent, about as high as telopodites, with three longitudinal lamellae on caudal face, median lamella micropilose; telopoditomere 1 particularly strongly setose on posterior face, telopoditomere 2 infuscate and with a vestigial segment apically.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9776A48FC86FCDA1DB1C3CF.taxon	discussion	REMARKS	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	description	(Figs 1 D-G; 4)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	materials_examined	TYPE MATERIAL. — Holotype: China, Yunnan Prov., Zheng Xiong County, Cave Bai Yin Dong, Nos. 523 & 524, 18. VIII. 1999, leg. J. & B. Lips, ♂ (MNHN DB 42). Paratypes: same locality, 1 ♂ (MNHN DB 42); 1 ♀ (MNHN DB 42); 1 ♀ (MNHN DB 42); 6 juvs (MNHN DB 42).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	etymology	ETYMOLOGY. — To emphasize the particularly large size.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by the particularly large size, as well as certain details of leg and gonopod structure. The unusually strong colpocoxites, surmounted by a pseudosegmented head-like structure, also make N. grandis n. sp. easily distinguishable.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	description	DESCRIPTION Length c. 40 (♂) to 42 mm (♀), maximum width 2.8 (♂) to 2.9 - 3.0 mm (♀). Coloration entirely pallid. Head width = segment 6 = 20 >> collum> segment 2> 3> 5 <7; after segment 20 or 21, body very gradually tapering toward telson. Body with 30 segments. Head densely setose, clypeolabral region slightly convex. Eye patches triangular, each composed of 10 (♀) or 15 (♂) rather convex ocelli. Antennae extremely long and slender (Figs 1 D; 4 A), reaching beyond body segment 8 dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third. Gnathochilarium without promentum. Collum usual, obcordate in shape, with rudimentary paraterga (Fig. 1 A). Tegument smooth, shining, translucid, only prozona distinctly and densely striolate transversely. Metatergal setation 3 + 3, typical, macrochaetae medium-sized, rather thick but pointed, positioned on minute knobs; stricture between pro- and metazona very shallow, inconspicuous (Fig. 1 D-F). Paraterga small dorsolateral swellings / shoulders, not set off by peritremata, rather regularly rounded in dorsal view, only more abruptly so after caudal corner (Fig. 1 D-F). CIX (15) = 0.46; MIX (15) = 2.08; MA (15) ≈ 165 °; PIX impossible to evaluate as paraterga too poorly developed. Axial suture distinct, usual; segment 7 only very slightly broader than adjacent ones, conspicuous due to a clearly visible and frontally projecting anterior lobe of pleurite, latter without ventral apophyses. Limbus barely visible, microdentate, traceable only near pleurites ventrolaterally. Legs very long and slender, slightly longer in males than in females, c. 1.8 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes, but without papillae; male coxa 2 perforated by gonopore orifice. All tarsi slightly bulging immediately at base, male prefemora microgranulate ventrally (Fig. 4 E). Male legs 3 - 7 somewhat incrassate, pairs 3 and 4 especially so, each conspicuously papillate over caudal face of distoventral third (Fig. 4 B). Male legs 10 and 11 with coxal glands; prefemur 10 conspicuously enlarged at base (Fig. 4 C); prefemur 11 with a strong, microgranulate, parabasal process ventrally (Fig. 4 D). Claws invariably long, sabre-shaped, simple. Anterior gonopods (leg-pair 8) as usual very strongly reduced, sternum with a median protuberance and two lateral lobules, coxites spike-like (Fig. 4 F). Posterior gonopods (leg-pair 9) hypertophied (Fig. 4 G, H), with two ear-shaped folds at base on anterior face of colpocoxites and two large flagella beset with simple pilosity. Bases of coxite with two small, roundish bulges beset with minute plumose setae on caudal face. Colpocoxites especially prominent, higher than telopodites, distal one-third gently curved and directed distocaudally, delimited at base by a transverse sulcus on front face, with a relatively small, recurved, spoon-shaped process at base of caudal face, and an inner, gently ribbed lobe with a minute solenomere at tip. Telopoditomere 1 more strongly setose on front face, telopoditomere 2 with two vestigial segments apically. Vulva (Fig. 1 G) rather short, densely pilose, receptacle process on operculum small but evident, curved, digitiform, slender.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA9706A45FCACFD591B6DC465.taxon	discussion	REMARKS Nepalella grandis n. sp. is amongst the largest species in the order Chordeumatida, and certainly the largest in Nepalella. This can also be associated with cave gigantism, as this animal is definitely troglobitic, showing such typical troglomorphisms as unpigmented ocelli and tegument, extremely long legs and antennae. The number of ocelli per eye patch is also reduced, especially in the female. However, because very long and slender antennae are characteristic of many, often definitely epigeic congeners, their value as a troglomorphic trait is highly limited. This genus is well defined (Shear 2002) and has hitherto been known to comprise 20 species or subspecies from Nepal (10), Thailand (2), Myanmar (2), Vietnam (1) and southern China (5). The only previous key dealt with 14 of these (Mauriès 1988). With the addition of a further two congeners from Chinese caves, Nepalella can now be regarded as one of the most speciose genera among the oriental Chordeumatida. Considering all available descriptions (Shear 1979, 1987, 1999, 2002; Golovatch 1983; Mauriès 1988), the following refined and updated key to Nepalella species is proposed. This will hopefully encourage further work on this remarkable genus, in which many other new species undoubtedly remain to be discovered.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97F6A46FF67FB651B14C08D.taxon	materials_examined	TYPE SPECIES. — Lipseuma josianae n. sp.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97F6A46FF67FB651B14C08D.taxon	etymology	ETYMOLOGY. — Dedicated to Josiane et Bernard Lips, - euma being a common generic end suffix in Chordeumatida. Gender neuter. OTHER SPECIES INCLUDED. — Lipseuma bernardi n. sp.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97F6A46FF67FB651B14C08D.taxon	diagnosis	DIAGNOSIS. — Differing from both other genera of Kashmireumatidae, Kashmireuma Mauriès, 1982 and Vieteuma Golovatch, 1984, as follows: – a completely unpaired, undivided colpocoxite of anterior gonopods (male leg-pair 9) (versus paired, fully or partly independent in the other two genera); – posterior gonopods (male leg-pair 9) unusually simple, telopodite voluminous but 1 - segmented (versus strongly differentiated and 2 - segmented, regardless of a vestigial segment 3 sometimes visible in the other two genera); – male legs 6 particularly strongly hypertrophied in relation to any other leg-pair (versus normal to moderately incrassate male legs 3 - 6 (7) in the other two genera); – normal female legs 2 (versus normal or vestigial in the other two genera); – bursa of vulva particularly short and wide (versus elongate in the other two genera).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97F6A46FF67FB651B14C08D.taxon	discussion	REMARKS The small Oriental family Kashmireumatidae has hitherto been known from six species, three each in Kashmireuma (Himalayas of India and Nepal) and Vieteuma (Vietnam and southern China). With the addition of two species of Lipseuma n. gen., it becomes possible to better outline the genera included in the family. Shear (2002) has recently provided a brief summary concerning the main traits distinguishing all six kashmireumatid species known to date. These are Kashmireuma nielseni Mauriès, 1982, from Kashmir, India; K. schawalleri Shear, 1987 and K. nepalense Mauriès, 1988 (nom. emend. pro nepalensis), both from Nepal; Vieteuma topali Golovatch, 1984, from North Vietnam; V. hubeiense Mauriès & Nguyen Duy-Jacquemin, 1997 (nom. emend. pro hubeiensis), a troglobite from Hubei Prov., China; and V. longi Shear, 2002, from Yunnan Prov., China (Mauriès 1982, 1988; Golovatch 1984; Shear 1987, 2002; Mauriès & Nguyen Duy-Jacquemin 1997). The main differences between Kashmireuma and Vieteuma are as follows: – Vieteuma has the posterior gonopods placed on a thick plate-like sternum, but telopoditomere 1 is very strongly modified, supplied with conspicuous lobes / outgrowths, versus sternite almost absent medially, but telopoditomere 1 only slightly modified in Kashmireuma;	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97F6A46FF67FB651B14C08D.taxon	description	– Vieteuma has 26 or 28 body segments, versus 28 in Kashmireuma; – Vieteuma has female legs 2 normal or vestigial, versus vestigial in Kashmireuma. As one can see, the above distinctions are relatively shaky, with only the posterior gonopod structure so far serving to reliably separate these still oligotypic genera. When more kashmireumatids become known, it is possible that Vieteuma will have to be sunk as a synonym of Kashmireuma. The characters distinguishing Lipseuma n. gen. appear to be much stronger, especially the unusually simple anterior and posterior gonopods. In fact these are the simplest among the Chordeumatida. However, in no way can Lipseuma n. gen. be regarded as basal within the family in relation to Kashmireuma and Vieteuma. The gonopods of Lipseuma n. gen. species are simple, but the new genus itself is evolutionarily advanced. This situation resembles that observed in the North American genus Pseudotremia Cope, 1869, in which several species highly troglomorphic in body form show strongly simplified gonopods. The simplification results most probably because of isolation in caves, which removes any selective pressure. Indeed, the anterior gonopods of Lipseuma n. gen. species show a complete fusion of the colpocoxites, these being fused only partly (parabasally) in Kashmireuma nepalense and entirely free in the other species of the family. Further apomorphies of Lipseuma n. gen. include telopoditomere 2 of the posterior gonopods being totally suppressed, and the vulvae very strongly transverse. The same holds for the hypertrophied and very special male legs 6, which apparently serve the male as claspers during copulation.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	description	(Figs 5; 6)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	materials_examined	TYPE MATERIAL. — Holotype: China, Hubei Prov., Banqiao County, Cave Chuan Dong Zi, Nos. 482, 485, 494, 9. VIII. 1999, leg. J. & B. Lips, ♂ (MNHN DA 155). Paratypes: same locality, 1 ♂ (MNHN DA 155); 1 ♂, 2 juvs (MNHN DA 155); ♀ (MNHN DA 155); 2 juvs (MNHN DA 155).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	etymology	ETYMOLOGY. — To honour Josiane Lips.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	diagnosis	DIAGNOSIS. — Differs from the only congener (L. bernardi n. gen., n. sp.) by the slightly larger size, as well as in certain details of leg and gonopod structure.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	description	DESCRIPTION Length 9.3 - 9.8 (♂) to 10.1 mm (♀); width of midbody segments, 0.75 - 0.8 (♂) to 0.9 mm (♀), maximum width of head, 0.95 mm (holotype). Vertical diameter of midbody segments, 0.75 (♂) to 0.88 mm (♀). Coloration entirely pallid. Head width >> collum> segment 2> 3> 4> 5> 6> 7> 8> segment 9 = 18; thereafter body very gradually tapering toward telson. Body with 28 segments (46 leg-pairs). Head sparsely setose, clypeolabral region slightly convex. Eyes completely reduced. Antennae very long and slender (Figs 5 A; 6 C), reaching beyond body segment 6 (♂) or 5 (♀) dorsally; antennomere 7 with a peculiar flattened seta dorsally in distal third; antennomere 8 with four unusually long and slender sensory cones. Gnathochilarium without promentum. Collum usual, obcordate in shape, with rudimentary paraterga. Tegument smooth, modestly shining, non-areolated, translucid. Metatergal setation 3 + 3, typical, macrochaetae very long, pointed, positioned on minute knobs; exterior macrochaeta longest, c. 0.7 times as short as body width and nearly three times longer than median or interior one; stricture between pro- and metazona very shallow, inconspicuous (Fig. 5 B, C). Paraterga very small dorsolateral shoulders, rather regularly rounded in dorsal view (Fig. 5 C), slightly better developed on anterior body third. CIX (15) ≈ 0.9; MIX (15) = 1.25; MA (15) ≈ 175 °; PIX impossible to evaluate as paraterga too poorly developed. Axial suture distinct, usual; segment 7 inconspicuous, pleurite without ventral apophyses. Pleural crests evident. Limbus invisible. Legs generally very long and slender, c. 1.8 times as long as midbody height. Legs 1 and 2 slightly reduced as usual, tarsi with usual ventral brushes, but without papillae; male coxa 2 perforated by gonopore orifice. Tarsal papillae absent. Male legs 3 - 5 unmodified. Male legs 6 (Fig. 5 D) exceptionally incrassate, with a distoventral process both on coxa and prefemur; acropodite strongly clasper-like, tarsus with two bulges subventrally and a modified claw. Male legs 7 (Fig. 5 E) slightly incrassate, with evidently enlarged coxa and prefemur; coxa with a pointed lobe ventrally. Male legs 10 (Fig. 6 A) and 11 normal, both pairs with coxal glands. Claws very long, sabre-shaped, simple. Anterior gonopods (leg-pair 8) unusually simple, small; coxites with traces of median division basally on caudal face, completely fused on front face; distally subsecuriform telopodites laterocaudal, with a few setae distally, nearly as high as the unpaired, linguiform, distally pilose syncolpocoxite, but somewhat higher than the voluminous angiocoxal sacs with a ridge on top; sternum plate-like, with each lateral bulge on front face supporting a strong seta (Fig. 5 F-H). Posterior gonopods (leg-pair 9) also very simple (Fig. 6 B), placed on a plate-like sternum; coxae contiguous medially, each crowned with a simple, long, slender, erect process ventrally; telopodite reduced to a single voluminous segment filled with glandular material. Female legs 2 normal. Vulva (Fig. 6 D, E) very broad; operculum small, devoid of outgrowths.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97E6A42FCB7FD9F1D30C7C1.taxon	discussion	REMARKS In one of the male paratypes, leg 6 is normal, not modified, whereas the anterior gonopod syncolpocoxite shows a small median notch apically, but there are no lateral setae on the coxite. Some of these traits are probably anomalous, likely neotenic.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	description	(Figs 7; 8)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	materials_examined	TYPE MATERIAL. — Holotype: China, Sichuan Prov., Xin Long County, Three Eyes Cave, No. 438, 7. VIII. 1999, leg. B. & J. Lips, ♂ (MNHN DA 156). Paratypes: same locality, 1 ♂, incomplete (MNHN DA 156); 1 ♂, subadult (MNHN DA 156).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	etymology	ETYMOLOGY. — To honour Bernard Lips.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	diagnosis	DIAGNOSIS. — Differs from the only congener (L. josianae n. gen., n. sp.) by the slightly smaller size, as well as minor details of leg and gonopod structure.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	description	DESCRIPTION Length 7.2 (holotype) or 6.8 mm (subadult ♀); width of body segment 6, 0.72 mm, of midbody segments, 0.78 mm, maximum breadth of head, 0.9 mm (holotype). Antennae (Fig. 7 A) with even more exposed, highly elongate, pointed apical cones on stalks. All peripheral characters like in L. josianae n. gen., n. sp. except as follows: exterior macrochaetae on midbody segments 0.6 mm long, i. e. nearly as wide as segment itself (Fig. 7 B). Male legs 6 (Fig. 7 C, D), 10 (Fig. 8 A) and 11 virtually the same as in L. josianae n. gen., n. sp., but male legs 7 (Fig. 7 E) with a characteristic protuberance on tarsus ventrally at about midway. Anterior gonopods (Fig. 7 F, G) also virtually the same as in L. josianae n. gen., n. sp., but posterior gonopods (Fig. 8 B) with traces of a second telopoditomere distally on caudal face. Subadult male (26 body segments) with small but evident gonopod promordia, legs 7 almost as strongly incrassate as in adult male, legs 7 also slightly enlarged.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
03FEF24BA97A6A5DFCB4FC3A1DD3C4A7.taxon	discussion	REMARKS Both Lipseuma n. gen. species are remarkable in being completely unpigmented and eyeless, as well as in bearing strongly elongated antennae, legs (especially tarsi) and exterior macrochaetae. All these features leave no doubt that we are dealing with troglobites. It is noteworthy that interspecific variation in Lipseuma n. gen. is very modest, in contrast to that observed in the other two kashmireumatid genera. In fact, L. josianae n. gen., n. sp. and L. bernardi n. gen., n. sp. are barely distinguishable. The very small differences noted between them suggest either a quite recent colonization of caves by their common ancestor or a fairly recent vicariance event with speciation within their respective cave systems. The latter scenario seems more plausible, given the profound troglomorphism of both species, which would require a geological time-scale to develop, along with the geographical proximity of Chuan Dong Zi and Three Eyes caves (both situated in the Xin Long geographical zone just at the border between Hubei and Sichuan provinces [Degouve et al. 1997]). In other words, the old geological age of the genus Lipseuma n. gen. seems unquestionable, whereas the isolation of both hitherto known congeners probably occurred only relatively recently.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul (2006): Four new Chordeumatida (Diplopoda) from caves in China. Zoosystema 28 (1): 75-92, DOI: 10.5281/zenodo.5393502
