taxonID	type	description	language	source
03F187B6B317FF83FF298C56FA9A0E8C.taxon	diagnosis	Diagnosis. Number of species: 55. Distribution: Ecuador (25), Colombia (15), Brazil (5), Costa Rica (4), Panama (4), Venezuela (2), Bolivia (1), Peru (1), Argentina (1). In some cases, the species are distributed in several countries, for example G. nemoralis (Ecuador, Costa Rica, Panama), G. parecis (Argentina, Brazil), G. crucifer (Ecuador, Peru). This list does not include undescribed species listed in Brown & Fragoso (2007). Genus diagnosis, modified, following Righi (1995): Eight setae per segment. One pair of intra-clitellar male pores. One pair of calciferous glands in segments 11 and 12 of intertwined-tubular structure, without glandular and membranous division. Three or two pairs of lateral hearts in segments 7 – 9 and two pairs of intestinal hearts in segments 10, 11. One pair of testes and male funnels present in segment 11. Seminal vesicles generally long. Copulatory chambers absent. Spermathecae mostly present, usually without seminal chambers or diverticula.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B314FF85FF298D9FFEB10D45.taxon	materials_examined	Material examined. Holotype: Clitellate specimen (CNLT-UTP- 0316). Locality: Colombia, department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain; intercropped coffee / plantain. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 20 cm. Collection date: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, and L. F. Zuluaga. Paratypes: 4 adults, 1 juvenile (CNLT-UTP- 0317). Locality: Colombia, Department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain; intercropped coffee / plantain. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 20 cm. Collection date: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, L. F. Zuluaga, and L. F. Arango.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B314FF85FF298D9FFEB10D45.taxon	etymology	Etymology. The new species name is given in honor of the chiefdom of the Quimbaya culture, where the chief was named " chaguala ".	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B314FF85FF298D9FFEB10D45.taxon	description	Description. Endogeic species, with a cylindrical, non-pigmented body. Holotype: length (fixed and preserved in 95 % alcohol) 19.1 mm, diameter 1.23 mm in preclitellar region, 1.15 mm at clitellum, 1.12 mm in postclitellar region. Number of segments: 74. Paratype (three adults): Adult 1: 21.9 mm long, width 0.93 mm (segment 10), 1.3 mm (segment 21), 0.99 mm (segment 40); number of segments: 87. Adult 2: 19.4 mm long, width 0.93 mm (segment 10), 0.91 mm (segment 21), 0.89 (segment 40); number of segments: 79. Adult 3: 15.3 mm long, width 0.78 mm (segment 10), 0.8 mm (segment 21), and 0.77 mm (segment 40); number of segments: 85. The prostomium small, epilobic. Setae arranged in four pairs of longitudinal rows. Setae sigmoid with nodulus, in the median zone with ornate nodule curve, length 0.05 mm (Fig. 2 A). Setae a and b of 10 and 16 modified into genital setae (Fig. 2 B). Distance between setae in segment 30 aa: ab: bc: cd: dd = 4: 1: 3: 2: 14. The saddle-shaped clitellum lies in segments 15 – 20, 21 (= 6, 7) (Fig. 2 C), with tubercula pubertatis located between segments 16 – 18, ½ 19 (= 3, 3.5 segments), they are shaped like a discontinuous thick line that closes in the ventral region, next to the line setae cd. In the ventral region of segment 13, a pair of round-shaped papillae. Male pores in intersegment 16 / 17, female pores in ½ 14. Nephropores between setae c and d. Two pairs of spermathecal pores were observed in intersegments 8 / 9 and 9 / 10. Internal anatomy. Septa 6 / 7 – 10 / 11 very thick, conical, interpenetrated. Gizzard in segment 6. One pair of calciferous glands in segments 11, 12 of intertwined-tubular structure, without glandular and membranous division. Esophagus-intestine transition in segment 15, typhlosole from segment 15. Absence of caeca. Three pairs of lateral hearts in segments 7 – 9, two pairs of intestinal hearts in segments 10 and 11. Absence of supraesophageal and subneural vessels. One pair of vesicular holonephridia per segment. The two large testis sacs coalesce ventrally in segment 11. The large seminal vesicles in segment 12 continue back to segment 26 (Fig. 2 D). The male ducts run ventrally along the parietal region to intersegment 16 / 17. Ovaries in segment 13, female pores in segment ½ 14. Two pairs of spermathecae in segments 9 and 10, bean-shaped, of different size, the second pair are larger. The duct thin and separate from the ampulla (Fig. 2 D).	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B314FF85FF298D9FFEB10D45.taxon	discussion	Remarks. Glossodrilus chaguala sp. nov. is a species closely related to G. baloghi Zicsi, 1988, Glossodrilus kalmari Zicsi, 1995, Glossodrilus kaszabi Zicsi, 1988, Glossodrilus mahnerti, Zicsi 1989, and Glossodrilus parecis Righi & Ayres, 1975 because of the presence of clitellum in segments 15 – 20, 21. The following characteristics of the new species allow its differentiation from the others: tubercula pubertatis in segments 16 – 18, ½ 19; male pores located in intersegment 16 / 17; large seminal vesicles in segment 12 that continue to the end of segment 26, and two pairs of bean-shaped spermathecae in segments 9 and 10. Distinctive characteristics of the other species are as follows: G. baloghi: clitellum annular; tubercula pubertatis in segments 16 – 1 / 4 19; male pores in segments 16; in the ventral region of segment 13, a pair of round-shaped papillae; seminal vesicles in segments 16 – 19. G. kalmari: tubercula pubertatis in segments 18 – ½ 20; male pores in segments 18 / 19; seminal vesicles in segment 11. G. kaszabi: tubercula pubertatis in segments 16, ½ 16 – 18; male pores in segments 16 / 17; not clear if; one pair of spermathecal pores in segments 8 / 9. G. mahnerti: tubercula pubertatis in segments 16 – 18; male pores in segments 17 / 18; seminal vesicles in segment 11. G. parecis: tubercula pubertatis with quadrangular-shaped glandular area in segments 2 / 3 16 – 2 / 3 18; male pores in segments 16 / 17; seminal vesicles in segments 11 – 14, 15; absence of spermathecal pores.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF85FF29887AFF380A12.taxon	materials_examined	Type species. Rhinodrilus ecuadoriensis Benham, 1892.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF85FF29887AFF380A12.taxon	diagnosis	Diagnosis. Number of species: 15. Distribution: Colombia (6), Ecuador (4), Peru (2), Guyana (2), Surinan (1). In some cases, the species are distributed in several countries, for example M. (Mp.) agricola, M. (Mp.) ecuadoriensis, and M. (Mp.) savanicola, in Colombia and Ecuador and M. (Mp.) micrurus and M. (Mp.) ecuadoriensis in Ecuador and Peru (Feijoo & Celis 2011). Genus diagnosis, modified, following Righi (1995), Zicsi (2001): Species with green, red, violet, brown or blueish pigmentation. Eight rows of regular, closely paired setae. Absent or very thin membranous septa in the gizzard area; 7 – 8 pairs of calciferous glands between segments 7 – 13 or 8 – 14 with a composite, reticulate tubular structure. Holandric and metagynic sexual apparatus with testes in segments 10 and 11 and hypo- or periesophageal testis sacs. Short seminal vesicles limited to segments in 11 and 12. Four pairs of spermathecae present in segments 6, 7, 8, 9.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF87FF298EA5FB930E8D.taxon	materials_examined	Material examined. Holotype: Clitellate specimen (CNLT-UTP- 0320). Locality: Colombia, Department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercropped coffee / plantain crops. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 10 cm litter. Collection dates: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo and L. J. Molina. Paratypes: Three adults (CNLT-UTP- 0321). Locality: Colombia, Department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole croppd plantain, intercropped coffee / plantain crops. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 10 cm litter. Collection dates: 26 January 2015, 26 April 2016 and 31 June 2017. Collectors: A. Feijoo (2015) and A. Feijoo and L. J. Molina, other dates. Other materials: three juveniles and 7 adults (CNLT-UTP- 0322), same collection data as holotype.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF87FF298EA5FB930E8D.taxon	etymology	Etymology. The name of the species is given because of an ancient indian group dominant in the Colombian coffee-growing region.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF87FF298EA5FB930E8D.taxon	description	Description. Epigeic species, with a light-blue to Bermuda grey-colored dorsum, milky white ventrum, and cylindrical body. Holotype: length (preserved) 74.2 mm, diameter 2.3 mm in preclitellar region (segment 8), 3.3 mm at clitellum, 2.9 mm in mid-posterior region. Number of segments: 103. Paratype (three adults): Adult 1: 56.6 mm long, diameter 3 mm in preclitellar region, 2.8 mm at clitellum, 2.6 mm in mid-posterior region, number of segments: 70. Adult 2: 88.7 mm long, diameter 2.8 mm in preclitellar region, 3.2 mm at clitellum, 2.8 mm in midposterior region; number of segments: 119. Adult 3: 94.8 mm long, diameter 2.8 mm in preclitellar region, 3.94 mm at clitellum, 2.3 mm in mid-posterior region; number of segments: 115. The prostomium short and lobeshaped. Eight rows of closely paired setae are visible (a, b, c, d) as of segment 3. Common setae slightly sigmoid with a slightly pronounced central nodule, tips at both ends almost rounded. Distance between setae in mid-body region (segments 40 – 50); aa: ab: bc: cd: dd = 4.4: 1: 4.9: 0.6: 20.9. Genital setae in segments 5 – 9, 15 – 23, with transverse excavations in proximal 2 / 3, in distal 1 / 3 semilunar marks; distally with a spear-shaped tip (Fig. 3 A). Clitellum whitish (preserved), saddle-shaped, in segments 14 – 1 / 2 23 (= 9.5 segments); tubercula pubertatis between segments ½ 19 – ½ 26 (Fig 3 B). Male pores in segment 20 / 21, female pores in segment 13 / 14. Nephropores aligned with setae c. Four pairs of spermathecal pores observed in intersegments 5 / 6, 6 / 7, 7 / 8, 8 / 9. Internal anatomy. Anterior septa very fragile: only tiny membranous threads representing the septa are observed between segments 5 and 12. Gizzard rounded, in segment 6. Seven pairs of small extramural, tubularshaped calciferous glands, opening into dorso-ventral region of esophagus in segments 7 – 13; all glands of similar size, with a small appendage in the distal region highly irrigated with blood; gland structure of composite-tubular type, consisting of short parietal tubules that flow into each other. In the center, the tubules merge together in the lumen from where they branch out toward the tip of the calciferous gland. The lumen narrows in the submedial region and tubules close irregularly. Transition esophagus-intestine at 17 / 18, typholosole beginning in segments 26 – 27. No caeca. Three pairs of thin lateral hearts in segments 7 – 9, separating the calciferous glands, and two pairs of voluminous intestinal hearts in segments 10 and 11. One pair of holonephridia per segment, nephridiopores located just above the line of setae c. Two pairs of large hypo-esophageal testis sacs in segments 10 and 11, two pairs of seminal vesicles extending forward in segments 11 and 12. One pair of ovaries in segment 13. Four pairs of spermathecae that increase in size, 9> 8> 7> 6, near setae c. Each spermatheca with conspicuous duct and ampulla visible from the outside; duct short, ampulla an elongate mass; no seminal chambers or diverticles (Fig. 3 C).	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B312FF87FF298EA5FB930E8D.taxon	discussion	Remarks. Martiodrilus (Maipure) quimbayaensis sp. nov. is a species closely related to M. (Mp.) ecuadoriensis (Benham, 1892), M. (Mp.) grandis Zicsi, 2001, M. (Mp.) huitoto Feijoo & Celis, 2011, M. (Mp.) palmirus Zicsi & Feijoo, 1994 and M. (Mp.) savanicola (Michaelsen 1900), because of the presence of seven pairs of calciferous glands (in segments 7 – 13 or 8 – 14, depending on the species) and four pairs of spermathecae. However, these species differ from the new species in the following characteristics: M. (Mp.) ecuadoriensis: clitellum in segments 15 – ½ 25 (vs. 14 – ½ 23), tubercula pubertatis in segments 21 – 27 or ½ 20 – ½ 27 (vs. ½ 19 – ½ 26); spermathecae bag-shaped and without distinction between duct and ampulla; shape and size of normal setae, and ornamentation of genital setae. M. (Mp.) grandis: clitellum in segments 14 – ½ 27, 27, (vs. 14 – ½ 23), tubercula pubertatis in segments ½ 20 – ½ 27 (vs. ½ 19 – ½ 26), shape of spermathecae. M. (Mp.) huitoto: clitellum in segments 14, 15 – 28 (vs. 14 – ½ 23), tubercula pubertatis in segments ½ 19 – ½ 26 (vs. ½ 19 – ½ 26), as well as shape of spermathecae. M. (Mp.) palmirus: clitellum in segments 15 – ½ 25 (vs. 14 – ½ 23), tubercula pubertatis in segments 20 – ¼ 25 (vs. ½ 19 – ½ 26), calciferous glands in segments 8 – 14 (vs. 7 – 13). M. (Mp.) savanicola: clitellum in segments 15 – 24 (vs. 14 – ½ 23), tubercula pubertatis in segments 20 – ¼ 27 (vs. ½ 19 – ½ 26), nephropores aligned with setae d (vs. aligned with setae c), calciferous gland in segments 8 – 14 (vs. 7 – 13).	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B310FF87FF298F64FE7B0A32.taxon	materials_examined	Material examined. Native, endogeic. Locality: Colombia, department of Quindío, La Cristalina farm (4 ° 27 ' 56 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercropped coffee / plantain crops. Altitude: 1227 m. a. s. l. Material collected: 8 adults. Sampling depth: 10 – 20 cm. Collection date: 26 April 2016. Collectors: A. Feijoo, L. J. Molina, and L. F. Zuluaga.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B310FF87FF298E85FB470983.taxon	materials_examined	Material examined. Native, endogeic. Clitellate specimen (CNLT-UTP- 0318). Locality: Colombia, department of Quindío, municipality of Armenia, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercropped coffee / plantain crops. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 20 cm. Collection dates: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, and L. F. Zuluaga. Clitellate specimen (CNLT-UTP- 0319), same collection data as previous specimen. Material collected: 9 adults, 3 juveniles. Locality: Colombia, department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercopped coffee / plantain crops. Altitude: 1479 m. a. s. l. Sampling depth: 0 – 20 cm. Collection dates: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, L. F. Zuluaga, and L. F. Arango.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B310FF87FF298995FBA10B54.taxon	materials_examined	Material examined. Native, endogeic. Locality: Colombia, department of Quindío, municipality of Armenia, Niagara rural community, La Catalina farm (4 ° 32 ' 58.48 " N, 75 ° 41 ' 32.98 " W). Altitude: 1436 m. a. s. l. Material collected: 3 adults. Collection date: 27 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Villa Sofia farm (4 ° 30´34.36 ´´ N, 75 ° 45´23.50 ´´ W). Altitude: 1222 m. a. s. l. 13 adults, 6 juveniles. Collection date: 27 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindio, municipality of Calarcá, Quebrada Negra rural community, Madrigal farm (4 ° 21 ' 07 " N, 75 ° 39 ' 39 " W). Altitude: 1576 m. a. s. l. 1 adult. Collection date: 28 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B310FF86FF298C8FFC050F81.taxon	materials_examined	Material examined. Native, endogeic. Locality: Colombia, department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercropped coffee / plantain crops. Altitude: 1479 m. a. s. l. Material collected: 9 adults, 3 juveniles. Sampling depth: 10 – 20 cm. Collection dates: 26 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, L. F. Zuluaga, and L. F. Arango.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B311FF86FF298EB9FD7B09CF.taxon	materials_examined	Material examined. Exotic, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Villa Sofia farm (4 ° 30 ' 34.36 " N, 75 ° 45 ' 23.50 " W). Altitude: 1222 m. a. s. l. 5 adults, 3 juveniles. Collection date: 27 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, Marmato rural community, La Cristalina farm (4 ° 27 ' 56.97 " N, 75 ° 44 ' 50.29 " W). Altitude: 1227 m. a. s. l. Material collected: 22 adults, 10 juveniles. Collection date: 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B311FF86FF298C12FF3308AC.taxon	materials_examined	Materials examined. Exotic, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Villa Sofia farm (4 ° 30 ' 34.36 " N, 75 ° 45 ' 23.50 " W). Altitude: 1222 m. a. s. l. Material collected: 3 adults, 2 juveniles. Collection date: 25 – 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B311FF86FF298A57FCC50DDE.taxon	materials_examined	Material examined. Native, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Bella Marina farm (4 ° 30 ' 37.69 " N, 75 ° 45 ' 28.36 " W). Altitude: 1214 m. a. s. l. Material collected: 2 adults. Collection dates: 25 – 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, El Rhin rural community, La Cabaña farm (4 ° 29 ' 36.80 " N, 75 ° 43 ' 38.15 " W). Altitude: 1270 m. a. s. l. Collection dates: 25 – 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Circasia, Barcelona rural community, La Sofe farm (4 ° 33 ' 36 " N, 75 ° 41 ' 5 " W). Altitude: 1479 m. a. s. l. 1 adult. Collection dates: 25 – 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B311FF86FF2989E1FB770CDB.taxon	materials_examined	Material examined. Native, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La India rural community, La Ermita farm (4 ° 31 ' 08.29 " N, 75 ° 43 ' 47.36 " W). Altitude: 1285 m. a. s. l. Material collected: 1 adult. Collection dates: 25 – 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B311FF86FF2988EEFCC50A71.taxon	materials_examined	Material examined. Exotic, endogeic. Locality: Colombia, department of Quindío, municipality of Armenia, Niagara rural community, La Catalina farm (4 ° 32 ' 58.48 " N, 75 ° 41 ' 32.98 " W). Altitude: 1436 m. a. s. l. 27 adults, 64 juveniles. Collection date: 28 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Circasia, Barcelona Baja rural community, Buenos Aires farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Altitude: 1463 m. a. s. l. Material collected: 32 adults, 70 juveniles. Collection dates: 27 and 28 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B31EFF89FF2989E2FCC50BB9.taxon	materials_examined	Material examined. Exotic, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, Niagara rural community, La Catalina farm (4 ° 32 ' 58.48 " N, 75 ° 41 ' 32.98 " W). Altitude: 1436 m. a. s. l. Material collected: 8 adults, 2 juveniles. Collection date: 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Villa Sofía farm (4 ° 30 ' 34.36 " N, 75 ° 45 ' 23.50 " W). Altitude: 1222 m. a. s. l. Material collected: 4 adults. Collection date: 25 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Circasia, Barcelona rural community, La Sofe farm (4 ° 33´36 ´´ N, 75 ° 41´5 ´´ W). Altitude: 1479 m. a. s. l. Material collected: 2 adults. Collection date: 30 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Circasia, Barcelona rural community, Buenos Aires farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Altitude: 1410 m. a. s. l. Material collected: 1 adult, 2 juveniles. Collection date: 27 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B31EFF89FF298E0FFC050AA6.taxon	materials_examined	Material examined. Exotic, endogeic. Locality: Colombia, department of Quindío, La Sofe farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Type of vegetation: sole cropped plantain, intercropped coffee / plantain crops. Altitude: 1479 m. a. s. l. 10 adults, 4 juveniles. Sampling depth: 0 – 10 cm. Collection dates: Material collected: 29 April 2016 and 1 June 2017. Collectors: A. Feijoo, L. J. Molina, L. F. Zuluaga, and L. F. Arango.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B31EFF89FF298B8CFDDF0D26.taxon	materials_examined	Material examined. Exotic, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Bella Marina farm (4 ° 30 ' 37.69 " N, 75 ° 45 ' 28.36 " W). Altitude: 1214 m. a. s. l. 2 adults. Collection date: 29 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, La India rural community, La Miranda farm (4 ° 30 ' 58.42 " N, 75 ° 43 ' 40.07 " W). Altitude: 1320 m. a. s. l. Material collected: 5 adults. Collection date: 25 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Armenia, La Patria rural community, La Brasilia farm (4 ° 32 ' 21.36 " N, 75 ° 43 ' 45.36 " W). Altitude: 1373 m. a. s. l. Material collected: 4 adults, 2 juveniles. Collection date: 29 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo. Locality: Colombia, department of Quindío, municipality of Circasia, Barcelona Baja rural community, Buenos Aires farm (4 ° 33 ' 30 " N, 75 ° 41 ' 10 " W). Altitude: 1463 m. a. s. l. Material collected: 3 adults. Collection date: 29 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
03F187B6B31EFF89FF298D29FAB009A3.taxon	materials_examined	Material examined. Exotic, epigeic. Locality: Colombia, department of Quindío, municipality of Armenia, La Revancha rural community, Villa Sofia farm (4 ° 30 ' 34.36 " N, 75 ° 45 ' 23.50 " W). Altitude: 1222 m. a. s. l. Material collected: 1 adult, 2 juveniles. Collection date: 25 April 2016. Collectors: A. Feijoo, L. Molina, and B. Murillo.	en	Feijoo, Alexander, Zuluaga, Luis Fernando, Molina, Ligia Janneth (2018): New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region. Zootaxa 4496 (1): 448-458, DOI: 10.11646/zootaxa.4496.1.34
