identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F36C399A1ED467D9FBFBD06694FCC0.text	03F36C399A1ED467D9FBFBD06694FCC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bouchardium Kamiński 2024	<div><p>Bouchardium gen. nov.</p><p>Type species. Bouchardium chillygonzalesi sp. nov.; here designated.</p><p>Diagnosis. The specific structure of the prosternum (elongate and projecting tip of prosternum, in lateral view, Figs 1A, 3F, K), elytra (presence of sharp lateral costae, Figs 1–3), and scutellum (apex with deep transverse grooves, Fig. 3A–C) place Bouchardium close to the following genera (Kamiński et al. 2021, Gearner et al. 2022): Amiantus Fåhraeus, 1870, Arturium Koch, 1951, Brachyphrynus Fairmaire, 1882, Dichtha, Distretus Haag-Rutenberg, 1871, Euphrynus Fairmaire, 1897, Glyptophrynus Fairmaire, 1899, Melanolophus Fairmaire, 1882, Phrynocolus Lacordaire, 1859, Phrynophanes Koch, 1951, and Physophrynus Fairmaire, 1882 . Among these, Bouchardium displays the greatest affinity towards Distretus and Dichtha as all share a similar structure of elytra (sharply carinate humeral edge and flattened surface of elytra). Nevertheless, Bouchardium can be differentiated from all of the abovementioned genera, including Dichtha and Distretus, by a unique arrangement of pronotal punctures (sides of the pronotal disc with large, confluent punctures; center of the disc with small punctures, 4–5 diameters apart vs pronotal disc evenly covered with large sometimes confluent punctures— Figs 2A, F, 3D, F, G, I) and subquadrate shape of the body (frontal/posterior views) (Fig. 1C, 2C, D, 3E).</p><p>Notes. According to Kamiński et al. (2022), the morphology of female terminalia is relatively invariable within the ‘ Dichtha clade’. All analyzed members of this grouping, including Bouchardium chillygonzalesi sp. nov., B. cubicum (Guérin-Méneville, 1845) comb. nov. and B. mariae sp. nov. possess 3-lobed ovipositors, where the merged lobes 1 (valvifer) and 2 are strongly emarginate basally. Proctigers were reported to be widely indented medially, while bursae were simple (no sclerites were detected) and accompanied by multibranched spermatheca. Finally, spicula ventrali were reported to be Y-shaped (Fig. 4C). Nevertheless, some variability of the apical coxites (c4 plates) has been recovered for different Bouchardium species. In particular, the c4 plates of the newly described B. chillygonzalesi sp. nov. are relatively short and only slightly projected above the outline of the c3 plate (Fig. 4A, B). In the case of B. cubicum and B. mariae sp. nov. the c4 plates are elongated (Fig. 4D).</p><p>The aedeagal morphology is relatively invariable within the subtribe Molurina (Kamiński et al. 2021) . Up to now, the shape of male terminalia has not been used for diagnostic purposes at the species or generic levels. Morphology of aedeagal tegmen and spiculum gastrale of Bouchardium mariae sp nov. is illustrated in Fig. 2I, J. Present investigation did not reveal any substantial differences in male terminalia morphology between that species and its other congeners.</p><p>Distribution. Botswana (Eastern part of the Country), Mozambique, South Africa (Provinces: Gauteng, Limpopo, Mpumalanga, North West), Zimbabwe (Southern part of the Country) (Fig. 5).</p><p>Etymology. This genus is named in honor of Dr. Patrice Bouchard (Canadian National Collection of Insects, Arachnids and Nematodes,Agriculture andAgri-Food Canada,Ottawa,Ontario,Canada), a specialist in Tenebrionidae and zoological nomenclature in gratitude for his professional support during the first years of my career.</p><p>Species included (3). B. chillygonzalesi sp. nov., B. cubicum (Guérin-Méneville, 1845) comb. nov., B. mariae sp. nov.</p><p>Key to the species of Bouchardium</p><p>1. Prosternal process with elongate appendages (Fig. 1E). Elytral disc relatively narrow leaving sides of the epipleura exposed in dorsal view (Fig. 1B). Ovipositor with short c4 plates (not extending beyond the tip of c3) (Fig. 4A, B)................................................................................................. B. chillygonzalesi sp. nov.</p><p>-. Prosternal process rounded (in lateral view) (Fig. 2G). Elytral disc relatively wide covering sides of epipleura in dorsal view (Fig. 1B). Ovipositor with elongate c4 plates (extending beyond the tip of c3) (Fig. 4D)............................. 2</p><p>2. Elytra entirely black. Elytral disc relatively flat (Fig. 3E).................................... B. cubicum comb. nov.</p><p>-. Elytra with two red stripes along the sides of the disc. Elytral disc convex (Fig. 2A, C, D).............. B. mariae sp. nov.</p></div>	https://treatment.plazi.org/id/03F36C399A1ED467D9FBFBD06694FCC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kamiński, Marcin Jan	Kamiński, Marcin Jan (2024): New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel. Zootaxa 5446 (1): 77-87, DOI: 10.11646/zootaxa.5446.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5446.1.4
03F36C399A1DD466D9FBFC7560DBF9E7.text	03F36C399A1DD466D9FBFC7560DBF9E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bouchardium chillygonzalesi Kamiński 2024	<div><p>Bouchardium chillygonzalesi sp. nov.</p><p>(Figs 1, 4A, B, 5)</p><p>Type material. Holotype (NHMB), male: “Hartebeestpoort Dom m. Pretoria S.Afr.Leg.Zumpt —10.49” . Paratypes (NHMB): two specimens same data as holotype, single specimen: “Syringa S Rhodesia R. West [unreadable] 08”, female (MLPC): “ ZIMBABWE SE, Matobo National Park, 10–15.III.1999, leg. St. Roth ”, “coll. Lillig ex Naturkundemuseum Erfurt”, single specimen (MLPC) : “ Central Zimbabwe, Mvuma, route Gutu-Chatsworth, 24.XI.1998, M. Snizek leg.”, “coll. Lillig ex Zoologische Staatssammlung München” .</p><p>Diagnosis. Due to the similar color pattern of elytra (presence of two red stripes along the sides of the disc) this new species superficially resembles Bouchardium mariae sp. nov. (see diagnosis below). Nevertheless, Bouchardium chillygonzalesi sp. nov. can be distinguished from all its congeners by the presence of posteriorly projecting appendages on the prosternal process (Fig. 1E) and by having short c4 plates of ovipositor, which do not extend beyond the tip of c3 (Fig. 4A, B).</p><p>Description. Length 20.0–24.0 mm, width of pronotum 5.0 mm, and elytra 10.0–13.0 mm.</p><p>Head: Hypognathous. Frons finely punctate (~6 diameters apart); frontoclypeal suture coarse, presenting as deep groove in middle; apical clypeal margin not emarginate; labrum coarsely punctate laterally with shallow groove along apical margin bearing yellow, acuminate setae. Eyes comma-shaped, with ventral portion reduced, strongly emarginate around epistomal base. Antennae slender, relatively short (only slightly reaching beyond base of pronotum), bearing acuminate yellow setae; length of antennomere 2 equal to ~0.2 of antennomere 3; length of antennomere 4 about 0.6 of length of antennomere 3. Prothorax: Pronotum laterally with coarse, confluent punctures; widest in anterior half. Disc dull, finely punctate (4–5 diameters apart); anterior angles slightly produced. Hypomeron convex, dull, and impunctate. Prosternal process with pair of appendages, with wide bordering (ventral view). Pterothorax: Scutellum apically covered with dense setation. Elytra widest in half, disc relatively flat, sharply angled below margin; disc dark brown, impunctate, without tubercles; edges of disc red, with corrugated borders. Elytral slope steep, impunctate, elytral apex rounded. Epipleura impunctate and without tubercles, clearly differentiated from neighbouring portion of elytra and encircling ventrite 5. Lateral regions of metaventrite (between coxae) extremely short. Metaepisternal suture abbreviated posteriorly. Legs: Densely covered in golden setae, slender. Tarsi narrow, covered with golden setation. Abdomen: Ventrites 1–4 extremely finely and sparsely punctate laterally (&gt;5 diameters apart); ventrite 5 with denser and coarser punctuation (3–4 diameters apart).</p><p>Etymology. This species is named in honor of Jason Charles Beck (20 March 1972)—professionally known as Chilly Gonzales—Grammy-winning pianist and composer from Canada.</p><p>Distribution. South Africa (Provinces: Gauteng), Zimbabwe (Fig. 5).</p></div>	https://treatment.plazi.org/id/03F36C399A1DD466D9FBFC7560DBF9E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kamiński, Marcin Jan	Kamiński, Marcin Jan (2024): New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel. Zootaxa 5446 (1): 77-87, DOI: 10.11646/zootaxa.5446.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5446.1.4
03F36C399A1CD463D9FBF91167F9FED0.text	03F36C399A1CD463D9FBF91167F9FED0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bouchardium mariae Kamiński 2024	<div><p>Bouchardium mariae sp. nov.</p><p>(Figs 2, 4C, 5)</p><p>Type material. Holotype (NHMB), male: “ Francistown Bechuanaland Zumpt XII.55 ” . Paratypes (NHMB): four specimens: “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.934166&amp;materialsCitation.latitude=-20.2725" title="Search Plazi for locations around (long 30.934166/lat -20.2725)">Kanye Bechuanaland Zumpt</a> XII.55 ” , single specimen: “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.934166&amp;materialsCitation.latitude=-20.2725" title="Search Plazi for locations around (long 30.934166/lat -20.2725)">Zoutpan, Pta</a> 9/XII/1926 G. v. Son, Dichtha incantatoris” , female: “Museum Paris Rhodesia du Sud Selukwe, A. Ellenberger 1915”, single specimen: “Museum Paris Matebele Penda-ma-tenka E. Holub 170-94”, male a1 “ Rhodesia ” , male (MIZ PAN): “ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.934166&amp;materialsCitation.latitude=-20.2725" title="Search Plazi for locations around (long 30.934166/lat -20.2725)">Mozambique</a> ”, “ MIZ PAN COL011552 ”, “ Dichtha cubica guer ”, Dichtha cubica Guer. H. Gebien det. 1939” , male, female, and single specimen (MLPC): “ ZIMBABWE Great Zimbabwe ca. 20°16’21”S 30°56’03”E, 20.IV.1990, leg. Martin Lillig” coll. Lillig ” , two specimens (MLPC): “ ZIMBABWE centr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.934166&amp;materialsCitation.latitude=-20.2725" title="Search Plazi for locations around (long 30.934166/lat -20.2725)">MASVINGO lake MUTIRIKWI</a>, 25.11.2998, M. Snížek leg.”, “coll. Lillig ex Zoologische Staatssammlung München” , single specimens (MLPC): “RSA (Limpopo) Farm Zulani 1000 m ca. 20 km s. Vivo 3/ 5.II.2014, Heinz leg.”, “coll. Lillig ex Naturkundemuseum Erfurt”, “collection NATURKUNDE-MUSEUM ERFURT” .</p><p>,</p><p>“</p><p>Diagnosis. Due to the similar color pattern of elytra (presence of two red stripes along the sides of the disc) this new species resembles Bouchardium chillygonzalesi sp. nov. These two species can be easily distinguished by the different structure of prosternal process (rounded in lateral view in B. mariae; with appendages in B. chillygonzalesi — Figs. 1E, 2G). Furthermore, both species differ in the elytral shape and structure (elytra wide with almost smooth borders of the disc in B. mariae, elytra narrow with corrugated disc borders in B. chillygonzalesi —</p><p>Figs 1A–C, 2A–D). Besides the different coloration of elytra, B. mariae differs from B. cubicum by having a more prominently convex elytral disc (Figs 2, 3E–F).</p><p>Description. Length 19.0–25.0 mm, width of pronotum 7.0 mm, and elytra 12.0–14.0 mm. Head: Hypognathous. Frons finely punctate (~6 diameters apart); frontoclypeal suture coarse, presenting as deep groove in middle; apical clypeal margin not emarginate; labrum coarsely punctate laterally with shallow groove along apical margin bearing yellow, acuminate setae. Eyes comma-shaped, with ventral portion reduced, strongly emarginate around epistomal base. Antennae slender, elongate (reaching beyond base of pronotum), bearing acuminate yellow setae; length of antennomere 2 equal to ~0.2 of antennomere 3; length of antennomere 4 about half antennomere 3. Prothorax: Pronotum laterally with coarse, confluent punctures; widest in anterior half. Disc dull, finely punctate (4–5 diameters apart); anterior angles slightly produced. Hypomeron convex, dull, and impunctate. Prosternal process rounded in lateral view, with wide bordering (ventral view). Pterothorax: Scutellum apically covered with dense setation. Elytra widest in basal third, disc convex, sharply angled below margin; disc black, impunctate, without tubercles; edges of disc red. Elytral slope steep, impunctate, elytral apex rounded. Epipleura impunctate and without tubercles, clearly differentiated from neighbouring portion of elytra and encircling ventrite 5. Lateral regions of metaventrite (between coxae) extremely short. Metaepisternal suture abbreviated posteriorly. Legs: Densely covered in golden setae, slender. Tarsi narrow, covered with golden setation. Abdomen: Ventrites 1–4 extremely finely and sparsely punctate laterally (&gt;5 diameters apart); ventrite 5 with denser and coarser punctuation (3–4 diameters apart).</p><p>Etymology. This species is named in honor of my daughter, Maria Antonina Kamińska, born on June 9, 2016 (Warsaw, Poland).</p><p>Distribution. Botswana, South Africa (Provinces: Gauteng), Mozambique, Zimbabwe (Fig. 5).</p></div>	https://treatment.plazi.org/id/03F36C399A1CD463D9FBF91167F9FED0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kamiński, Marcin Jan	Kamiński, Marcin Jan (2024): New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel. Zootaxa 5446 (1): 77-87, DOI: 10.11646/zootaxa.5446.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5446.1.4
03F36C399A19D463D9FBF98A6186F823.text	03F36C399A19D463D9FBF98A6186F823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toktokkus Kaminski & Gearner 2021	<div><p>Toktokkus Kamiński &amp; Gearner, 2021</p><p>Notes. The most recent taxonomic contributions concerning Toktokkus were published by Kamiński et al. (2021), Gearner et al. (2022), and Kamiński (2022).</p><p>Species and subspecies included (16). T. barclayi Kamiński &amp; Gearner, 2021, T. congolensis Kamiński &amp; Gearner, 2021, T. herero Gearner, 2021, T. makuya Gearner, 2021, T. mariae Kamiński, 2022, T. mashunus (Péringuey, 1896), T. mulleri (Péringuey, 1899), T. o-serraferrus Gearner, Lumen, Kamiński, 2022, T. schultzei (Péringuey, 1908), T. tschinkeli Kamiński &amp; Gearner, 2021, T. tuberculipennis (Haag-Rutenberg, 1871), T. vialis vialis (Burchell, 1822), T. vialis sellatus (Haag-Rutenberg, 1875), T. vialis tuberculifer (Haag-Rutenberg, 1871), T. waclawae Kamiński, 2021, T. zofiae sp. nov.</p></div>	https://treatment.plazi.org/id/03F36C399A19D463D9FBF98A6186F823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kamiński, Marcin Jan	Kamiński, Marcin Jan (2024): New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel. Zootaxa 5446 (1): 77-87, DOI: 10.11646/zootaxa.5446.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5446.1.4
03F36C399A18D46DD9FBFACE6745F843.text	03F36C399A18D46DD9FBFACE6745F843.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Toktokkus zofiae Kamiński 2024	<div><p>Toktokkus zofiae sp. nov.</p><p>(Fig. 6A–F)</p><p>Type material. Holotype (NHMB), female: “ V.-L. Kai. Exp. Gemsbok Pan. 23/4–5/5/30.”, “ Psammodes peezi K.”.</p><p>Diagnosis. The species is most similar to T. herero Gearner, 2021 by having prominent punctures on the pronotal disc and a golden setation of the elytral disc (Kaminski et al. 2021). Both species can be distinguished by different body size ( T. zofiae: 18.5 mm, T. herero: 23.0–25.0 mm), basal margination of pronotal disc ( T. zofiae: complete, T. herero: interrupted in the middle) (Fig. 6D), structure of prosternal process ( T. zofiae: rounded in lateral view, T. herero: angular in lateral view) (Fig. 6H, E), and tuberculation of elytral disc ( T. zofiae: microtubercles present, T. herero: microtubercles absent) (Fig. 6A, G).</p><p>Description (based on holotype). Length 18.5 mm, width of pronotum 7.0 mm, and elytra 11.0 mm.</p><p>Head: Hypognathous. Frons finely and irregularly punctate; frontoclypeal suture coarse, with deep groove in middle; apical clypeal margin broadly shallowly emarginate; clypeus projected toward front of body. Eye comma-shaped, with reduced ventral part, strongly emarginate around epistomal base; with deep groove on temporal side. Mentum and submentum missing due to dermestid damage. Antennae broken off at level of 4th segments; slender, moderately covered in recumbent acuminate goldish setae; antennomere 2 short, equal to 0.2 of antennomere 3 length; antennomere 4 about half of antennomere 3 length.</p><p>Prothorax: Pronotal lateral margin rounded, well visible. Pronotum widest above middle. Disc dull, finely punctate (4–5 diameters apart); anterior and basal margins complete, anterior apices slightly produced. Hypomeron convex, without submarginal groove, impunctate. Prosternal process rounded in lateral view, longitudinally depressed in middle (ventral view). Anterior margin of prosternum straight, with gold setae.</p><p>Pterothorax: Scutellum densely covered with microtubercles. Elytra widest in half, slightly rounded, covered in scattered gold setae; covered with dense tubercles (1–3 diameters apart) and microtubercles (2–4 diameters apart). Laterally situated tubercles very pronounced. Elytral slope relatively steep, elytral apex flattened. Epipleura impunctate, not tuberculated, clearly distinguishable from neighbouring portion of elytra, widely enfolding fifth ventrite. Mesoventrite with deep median groove and elevated sides. Metaventrite impunctate, densely setose. Lateral regions of metaventrite (between coxae) extremely short. Metaepisternal suture abbreviated posteriorly.</p><p>Legs: Covered with dense gold setae. Procoxa exposed basally. Apex of protibia with denticle on outer margin, lateral carina terminating in basal third. Spurs on meso- and metatibiae of equal length. Tarsi narrowed laterally.</p><p>Abdomen: Ventrites 1–3 medially covered with goldish setae, moderately punctate (2–3 diameters apart) and weakly rugulose; ventrites 4–5 finely punctate (1–3 diameters apart); ventrite 5 without submarginal sulcus.</p><p>Terminalia: Due to scarcity of materials, terminalia were not dissected. Nevertheless, the tip of the ovipositor was visible on the analyzed specimen. The morphology of coxites is consistent with the bauplan reported for the genus (Kamiński et al. 2021).</p><p>Etymology. This species is named in honor of my daughter, Zofia Irena Kamińska, born on November 3, 2018 (Flagstaff, USA).</p><p>Distribution. South Africa —only known from the type locality (Gemsbok Pan).</p></div>	https://treatment.plazi.org/id/03F36C399A18D46DD9FBFACE6745F843	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kamiński, Marcin Jan	Kamiński, Marcin Jan (2024): New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel. Zootaxa 5446 (1): 77-87, DOI: 10.11646/zootaxa.5446.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5446.1.4
