taxonID	type	description	language	source
03F38788FFD3FFA3FF3FFE294ED02815.taxon	type_taxon	TYPE SPECIES. — Potamon villosum Yeo & Ng, 1998, by original designation; gender neuter.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD3FFAFFF20FDEB4AEA2DDD.taxon	description	(Figs 2; 3; 4)	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD3FFAFFF20FDEB4AEA2DDD.taxon	materials_examined	TYPE MATERIAL. — Holotype. Myanmar • ♂ (29.3 × 23.6 mm); Kayin state: Myawaddy district: “ Sukli ” [Su Ka Li], eastern side of Dawna Hills, near Myanmar-Thailand border; [c. 16 ° 9 ’ 18 ” N, 98 ° 35 ’ 52 ” E]; alt. 366 m; no date; F. H. Gravely leg.; NHM 1934.1.15.1 (“ Exch: Indian Museum ” “ 9770 / 10 ”). OTHER MATERIAL. — Myanmar • ♀ (31.3 × 25.2 mm); same data as for holotype; NHM 1934.1.15.2 (“ Exch: Indian Museum ” “ 9770 / 10 ”).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD3FFAFFF20FDEB4AEA2DDD.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3), relatively low (CH / CW = 0.4); dorsal surface gently convex in frontal view; anterolateral margins cristate, with distinct granules; front with broad anterior margin (FW / CW = 0.3); epigastric cristae well-developed, rugose, anterior to postorbital cristae, separated from postorbital cristae by short groove; postorbital cristae well-developed, relatively sharp, not reaching lateral margin; external orbital angle triangular, outer margin short; epibranchial tooth low, blunt; cervical grooves shallow, narrow, discontinuous, not reaching to level of postorbital cristae; epibranchial region with closely spaced granules; suborbital margin joining with supraorbital margin; epistome posterior margin with well-developed, triangular median tooth (Fig. 2 A-C). Third maxilliped exopod with short flagellum, shorter than width of merus (Fig. 3 A). Chelipeds rugose, subequal, carpus with prominent, narrow, triangular inner distal major tooth and low subbasal tooth (Figs 2 A; 3 C). Ambulatory legs short, stout, with short setae on margins (Fig. 2 A). Male s 2 / s 3 deep, reaching lateral margins; male s 3 / s 4 indiscernible (Figs 2 D; 3 D). Male sternopleonal cavity long, reaching to imaginary line joining medial part of cheliped coxae (Figs 2 D; 3 D). Male pleon relatively narrow; pleonal somite 6 trapezoidal, broader than long (proximal width c. 1.8 × medial length), longer than pleonal somite 5, with almost straight lateral margins (Figs 2 D; 3 B). Male telson triangular, slightly longer than pleonal somite 6, relatively narrow (proximal width c. 1.2 × medial length), with gently concave lateral margins (Figs 2 D; 3 B). G 1 slender, with ultimate article gently curved outwards at angle of about 25 ° from longitudinal axis, tip blunt, reaching pleonal locking structure; ultimate article relatively slender, short, c. 0.3 × combined length of flexible zone and penultimate article, subcylindrical, distally abruptly narrow, lacking dorsal flap, penultimate article relatively slender, gently sinuous (Figs 3 D, E; 4 A-C). G 2 long, c. 1.2 × length of G 1; ultimate article long, c. 0.6 × length of penultimate article (Figs 3 F; 4 D). Female pleon together with telson broadly ovate (Fig. 3 G). Vulvae on s 6 positioned close to each other, ovate, opening mesially, relatively small, occupying 0.6 × length of s 6, touching s 5 / s 6, some distance from s 4 / s 5, laterally partially covered by protruding sternal cover, visible in ventral view (Fig. 3 H).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD3FFAFFF20FDEB4AEA2DDD.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Indochinamon beieri is known only from the type locality, i. e., Su Ka Li, eastern side of Dawna Hills, Myawaddy district, Kayin state, Myanmar (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD3FFAFFF20FDEB4AEA2DDD.taxon	discussion	REMARKS Pretzmann (1966 a: 5, 6), during his visit to the NHM between 1963 and 1964, briefly diagnosed a new species, Potamon beieri, from a single male specimen (23.5 mm CL) from “ Sukli, Dawane Hills, 1200 ft., 1934 coll. ”, characterizing it by a G 1 that was described as follows: “ Gonopoden schlank, Subterminalglied am Ende fast rechtwinkelig nach aussen abgewinkelt. Terminalglied setzt diese Richtung fort. Innenkante stark ausgebuchtet, so dass der Umriss des Gonopoden an einen Vogelkopf erinnert. Innenkante des Subterminalgliedes dicht behaart bis zur Hälfte, Aussenkante mit einer Reihe langer Borsten. ” [Gonopods slender, subterminal segment angled outwards almost at right angles at tip. Terminal member continues this direction. Inner edge strongly bulged, so that the outline of the gonopod resembles a bird’s head. Inner edge of subterminal segment densely haired up to halfway, outer edge with a row of long bristles.] (translation ours). No figures or catalogue number was provided in the original publication. Pretzmann (1966 b: 301, pl. 4, figs 12 - 15) subsequently described the species in greater detail with figures of the specimen and its supposed G 1. The G 1 figured (Pretzmann 1966 b: pl. 4, fig. 12) matches his original description (Pretzmann 1966 a: 5, 6). No catalogue number was provided by Pretzmann (1966 b), but the figures he provided of the dorsal, frontal and ventral views of the specimen (see Pretzmann 1966 b: pl. 4, figs 13 - 15) perfectly match a male specimen in a jar with the catalogue number NHM 1934.1.15.1 - 2, which also contains a female (31.3 × 25.2 mm) collected at the same time. There are no labels inside the bottle indicating the specimen (s) was examined by Pretzmann or that the male is a type. This male specimen, measuring 29.3 × 23.6 mm agrees well in the measurements, and has a distinctive small round scar on the right side of the cardiac region (Fig. 2 A, C; also see Pretzmann 1966 b: pl. 4, fig. 15). As such, we have no doubt this is the specimen Pretzmann (1966 a, b) designated and figured as the holotype male specimen of Potamon beieri. We here select the catalogue number NHM 1934.1.15.1 for the holotype male of Potamon beieri, with the number NHM 1934.1.15.2 assigned to the female. The female specimen is a not a type as it was not mentioned by Pretzmann (1966 a, b). The type locality of the specimen should be “ Sukli ” [Su Ka Li], eastern side of Dawna Hills (not Dawane Hills), and in the original label, it states that this site is near the Myanmar-Thailand border. Pretzmann (1966 b: 303, pl. 5, figs 16 - 18) also figured a specimen of “ Potamon andersonianum andersonianum ” (dorsal, frontal and ventral views only, no G 1) in his paper treating specimens from NHM, but no data was provided. The species, however, was not even mentioned by Pretzmann (1966 b) in the main text. We are certain this specimen is the one now catalogued in the NHM as “ Potamon andersonianum andersonianum ” under the catalogue number NHM 1909.5.1.1. Alcock (1910: 35) had originally identified this specimen as “ Potamon (Potamon) andersonianum var. rangoonense ”, who listed four males and two females under the “ Indian Museum ” [ZSIK] catalogue number 4115 / 4 from an unknown locality collected by “ Captain Butler ”. One of these male specimens was gifted to NHM and was catalogued as “ 1909.5.1.1 ”. The labels with the NHM specimen indicate it was determined by “ A. Alcock ”, “ ex 4115 / 4 ” and presented by the Indian Museum, and the locality was stated as “ Burma ” [present day Myanmar]. The same specimen (NHM 1909.5.1.1) was later examined and listed as “ Potamon beieri ” byBrandis (2000: 75) as being collected from Myanmar. Specimen NHM 1909.5.1.1, however, is unlikely to have been collected from Myanmar. The collector, “ Captain Butler ”, was almost certainly the Permanent Official assigned by the Lieutenant-Governor of Bengal in 1869 to manage Nagaland in northeastern India (see Johnstone 1896; Bhatia 2004), and likely the specimen was obtained from Nagaland instead. The specimen Pretzmann (1966 b: pl. 5, figs 16 - 18) figured matches NHM 1909.5. 1.1 in all respects. The problem with the identity of Potamon beieri is that the G 1 figured by Pretzmann (1966 b: pl. 4, fig. 12) does not belong to the holotype. The actual G 1 of the holotype of Potamon beieri (NHM 1934.1.15.1) is more slender and much straighter, with the ultimate article cylindrical and gently curved outwards (not bent at almost right angle to the penultimate article) (Figs 3 E; 4 A-C). The G 1 he figured (Pretzmann 1966 b: pl. 4, fig. 12), actually belongs to NHM 1909.5.1.1 (Fig. 8 C). Although the left G 1 s of both specimens had been detached and placed in separate vials in their respective jars (leading to the possibility they had been accidentally misplaced), it is fortunate that the right G 1 s are still attached to the specimens (Figs 3 D; 8 E) and confirm our interpretation. Pretzmann (1966 a, b), therefore, had confused the G 1 s of the two specimens. This observation is also aligned with the discussion earlier that specimen NHM 1909.5.1.1 was likely to have been collected from Nagaland or its environs rather than Myanmar; all the species we have seen with this distinctive form of G 1 have been from that area. The incorrect G 1 figured by Pretzmann (1966 b) for Potamon beieri has caused substantial confusion in the identity of this species. Brandis (2000) discussed the taxonomy of this species, identified three specimens each from Assam (SMF 2807), Naga Hills (NMB 951 a) and Myanmar (NHM 1909.5.1.1). Brandis (2000) followedPretzmann (1966 a, b) in characterizing the species as one with a sharply bent G 1 ultimate article and provided figures for the SMF 2807 male specimen, including its G 1 (see Brandis 2000: pl. 10, fig. 2 a-c). Brandis (2000: 78) observes: “ Especially characteristic for P. beieri is the terminal joint of the G 1. Here a true torsion of the terminal joint turns the whole joint at 180 ° in relation to the subterminal joint. Due to this torsion the original dorsal side appears ventrally. This special character does not occur in any other potamid crab as far as known. It must have a special function in copulation process that only could be studied with live specimens. ” Brandis (2000) noted that Bott (1966: 481, fig. 15; 1970: 163, pl. 38, fig. 35; pl. 47, fig. 31) – who also provided figures for the SMF 2807 male specimen – had misidentified the species as “ Ranguna rangoonense (Rathbun, 1904) ”. Türkay & Naiyanetr (1987), while citing Pretzmann (1966 b: pl. 4, fig. 12), observed that the G 1 of the SMF 2807 male specimen is very similar to that of Potamon beieri, with only difference in the angle between the ultimate and penultimate articles. Türkay & Naiyanetr (1987) argued that the said difference could be due to size differences and treated the SMF 2807 specimen as Ranguna beieri (Pretzmann, 1966), or possibly as Potamon (Potamon) pruinosum Alcock, 1909. Potamon (Potamon) pruinosum is entirely different from Potamon beieri or SMF 2807 specimen in G 1 structure, and the species has already been assigned to Stelomon Yeo & Naiyanetr, 2000 (see Yeo & Naiyanetr 2000: 1630, fig. 3 c). Neither Türkay & Naiyanetr (1987) nor Brandis (2000) correctly identified Potamon beieri as they were referring to the G 1 described or illustrated in Pretzmann (1966 a, b), which belonged to the NHM 1909.5.1.1 specimen and was not that of the holotype (NHM 1934.1.15.1). In fact, Brandis (2000: 68, 71) incorrectly included the NHM 1934.1.15.1 - 2 specimens from “ Sukli ” under “ Potamiscus andersonianus (Wood-Mason, 1871) ”. Recently, Mitra (2017) identified several specimens from Mizoram state of northeastern India as I. beieri. Those specimens from Mizoram also differ in G 1 morphology from that of the holotype of Potamon beieri. All the confusion regarding the identity of Potamon beieri was probably because the NHM 1934.1.15.1 specimen was not properly labelled as a type in the first place. The incorrect G 1 figure in Pretzmann (1966 b: pl. 4, fig. 12) is another major reason for the previous misunderstanding of what the species is. The actual identities of these misidentified specimens are now revealed. The NMB 951 a male specimen from Naga Hills has already been refereed to Ranguna rangoonensis (Rathbun, 1904) byNg & Yeo (2023). The NHM 1909.5.1.1 male probably from Nagaland, the SMF 2807 male from Assam and the specimens examined by Mitra (2017) from Mizoram are all assigned herein to a new genus, Capitamon n. gen. (see Remarks for Capitamon n. gen.). In the G 1 structure, I. beieri s. str. most closely resembles I. andersonianum (Wood-Mason, 1871) [Yunnan, southwestern China], I. chinghungense (Dai, Song, He, Cao, Xu & Zhong, 1975) [Yunnan, southwestern China], I. gengmaense (Dai, 1995) [Yunnan, southwestern China], I. khinpyae Ng & Win Mar, 2018 [northern Myanmar], I. kimboiense (Dang, 1975) [northern Vietnam], I. orleansi (Rathbun, 1904) [northern Vietnam], I. phongnha Naruse, Nguyen & Yeo, 2011 [central Vietnam], and I. prolatum (Brandis, 2000) [northern Thailand] due to the relatively shorter and slenderer ultimate article (Figs 3 E; 4 A-C; see Yeo & Ng 1998: fig. 2 i; Dai 1999: figs 86 (4); 98 (4); Brandis 2000: pl. 13, fig. 2 c; Naruse et al. 2011: figs 3 b; 9 b; Ng & Win Mar 2018: fig. 4 b; unpublished data on I. andersonianum). Indochinamon beieri is nevertheless distinguished from all the above species mainly by the gently curved ultimate article, i. e., curved outwards at an angle of about 25 ° from longitudinal axis of the G 1 (Figs 3 E; 4 A) (vs G 1 ultimate article relatively strongly bent at an angle of about 45 - 60 ° from longitudinal axis; see Yeo & Ng 1998: fig. 2 i; Dai 1999: figs 86 (4); 98 (4); Brandis 2000: pl. 13, fig. 2 c; Naruse et al. 2011: figs 3 b; 9 b; Ng & Win Mar 2018: fig. 4 b; unpublished data on I. andersonianum). In addition, I. beieri can be also distinguished from all of them, except for I. khinpyae, by the abruptly narrow tip of the G 1 ultimate article (Figs 3 E; 4 A-C; see Ng & Win Mar 2018: fig. 4 b) (vs G 1 ultimate article with a gradually narrow tip; seeYeo & Ng 1998: fig. 2 i; Dai 1999: figs 86 (4); 98 (4); Brandis 2000: pl. 13, fig. 2 c; Naruse et al. 2011: figs 3 b; 9 b; unpublished data on I. andersonianum). Indochinamon beieri can still be differentiated from I. khinpyae by the relatively slender penultimate article of the G 1 (Figs 3 E; 4 A-C) (vs G 1 penultimate article relatively stouter; see Ng & Win Mar 2018: fig. 4 b).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	description	(Figs 5; 6; 7; 8; 9; 10; 11; 12; 13; 14; 15; 16; 17) urn: lsid: zoobank. org: act: 4 E 204 ABF- 5 B 88 - 4 A 72 - A 41 E- 44571 B 31469 C	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	type_taxon	TYPE SPECIES. — Capitamon capitatum n. sp., by present designation; gender neuter.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3 - 1.4), relatively low (CH / CW = 0.4 - 0.5); dorsal surface smooth except for epigastric cristae, postorbital cristae, epibranchial region and lateral surfaces; anterolateral margins cristate, with distinct granules; front with broad anterior margin (FW / CW = 0.3); epigastric cristae well-developed, rugose, anterior to postorbital cristae, separated from postorbital cristae by short groove; postorbital cristae well-developed, relatively sharp, not reaching lateral margin; external orbital angle triangular, with short outer margin; epibranchial tooth low, blunt; cervical grooves deep, narrow; epibranchial region with closely spaced granules; suborbital margin not joining with supraorbital margin; epistome posterior margin with well-developed, triangular median tooth (Figs 5 A, B; 8 A; 9 A, C; 12 A, B; 15 A, B; see Pati et al. 2020: figs 1 a; 2 a, b). Antennae shorter than eye stalk; antennules long, folded in longitudinally broad fossae (Figs 5 B; 9 C; 12 B; 15 B; see Pati et al. 2020: fig. 2 b). Mandibular palp with three articles; terminal article simple, undivided (Figs 6 A; 13 A; 16 A). First, second maxillipeds each with long flagellum on exopod (Figs 6 A; 13 A; 16 A). Third maxilliped exopod slender, lacking flagellum or with relatively shorter flagellum (shorter than merus width) (Figs 6 B; 9 B; 13 B; 16 B; see Pati et al. 2020: fig. 2 c). Chelipeds rugose; carpus with prominent, sharp, triangular inner distal major tooth and low subbasal tooth (Figs 5 A; 6 C; 8 A; 9 E; 12 A; 13 C; 15 A; 16 C; see Pati et al. 2020: fig. 1 a). Ambulatory legs short, stout, with short setae on margins (Figs 5 A, C; 8 A; 12 A, C; 15 A, C; see Pati et al. 2020: fig. 1 a). Male s 2 / s 3 deep, reaching lateral margins; male s 3 / s 4 indiscernible or with shallow groove; male s 7 / s 8 interrupted by longitudinal medial groove of s 7 and s 8, lacking transverse ridge (Figs 5 C; 6 D; 8 B, E; 12 C; 13 D; 15 C; 16 D; see Pati et al. 2020: figs 1 b; 2 e). Pleonal locking mechanism with prominent tubercle on posterior submedial part of s 5 (Figs 6 D; 8 E; 13 D; 16 D; see Pati et al. 2020: fig. 2 e). Male sternopleonal cavity long, reaching to imaginary line joining medial part of cheliped coxae (Figs 5 C; 6 D; 8 B, E; 12 C; 13 D; 15 C; 16 D; see Pati et al. 2020: figs 1 b; 2 e). Male pleon relatively broad; pleonal somite 6 trapezoidal, broader than long (proximal width c. 1.9 - 2.2 × medial length) (Figs 5 C; 6 E; 8 B; 9 D; 12 C; 13 E; 15 C; 16 E; see Pati et al. 2020: figs 1 b; 2 d). Male telson triangular, slightly longer than pleonal somite 6 (Figs 5 C; 6 E; 8 B; 9 D; 12 C; 13 E; 15 C; 16 E; see Pati et al. 2020: figs 1 b; 2 d). G 1 moderately stout, with ultimate article strongly bent outwards at angle of about 60 - 110 ° from longitudinal axis, tip acute, not reaching pleonal locking structure; flexible zone large; ultimate article stout, short, c. 0.35 × combined length of flexible zone and penultimate article, with distinct, narrow to broadly triangular dorsal flap, entire structure resembling bird’s head in shape; penultimate article sinuous (Figs 6 D, F; 7 A, C-E; 8 C, E; 10 A, B, D; 11 A, C-E; 13 D, F; 14 A, C-E; 16 D, F; 17 A, C-E; see Pati et al. 2020: figs 2 e; 4 a-d; 5 a-c). G 2 longer than G 1; ultimate article long, c. 0.5 - 0.6 × length of penultimate article (Figs 6 F, G; 7 B, D; 8 C, D; 10 A, C; 11 B, D; 13 F, G; 14 B, D; 16 F, G; 17 B, D; see Pati et al. 2020: figs 4 a, e; 5 a, d). Female pleon together with telson broadly ovate (Figs 6 H; 13 H; 16 H; see Pati et al. 2020: fig. 3 b). Vulvae on s 6 positioned close to each other, ovate, opening mesially, longitudinally broad, large, occupying 0.5 - 0.6 × length of s 6, touching s 5 / s 6, almost reaching s 4 / s 5 (Figs 6 I; 13 I; 16 I; see Pati et al. 2020: fig. 3 c).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	etymology	ETYMOLOGY. — All the known congeners of the new genus have an ultimate article of the male first gonopod that resembles a bird’s head. The genus name is therefore derived from caput, Latin for head, in arbitrary combination with the genus name Potamon. The gender of the generic name is neuter.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Capitamon n. gen. is currently known from northeastern India, with its nominal species recorded from Arunachal Pradesh, Manipur, Mizoram and Nagaland states (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDCFFADFEF6FF494BA32F38.taxon	discussion	REMARKS Capitamon n. gen. is undoubtedly a potamiscine genus (sensu Yeo & Ng 2004), with the s 7 / s 8 being interrupted by the longitudinal medial groove of s 7 and s 8, and lacking a transverse ridge (Figs 6 D; 8 E; 13 D; 16 D; see Pati et al. 2020: fig. 2 e). Capitamon n. gen. can be differentiated from other potamiscine genera by the flagellum on the third maxilliped exopod being relatively shorter (shorter than the width of the merus) or altogether absent, and significantly, the “ bird’s head-like ” ultimate article of the G 1, which is short, relatively stout, strongly bent outwards at an angle of about 60 - 110 ° from longitudinal axis of the G 1 and possesses a distinct dorsal flap (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; 13 F; 14 A, C-E; 16 F; 17 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c). As compared to that of Capitamon n. gen., the flagellum on the third maxilliped exopod is relatively longer (equal to or longer than the width of the merus), and the dorsal flap on the G 1 ultimate article is absent or relatively low in Indochinamon (cf. Ng & Win Mar 2018). The well-developed epigastric and postorbital cristae of Capitamon n. gen. (Figs 5 A; 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a; 2 a) are also useful to distinguish it from other potamiscine genera with similar conditions of the G 1 and the flagellum on the third maxilliped exopod. For example, Gempala Ng & Ahmad, 2016, from Peninsular Malaysia has a superficially similar short, relatively stout and strongly bent G 1 ultimate article with a distinct dorsal flap in addition to the absence of the flagellum on the third maxilliped exopod (see Ng & Ahmad 2016: fig. 7 a, e). Capitamon n. gen. cannot be confused with Gempala, however, because its epigastric and postorbital cristae are well-developed (Figs 6 A; 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a; 2 a) (vs poorly developed; see Ng & Ahmad 2016: fig. 3 a); the dorsal flap on G 1 ultimate article is comparatively low (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; 13 F; 14 A, C-E; 16 F; 17 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c) (vs conspicuously high; see Ng & Ahmad 2016: fig. 7 e); and G 1 penultimate article is relatively stouter (Figs 6 F; 7 D, E; 8 C; 10 A; 11 D, E; 13 F; 14 D, E; 16 F; 17 D, E; Pati et al. 2020: figs 4 a, d; 5 a, c) (vs relatively slender; see Ng & Ahmad 2016: fig. 7 e). Among the Indian potamiscines, Abormon Mitra, Pati & Ng, 2021, and Teretamon Yeo & Ng, 2007, also have a G 1 ultimate article with a distinct dorsal flap (see Yeo & Ng 2007: fig. 13 d; Absar et al. 2017: fig. 3 a; Mitra 2017: fig. 116; Mitra et al. 2018: fig. 4 a; 2021: figs 3 a; 7 a; Mitra & Pati 2021: fig. 4 a). Capitamon n. gen., however, is differentiated from Abormon and Teretamon mainly by the well-developed epigastric and postorbital cristae (Figs 5 A; 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a; 2 a) (vs epigastric and postorbital cristae poorly developed; see Yeo & Ng 2007: fig. 13 a; Absar et al. 2017: fig. 2 a; Mitra 2017: fig. 58; Mitra et al. 2018: fig. 2 a; 2021: figs 2 a; 6 a; Mitra & Pati 2021: fig. 2 a), and the relatively shorter G 1 ultimate article (Figs 6 F; 7 D, E; 8 C; 10 A; 11 D, E; 13 F; 14 D, E; 16 F; 17 D, E; see Pati et al. 2020: figs 4 a, d; 5 a, c) (vs G 1 ultimate article relatively longer; see Yeo & Ng 2007: fig. 13 d; Absar et al. 2017: fig. 3 a; Mitra 2017: fig. 116; Mitra et al. 2018: fig. 4 a; 2021: figs 3 a; 7 a; Mitra & Pati 2021: fig. 4 a). Moreover, the crabs of Capitamon n. gen. are relatively large in size (adult CW> 40 mm) than those of Abormon (adult CW <15 mm; see Mitra et al. 2021) and Teretamon (adult CW <27 mm; see Yeo & Ng 2007; Absar et al. 2017; Mitra 2017; Mitra et al. 2018; Mitra & Pati 2021; Pan et al. 2021). Some Indian species of Potamiscus Alcock, 1909 [P. chizami Pati, 2021, P. mima Pati, 2021, and P. takedai Pati, Mitra & Ng, 2020], like Capitamon n. gen., possess well-developed epigastric and postorbital cristae (Figs 5 A; 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a, c; 2 a, f; Pati 2021: fig. 2 a, g) in addition to a distinct but low dorsal flap on the G 1 ultimate article (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; 13 F; 14 A, C-E; 16 F; 17 A, C-E; see Pati et al. 2020: figs 4 a-d, f-i; 5 a-c, e-g; Pati 2021: fig. 3 a, b, e, f). Capitamon n. gen., however, is immediately differentiated from Potamiscus chizami and Potamiscus mima by the low carapace (Figs 5 B; 9 C; 12 B; 15 B; see Pati et al. 2020: fig. 2 b) (vs carapace conspicuously high; see Pati 2021: fig. 2 b, h) and the short ultimate article of the G 1 (Figs 6 F; 7 D, E; 8 C; 10 A; 11 D, E; 13 F; 14 D, E; 16 F; 17 D, E; see Pati et al. 2020: figs 4 a, d; 5 a, c) (vs G 1 ultimate article elongated; see Pati 2021: fig. 3 a, e). On the other hand, Capitamon n. gen. can easily be confused with Potamiscus takedai due to several characters in common, including the low carapace (CH / CW = 0.4 - 0.5) (Figs 5 B; 9 C; 12 B; 15 B; see Pati et al. 2020: fig. 2 b, g), the well-developed epigastric and postorbital cristae (Figs 5 A; 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a; 2 a, f), the deep male s 2 / s 3 reaching lateral margins (Figs 5 C; 6 D; 8 B, E; 12 C; 13 D; 15 C; 16 D; see Pati et al. 2020: figs 1 b, d; 2 e, j), the relatively broader male pleon with a broader pleonal somite 6 (Figs 5 C; 6 E; 8 B; 9 D; 12 C; 13 E; 15 C; 16 E; see Pati et al. 2020: figs 1 b, d; 2 d, i), the moderately stout G 1 with short, stout, strongly bent ultimate article (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; 13 F; 14 A, C-E; 16 F; 17 A, C-E; see Pati et al. 2020: figs 4 a-d, f-i; 5 a-c, e-g), the longer G 2 (Figs 6 F, G; 7 B, D; 8 C, D; 10 A, C; 11 B, D; 13 F, G; 14 B, D; 16 F, G; 17 B, D; see Pati et al. 2020: figs 4 a, e, f, j; 5 a, d, e, h), and the ovate, closely located vulvae touching the s 5 / s 6 and almost reaching the s 4 / s 5 (Figs 7 I; 13 I; 16 I; see Pati et al. 2020: fig. 3 c, f). Capitamon n. gen. is nevertheless separated from Potamiscus takedai by the narrow to broadly triangular dorsal flap on the G 1 ultimate article (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; 13 F; 14 A, C-E; 16 F; 17 A, C-E; see Pati et al. 2020: figs 4 a, b; 5 a, b) (vs G 1 ultimate article with a semicircular dorsal flap; see Pati et al. 2020: figs 4 f, g; 5 e, f), and the transversely broader and relatively small vulvae, occupying 0.5 - 0.6 times the length of the s 6 (Figs 6 I; 13 I; 16 I; see Pati et al. 2020: fig. 3 c) (vs vulvae longitudinally broader and relatively large, occupying 0.8 times the length of the s 6; see Pati et al. 2020: fig. 3 f). The specimens with “ bird’s head-like ” ultimate article of the G 1, hitherto incorrectly referred to Potamon beieri, Ranguna beieri, Potamiscus beieri (Pretzmann, 1966), or Indochinamon beieri by previous workers (see Remarks for Indochinamon beieri) actually belong to different species of Capitamon n. gen. The NHM 1909.5.1.1 male specimen probably from Nagaland illustrated in Pretzmann (1966 b: pl. 4, fig. 12; pl. 5, figs 16 - 18) is assigned here to a new species, C. clarki n. gen., n. sp. The specimens of I. beieri from Mizoram recorded in Mitra (2017) is recognized in the present study as C. mizoramense n. gen., n. sp. The SMF 2807 male specimen from Assam reported in Bott (1966: fig. 15; 1970: pl. 38, fig. 35; pl. 47, fig. 31) and Brandis (2000: pl. 10, fig. 2 a-c) is similar to Capitamon mizoramense n. gen., n. sp. in G 1 structure, but its dorsal flap on the ultimate article is comparatively low and broadly triangular (vs comparatively high and narrowly triangular; Figs 16 F; 17 A, C-E). As the SMF 2807 male could not be examined in this study, we are not sure about its conspecificity with C. mizoramense n. gen., n. sp. Furthermore, Indochinamon manipurense from Manipur is also assigned to Capitamon n. gen. since it has a distinct dorsal flap on the G 1 ultimate article that resembles a bird’s head (Fig. 11 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c). In addition, two more new species of Capitamon n. gen. are recognized herein, i. e., C. capitatum n. gen., n. sp. and C. meitei n. gen., n. sp. Capitamon n. gen. currently has five nominal species: C. capitatum n. gen., n. sp. (type species); C. clarki n. gen., n. sp.; C. manipurense (Alcock, 1909) n. comb.; C. meitei n. gen., n. sp.; and C. mizoramense n. gen., n. sp.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	description	(Figs 5; 6; 7) urn: lsid: zoobank. org: act: 66331159 - 75 CC- 4 D 8 E-B 15 A-B 65 CCD 51 A 361	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (49.5 × 37.3 mm); Nagaland: Tuensang district: Konya; 26 ° 15 ’ 36 ” N, 94 ° 51 ’ 35 ” E; alt. 1660 m; 20. VII. 2017; Khampong leg.; ZSI-WRC C. 2400. Paratypes. India • 2 ♂ (47.7 × 36.1 mm, 47.3 × 35.7 mm), 2 ♀ (49.6 × 38.4 mm, 45.1 × 34.9 mm); same data as for holotype; ZSI-WRC C. 2401. OTHER MATERIAL. — India • ♂, 2 ♀; Arunachal Pradesh: Anjaw district: Manchila; 28 ° 5 ’ 20 ” N, 96 ° 27 ’ 32 ” E; alt. 931 m; 22. XI. 2022; S. D. Gurumayum leg.; ZSI-WRC C. 2402 • ♂, ♀; Arunachal Pradesh: Anjaw district: Khupa, Hayuliang; 28 ° 4 ’ 26 ” N, 96 ° 28 ’ 58 ” E; alt. 690 m; 23. XI. 2022; S. D. Gurumayum leg.; ZSI-WRC C. 2403 • 5 ♂, 5 ♀; Arunachal Pradesh: Anjaw district: Knyao Stream, Hayuliang; 28 ° 4 ’ 15 ” N, 96 ° 34 ’ 8 ” E; alt. 677 m; 24. XI. 2022; S. D. Gurumayum leg.; ZSI-WRC C. 2404.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.4), relatively low (CH / CW = 0.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, straight in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves continuous, reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards laterally (Fig. 5 A, B). Third maxilliped exopod lacking flagellum or with vestigial flagellum (Fig. 6 B). Chelipeds rugose, unequal, inner distal tooth on carpus relatively broad (Figs 5 A, C; 6 C). Male s 3 / s 4 shallow, running from sternopleonal cavity to lateral edges of sternum (Figs 5 C; 6 D). Male pleonal somite 6 relatively broader (proximal width c. 2.0 × medial length), equal in length to pleonal somite 5, with almost straight lateral margins (Figs 5 C; 6 E). Male telson relatively broader (proximal width c. 1.3 × medial length), with strongly concave lateral margins (Figs 5 C; 6 E). G 1 ultimate article relatively less strongly bent at angle of about 75 ° from longitudinal axis, with sinuous outer margin, dorsal flap relatively low, broadly triangular (Figs 6 F; 7 A, C-E). G 2 c. 1.1 × length of G 1; ultimate article long, c. 0.5 × length of penultimate article (Figs 6 F, G; 7 B, D). Vulvae laterally completely covered by protruding sternal cover, invisible in ventral view (Fig. 6 I).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	etymology	ETYMOLOGY. — The specific epithet is an adjective in the Latin nominative singular meaning having a head, referring the head-like ultimate article of the male first gonopod of the type species.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Capitamon capitatum n. gen., n. sp. is known only from Arunachal Pradesh and Nagaland states of northeastern India (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFDDFFA9FC1FFADB4DAE2999.taxon	discussion	REMARKS Capitamon capitatum n. gen., n. sp. is unique among the nominal species of the genus because of the straight postorbital cristae in dorsal view (Fig. 5 A) (vs postorbital cristae oblique in dorsal view; Figs 8 A; 9 A; 12 A; 15 A; see Pati et al. 2020: figs 1 a; 2 a) and the vestigial or missing flagellum on the third maxilliped exopod (Fig. 6 B) (vs flagellum on the third maxilliped exopod relatively longer, equal to or more than half the width of the merus; Figs 9 B; 13 B; 16 B; see Pati et al. 2020: fig. 2 c). The G 1 ultimate article of C. capitatum n. gen., n. sp. is similar in structure to that of C. clarki n. gen., n. sp. in that it is bent at an angle of about 75 ° from longitudinal axis of the G 1, with the dorsal flap being relatively low and broadly triangular (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D). The outer margin of the G 1 ultimate article, however, is sinuous in C. capitatum n. gen., n. sp. (Figs 6 F; 7 A, C-E), whereas the outer margin of the said structure is straight in C. clarki n. gen., n. sp. (Figs 8 C; 10 A, B, D). The following differences between them can also be noted: the carapace is relatively low (CH / CW = 0.4) in C. capitatum n. gen., n. sp. (Fig. 5 B) against the relatively high carapace (CH / CW = 0.5) in C. clarki n. gen., n. sp. (Fig. 9 C); the male pleonal somite 6 is equal in length to the pleonal somite 5 in C. capitatum n. gen., n. sp. (Figs 5 C; 6 E) against the longer male pleonal somite 6 than the pleonal somite 5 in C. clarki n. gen., n. sp. (Figs 8 B; 9 D); and the lateral margins of the male telson is strongly concave in C. capitatum n. gen., n. sp. (Figs 5 C; 6 E) against the gently concave lateral margins of the male telson in C. clarki n. gen., n. sp. (Figs 8 B; 9 D).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	description	(Figs 8; 9; 10) urn: lsid: zoobank. org: act: B 9049 FED- 7231 - 43 FC-B 80 D- 7 ACFDF 268 F 42	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	materials_examined	TYPE MATERIAL. — Holotype. ♂ (42.1 × 32.9 mm); “ Burma ” [probably from Nagaland, India]; [c. 26 ° 0 ’ 0 ” N, 95 ° 0 ’ 0 ” E]; [alt. c. 1200 m]; no date; Captain Butler leg.; NHM 1909.5.1.1 (“ Presd: Indian Museum ” “ 4115 / 4 ”).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3), relatively high (CH / CW = 0.5); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves continuous, reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards laterally (Figs 8 A; 9 A, C). Third maxilliped exopod with short flagellum, equal to half width of merus (Fig. 9 B). Chelipeds rugose, subequal, inner distal tooth on carpus relatively broad (Figs 8 A; 9 E). Male s 3 / s 4 shallow, running from sternopleonal cavity to lateral edges of sternum (Fig. 8 B, E). Male pleonal somite 6 relatively narrower (proximal width c. 1.9 × medial length), longer than pleonal somite 5, with almost straight lateral margins (Figs 8 B; 9 D). Male telson relatively broader (proximal width c. 1.3 × medial length), with gently concave lateral margins (Figs 8 B; 9 D). G 1 ultimate article relatively less strongly bent at angle of about 75 ° from longitudinal axis, with straight outer margin, dorsal flap relatively low, broadly triangular (Figs 8 C; 10 A, B, D). G 2 c. 1.2 × length of G 1; ultimate article long, c. 0.5 × length of penultimate article (Figs 8 C, D; 10 A, C).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	etymology	ETYMOLOGY. — The species is named after English carcinologist Dr Paul Clark, the curator of Crustacea in the NHM, whose help with the collections was important in helping the authors solve the confusion with I. beieri. The species name is conceived as a noun in the genitive singular.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — The precise geographical distribution of C. clarki n. gen., n. sp. is not known (see Remarks for I. beieri). The species is possibly originated from Nagaland state of northeastern India (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFA9FE82FC6A4B932F1B.taxon	discussion	REMARKS Capitamon clarki n. gen., n. sp. has a relatively high carapace (CH / CW = 0.5) (Fig. 9 C) among the nominal species of the genus with the relatively low carapace (CH / CW = 0.4) (Figs 5 B; 12 B; 15 B; see Pati et al. 2020: fig. 2 b). While C. clarki n. gen., n. sp. most resembles C. capitatum n. gen., n. sp. in the structure of the G 1 ultimate article (see Remarks for C. capitatum n. gen., n. sp.), C. clarki n. gen., n. sp. also resembles C. manipurense n. comb. as both have a G 1 ultimate article with the straight outer margin and the relatively low, broadly triangular dorsal flap (Figs 8 C; 10 A, B, D; 11 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c). The G 1 ultimate article, however, is little more strongly bent at an angle of about 75 ° from longitudinal axis of the G 1 in C. clarki n. gen., n. sp. (Figs 8 C; 10 A), while it is little less strongly bent at an angle of about 60 ° from longitudinal axis of the G 1 in C. manipurense n. comb. (Fig. 11 D, E; see Pati et al. 2020: figs 4 a, d; 5 a, c). Capitamon clarki n. gen., n. sp. can be also distinguished from C. manipurense n. comb. by the shallow male s 3 / s 4, running from the sternopleonal cavity to the lateral edges of the sternum (Fig. 8 B, E) (vs male s 3 / s 4 indiscernible except for two short lateral clefts; see Pati et al. 2020: figs 1 b; 2 e), and the relatively broader male telson, proximal width c. 1.3 times the medial length, with the lateral margins gently concave (Figs 8 B; 9 D) (vs male telson relatively narrower, proximal width c. 1.2 times the medial length, with the lateral margins strongly concave; see Pati et al. 2020: figs 1 b; 2 d).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	description	(Fig. 11)	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	materials_examined	TYPE MATERIAL. — Lectotype. India • ♂ (39.9 × 30.0 mm); Manipur state: Manipur Hills; [c. 24 ° 50 ’ 41 ” N, 93 ° 54 ’ 14 ” E]; [c. alt. 986 m]; no date; H. H. Godwin-Austen leg.; ZSIK C. 6923 / 3.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	materials_examined	Paralectotype. India • ♀ (40.1 × 30.4 mm); same data as for lectotype; ZSIK C. 6923 / 3.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3), relatively low (CH / CW = 0.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves discontinuous, not reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards laterally (see Pati et al. 2020: figs 1 a; 2 a, b). Third maxilliped exopod with short flagellum, shorter than width of merus (see Pati et al. 2020: fig. 2 c). Chelipeds rugose, unequal, inner distal tooth on carpus relatively broad (see Pati et al. 2020: fig. 1 a). Male s 3 / s 4 indiscernible except for two short lateral clefts (see Pati et al. 2020: figs 1 b; 2 e). Male pleonal somite 6 relatively broader (proximal width c. 2.0 × medial length), longer than pleonal somite 5, with almost straight lateral margins (see Pati et al. 2020: figs 1 b; 2 d). Male telson relatively narrower (proximal width c. 1.2 × medial length), with strongly concave lateral margins (see Pati et al. 2020: figs 1 b; 2 d). G 1 ultimate article relatively less strongly bent at angle of about 60 ° from longitudinal axis, with straight outer margin, dorsal flap relatively low, broadly triangular (Fig. 11 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c). G 2 c. 1.3 × length of G 1; ultimate article long, c. 0.5 × length of penultimate article (Fig. 11 B, D; see Pati et al. 2020: figs 4 a, e; 5 a, d). Vulvae laterally partially covered by protruding sternal cover, visible in ventral view (see Pati et al. 2020: fig. 3 c).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Capitamon manipurense n. comb. is known only from Manipur state of northeastern India (see Pati et al. 2020) (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFD9FFB4FC95FAF64B012CBB.taxon	discussion	REMARKS Capitamon manipurense n. comb. can easily be distinguished from the nominal species of the genus by the relatively narrower male telson, proximal width c. 1.2 times the medial length (see Pati et al. 2020: figs 1 b; 2 d) (vs male telson relatively broader, proximal width c. 1.3 - 1.4 times the medial length; Figs 5 C; 6 E; 8 B; 9 D; 12 C; 13 E; 15 C; 16 E) and the relatively less strongly bent G 1 ultimate article, which is bent at an angle of about 60 ° from longitudinal axis of the G 1 (Fig. 11 D, E; see Pati et al. 2020: figs 4 a, d; 5 a, c) (vs G 1 ultimate article relatively more strongly bent at an angle of about 75 - 110 ° from longitudinal axis of the G 1; Figs 6 F; 7 D, E; 8 C; 10 A; 11 D, E; 13 F; 14 D, E; 16 F; 17 D, E). Otherwise, C. manipurense n. comb. is most similar to C. capitatum n. gen., n. sp. and C. clarki n. gen., n. sp. as their G 1 ultimate article has the relatively low and broadly triangular dorsal flap (Figs 6 F; 7 A, C-E; 8 C; 10 A, B, D; 11 A, C-E; see Pati et al. 2020: figs 4 a-d; 5 a-c).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	description	(Figs 12; 13; 14) urn: lsid: zoobank. org: act: 109 C 2 D 01 - A 922 - 4 EE 1 - ABCE- 710 D 7 E 91 F 2 F 2	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (51.7 × 38.8 mm); Manipur state: Thoubal district: Yairipok; 24 ° 39 ’ 53 ” N, 94 ° 3 ’ 50 ” E; alt. 784 m; 24. XI. 2018; Thangpurba Devi leg.; ZSI-WRC C. 2405. Paratypes. India • ♂ (39.9 × 30.4 mm), ♀ (36.0 × 27.4 mm); Manipur state: Kangpokpi district: Leimakhong; 24 ° 56 ’ 31 ” N, 93 ° 50 ’ 27 ” E; alt. 884 m; 27. V. 2019; Thangpurba Devi leg.; ZSI-WRC C. 2406.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3 - 1.4), relatively low (CH / CW = 0.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves continuous, reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards laterally (Fig. 12 A, B). Third maxilliped exopod with short flagellum, shorter than width of merus (Fig. 13 B). Chelipeds rugose, unequal in adult males, inner distal tooth on carpus relatively broad (Figs 12 A, C; 13 C). Male s 3 / s 4 indiscernible (Figs 12 C; 13 D). Male pleonal somite 6 relatively broader (proximal width c. 2.1 - 2.2 × medial length), equal in length to pleonal somite 5, with convex lateral margins (Figs 12 C; 13 E). Male telson relatively broader (proximal width c. 1.3 - 1.4 × medial length), with gently to strongly concave lateral margins (Figs 12 C; 13 E). G 1 ultimate article relatively less strongly bent at angle of about 75 ° from longitudinal axis, with sinuous outer margin, dorsal flap relatively high, narrowly triangular (Figs 13 F; 14 A, C-E). G 2 c. 1.2 × length of G 1; ultimate article long, c. 0.5 × length of penultimate article (Figs 13 G; 14 B, D). Vulvae laterally completely covered by protruding sternal cover, invisible in ventral view (Fig. 13 I).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	etymology	ETYMOLOGY. — The species name, meitei, means Manipuri, the people / language of Manipur. The name is used as a Latin noun in apposition.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Capitamon meitei n. gen., n. sp. is known only from Manipur state of northeastern India (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC4FFB0FC21F9204DBD2FDB.taxon	discussion	REMARKS Among the nominal species of the genus, C. meitei n. gen., n. sp. most resembles C. mizoramense n. gen., n. sp. mainly by the G 1 ultimate article, which has a sinuous outer margin, and the relatively high and narrowly triangular dorsal flap (Figs 13 F; 14 A, C-E; 16 F; 17 A, C-E). Capitamon meitei n. gen., n. sp. is nevertheless differentiated from C. mizoramense n. gen., n. sp. by the relatively less strongly bent G 1 ultimate article, which is bent at an angle of about 75 ° from longitudinal axis of the G 1 (Figs 13 F; 14 D, E) (vs G 1 ultimate article more strongly bent at an angle of about 110 ° from longitudinal axis of the G 1; Figs 16 F; 17 D, E). In addition, the vulvae are completely covered laterally by the protruding sternal cover and invisible in the ventral view in C. meitei n. gen., n. sp. (Fig. 13 I), whereas the vulvae are partially covered laterally by the protruding sternal cover and visible in the ventral view in C. mizoramense n. gen., n. sp. (Fig. 16 I).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	description	(Figs 15; 16; 17) urn: lsid: zoobank. org: act: 325 C 0 E 17 - 991 F- 43 A 7 - 9 D 11 - 48 CD 57 D 41299	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	materials_examined	TYPE MATERIAL. — Holotype. India • ♂ (46.3 × 35.7 mm); Mizoram state: Champhai district: Zokhawthar, c. 2 km from Indo-Myanmar International Border, collected from small stream connected to Tiau River; 23 ° 21 ’ 53 ” N, 93 ° 23 ’ 9 ” E; alt. 728 m; 23. IX. 2013; S. Mitra leg.; ZSIK C. 6122 / 2 A. Paratypes. India • ♂ (31.4 × 24.9 mm), ♀ (51.3 × 37.6 mm); same data as for holotype; ZSIK C. 6122 / 2 B • 7 ♂ (largest male, 60.0 × 44.6 mm), 4 ♀ (largest female, 52.0 × 39.6 mm); Mizoram state: Lunglei district: Tui-pui village, collected from small stream connected to Tui-pui River; 22 ° 52 ’ 47 ” N, 92 ° 56 ’ 9 ” E; alt. 228 m; 24. IX. 2013; S. Mitra leg.; ZSIK C. 6131 / 2 • 5 ♂ (largest male, 37.5 × 29.2 mm), 4 ♀ (largest female, 37.7 × 28.8 mm); Mizoram state: Serchhip district: near Vantawng waterfalls, about 9 km from Thenzawl market, collected from small water channel; 23 ° 14 ’ 2 ” N, 92 ° 45 ’ 3 ” E; alt. 720 m; 26. IX. 2013; S. Mitra leg.; ZSIK C. 6125 / 2.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.3), relatively low (CH / CW = 0.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves discontinuous, not reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards (Fig. 15 A, B). Third maxilliped exopod with short flagellum, shorter than width of merus (Fig. 16 B). Chelipeds rugose, unequal, inner distal tooth on carpus relatively narrow (Figs 15 A, C; 16 C). Male s 3 / s 4 indiscernible except for two short lateral clefts (Figs 15 C; 16 D). Male pleonal somite 6 relatively broader (proximal width c. 2.1 × medial length), equal in length to pleonal somite 5, with gently convex lateral margins (Figs 15 C; 16 E). Male telson relatively broader (proximal width c. 1.3 × medial length), with concave lateral margins (Figs 15 C; 16 E). G 1 ultimate article relatively more strongly bent at angle of about 110 ° from longitudinal axis, with sinuous outer margin, dorsal flap relatively high, narrowly triangular (Figs 16 F; 17 A, C-E). G 2 c. 1.3 × length of G 1; ultimate article long, c. 0.6 × length of penultimate article (Figs 16 F, G; 17 B, D). Vulvae laterally partially covered by protruding sternal cover, visible in ventral view (Fig. 16 I).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	etymology	ETYMOLOGY. — The specific epithet refers to Mizoram state in northeastern India, where the new species is found.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	description	COLOUR IN LIFE. — Crabs are olive green or brown, with paler ventral side and blood red joints of chelipeds (see Mitra 2017: fig. 71).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	biology_ecology	ECOLOGICAL NOTES (modified from Mitra 2017). — Capitamon mizoramense n. gen., n. sp. is a hill stream crab that prefers rocky habitats. This species is very common in the Champhai district of Mizoram. Local fishermen catch these crabs at night and sell them alive in the early morning for food. The population of this species may decline in the near future if these crabs are over-harvested.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	distribution	GEOGRAPHICAL DISTRIBUTION. — Capitamon mizoramense n. gen., n. sp. is known only from Mizoram state of northeastern India (Fig. 1).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFC0FFBFFED7F9AC4B2E2DDD.taxon	discussion	REMARKS Capitamon mizoramense n. gen., n. sp. is unique among congeners due to the G 1 ultimate article, which is relatively more strongly bent at an angle of about 110 ° from longitudinal axis of the G 1, and possesses the relatively high and narrowly triangular dorsal flap (Figs 16 F; 17 A, C-E). The remaining nominal species of Capitamon n. gen. have a G 1 ultimate article relatively less strongly bent at an angle of about 60 - 75 ° from longitudinal axis of the G 1 (Figs 6 F; 7 D, E; 8 C; 10 A; 11 D, E; 13 F; 14 D, E; see Pati et al. 2020: figs 4 a, d; 5 a, c). Capitamon mizoramense n. gen., n. sp. shares the following features of the G 1 with C. meitei n. gen., n. sp.: sinuous outer margin of the G 1 ultimate article and the relatively high, narrowly triangular dorsal flap on the G 1 ultimate article (Figs 13 F; 14 A, C-E; 16 F; 17 A, C-E). Other features of G 1 and vulvae, however, distinguish them (see Remarks for C. meitei n. gen., n. sp.).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFCCFFBCFEE3FF494BA9294A.taxon	materials_examined	MATERIAL EXAMINED. — None.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFCCFFBCFEE3FF494BA9294A.taxon	diagnosis	DIAGNOSIS. — Carapace ovate, broader than long (CW / CL = 1.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin, c. 2 × inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves continuous, reaching to level of postorbital cristae (see Brandis 2000: pl. 10, fig. 2 a). Chelipeds rugose, unequal, inner distal tooth on carpus relatively broad (see Brandis 2000: pl. 10, fig. 2 a). G 1 ultimate article relatively more strongly bent at angle of about 110 ° from longitudinal axis, with sinuous outer margin, dorsal flap relatively low, broadly triangular (see Brandis 2000: pl. 10, fig. 2 b, c).	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
03F38788FFCCFFBCFEE3FF494BA9294A.taxon	discussion	REMARKS The male specimen (SMF 2807) of an unknown species of Capitamon n. gen. reported by Bott (1966, 1970) and Brandis (2000) also possesses a G 1 ultimate article as strongly bent as that of C. mizoramense n. gen., n. sp. The dorsal flap on the G 1 ultimate article of the SMF 2807 male, however, is relatively low and broadly triangular (see Brandis 2000: pl. 10, fig. 2 b, c) against the relatively high and narrowly triangular dorsal flap of C. mizoramense n. gen., n. sp. (Figs 16 F; 17 A, C-E). In addition, the cervical grooves of the SMF 2807 male are continuous, which reach to the level of the postorbital cristae (see Brandis 2000: pl. 10, fig. 2 a); whereas those of C. mizoramense n. gen., n. sp. are not continuous and do not reach to the level of the postorbital cristae (Fig. 15 A). The SMF 2807 male specimen has some affinities with C. mizoramense n. gen., n. sp., but it is certainly not that species. The said SMF material is a distinct species of Capitamon n. gen. but is not described here as we could not access the actual material during this study. Fresh specimens of this species will also need to be collected.	en	Pati, Sameer K., Mitra, Santanu, Ng, Peter K. L. (2024): Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae). Zoosystema 46 (23): 589-615, DOI: 10.5252/zoosystema2024v46a23, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf
