taxonID	type	description	language	source
03F4C378A22C304BFD7E062AFBD4FD14.taxon	description	Trachyiulidae Silvestri, 1896 (recte: Trachyjulidae): 52. Glyphiulinae Chamberlin, 1922: 2.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A22C304CFC96058EFB67FB4D.taxon	materials_examined	TYPE SPECIES. — Iulus (recte: Julus) granulatus Gervais, 1847. SPECIES INCLUDED. — Below is a checklist of the species currently attributed to Glyphiulus, arranged in alphabetical order (after Jeekel 2004). In contrast, Mauriès & Nguyen Duy-Jacquemin (1997) referred to only 16 or 17 species as belonging to this genus, depending on synonymy. 1. G. adeloglyphus Zhang & Li, 1982, described from Xingping, Guangxi Province, China; still only known from the original description (Zhang & Li 1982). 2. G. anophthalmus (Loksa, 1960), described as Trogloglyphus anophthalmus Loksa, 1960 from Nyu-Jie Cave near Pulung (= Fulong, near the frontier with Vietnam), Guangxi Province, China; still only known from the original description (Loksa 1960). 3. G. balazsi (Loksa, 1960), described as Trogloglyphus balazsi Loksa, 1960 from Lodjen (= Luodian, or Longping), southern Guizhou Province, China; still only known from the original description (Loksa 1960). 4. G. capucinus Attems, 1938, described from Mt. Bana, Danang Province, central Vietnam (Enghoff et al. 2004); still only known from the original description (Attems 1938). 5. G. formosus (Pocock, 1895), described as Cambalomorpha formosa Pocock, 1895 from Hong Kong, China; known from the original description (Pocock 1895) and a partial redescription (Mauriès 1970). 6. G. granulatus (Gervais, 1847), described as Iulus (recte: Julus) (Glyphiulus) granulatus Gervais, 1847 from Mauritius and Réunion islands, Indian Ocean; currently known as a pantropical “ tramp ” species from Hawaii, Hong Kong, Marquesas, Society Islands, Cook Islands, Samoa, Loyalty Islands and New Caledonia in the Pacific Ocean, from the Mascarenes, Seychelles and Comoro Islands in the Indian Ocean, and from St. Helena in the Atlantic Ocean (Jeekel 2004). Cambala nodulosa Butler, 1876, from Rodrigues Island, Mascarene Islands, Formosoglyphius tuberculatus Verhoeff, 1936, from Taiwan, and Glyphiulus vulgatus Zhang & Li, 1982, from Longzhou, Guangxi Province, China, are junior subjective synonyms of this species (see below). 7. G. javanicus Carl, 1911, described from Passaroean, Java, Indonesia; still only known from the original description (Carl 1911). 8. G. lipsorum Mauriès & Nguyen Duy-Jacquemin, 1997, described and only known from a cave not in Hubei Province, China, as reported originally (Mauriès & Nguyen Duy-Jacquemin 1997), but in Guangxi Province (Fig. 30). 9. G. mediator Attems, 1938, described from Mt. Bana, Danang Province, central Vietnam (Enghoff et al. 2004); still only known from the original description (Attems 1938). 10. G. melanoporus Mauriès & Nguyen Duy-Jacquemin, 1997, described and only known from a cave near Guilin, Guangxi Province, China (Mauriès & Nguyen Duy-Jacquemin 1997). 11. G. pulcher (Loksa, 1960), described as Octoglyphus pulcher Loksa, 1960 from Nyu-Jie Cave near Pulung (= Fulong, near the frontier with Vietnam), Guangxi Province, China; still only known from the original description (Loksa 1960). 12. G. quadrohamatus Chen & Meng, 1991, described from several caves in the Zhenning Bouyzu Miaozu Autonomous County, Guizhou Province, China (Chen & Meng 1991). 13. G. rayrouchi Mauriès & Nguyen Duy-Jacquemin, 1997, described and only known from Hei Yan Cave near Maguan, Guizhou Province, China (Mauriès & Nguyen Duy-Jacquemin 1997). 14. G. recticullus Zhang & Li, 1982, described and still only known from Qinyuan, Zhejiang Province, China (Zhang & Li 1982). 15. G. septentrionalis Murakami, 1975, described from Okinawa Island, Ryukyus, Japan (Murakami 1975), but apparently introduced there from continental China, given that G. multicarinus Zhang & Li, 1982, described from near Guilin, Guangxi Province, China (Zhang & Li 1982), is its junior synonym (see below). 16. G. siamensis Mauriès, 1983, described and still only known from Doi Sutep, northern Thailand (Mauriès 1983). Enghoff (2005) records further material from the type locality. 17. G. superbus Silvestri, 1923, described from Dalat, Lamdong Province, central Vietnam (Attems 1938), later recorded from Kampot, Cambodia (Attems 1953), although the latter record needs confirmation (Enghoff et al. 2004). The carinotaxy formula as given in the original description is somewhat ambiguous (Silvestri 1923) and difficult to interpret (see key below). 18. G. vietnamicus Mauriès, 1977, described and only known from a cave in Ke Bang karst, Quang-Binh Province, Vietnam (Mauriès 1977; Enghoff et al. 2004). 19. G. zorzini Mauriès & Nguyen Duy-Jacquemin, 1997, described and still only known from Anjia Yan Cave, Shuichang County, Guizhou Province, China (Mauriès & Nguyen Duy-Jacquemin 1997).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A22B3046FCA90270FEDAFB79.taxon	discussion	REMARKS Species of the granulatus- group are distinguished by the following characters: 1. Male legs 1 are usually very strongly reduced, represented solely or mainly by a sternum lacking any median structures but bearing strongly separated, evidently curved prongs with a leg vestige on each side at base (usually represented by just a few setae, more seldom also one or two rudimentary segments). Very seldom, a nearly complete telopodite persists, but the sternum still supports a pair of widely separated and curved prongs. [Male legs 1 with central structures on the sternum in other groups]. 2. Other male legs normal or nearly so, not enlarged. [Male legs 2 sometimes incrassate in other groups]. 3. A typical carinotaxy pattern of the collum is I- VI + 7 a + pc + ma + pc + 7 a + VI-I. Quite often the pattern is different, either due to crest reduction (especially anteriorly) or hypertrophy, but a median crest is always traceable, at least near the caudal margin. [Usually typical carinotaxy formulae of the collum are different in other groups]. 4. A typical carinotaxy pattern of the metaterga is 3 (2) + I + 3 + I + 3 (2). The crests usually being divided transversely into two halves, while the median crest is often doubled anteriorly, the formula becomes 3 (2) + I + 4 + I + 3 (2) and 3 (2) + i + 3 + i + 3 (2). If the pattern is different, it is usually readily derived from the typical one (or vice versa), i. e. the lateral crests are reduced to two or the crests are undivided, or some of the crests are divided into three, rather than two, transverse rows of tubercles (see also below), etc. A median crest, even when strongly reduced, is always present as well. [Usually typical carinotaxy formulae of the metaterga are different in other groups]. 5. The anterior gonopods are reduced to a plate-like coxosternum with moveable, lateral, 1 - segmented telopodites. [Shared with some other species groups]. 6. The posterior gonopods are highly compressed, showing a subflagelliform (rarely subspatuliform), often plumose, distal process. [Shared with some other species groups]. 7. The pleural flaps behind the gonopod opening on male segment 7 usually do not form an apparent transverse ventral ridge. [Shared with some other species groups]. The group thus roughly corresponds to the concept of Glyphiulus in the sense of Verhoeff (1936) and Loksa (1960). At present it contains the following species: G. adeloglyphus, G. anophthalmus, G. balazsi, G. capucinus, G. granulatus, G. lipsorum,	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A22B3046FCA90270FEDAFB79.taxon	description	G. melanoporus, G. quadrohamatus, G. rayrouchi, G. septentrionalis, G. superbus, as well as all nine new species described below. The remaining known cambalopsid species are considered as belonging to other groups and will be treated elsewhere. Below a description is given of the available material, followed by a key to all of the constituent species of the granulatus - group, as well as a brief analysis of their variation and distribution.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A22B3046FCA90270FEDAFB79.taxon	materials_examined	TYPE MATERIAL. — Réunion (= Île Bourbon). La Bonite, 1838, leg. Eydoux, lectotype ♀ (incomplete, lacking caudal body end), 5 ♀♀ paralectotypes (all fragmented), all here designated (MNHN GA 013); material in rather poor condition (most legs missing), apparently dried before being placed in alcohol. Mauriès (1983) mentioned that only part of the type material of this species is still preserved in the MNHN, with the series from Île de France (= Mauritius) presumably being lost. The lectotype selection made here is necessary to fix the exact type locality (Réunion Island). TYPE LOCALITY. — Réunion. OTHER MATERIAL EXAMINED. — Réunion. 10. II. 1972, leg. J. Travé, strict topotype juv. ♀ (MNHN GA 022). Mauritius. Near Mahébourg, sugar cane debris, 6. II. 1972, leg. J. Travé, 1 ♀ (det. J. - P. Mauriès, MNHN GA 022). Comoros. Mayotte Island, Mt. Combani, 12 ° 48 ’ S, 045 ° 08 ’ E, 470 m, Winkler extraction of forest litter, 22 - 24. II. 1999, leg. Jocqué & De Smet, 3 ♂♂, 1 ♀, 1 juv. (ZMUM, ex coll. MRAC No. 017.940), several ♂♂ and ♀♀ (SEM). China. Hong Kong, leg. M. Crosland, det. J. - P. Mauriès, 2 ♀♀ (MNHN GA 013). Fiji. Viti Levu Island, Suva, park, in litter, 5. VI. 1980, leg., det. & don. S. Golovatch, 1 ♂ (MNHN GA 013). Seychelles. Mahé Island, 1892, leg. C. Alluaud, det. H. W. Brölemann, 1 complete ♀, 1 incomplete ♀ (MNHN GA 013). New Caledonia. Dumbea, leg. Mme Pruvot, det. H. W. Brolemann, 4 ♂♂, 2 ♀♀ (MNHN GA 013). Loyalty Islands. Lifou, leg. Mme Pruvot, det. H. Brolemann, 1 incomplete ♀ (MNHN GA 013). DIAGNOSIS. — Differs from congeners primarily by the peculiar anterior gonopods, each showing an unusually high median outgrowth of the coxosternum, combined with the typical patterns of carinotaxy. Review of Glyphiulus: the granulatus - group (Diplopoda, Cambalopsidae) DESCRIPTION Length of adults of both sexes 10 - 18 mm, midbody segments round in cross-section (Fig. 3 B), their width (horizontal diameter) and height (vertical diameter) similar, 0.6 - 0.9 mm. Coloration yellowbrown; non-faded specimens variegated, with a dark brown vertex, blackish ocellaria, mainly brown crests on collum and brownish lateral longitudinal stripes beginning from dark brown ozoporiferous tubercles. Sometimes a thin axial line traceable due to darker median crests. Adult body with 30 - 53 podous segments + 4 - 2 apodous ones + telson (formula 30 - 38 p + 4 a + T in smaller specimens, up to 44 - 53 p + 2 a + T in larger ones; formulae generally following Enghoff et al. 1993). Clypeus with 3 - 6 teeth anteromedially. Eye patches transversely ribbon-shaped (smaller specimens) to ovoid (larger individuals), each composed of 11 - 18 rather flat ocelli in 5 or 6 irregular longitudinal rows (Fig. 4 A). Antennae short and clavate (Figs 2 B; 3 A; 4 A), antennomeres 6 and 7 with a small distodorsal field or corolla of bacilliform sensilla (Fig. 4 B, C). Gnathochilarium usually, but not always (n = 10: 1), with a separate promentum (Fig. 3 A). Head width = segment 2 <collum = midbody segments (close to 18 th to 20 th)> segment 3 = 6> 4 = 5 <7 <8 = 10; body abruptly tapering toward telson on a few posteriormost segments. Postcollar constriction very evident (Fig. 2 C). Collum with 7 + 7 longitudinal crests starting from fore edge, but both median crests interrupted in about caudal 1 / 3, being replaced there by similar 1 + 1 + 1 crests (Fig. 2 A-C) (formula I- VI + 7 a + pc + ma + pc + 7 a + VI-I). Subsequent metaterga similarly strongly crested (Fig. 2 B-E), especially so from segment 5 onwards, whence enlarged pore-bearing tubercles commence, less evident on legless segments in front of telson due to loss of ozopores (Fig. 2 E). Ozoporiferous tubercles round, wider than high; midbody metatergal crests divided into two about midway, each half neither especially high nor sharp (Fig. 2 B-F). Carinotaxy formulae 3 + I + 4 + I + 3 and 3 + i + 3 + i + 3, the former standing for front row of crests, the latter for caudal one, both fairly independent (Figs 2 D-F; 3 B). Tegument delicately alveolate-areolate, dull throughout. Fine longitudinal striations in front of stricture between pro- and metazona, remaining surface of prozona very delicately shagreened. Metatergal setae absent. Segment 2 with long pleural flaps (Fig. 2 B). Limbus extremely finely and more or less regularly denticulate. Epiproct simple, devoid of tuberculation, like a transverse rounded ridge in caudal part (Figs 2 E, F; 3 C, D). Paraprocts rather regularly convex, each with a row of several setae at medial margin and 2 + 2 setae more laterally (Fig. 3 C, D). Hypoproct rather evidently but broadly emarginate caudally to receive ventral edges of paraprocts, with 1 + 1 strongly separated setae near caudal margin (Fig. 3 C). Ventral flaps behind gonopod opening on male segment 7 barely distinguishable as low swellings, not forming a marked transverse ridge (Fig. 3 E, F). Legs short, on midbody segments about 3 / 4 length of segment height (Figs 2 B, D, E; 3 B). Claw at base with a strong accessory spine almost half as long as claw itself (Fig. 5 D). Tarsi characteristically fringed terminally, some of terminal setae with scattered denticles (Fig. 5 D). Male legs 1 highly characteristic (Figs 4 D; 6 A) in being very strongly reduced, represented only by a sternum devoid of any median or paramedian structures but carrying 1 + 1 strongly separated prongs, both evidently curved posteriad and bearing several strong setae and a minute tubercle (vestige of legs) at base on caudal face. Male legs 2 very slightly hypertrophied, only claw and, anteriorly, coxa somewhat reduced; penes broad, oblong-subtrapeziform, each with 2 or 3 strong setae distolaterally (Figs 4 E; 6 B). Male legs 3 modified in having coxa especially slender and elongate (Fig. 4 F). Anterior gonopods (Figs 5 A, B; 6 C) with a typical shield-like coxosternum which is modestly setose on caudal face and provided with a pair of con- spicuous, high, terminally coiled, mesal processes (visible also in situ, Fig. 3 E). Telopodite small but movable, 1 - segmented, lateral in position, with 2 or 3 strong apical setae and a field of small setae at base, slightly longer than adjacent lateral corner of coxosternum. Posterior gonopods (Figs 5 C; 6 D) very compact, coxite medio-apically with a plumose distal process and a hyaline lobe, telopodite setose both laterally and medially, lower than both coxal process and lobe. Vulvae very simple, bare, modestly emarginate medially (Fig. 6 G, H). Female legs 1 and 2 as in Figure 6 E, F, female coxa 3 as slender as in male (Fig. 6 G). Early juvenile stadia recognised by normal, large ozoporiferous tubercles on segments 5 and 6, these tubercles being considerably reduced in size on subsequent segments. REMARKS The above is the first formal record of G. granulatus in Fiji. Based on the original description and illustrations of G. vulgatus alone (Zhang & Li 1982), it is clear that this taxon does not differ in any way from G. granulatus, thus justifying the above new synonymy. The far-inland record of G. granulatus (= G. vulgatus) in Guangxi Province, southern China suggests a continental Chinese origin of this species, which has since become established on numerous tropical islands all over the globe. The port of Hong Kong seems to have been a plausible outlet for the recent expansion of G. granulatus through human agency. The fact that, of all the numerous congeners, only this species that has attained such a vast distribution seems to be rooted in its biology and ecology, the most relevant traits being its fairly small size (on average the smallest) and, apparently, a relatively rapid development, suggested indirectly by the observations of Enghoff (1993) and Mauriès & Nguyen Duy-Jacquemin (1997). The ontogeny of certain cambalopsids is unique among Diplopoda in containing a haplopodous early juvenile stage (third?). At this stage, all body segments after the sixth are only equipped with a single pair of legs! In G. granulatus, the total number of such segments is only up to 13, as opposed to 18 in G. subgranulatus n. sp. and G. semigranulatus n. sp. (see below), 19 in Trachyjulus tjampeanus (Attems, 1903), from Java, Indonesia (NB: in the abundant material of cambalopsids taken from numerous caves from all over Java and accumu- lated at MNHN, we have only encountered this species!), and 28 in Glyphiulus zorzini, a southern Chinese cavernicole. In addition, in the Hawaiis, G. granulatus has been recorded as a myrmecophile (Crosland 1994). Similarly, based solely on the original descriptions of G. septentrionalis and G. multicarinus (cf. Murakami 1975; Zhang & Li 1982), it is clear that these species cannot be separated, agreeing perfectly with each other in every character. Hence the new synonymy advanced: G. septentrionalis Murakami, 1975 = G. multicarinus Zhang & Li, 1982. The occurrence of this species on Okinawa, Ryukyus, both in caves and epigeically (Murakami 1975), seems only to represent a recent introduction through human agency, with southern China definitely having served as the source area. Much like G. granulatus, G. septentrionalis is a relatively small species (adults 17 - 30 mm long), but the number of segments is usually considerably higher (43 - 66 p + 4 - 1 a + T), implying a longer development, which apparently reduces its vagility.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	description	(Figs 7 - 9)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	materials_examined	TYPE MATERIAL. — China. Yunnan Prov., Mengzi County, cave near footpath to plateau, 5. I. 1989, leg. P. Beron, holotype ♂ (NMNHS); paratypes 7 ♂♂, 14 ♀♀, 11 juv. (NMNHS); 2 ♂♂, 2 ♀♀ (MNHN GA 035); 2 ♂♂, 2 ♀♀ (ZMUM); 1 ♂, 1 ♀ (ZMUC); 1 ♂, 1 ♀ (MCSNV); 1 ♂, 1 ♀ (SEM). — Yunnan Prov., Mengzi County, Pothole No. 2 (Ma Fa Tiao Dong), 6. I. 1989, leg. P. Beron, paratype 1 ♀ (NMNHS).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	etymology	ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	diagnosis	DIAGNOSIS. — Differs from the most similar congeners by the crests on the collum being underdeveloped (especially so anteriorly), the body segments somewhat compressed laterally, the anterior gonopods with unusually slender telopodites, etc.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	description	DESCRIPTION Length of adults of both sexes 19 - 28 mm, width 1.0 - 1.3 mm; males usually a little shorter and more slender than females. Coloration much as in G. granulatus, variegate, yellow-brown, with a dark brown vertex, blackish ocellaria, mainly brown crests on collum and grey-brownish lateral longitudinal stripes running down from dark brown ozoporiferous tubercles; dorsum generally light, but a thin axial line mostly present due to darker median crests. Adults with 38 - 52 p + 5 - 2 a + T in males, 39 - 63 p + 5 - 1 a + T in females; largest juvenile ♂ with 31 p + 3 a + T; two smallest juveniles with 17 or 18 haplopodous segments after 6 th, plus 2 a + T. Holotype about 23 mm long, 1.0 mm wide, with 47 p + 2 a + T. Other characters as in G. granulatus, including mentum either divided (n = 1) (Fig. 9 B) or undivided (n = 1) (Fig. 7 A), 11 - 18 ocelli, antennae (Figs 8 A; 9 A), the carinotaxy formulae (Fig. 7 B-F), claw, posterior gonopod structure (Fig. 9 F, G), male legs 1 and 2 (Figs 8 D; 9 C, D), telson (Fig. 8 B), etc., but body size much larger (see above); segments compressed laterally (Fig. 8 C); crests on collum (7 + m + 7 at caudal margin) evidently reduced in size, almost obsolete anteromedially (Figs 7 B; 8 A); legs somewhat longer (about 4 / 5 as long as body height); anterior gonopods with telopodites more slender, while the coxosternum has much shorter distomedian outgrowths and acute distolateral corners (Fig. 9 E).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2213058FF450270FB77F935.taxon	discussion	REMARKS In the absence of evident troglomorphic traits, this species can only be considered as troglophilic.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23E305AFF5E048CFD9EF935.taxon	materials_examined	TYPE MATERIAL. — China. Yunnan Prov., Jianshui County, Yan Dong Cave, 12. I. 1989, leg. P. Beron, holotype ♂ (NMNHS); paratypes 2 ♀♀, 1 fragm. juv. (NMNHS); 1 ♂, 1 ♀ (MNHN GA 036); 1 ♂ (ZMUM); 1 ♂, 1 ♀ (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23E305AFF5E048CFD9EF935.taxon	etymology	ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23E305AFF5E048CFD9EF935.taxon	diagnosis	DIAGNOSIS. — Differs from the most similar congeners by the antennae being more slender, the body segments somewhat compressed laterally, the ocellaria only slightly reduced, the accessory spine at the claw base especially prominent, the anterior gonopod coxosternum with small but evident distomedial outgrowths.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23E305AFF5E048CFD9EF935.taxon	description	DESCRIPTION Length of adults of both sexes 18 - 25 mm, width 1.0 - 1.2 mm. General coloration pallid, pattern much as in G. granulatus, but ocellaria light brown while lateral stripes level to ozoporiferous tubercles narrow. Adults with 39 - 45 p + 2 - 1 a + T. Holotype about 25 mm long, 1.2 mm wide, with 44 p + 2 a + T. Other characters as in G. granulatus, including gnathochilarium (divided into pro- and eumentum, n = 1) (Fig. 12 B), the carinotaxy formulae (Figs 10 A-D; 11 B), telson (Figs 10 E, F; 11 A), male legs 1 - 3 (Fig. 12 C-E), posterior gonopod structure (Figs 11 E; 12 G, H), etc., but body size much larger (see above); segments compressed laterally (Fig. 11 B); antennae slightly elongate (Figs 10 A, B; 12 A); claw with a particularly prominent spine at base (Fig. 11 D); only some 9 - 11 ocelli in adults; legs somewhat longer (about as long as body height) (Figs 10 D; 11 C); anterior gonopod coxosternum with small but curved distomedian outgrowths and acute distolateral corners (Figs 11 F; 12 F).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23E305AFF5E048CFD9EF935.taxon	discussion	REMARKS Even though there are some indications of troglomorphism in this species (pallid general coloration, somewhat elongated antennae and legs), its troglophily seems more likely than troglobiosis. The morphological evidence being inconclusive, without further observations this problem can never be solved. On the other hand, Bollmania beroni Stoev & Enghoff, 2005 (Diplopoda, Callipodida, Caspiopetalidae Lohmander, 1931), a presumed troglobitic millipede, occurs in the same cave (Stoev & Enghoff 2005).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23D305DFD6A0156FDCEFA37.taxon	materials_examined	TYPE MATERIAL. — China. Yunnan Prov., Meele (= Mile) County, Bai Long Dong Cave (= White Dragon Cave), 2. I. 1989, leg. P. Beron, holotype ♂ (NMNHS); paratypes 1 ♂, 1 ♀ (NMNHS); 1 ♀ (SEM); 1 ♂ (MNHN GA 037). — Yunnan Prov., Jianshui County, Yan Zi Dong Cave, 10. I. 1989, leg. P. Beron, paratypes 5 ♂♂, 3 ♀♀, 4 juv. (NMNHS); 4 ♂♂, 2 ♀♀ (MNHN GA 037); 4 ♂♂, 2 ♀♀ (ZMUM); 2 ♂♂, 2 ♀♀ (ZMUC); 1 ♂, 1 ♀ (MCSNV); 1 ♂ (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23D305DFD6A0156FDCEFA37.taxon	etymology	ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23D305DFD6A0156FDCEFA37.taxon	diagnosis	DIAGNOSIS. — Differs by a position somewhat intermediate between the most similar congeners G. granulatus and G. paragranulatus n. sp., i. e. coloration, carinotaxy, antennae, claws, etc., as in G. granulatus, but size, segment shape, leg length and anterior gonopods more like in G. paragranulatus n. sp.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23D305DFD6A0156FDCEFA37.taxon	description	DESCRIPTION Length of adults of both sexes 12 - 24 mm, width 0.8 - 1.2 mm. Adults of both sexes with 31 - 64 p + 5 - 1 a + T; largest juvenile ♂ with 28 p + 5 a + T, largest juvenile ♀ with 27 p + 5 a + T; two smallest juveniles with 15 or 18 haplopodous segments after 6 th, plus 2 a + T or 1 a + T, respectively. Length of holotype about 22 mm, width 1.0 mm, with 45 p + 3 a + T. Coloration, carinotaxy (Fig. 13 A-E), antennae (Figs 13 A; 14 A), gnathochilarium, claws, etc. (Fig. 14 B-D, F, G) basically as in G. granulatus, but body size (usually much larger), segment shape (slightly compressed laterally) (Fig. 13 C), telson (Fig. 13 E, F) with peculiar flaps on paraprocts near hypoproct (Fig. 13 F), legs (somewhat longer) and anterior gonopods (Fig. 14 E) more like those of G. paragranulatus n. sp. Adults with 11 - 15 ocelli.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23D305DFD6A0156FDCEFA37.taxon	discussion	REMARKS In the absence of any evident troglomorphic traits, this species can only be considered as troglophilic at most. This is supported by G. semigranulatus n. sp. being present in several relatively strongly isolated caves, sometimes in significant numbers.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	description	(Figs 15; 16)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	materials_examined	TYPE MATERIAL. — China. Yunnan Prov., Jianshui County, Cave Ba Guo Dong, 11. I. 1989, leg. P. Beron, holotype ♂ (NMNHS); paratypes 2 ♂♂, 6 ♀♀, 3 juv. (NMNHS); 2 ♂♂, 2 ♀♀ (MNHN GA 038); 1 ♂, 1 ♀ (ZMUM); 1 ♂, 1 ♀ (ZMUC); 1 ♀ (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	etymology	ETYMOLOGY. — Honours Petar Beron, the collector.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	diagnosis	DIAGNOSIS. — Differs by the crests just below and above the ozopore-bearing crests being divided into three, not two, transverse rows of tubercles, coupled with an evident mediodorsal tubercle present in front of the caudal edge of the epiproct.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	description	DESCRIPTION Length of adults 11 - 17 (♂) or 12 - 24 mm (♀), width 0.8 - 1.0 (♂) or 0.8 - 1.3 mm (♀). Coloration nearly entirely light yellowish, only ocellaria brown, as well as sometimes a vague lateral line level to ozoporiferous crests. Adults with 28 - 40 p + 5 - 2 a + T (♂) or 28 - 53 p + 5 - 1 a + T (♀); largest juvenile ♂ with 27 p + 5 a + T, largest juvenile ♀ with 27 p + 4 a + T. Length of holotype about 17 mm, width 1.0 mm, with 40 p + 2 a + T. Carinotaxy of collum typical (Fig. 15 A, B), that of trunk segments generally typical but, starting from segment 7 until 6 or 7 penultimate segments, crests just below as well as between ozopores divided into three, not two, transverse rows of tubercles (Fig. 15 C, D). An evident middorsal tubercle just above caudal edge of epiproct (Fig. 15 E). Antennae (Figs 15 A, B; 16 A), legs (Fig. 16 B, C), claws, posterior gonopods (Fig. 16 E, F), etc., as in G. granulatus, but body size (often much larger), segment shape (slightly compressed laterally) and anterior gonopods (Fig. 16 D) more like those of G. semigranulatus n. sp. Adults with 5 - 13 ocelli. Male legs 1 with tubercular leg vestiges near base of prongs (Fig. 16 B).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A23A305FFF6E02B6FCCAFAD6.taxon	discussion	REMARKS In the absence of any clearly troglomorphic traits, except perhaps for its nearly completely unpigmented body and the somewhat reduced ocellaria, this species can hardly be considered as more than troglophilic.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2383051FCB302D1FB9AFA99.taxon	description	(Figs 17; 18)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2383051FCB302D1FB9AFA99.taxon	materials_examined	TYPE MATERIAL. — Laos. Luang Prabang Prov., Nong Kiaw (= Muang Ngoy), Tham Pha Kouang Cave, 29. XII. 2000, leg. A. Bedos & L. Deharveng (LAO- 099), holotype ♂ (MNHN GA 039); paratypes 1 ♂ (MNHN GA 039), 1 ♀ (SEM). ETYMOLOGY. — Honours Anne Bedos, one of the collectors.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2383051FCB302D1FB9AFA99.taxon	diagnosis	DIAGNOSIS. — Differs from other species by the paraprocts showing a clear median marginal ridge, combined with a plumose spatuliform distal process of the posterior gonopods and a subunciform distomedian corner of the anterior gonopods.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2383051FCB302D1FB9AFA99.taxon	description	DESCRIPTION Length of adults of both sexes 16 - 20 mm, width 1.0 - 1.1 mm. Holotype 16 mm long, 1.0 mm wide, with 40 p + 3 a + T and 11 ocelli; paratype ♂ about 20 mm long, 1.1 mm wide, with 54 p + 2 a + T and 14 ocelli. Coloration, antennae (Figs 17 A; 18 A), gnathochilarium (mentum divided, n = 1) (Fig. 18 B), carinotaxy (Fig. 17 A-E), segment shape, legs (Fig. 18 C-F), etc., generally as in G. granulatus, but paraprocts with a low, but evident, median marginal ridge (Fig. 17 F). Anterior gonopods like those of G. semigranulatus n. sp. (Fig. 18 G), but posterior gonopods with a plumose spatuliform distal process (Fig. 18 H), i. e. much as in G. quadrohamatus or G. lipsorum.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2383051FCB302D1FB9AFA99.taxon	discussion	REMARKS In the absence of any troglomorphic traits, this species can only be considered troglophilic at most.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	description	(Figs 19; 20)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	materials_examined	TYPE MATERIAL. — China. Hunan Prov., Longshan: Huoyan, Feihu Dong Cave (Flying Tiger Cave), 11. VIII. 1995, leg. A. Bedos & L. Deharveng (CHI- 022), holotype ♂ (MNHN GA 040); paratype 1 ♂ (MNHN GA 040). — Hunan Prov., Longshan, Huoyan, Baiyan Dong Cave (White Rocks Cave), 15. VIII. 1995, leg. A. Bedos & L. Deharveng (CHI- 032), paratypes 1 ♂, 1 ♀, 1 ♀ juv. (MNHN GA 040); 1 ♂ (ZMUM). — Hunan Prov., Longshan: Huoyan, Renmi Dong Cave, 19. VIII. 1995, leg. A. Bedos & L. Deharveng (CHI- 043); paratypes 1 ♂ (MNHN GA 040); 1 ♀ (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	etymology	ETYMOLOGY. — Honours Louis Deharveng, one of the collectors.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	diagnosis	DIAGNOSIS. — Differs by the relatively large size, the nearly completely smooth collum, the complete depigmentation (even of ocellaria), the elongated legs (with claws devoid of a spine at base from leg 4 onwards), the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7, the bare but flagelliform distal process of the posterior gonopods, etc.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	description	DESCRIPTION Length of adults of both sexes 40 - 55 mm, width 2.0 - 2.3 mm. Coloration usually uniformly dark brown due to heavy sclerotization, tegument being coloured only due to chitin; sometimes ozoporiferous tubercles reddish. Adults with 55 - 68 p + 4 - 1 a + T; juvenile ♀ with 40 p + 5 a + T, recognised by considerably reduced ozoporiferous tubercles on segments 7 - 33, plus 5 a + T. Length of holotype about 55 mm, width 2.3 mm, with 67 p + 1 a + T. Antennae slightly more elongate than usual (Figs 19 A; 20 A). Ocellaria unpigmented, barely traceable, with about 10 or 11 ocelli in largest specimens. Gnathochilarium (Fig. 20 B) with an undivided mentum (n = 1). Carinotaxy of collum unusual in that all crests are nearly obliterated (Fig. 19 A, B), though a typical pattern is traceable. Carinotaxy of subsequent metaterga typical (Fig. 19 A-C, E), but crests low, especially dorsal ones on segments 2 - 6; only two lateral crests below ozoporiferous ones, divided from segment 7 onwards, followed by a dozen striae. A low, but evident, transverse pleural ridge behind gonopod opening on male segment 7. Epiproct slightly elongate apically, more narrowly rounded than usual; hypoproct more narrowly emarginate at caudal margin (Fig. 19 D). Legs elongate (Fig. 20 F), claws strongly elongate, without a spine near base starting from leg 4 (Fig. 19 F). Male legs 1 with vestigial tubercular coxites (Fig. 20 C). Penes with 4 or 5 long setae distolaterally (Fig. 20 D), male legs 3 elongate, much as in G. granulatus (Fig. 20 E). Anterior gonopods with small subunciform protuberances at median corners of coxosternum (Fig. 20 G). Posterior gonopods with a flagelliform but bare distal process (Fig. 20 H).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2363053FC800210FE2FF935.taxon	discussion	REMARKS Based on its evident troglomorphic traits, this species seems to be a troglobite.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2343054FC9103D2FC67FA57.taxon	description	(Figs 21 - 23)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2343054FC9103D2FC67FA57.taxon	materials_examined	TYPE MATERIAL. — China. Guizhou Prov., Guanling County, Huajiang, Da Dong Cave, 4. IV. 2004, leg. L. Latella & Ghen Hu, holotype ♂ (MNHN GA 041); paratypes 1 ♂ (MNHN GA 041); 1 ♀ (SEM). — Guizhou Prov., Guanling County, Huajiang, An Jia Da Dong Cave, 4. IV. 2004, leg. L. Latella & R. Zorzin, paratype 1 ♀ (MNHN GA 041).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2343054FC9103D2FC67FA57.taxon	etymology	ETYMOLOGY. — To emphasize the close affinities to G. granulatus.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2343054FC9103D2FC67FA57.taxon	diagnosis	DIAGNOSIS. — Characterized by the somewhat larger ocellaria, the underdeveloped tergal crests (becoming almost obliterated in the median part of the collum), coupled with the medium body size, slightly elongated legs, simple anterior gonopods, etc. DESCRIPTION Length of holotype and of paratype ♀ from An Jia Da Dong about 30 mm, of paratype ♂ about 33 mm, width 1.5, 1.5 and 1.7 mm, with 56 p + 2 a + T, 67 p + 1 a + T and 57 p + 2 a + T, respectively. Coloration, antennae (Figs 21 A; 23 A), segment shape (Fig. 22 D), carinotaxy patterns (Fig. 21), male legs 1 - 3 (Fig. 23 B-D), etc., as in G. granulatus, but tergal crests somewhat underdeveloped (Figs 21; 22 B, D), almost obliterated in median part of collum (Fig. 21 A, B). Ocellaria rounded-triangular, black, each with 23 - 25 poorly convex ocelli. Claws slightly longer, spine at base slightly shorter (Fig. 22 C); legs about 4 / 5 as long as midbody height (Fig. 23 E). Epiproct with a faint but evident middorsal tubercle (Fig. 22 B); hypoproct more narrowly emarginate at caudal margin (Fig. 22 A). Anterior gonopod coxosternum particularly simple (Fig. 23 F). Posterior gonopods with a typical flagelliform and plumose distal process (Fig. 23 G, H).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2343054FC9103D2FC67FA57.taxon	discussion	REMARKS Even though taken from caves, this species seems to represent only a troglophile at most. However, this cave harbours another, apparently troglobitic congener from a different species group, to be described elsewhere.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2333057FCBE0156FF34F9D4.taxon	description	(Figs 24 - 26)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2333057FCBE0156FF34F9D4.taxon	materials_examined	TYPE MATERIAL. — China. Guizhou Prov., Gianxi County, Hong Lin Village, Chang Tu Dong Cave, 13. XI. 2003, leg. L. Latella, holotype ♂ (MNHN GA 042); paratypes 2 ♂♂, 1 ♀ (MNHN GA 042), 1 ♀ (SEM). — Same locality, 3. IV. 2004, leg. L. Latella, paratype 1 ♀ (MCSNV). — Guizhou Prov., Gianxi County, Hong Lin Village, Xiao Dong Cave, 16. XI. 2003, leg. Latella & Cioperelli-Repetto, paratype 1 ♂ (ZMUM). — Guizhou Prov., Gianxi County, Hong Lin Village, Xi Xiang Dong Cave, 14. XI. 2001, leg. L. Latella, paratype 1 ♀ (MNHN GA 042). — Guizhou Prov., Gianxi County, Hong Lin Village, Dayan Dong Cave, 22. XI. 2003, leg. L. Latella, paratype 1 ♀ (ZMUC). — Guizhou Prov., Gianxi County, Hong Lin Village, Tiao Shuz Dong Cave, 18. XI. 2001, leg. L. Latella, D. Avesani & G. Rossi, paratypes 1 ♀, 3 juv. (MNHN GA 042); 1 ♀ (SEM). — Guizhou Prov., Gianxi County, Hong Lin Village, Lu Diao Ai Dong Cave (= Cow Fell Down Cave), XI. 2003, leg. L. Latella, paratype 1 ♂ (NMNHS). — Guizhou Prov., Gianxi County, Hong Lin Village, Shu Jia Yan Dong Cave, 17. XI. 2001, leg. Latella & Berzacola, paratypes 2 ♂♂, 1 ♀, 1 juv. (MNHN GA 042). — Guizhou Prov., Gianxi County, Ishui Luo Dong Cave, 20. XI. 2001, leg. L. Latella, paratype 1 ♂ (MNHN GA 042).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2333057FCBE0156FF34F9D4.taxon	etymology	ETYMOLOGY. — Honours Leonardo Latella, the collector.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2333057FCBE0156FF34F9D4.taxon	diagnosis	DIAGNOSIS. — Differs from congeners by the strongly reduced crests on the collum and metaterga, the completely undivided dorsal crests (the mediodorsal ones also undivided anteriorly), the complete depigmentation (even of ocellaria), the evidently elongated antennae and legs, the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7, etc.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A2333057FCBE0156FF34F9D4.taxon	description	DESCRIPTION Length of adults of both sexes 18 - 45 mm, width 1.3 - 2.0 mm. Coloration usually uniformly dark brown due to heavy sclerotization, sometimes ozoporiferous tubercles reddish; more rarely, coloration uniform yellowish. Adults of both sexes with 42 - 63 p + 6 - 1 a + T; largest juveniles (♀) with 35 p + 4 a + T or 36 p + 5 a + T. Length of holotype about 30 mm, width 1.3 mm, with 50 p + 3 a + T. Antennae much more elongate than usual (Figs 24 A; 26 A), with especially numerous and dense distodorsal sensilla on antennomeres 5 and 6 (Fig. 25 C). Ocellaria, if any, unpigmented, ocelli, when present, 1 - 11, irregularly spaced, barely traceable. Gnathochilarium usual, mentum divided (n = 2) (Fig. 26 B). Carinotaxy of collum rather typical, though hardly traceable because of crests being nearly obliterated, especially anteromedially (Fig. 24 A, B). Carinotaxy of subsequent metaterga deviating slightly from typical formula (Figs 24; 25 B) in that none of the dorsal crests is divided transversely or longitudinally; all crests somewhat reduced (Fig. 25 B), with only 2 or 3 very low lateral ones (formula 2 (3) + I + 3 + I + 2 (3 )). From segment 7 on, crest below “ I ” (i. e. below ozoporiferous tubercle) poorly divided at about midway; further ventrad, segments with 1 or 2 barely distinguishable crests followed by about a dozen fine striae (Fig. 24 A, D). A low, but evident, transverse pleural ridge present behind gonopod opening on male segment 7. Epiproct slightly elongate apically, more narrowly rounded than usual; hypoproct more narrowly emarginate at caudal margin (Fig. 25 A). Legs elongate (Fig. 25 B), about as long as midbody height; claws strongly elongate, with a small spine near base on legs of anterior 1 / 5 - 1 / 4 body (Fig. 26 F), thereafter devoid of any basal spines (Fig. 25 D). Male legs 1 with rudimentary tubercular coxites (Fig. 26 C); male legs 2 and 3 typical (Fig. 26 D, E).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	description	(Figs 27 - 29)	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	materials_examined	TYPE MATERIAL. — China. Sichuan Prov., Xin Long County, Gan Chuan Cave, Nos 414, 417 and 421, 6. VIII. 1999, leg. J. & B. Lips, holotype ♂ (MNHN GA 043); paratypes 1 ♂, 3 ♀♀ (MNHN GA 043); 1 ♀ (ZMUM); 1 ♀ (ZMUC); 1 ♀ (SEM).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	etymology	ETYMOLOGY. — To emphasize the presumed basal position of this species within the group.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	diagnosis	DIAGNOSIS. — Differs primarily by the retained telopodites of male legs 1, coupled with the large size, the crests on the collum and the spines at claw base both strongly reduced, the crests on metaterga reduced but the ozoporiferous tubercles high, the troglomorphic traits (complete depigmentation, the lack of ocelli, the evidently elongated antennae and legs), the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	description	DESCRIPTION Length of ♂, 30 - 32 mm, width 1.8 mm; of ♀, 23 - 42 and 1.5 - 2.2 mm, respectively. Length of holotype about 30 mm, width 1.3 mm. Coloration usually uniformly yellowish, sometimes dark red-brown due to heavy sclerotization. Holotype with 48 p + 3 a + T; ♂ paratype with 45 p + 4 + T; ♀ paratypes with 40 - 59 p + 4 - 1 a + T. Antennae much more elongate than usual (Figs 27 A; 29 A). Ocelli absent. Gnathochilarium with (n = 1) or without (n = 2) division of mentum (Fig. 29 B). Crests on collum, dorsad of a well-developed lateral carina, nearly completely obliterated, pattern visible only as (4) 5 + m + 5 (4) extremely faint undulations at caudal margin (Fig. 27 A, B). Carinotaxy of subsequent metaterga typical (Fig. 27 D; 28 D), but only two crests present below rather strikingly elongated ozoporiferous tubercles; all other crests distinctly underdeveloped (Figs 27 A, D; 28 A, C). From segment 7 or 8 on, median crest clearly doubled anteriorly (Fig. 27 D). Midbody segments round in crosssection (Fig. 28 C). A low, but evident, transverse pleural ridge present behind gonopod opening on male segment 7. Epiproct narrower caudally, hypoproct more narrowly emarginate at caudal margin (Fig. 28 B). Legs elongate (Figs 27 C; 28 C; 29 F), longer than midbody height; claws strongly elongate, with a very small spine at base (Fig. 28 D). Male legs 1 unusual in being fully leg-like, only very modestly reduced: a coxa, a 3 - segmented telopodite and a claw supplied with a spine at base are all very clearly expressed (Fig. 29 C); male legs 2 and 3 typical (Fig. 29 D, E), but penes devoid of strong setae distolaterally. Anterior gonopods with evident, subunciform protuberances at median corners of coxosternum (Fig. 29 G). Posterior gonopods with a flagelliform and micropilose distal process (Fig. 29 H).	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
03F4C378A20F3069FF6D0253FC4EF935.taxon	discussion	REMARKS Based on morphological evidence alone, this species seems to be a troglobite.	en	Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Spiegel, Didier Van Den (2007): Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group. Zoosystema 29 (1): 7-49, DOI: 10.5281/zenodo.5392079
