identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F487A3FFCFFFD7FCA8FBD592E3FBE1.text	03F487A3FFCFFFD7FCA8FBD592E3FBE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dryopidae	<div><p>DRYOPIDAE</p> <p>Helichus basalis LeConte. Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Clarke: South Whitebreast Cr., Weldon, TMM, 8/17/15; Davis: Fox R., Moulton, CTH, 7/17/06; Henry: Little Potomac Cr., Mt. Pleasant, CTH, 8/7/ 15; Jefferson: North Walnut Cr., Pleasant Plain, CTH, 8/14/15; Keokuk: Twin Grove Cr., Ollie, CTH, 6/2/15; Lee: Lick Cr., Shimek State Forest, CTH, 9/10/10; Lucas: Wolf Cr., Chariton, CTH, 8/ 10/05; Monroe: Sage Cr., Stephens Forest, CTH, 8/ 8/14; Poweshiek: Bear Cr., Brooklyn, CTH, 9/25/ 02; Ringgold: Middle Fk. Grand R., Mt. Ayr, KLS, 7/14/04; Taylor: Honey Cr., Bedford, KBK, 8/18/ 04; Van Buren: Coppers Cr., Keosauqua, CTH, 10/ 16/09; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Wayne: Wolf Cr., Millerton, CTH, 8/10/ 05. Distribution: Eastern USA, extending west to Iowa in the north and Texas in the south. Iowa forms the northwestern border in the USA with distribution restricted to southern Iowa. This species’ habitat preference is under rocky substrate at the edge of streams. In Iowa, H. basalis tends to be collected more frequently in smaller streams. New state record.</p> <p>Helichus fastigiatus (Say). Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Black Hawk: Black Hawk Cr., Popp County Access, CTH, 8/22/12; Cerro Gordo: Bailey Cr., Thornton, TMM, 8/30/07; Clayton: Volga R., Elkport, MWB, 9/15/08; Clinton: Brophy Cr., McCausland, CTH, 9/30/13; Davis: Soap Cr., Eldon, CTH, 10/4/10; Decatur: Little R., Leon, CTH, 9/23/02; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Harrison: Soldier R., Pisgah, KLS, 9/21/01; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/12/08; Ida: Battle Cr., Battle Cr., TFW, 9/11/02; Jefferson: North Walnut Cr., Pleasant Plain, CTH, 8/14/15; Johnson: Iowa City (Wickham 1911) Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Lick Cr., Farmington, CTH, 7/14/03; Linn: Wapsipinicon R., Central City, MWB, 9/27/06; Lucas: Whitebreast Cr., Lacona, TMM, 8/5/09; Mahaska: Muchakinock Cr., Eddyville, MWB, 9/2/11; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Ringgold: Lotts Cr., Lamoni, CTH, 8/26/96; Van Buren: Chequest Cr., Pittsburg, CTH, 7/21/11; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05. Distribution: Eastern USA, extending west to Iowa in the north and Texas in the south. Iowa forms the northwestern border. Habitat preference is under rocky substrate in small to large streams.</p> <p>Helichus lithophilus (Germar). Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/01; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wapsipinicon R., Dunkerton, CTH, 8/2/06; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: East Branch Wapsipinicon R., Tripoli, CTH, 8/27/97; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Butler: Coldwater Cr., Greene, CTH, 8/29/96; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/ 13/07; Cass: West Nodaway R., Grant, CTH, 8/ 19/98; Cedar: Cedar R., West Branch, CTH, 9/ 12/06; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 9/22/15; Chickasaw: Wapsipinicon R., Ionia, CTH, 10/ 12/00; Clarke: South Whitebreast Cr., Weldon, CTH, 9/28/95; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 10/9/10; Clayton: Volga R., Elkport, MWB, 9/15/08; Clinton: Brophy Cr., McCausland, CTH, 8/11/04; Crawford: Big Cr., Denison, TMM, 10/1/13; Dallas: Mosquito Cr., Panora, CTH, 7/15/96; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: Long Cr., Van Wert,JML, 10/9/01; Delaware: South Fk. Maquoketa R., Dundee, CTH, 7/12/05; Des Moines: Skunk R., Burlington, MWB, 9/7/06;</p> <p>Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Fayette: Nutting Cr., Clermont, CTH, 7/26/06; Floyd: Cedar R., Charles City, MWB, 9/9/08; Franklin: Maynes Cr., Hampton, CTH, 8/18/08; Fremont: Nishnabotna R, Hamburg, TMM, 10/ 1/13; Greene: Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hardin: Iowa R., Eldora, CTH, 10/11/ 05; Harrison: Mosquito Cr., Persia, TFW, 10/12/ 00; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Humboldt: West Fk. Des Moines R, Humboldt, TMM, 8/2/12; Ida: Maple R., Galva, CTH, 8/6/ 97; Iowa: Big Bear Cr., Victor, CTH, 9/20/99; Jackson: Maquoketa R., Canton, CTH, 9/7/05; Jasper: Indian Cr., Mingo, TMM, 9/30/03; Jefferson: Cedar Cr., Stockport, CTH, 7/20/ 05; Johnson: Iowa R., Iowa City, CTH, 9/22/ 05; Jones: Maquoketa R., Monticello, CTH, 9/6/ 05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: East Fk. Des Moines R., Seneca, TMM, 8/29/11; Lee: Skunk R., Augusta, CTH, 10/6/15; Linn: Buffalo Cr., Central City, CTH, 9/ 2/08; Louisa: Iowa R., Columbus Junction, MWB, 9/10/08; Lucas: Wolf Cr., Chariton, CTH, 8/10/05; Madison: North Branch North R., Winterset, CTH, 6/20/95; Mahaska: North Skunk R., Rose Hill, CTH, 7/10/96; Marion: North Cedar Cr., Bussey, CTH, 8/25/98; Marshall: South Minerva Cr., Clemons, TMM, 8/8/07; Mills: West Nishnabotna R., Malvern, TMM, 9/4/12; Mitchell: Wapsipinicon R., McIntyre, CTH, 8/27/08; Monona: Maple R., Mapleton, TMM, 9/5/12; Montgomery: Pilot Branch, Stennett, CTH, 9/9/96; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/14; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Keg Cr., McClelland, JML, 9/4/02; Poweshiek: Little Bear Cr., Brooklyn, CTH, 8/18/05; Ringgold: Lotts Cr., Lamoni, CTH, 8/26/96; Sac: North Raccoon R., Auburn, KBK, 9/13/05; Scott: Lost Cr., Princeton, CTH, 8/30/04; Shelby: East Branch West Nishnabotna R., Avoca, TMM, 8/ 16/07; Story: South Skunk R., Ames, CTH, 9/ 15/95; Tama: Richland Cr., Haven, CTH, 9/8/95; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Van Buren: Des Moines R., Farmington, CTH, 10/10/06; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, KLS, 10/8/04; Washington: English R., Riverside, MWB, 10/15/99; Wayne: Wolf Cr., Millerton, CTH, 8/10/05; Webster:</p> <p>Lizard Cr., Clare, CTH, 9/11/96; Winneshiek: Yellow R., Postville, CTH, 7/25/07. Distribution: Eastern USA, extending west to Minnesota in the north, Nebraska, and Texas in the south. Habitat preference is woody debris and rocky substrate in small to large streams.</p> <p>Helichus striatus LeConte. Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Little Paint Cr., Harpers Ferry, CTH, 9/11/07; Audobon: Unnamed trib. to East Nishnabotna R., Audobon, TMM, 8/23/11; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Boone: Unnamed trib. to Little Beaver Cr., Bouton, JML, 10/4/06; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/ 23/11; Buchanan: Lime Cr., Brandon, CTH, 5/ 15/96; Butler: Dry Cr., Clarksville, CTH, 5/29/ 07; Calhoun: Unnamed trib. to Purgatory Cr., Lohrville, KLS, 6/25/08; Cass: Sevenmile Cr., Grant, KLS, 7/26/06; Cedar: Sugar Cr., Tipton, CTH, 8/23/01; Cerro Gordo: Bailey Cr., Thornton, TMM, 8/30/07; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 5/15/12; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 8/26/14; Clarke: South Whitebreast Cr., Weldon, TMM, 9/29/10; Clay: Willow Cr., Royal, CTH, 9/13/99; Clayton: North Cedar Cr., Clayton, CTH, 8/22/94; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Crawford: Big Cr., Denison, CTH, 8/12/98; Dallas: Little Beaver Cr., Woodward, JML, 7/24/07; Decatur: Weldon R., Woodland, TMM, 8/8/13; Delaware: Fenchell Cr., Strawberry Point, CTH, 9/17/07; Dubuque: Hickory Cr., New Vienna, CTH, 8/11/ 99; Fayette: Nutting Cr., Clermont, CTH, 7/26/ 06; Floyd: Little Cedar R., Colwell, CTH, 8/20/ 02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/ 03; Greene: Buttrick Cr., Grand Junction, TMM, 7/30/14; Guthrie: South Raccoon R., Guthrie Center, KLS, 6/12/03; Hamilton: Long Dick Cr., Ellsworth, TMM, 10/6/08; Hancock: East Branch Iowa R., Goodell, TMM, 10/7/10; Hardin: Tipton Cr., Buckeye, CTH, 7/30/99; Harrison: Potato Cr., Persia, KLS, 7/12/05; Howard: Crane Cr., Saratoga, CTH, 10/10/00; Ida: Maple R., Galva, CTH, 8/6/97; Jackson: Storybook Hollow, Bellevue, CTH, 8/29/07; Jasper: North Skunk R., Kellogg, CTH, 8/7/ 01; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Bear Cr., Wyoming, CTH, 7/13/ 04; Keokuk: Troublesome Cr., South English, CTH, 7/7/03; Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Linn: Wapsipinicon R., Central City, CTH, 9/27/06; Louisa: Honey Cr., Conesville, CTH, 9/2/98; Lucas: Wolf Cr., Chariton, TMM, 9/14/15; Madison: North Branch North</p> <p>R., Winterset, CTH, 5/15/97; Mahaska: Buck</p> <p>Cr., Barnes City, MWB, 8/21/00; Marion: Whitebreast Cr., Knoxville, TMM, 8/27/12; Marshall: South Minerva Cr., Clemons, 8/8/ 07; Mills: Mud Cr., Hastings, JML, 8/2/05; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Monona: Jordan Cr., Soldier, TMM, 7/ 5/06; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Montgomery: Walnut Cr., Red Oak, CTH, 10/14/96; Muscatine: Mad Cr., Muscatine, CTH, 6/30/04; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01; Page: East Nishnabotna R., Shenandoah, TMM, 9/18/08; Plymouth: Perry Cr., Hinton, KLS, 7/20/04; Polk: Camp Cr., Mitchellville, TMM, 7/20/05; Pottawattamie: Mosquito Cr., Underwood, TMM, 8/17/04; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/ 17/08; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/ 03; Scott: Lost Cr., Princeton, CTH, 8/30/04; Shelby: West Nishnabotna R., Irwin, TMM, 8/ 25/03; Sioux: Dry Cr., Hawarden, JML, 7/12/06; Story: Ballard Cr., Cambridge, KJS, 7/18/07; Tama: Richland Cr., Haven, CTH, 9/8/95; Warren: North R., Norwalk, KLS, 10/8/04; Washington: English R., Riverside, CTH, 9/ 24/02; Wayne: Dick Cr., Corydon, TMM, 9/ 11/08; Webster: Allen Cr., Stratford, JML, 8/ 15/07; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08; Woodbury: West Fk. Little Sioux R., Bronson, JML, 8/9/05; Wright: Drainage Ditch #49, Eagle Grove, JML, 9/3/02. Distribution: Transcontinental in the USA. Habitat preference for H. striatus is on debris and under rocky substrate in small to large streams.</p> </div>	https://treatment.plazi.org/id/03F487A3FFCFFFD7FCA8FBD592E3FBE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFC8FFC2FEB4FBB39267FE84.text	03F487A3FFC8FFC2FEB4FBB39267FE84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dytiscidae Leach 1815	<div><p>DYTISCIDAE</p> <p>Acilius fraternus (Harris). Clinton: Brophy Cr., McCausland, CTH, 8/11/04; Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Jackson: South Big Mill Cr., Bellevue, CTH, 9/3/10; Lee: Bonnell Cr., Franklin, CTH, 7/16/02; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05. Distribution: Eastern USA, extending west to Iowa in the north and Texas in the south. Iowa forms the northwestern border in the USA. Beyond the single wetland collection, stream habitats varied from small and intermittent to a wider, slow-moving stream.</p> <p>Acilius mediatus (Say). Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Davis: Fox R., Moulton, CTH, 6/29/06; Delaware: puddle at Ram Hollow Spring, Colesburg, CTH, 10/16/00; Johnson: Iowa City (Wickham 1911) Lucas: Whitebreast Cr., Woodburn, KLS, 9/24/02; Ringgold: Gooseberry Cr., Mt.Ayr, TMM, 7/2/ 03; Van Buren: Town Branch, Pittsburg, CTH,</p> <p>9/14/15. Distribution: Eastern USA, extending</p> <p>west to Minnesota in the north, Nebraska, and Mississippi in the south. Collection habitats consisted of smaller, slow flowing streams, and from a puddle.</p> <p>Acilus semisulcatus Aubé. Cerro Gordo: Union Hills West Wetland, Thornton MTF, 7/20/07; Hancock: Pilot Knob Wetland, Forest City, MTF, 7/18/07; Winnebago: Private Wetland, Forest City, MTF, 7/18/07; Winneshiek: UV light, KR, 11/3/01; Worth: Pilot Knob Wetland, Lake Mills, MTF, 7/18/07. Distribution: Northern transcontinental in the USA. Iowa forms the southern border in the Midwest. Its habitat preference is lentic environments.</p> <p>Acilus sylvanus Hilsenhoff. Black Hawk: Dry Run Cr., Cedar Falls, CTH, 8/29/11; Buchanan: Bear Cr., Brandon, TFW, 8/8/96; Jackson: South Big Mill Cr., Bellevue, CTH, 9/17/14; Poweshiek: Walnut Cr., Brooklyn, CTH, 7/28/09; Winneshiek: Trout R., Decorah, CTH, 7/18/07. Distribution: Northeastern USA, with Iowa forming part of the southwestern border. Habitat preferences are generally standing waters, although all collections recorded in Iowa occurred in small to midsized streams.</p> <p>Agabus aeruginosus Aubé. Johnson: Iowa City (Wickham 1911). Distribution: Northeastern USA, with Iowa forming part of the northwestern border.</p> <p>Agabus ambiguus (Say). Emmet: Ditch 151, Wallingford, JML, 6/17/03. Distribution: Northern transcontinental in the USA. Iowa forms the southern border in the Midwest.Habitat preference for this species is depositional environments in lotic systems, generally small streams, springs, and stream fed ponds.</p> <p>Agabus antennatus Leech. Hamilton: Little Wall Lake, PJS (National Museum of Natural History), 6/14/52. Distribution: Northwestern USA, with Iowa forming part of the southeastern border.</p> <p>Agabus anthracinus Mannerheim. (Larson et al. 2000). Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Agabus bifarius (Kirby). Johnson: (Wickham 1911). Distribution: northern transcontinental in the USA with Iowa forming part of the southern border in the Midwest.</p> <p>Agabus disintegratus (Crotch). Kossuth: Private Wetland, Bancroft, MTF, 6/13/07; Pottawattamie: (Wickham 1911) Sioux: Deep Cr., Granville, KLS,</p> <p>8/2/06; Union: Sand Cr., Arispe, JML, 6/11/03. Distribution: Transcontinental in the USA. Its habitat preference is temporary ponds. However, collection sites include two streams, one intermittent, and one drainage ditch flowing through an agricultural field.</p> <p>Agabus punctatus Melsheimer. Story: Ames (Wickham 1911). Distribution: Eastern USA, with Iowa forming part of the western border.</p> <p>Agabus subfuscatus Sharp. A specimen is in the National Museum of Natural History and has no data on the collection label beyond IA (Larson et al. 2000). Distribution: Northeastern USA, extending west to Iowa.</p> <p>Boreonectes griseostriatus (De Geer). Hamilton: White Fox Cr., Webster City, CTH, 8/25/97; Johnson: (Wickham 1911); Kossuth: Buffalo Cr., Michaelson, JML, 8/24/00. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest. Its habitat preference is mineral substrate in disturbed, or erosive, environments.</p> <p>Celina angustata Aubé. Winneshiek: Upper Iowa R. (UV light), MJH, 6/18/12. Distribution: Primarily eastern USA, extending from Massachusetts in the north to Florida in the south and west to Texas in the south and Minnesota in the north. Iowa forms part of the western border of its distribution. New state record.</p> <p>Celina hubbelli Young. Johnson: North Liberty (UV light), MHB, 7/31/13. Distribution: Primarily eastern USA, extending west to Texas in the south and Iowa in the north. New state record.</p> <p>Coelambus dissimilis (Gemminger and Harold). Allamakee: Yellow R., Postville, CTH, 7/24/12; Clinton: Silver Cr., DeWitt, CTH, 7/21/98; Dubuque: Cloie Branch, Durango, CTH, 8/5/ 14; Hancock: Unnamed trib. East Branch of Iowa R., Goodell, TMM, 6/26/13; Johnson: Iowa City (Wickham 1911); Linn: Behrens Pond (UV light), DAV, 3/6/04; Story: Ames (Wickham 1911). Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border. This species’ habitat preference is shallow, often temporary ponds and marshes. The Iowa stream collection sites were small flowing streams with coarse substrate and sand mix.</p> <p>Coelambus laccophilinus (LeConte). Mitchell: Deer Cr., Carpenter, CTH, 8/25/08. Distribution: Northeastern USA. Iowa forms part of southwestern edge of the species’ distribution. Its habitat preference is small, but permanent, ponds, especially among emergent grasses and sedges on the margins. The Iowa collection site is a wooded, cobbled flowing stream, but has areas with slow velocity and aquatic vegetation. New state record.</p> <p>Coelambus nubilis (LeConte). Grundy: Middle Fk. South Beaver Cr., Ackley, CTH, 8/15/01; Ida: Battle Cr., Holstein, KLS, 6/29/04; Iowa: Matt Foley Wetland, Koszta, DPH, 5/31/02; Johnson: Iowa City (Wickham 1911); Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Nearly transcontinental in the USA, except for the extreme West. Its habitat preference tends to be standing bodies of water that are permanent in nature. Several of the Iowa collections were in small streams.</p> <p>Coelambus sylvanus (Fall). Linn: Cedar Rapids (UV light), ELF, 9/4/99. Distribution: Northeastern USA, extending west to Minnesota in the north. Iowa forms part of the southwestern border of its distribution. Its habitat preference is marshy areas dominated by Typha L. (Typhaceae) and sedges. New state record.</p> <p>Colymbetes exaratus LeConte. Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Story: South Skunk R., Ames, CTH, 9/18/95. Distribution: Western USA, extending east to Illinois and Wisconsin. Habitat preference is among dense emergent vegetation at the margins of standing waters.</p> <p>Colymbetes sculptilis Harris. Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/ 07; Cherokee: (Wickham 1911); Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Johnson: (Wickham 1911); Pottawattamie: (Wickham 1911); Story: (Wickham 1911). Distribution: Northern transcontinental in the USA. Iowa forms the southern border in the Midwest. Its habitat preference is emergent plants on the margins of lakes or longer lasting temporary ponds.</p> <p>Copelatus chevrolati Aubé. Allamakee: Hecker Cr., Postville, CTH, 8/14/00, French Cr., Churchtown, CTH, 3/12/07; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Clay: Montgomery Cr., Gillet Grove, JML, 8/3/04; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Hamilton: Lyons Cr., Webster City, TMM, 9/1/10; Kossuth: East Fk. Des Moines R., Seneca, TMM, 8/12/15; Lucas: Jordan Cr., Macedonia, TMM, 7/27/09; Mitchell: Burr Oak Cr., Osage, CTH, 8/2/11; Pottawattamie: Jordan Cr., Macedonia, TMM, 7/27/09; Story: Bear Cr., Ames, JML, 9/19/07; Van Buren: Chequest Cr., Pittsburg, CTH, 9/14/ 15; Winneshiek: UV light, KJL, 10/6/93. Distribution: Transcontinental in the southern USA, but extends more northerly in the Midwest. Iowa</p> <p>forms the northern border of its distribution in the Midwest. Its habitat preference is generally lentic with organic debris, but records occurr in flowing streams.</p> <p>Copelatus glyphicus (Say). Allamakee: Hecker Cr., Postville, CTH, 8/14/00; Black Hawk: Dry Run Cr., Cedar Falls, CTH, 8/29/13; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Cedar: Sugar Cr., Tipton, CTH, 8/23/01; Clay: Unnamed trib. to Elk Cr., Gillett Grove, TMM, 8/24/10; Clayton: North Cedar Cr., Clayton, MWB, 8/ 21/08; Crawford: Otter Cr., Deloit, CTH, 8/13/ 98; Fayette: Volga R., Vandalia, CTH, 8/31/10; Harrison: Potato Cr., Persia, KLS, 7/12/05; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Johnson: (Wickham 1911); Kossuth: Buffalo Cr., Titonka, JML, 8/24/00; Mitchell: Deer Cr., Carpenter, CTH, 8/18/10; Pottawattamie: (Wickham 1911); Webster: Lyons Cr., Webster City, TMM, 9/1/10. Distribution: Transcontinental in the USA. Its habitat preference tends to be lentic waters, but records occurr in flowing streams.</p> <p>Coptotomus longulus LeConte. Black Hawk: Cedar R., Waterloo, CTH, 9/9/14; Butler: West Fk. Cedar R., Lake Considine County Park, CTH, 10/2/12; Floyd: Cedar R., Floyd, CTH, 9/ 23/08; Ida: Battle Cr., Holstein, KLS, 6/29/04; Kossuth: Kettle Wetland-Union Slough NWR, Bancroft, CTH, 5/6/97; Mahaska: North Skunk R., Rose Hill, CTH, 8/22/11; Mitchell: Cedar R., Floyd, MWB, 9/23/08; Muscatine: Pike Run, Nichols, CTH, 7/26/04; Sac: North Raccoon R., Sac City, KLS, 7/10/00. Distribution: Transcontinental in the USA, exclusive of the Southwest. Its habitat preference is open, permanent ponds, although in Iowa it has also been collected in flowing waters (from small streams to large rivers).</p> <p>Coptotomus loticus Hilsenhoff. Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05, Upper Iowa R., Dorchester, CTH, 10/6/05; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: West Fk.Cedar R., Finchford, CTH, 9/15/08; Bremer: Cedar R., Janesville, CTH, 9/9/14; Buchanan: Mud Cr., Brandon, CTH, 9/29/11; Butler: Shell Rock R., Shell Rock, CTH, 9/8/14; Clayton: Hewitt Cr., Volga, MWB, 9/1/04; Clinton: Brophy Cr., McCausland, CTH, 8/11/ 04; Dickinson: Little Sioux R., Lake Park, TMM, 8/11/14; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Bell Cr., Clermont, CTH, 7/20/04; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 10/9/07; Jones: Maquoketa R., Monticello, CTH, 9/6/05; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Honey Cr., Conesville, CTH, 9/2/ 98, 8/24/04; Marshall: Iowa R., Marshalltown, TMM, 9/11/14; Muscatine: Pike Run, Nichols, CTH, 7/26/04; Polk: Des Moines R, Des Moines, TMM, 8/31/09; Scott: Lost Cr., Princeton, CTH, 8/ 30/04; Story: South Skunk R., Cambridge, TMM, 10/7/14; Webster: Des Moines R., Ft. Dodge, TMM, 9/29/14; Winnebago: Winnebago R, Lake Mills, TMM, 9/28/15; Winneshiek: Coldwater Cr., Bluffton, CTH, 9/22/04; Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Eastern USA, extending west to Texas in the south. Iowa forms part of the western border of its distribution. New state record.</p> <p>Cybister fimbriolatus (Say). Cass: Indian Cr., Lewis, KLS, 7/23/02; Cerro Gordo: Private Wetland, Clear Lake, MTF, 7/2/08; Greene: Snake Cr., Rippey, MTF, 8/14/07; Humboldt: East Fk. Des Moines R., St. Joseph, TMM, 9/13/ 11; Kossuth: East Fk. Des Moines R., St. Joseph, TMM, 9/16/11; Wright: PBG Control Wetland, Belmond, MTF, 7/15/08. Distribution: Eastern USA, extending west to Texas in the south, Kansas, and North Dakota in the north. Its habitat preference is deeper areas of standing waters.</p> <p>Desmopachria convexa (Aubé). Johnson: (Wickham 1911); Muscatine: Honey Cr., Conesville, CTH, 7/18/06; Story: (Wickham 1911). Distribution: Transcontinental in the USA, except the southwestern states. New state record.</p> <p>Dytiscus alaskanus Balfour-Browne. Winnebago: Private Wetland, Thompson, MTF, 7/18/07. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest. Its habitat preference is generally ponds or lakes. New state record.</p> <p>Dytiscus circumcinctus Ahrens (Larson et al. 2000). Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Dytiscus cordieri Aubé. Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/07; Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Winnebago: Private Wetland, Leland, MTF, 7/1/06. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest. Its habitat preference is standing waters, ranging from permanent to temporary bodies.</p> <p>Dytiscus dauricus Gebler. Cerro Gordo: Private Wetland, Clear Lake, MTF, 7/2/08. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest. Its habitat preference is permanent ponds in forested areas as well as beaver ponds. New state record.</p> <p>Dytiscus fasciventris Say. Greene: Snake Cr., Rippey, MTF, 8/14/07; Winnebago: Private Wetland, Lake Mills, MTF, 7/18/07. Distribution: Northeastern USA, extending west to Minnesota in the north and Iowa. Iowa forms the southwestern edge of the range. Its habitat preference includes ponds and marshes.</p> <p>Dytiscus harrisii Kirby. (Larson et al. 2000). Distribution: Northeastern USA, extending west to North Dakota in the north and Nebraska.</p> <p>Dytiscus hybridus Aubé. Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/07; Clay: Private Wetland, Gillet Grove, MTF, 7/18/ 08; Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Emmet: Private Wetland, Wallingford, MTF, 7/16/08; Floyd: Little Cedar R., Colwell County Park, CTH, 9/9/09; Hancock: Pilot Knob State Park, Forest City, MTF, 7/18/07; Hardin: Private Wetland, Buckeye, MTF, 8/14/ 07; Johnson: (Wickham 1911); Kossuth: East Fk. Des Moines R., Seneca, TMM, 8/12/15; Pottawattamie: (Wickham 1911); Winnebago: Private Wetland, Forest City, MTF, 7/18/07; Winneshiek: UV light, KR, 11/3/01. Distribution: Throughout the eastern and northern USA, extending to the Great Plains with limited collections on the West Coast. Its habitat preference is ponds and marshes, although two stream collections occurred in Iowa.</p> <p>Dytiscus verticalis Say. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/27/14. Distribution: Northeastern USA, extending west to Minnesota in the north and Colorado. New state record.</p> <p>Eretes explicitus Miller. Fremont: Hamburg, Bailey, and Davis, 7/19/36 (Snow Entomology Museum). Distribution: Southcentral USA, extending from Arizona to Texas in the south and north to Iowa, which forms the northeastern border. New state record.</p> <p>Graphoderus fascicollis (Harris). Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/ 07; Polk: Grimes, Unknown collector, 7/24/ 1919. Distribution: Northeastern USA, extending west to Montana in the north, although Iowa forms the southwestern border. Its habitat preference is emergent zones within permanent ponds, usually in areas with deciduous forests.</p> <p>Graphoderus liberus (Say). Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Johnson: Ciha Fen, Sutliff, CTH, 4/17/17; Story: (Wickham 1911); Winnebago: Private Wetland, Forest City, MTF, 7/18/07. Distribution: Primarily eastern USA, east of the Great</p> <p>Plains, although there are Pacific Northwest records. Its habitat preference is boggy ponds and lakes.</p> <p>Graphoderus manitobensis Wallis. Hancock: Private Wetland, Garner, MTF, 7/10/08. Distribution: Northcentral USA, with localized populations only collected from Wisconsin, Minnesota, and Iowa in the USA, thus Iowa forms the southern border. Its habitat preference is large sedge and cattail marshes/ponds.</p> <p>Graphoderus occidentalis Horn. Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Emmet: Private Wetland, Wallingford, MTF, 7/27/07; Hancock: Eagle Lake, Duncan, MTF, 7/ 20/07; Winnebago: Private Wetland, Thompson, MTF, 7/18/07; Wright: Lower Morse Lake, Belmond, MTF, 7/27/07. Distribution: Northwestern USA, extending east to Wisconsin and Illinois. Its habitat preference is open ponds, both permanent and temporary.</p> <p>Graphoderus perplexus Sharp. Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/ 07; Davis: Soap Cr., Eldon, CTH, 7/23/02; Dickinson: Jemmerson Slough, Spirit Lake, MTF, 7/24/07; Hancock: Pilot Knob Wetland, Forest City, MTF, 7/18/07; Kossuth: Kettle Wetland-Union Slough NWR, Bancroft, CTH, 5/ 6/97; Mahaska: North Skunk R., CTH, 8/14/02; Winnebago: Private Wetland, Thompson, MTF, 7/18/07. Distribution: Northern transcontinental in the USA. Iowa appears to be the southern edge of its distribution in the Midwest. New state record.</p> <p>Heterosternuta diversicornis (Sharp). Page: East Nishnabotna R., Shenandoah, TMM, 9/18/08. Distribution: Southcentral USA, having been collected from Texas, Oklahoma, and Kansas. Specimens have been collected from grassy vegetation overhanging streams in Texas to temporary ponds in Kansas. Iowa is on the northern edge of its distribution. New state record.</p> <p>Heterosternuta pulchra (LeConte). Clayton: Peck Cr., Osterdock, CTH, 7/26/11; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Fayette: Deep Cr., Wadena, CTH, 9/28/05; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 8/27/07; Winneshiek: Ten Mile Cr., Decorah, DPH, 8/23/05. Distribution: Eastern USA, exclusive of the bottom tier of southern states. Iowa is on the northwestern edge of its distribution with collections only from the northeastern quarter of the state. Its habitat preference is the mineral substrates on the margin of streams, as well as lake shores and quarry pools. New state record.</p> <p>Heterosternuta wickhami (Zaitzev). Allamakee: Hecker Cr., Postville, CTH, 7/24/07; Black Hawk: West Fk.Cedar R., Finchford, MWB, 9/ 15/08; Bremer: Horton Cr., Horton, CTH, 7/24/ 01; Buchanan: Lime Cr., Brandon, CTH, 9/4/07; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Cherokee: Maple R., Aurelia, CTH, 8/6/97; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 6/13/11; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Delaware: Fenchell Cr., Strawberry Point, CTH, 9/17/07; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08; Fayette: Nutting Cr., Clermont, CTH, 7/26/06; Floyd: Cedar R., Charles City, MWB, 9/ 9/08; Franklin: Bailey Cr., Sheffield, CTH, 7/ 22/96; Grundy: South Beaver Cr., Parkersburg, CTH, 7/30/96; Ida: Maple R., Ida Grove, CTH, 8/5/97; Iowa: Big Bear Cr., Victor, CTH, 9/20/ 99; Jackson: Storybook Hollow, Bellevue, CTH, 8/29/07; Jasper: North Skunk R., Newton, CTH, 8/8/01; Jones: Helmer Cr., Prairieburg, CTH, 8/ 31/05; Kossuth: Plum Cr., Wesley, TMM, 8/15/ 06; Lee: Lick Cr., Shimek State Forest, CTH, 9/ 10/10; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Shelby: East Branch West Nishnabotna R., Avoca, TMM, 8/16/07; Story: College Cr., Ames, TMM, 9/24/07; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/ 04; Warren: North R., Norwalk, KLS, 10/8/04; Winnebago: Little Buffalo Cr., Thompson, CTH, 7/27/04; Wright: Buttermilk Cr., Goldfield, KJS, 9/6/06. Distribution: Eastern USA, extending west to Kansas in the south, Iowa, and Minnesota in the north. Iowa forms part of the western border. Its habitat preference is the gravelly margins of medium to small streams.</p> <p>Hydaticus aruspex Clark. Cerro Gordo: Private Wetland, Clear Lake, MTF, 7/2/08; Dickinson: Lake Okoboji, LLB – Smithsonian Institute collection, 8/22/1916; Emmet: Drainage Ditch #40, Dolliver, TMM, 7/30/15; Hancock: Private Wetland, Garner, 7/10/08. Distribution: Transcontinental in the USA, exclusive of the southeastern states. Its habitat preference tends to be larger permanent bodies of standing water with deep detritus and emergent vascular plants.</p> <p>Hydaticus piceus LeConte. Clay: Trumball Lake East, Dickens, MTF, 8/8/07; Johnson: (Wickham 1911); Polk: Grimes, Unknown collector, 7/ 24/1919; Story: (Wickham 1911); Winnebago: Pilot Knob Wetland, Forest City, MTF, 8/8/07. Distribution: Northeastern USA, with Iowa forming part of the western border and Missouri</p> <p>the southwestern edge. Its habitat preference is characterized by woodland pools with emergent aquatic vascular plants and an organic substrate.</p> <p>Hydrocolus paugus (Fall). (Larson et al. 2000). Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Hydrocolus stagnalis (Gemminger and Harold). Johnson: (Wickham 1911); Story: (Wickham 1911). Distribution: Northeastern USA, with Iowa forming the southwestern border.</p> <p>Hydroporus dichrous Melsheimer. Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/ 14/01; Louisa: Honey Cr., Conesville, CTH, 8/ 24/04; Muscatine: Pike Run, Nichols, CTH, 8/1/ 16. Distribution: Northeastern USA with Iowa forming part of the western border and Missouri the southwestern edge. Its habitat preference is the margin of small ponds and very slow streams with abundant vegetation and detritus.</p> <p>Hydroporus niger Say. Guthrie: Unnamed trib. Middle Raccoon R., Panora, TMM, 8/5/13; Johnson: (Wickham 1911); Muscatine: Pike Run, Nichols, CTH, 8/20/12; Scott: (Wickham 1911). Distribution: Eastern USA, extending west to South Dakota in the north, Nebraska, Kansas, and Texas in the south. Its habitat preference is emergent vegetation in ponds, or slow flowing streams.</p> <p>Hydroporus notabilis LeConte. (Larson et al. 2000). Clayton: Mill Cr., Osterdock, CTH, 9/6/ 11. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Hydroporus signatus Mannerheim. Linn: Rock Island Preserve (UV light), DAV, 3/13/07. Distribution: Transcontinental in the USA, exclusive of southwestern states. Its habitat preference is small mineral substrate pools and vegetated peat land pools. New state record.</p> <p>Hydrovatus pustulatus (Melsheimer). Allamakee: Hecker Cr., Postville, CTH, 8/14/00; Clinton: Brophy Cr., McCausland, CTH, 8/14/12; Mahaska: Buck Cr., Barnes City, CTH, 7/26/ 12; Muscatine: Pike Run, West Liberty, CTH, 7/19/06. Distribution: Eastern USA, extending west to North Dakota in the north and Texas in the south. Its habitat preference is among vegetation and debris in shallow areas of warm ponds, although the Iowa records were from smaller streams. New state record.</p> <p>Hygrotus acaroides (LeConte). Johnson: Iowa City (Wickham 1911); (Downie and Arnett 1996;</p> <p>Larson et al. 2000). Distribution: Central USA,</p> <p>although records extend west to Montana and Oregon.</p> <p>Hygrotus impressopunctatus (Schaller). Winneshiek: UV light, KR, 11/3/01; (Larson et al. 2000). Distribution: Transcontinental in the USA, except the southeastern states.</p> <p>Hygrotus patruelis (LeConte). (Downie and Arnett 1996; Larson et al. 2000). Distribution: Western USA, extending east to Wisconsin and Illinois.</p> <p>Hygrotus sayi Balfour-Browne. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Clay: Willow Cr., Royal, CTH, 9/13/99; Clinton: Brophy Cr., McCausland, CTH, 8/19/ 14; Dickinson: Milford Cr., Milford, JML, 9/5/ 01; Dubuque: Cloie Cr., Durango, CTH, 8/13/ 13; Hancock: Plum Cr., Woden, TMM, 5/24/16; Jackson: South Big Mill Cr., Bellevue, CTH, 9/ 17/14; Johnson: Ciha Fen, Sutliff, CTH, 4/17/ 17; Kossuth: Little Buffalo Cr., Titonka, JML, 8/ 21/00; Mitchell: Deer Cr., Carpenter, CTH, 8/25/ 08; Muscatine: Pike Run, West Liberty, CTH, 7/ 19/06; Polk: Camp Cr., Mitchellville, TMM, 7/ 20/05; Winneshiek: UV light, KR, 11/3/01; Worth: Elk Cr., Kensett, TMM, 9/29/15; Wright: Treatment Wetland, Belmond, MTF, 5/9/08. Distribution: Transcontinental in the USA, exclusive of the southeastern states. Its habitat preference is sun-warmed margins of ponds and slow flowing streams.</p> <p>Ilybiosoma seriatum (Say). Allamakee: French Cr., Churchtown, CTH, 9/5/05; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/23/11; Cedar: Unnamed trib. to Bennett Cr., Tipton, CTH, 7/16/12; Cherokee: Unnamed trib. to Little Sioux R., Cherokee, TMM, 6/7/16; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 8/13/13; Clayton: South Cedar Cr., Garnavillo, CTH, 9/18/07; Clinton: Elk R., Teeds Grove, CTH, 8/12/13; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 9/9/04; Fayette: Mink Cr., Wadena, MWB, 7/13/06; Guthrie: Springbrook Cr., Guthrie Center, TMM, 7/16/13; Jackson: South Big Mill Cr., Bellevue, CTH, 8/6/14; Mitchell: Turtle Cr., St. Ansgar, CTH, 9/10/07; Monona: Unnamed trib. to Norway Cr., Soldier, JML, 6/16/03; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 6/24/13; Plymouth: Perry Cr., Hinton, KLS, 7/20/04; Pottawattamie: Unnamed trib. Silver Cr., Shelby, TMM, 8/19/14; Scott: Duck Cr., Blue Grass, CTH, 6/12/13; Sioux: Dry Cr., Sioux Center, JML, 7/2/05; Story: Unnamed trib. Willow Cr., Collins, TMM, 8/25/14; Tama:</p> <p>South Branch Salt Cr., Clutier, CTH, 8/7/14; Van</p> <p>Buren: Coppers Cr., Keosauqua, CTH, 6/7/11; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/ 29/08; Woodbury: Rock Cr., Anthon, TMM, 6/ 25/13. Distribution: Transcontinental in the USA, exclusive of extreme south-central and southeastern states. Its habitat preference is flowing water, usually springs and small creeks.</p> <p>Ilybius biguttulus (Germar). Allamakee: French Cr., Churchtown, CTH, 7/24/00; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Clayton: Wolf Cr., Volga, CTH, 9/10/12; Davis: Burr Oak Cr., Troy, CTH, 6/5/13; Johnson: (Wickham 1911); Lee: Bonnell Cr., Franklin, CTH, 7/16/02; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Ringgold: Middle Fk. Grand R., Mt. Ayr, TMM; Story: (Wickham 1911); Van Buren: Shimek State Forest (UV light), ELF, 5/18/12; Winneshiek: Coldwater Cr., Bluffton, CTH, 9/ 22/04; Woodbury: Rock Cr., Anthon, TMM, 6/ 25/13; Wright: Lower Morris Lake, Belmond, MTF, 7/27/07. Distribution: transcontinental in the USA, except most states west of the Rockies and extreme southcentral and southeastern states. Its habitat preference is mostly lentic habitats, sun warmed, and vegetated. The majority of collection sites in Iowa were from flowing streams.</p> <p>Ilybius fraterculus (LeConte). Cerro Gordo: Private Wetland, Ventura, MTF, 7/20/07; Clay: Unnamed trib. Little Sioux R., Peterson, TMM, 8/18/09; Clayton: Mill Cr., Osterdock, CTH, 7/ 25/11; Dickinson: Jemmerson Slough, Spirit Lake, MTF, 7/24/07; Emmet: Private Wetland, Wallingford, MTF, 7/27/07; Ida: Maple R., Galva, CTH, 8/6/97; Winnebago: Private Wetland, Forest City, MTF, 7/18/07. Distribution: Transcontinental in the USA, exclusive of the southeastern states, but collections are only common west of the Great Lakes. Its habitat preference is grassland ponds where vegetation is abundant, but Iowa records include some collections from flowing streams.</p> <p>Ilybius incarinatus Zimmermann. (Larson et al. 2000). Distribution: Northeastern USA, extending west to Minnesota and Iowa. Iowa forms the southwestern border of the range.</p> <p>Ilybius pleuriticus LeConte. Harrison: (Wickham 1911); Johnson: (Wickham 1911); Story: (Wickham 1911). Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border in the Midwest.</p> <p>Laccophilus fasciatus Aubé. Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Hecker Cr., Postville, CTH, 7/31/06;</p> <p>Appanoose: Cooper Cr., Centerville, CTH, 8/11/ 05; Clayton: Honey Cr., Littleport, CTH, 7/23/ 12; Decatur: Redmans Branch, Dekalb WMA, TMM, 9/15/15; Franklin: Beaver Cr., Ackley, CTH, 9/10/14; Guthrie: South Raccoon R., Stuart, TMM, 7/25/12; Jasper: Walnut Cr., Prairie City, CTH, 10/8/04; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Marion: Cedar Cr., Marysville, JML, 8/7/06; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Page: East Nishnabotna R., Shenandoah, TMM, 7/16/12; Pottawattamie: Mosquito Cr., Manawa, TMM, 9/24/12; Ringgold: Middle Fk. Grand R., Mt. Ayr, KLS, 7/14/04; Scott: Lost Cr., Princeton, CTH, 8/9/12; Van Buren: Coppers Cr., Keosauqua, CTH, 10/6/09; Wapello: Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/ 11/05; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 9/17/12. Distribution: Transcontinental in the USA, except the Rocky Mountains and Pacific Northwest. Its habitat preference is fine substrate in temporary ponds, although Iowa records are all flowing systems, some of which are intermittent.</p> <p>Laccophilus maculosus Say. Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Bremer: Horton Cr., Horton, CTH, 7/24/01; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cedar: Sugar Cr., Tipton, CTH, 8/23/01; Cerro Gordo: Winnebago R., Mason City, CTH, 9/4/13; Chickasaw: Wapsipinicon R., New Hampton, CTH, 10/6/04; Clay: Prairie Cr., Spencer, JML, 9/4/01; Clayton: Miners Cr., Guttenberg, CTH, 10/8/07; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Davis: North Fabius R., Moulton, CTH, 7/17/ 06; Decatur: Short Cr., Dekalb WMA, TMM, 9/ 15/15; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dickinson: Little Sioux R., Lake Park, TMM, 9/29/11; Dubuque: Hickory Cr., New Vienna, CTH, 9/18/08; Fayette: Deep Cr., Wadena, CTH, 9/28/05; Floyd: Cedar R., Charles City, MWB, 9/9/08; Franklin: Maynes Cr., Hampton, KLS, 8/23/01; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: Middle Fk. South Beaver, Wellsburg, CTH, 8/14/01; Hamilton: Boone R., Stratford, TMM, 8/16/12; Hancock: Winnebago R., Forest City, CTH, 9/18/06; Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/14/01; Harrison: Soldier R., Pisgah, CTH, 10/27/99; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Lourdes, CTH, 9/28/09; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 10/9/07; Jasper: North Skunk R., Kellogg, CTH, 8/7/01; Johnson: Iowa R., Iowa City, CTH, 9/22/05; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Honey Cr., Conesville, CTH, 8/24/04; Lyon: Little Rock R., George, TMM, 9/28/11; Madison: North Branch North R., Goeldner Woods, TMM, 7/18/12; Mahaska: North Skunk R., Rose Hill, CTH, 8/14/02; Marion: Cedar Cr., Marysville, JML, 8/7/06; Mills: West Nishnabotna R., Malvern, TMM, 9/4/12; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Muscatine: Pike Run, West Liberty, CTH, 7/21/05; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/14; Page: East Nishnabotna R., Shenandoah, TMM, 8/5/13; Palo Alto: Prairie Cr., Whittemore, CTH, 10/2/01; Plymouth: Big Sioux R., Akron, TMM, 10/10/12; Polk: Four Mile Cr., Ankeny, CTH, 9/30/99; Pottawattamie: Lapworth Cr., Crescent, KLS, 7/1/03; Poweshiek: Walnut Cr., Holiday Lake, CTH, 7/27/09; Ringgold: Middle Fk. Grand R., Mt. Ayr, KLS, 7/14/04; Scott: Lost Cr., Princeton, CTH, 8/30/04; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: Walnut Cr., Huxley, TMM, 8/22/11; Tama: Little Wolf Cr., Gladbrook, CTH, 7/1/04; Union: Sand Cr., Arispe, JML, 6/11/03; Van Buren: Des Moines R., Farmington, CTH, 9/6/ 06; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Warren: North R., Norwalk, KLS, 10/ 8/04; Washington: English R., Riverside, CTH, 9/24/02; Wayne: Jordan Cr., Millerton, TMM, 8/ 24/12; Webster: Holiday Cr., Coalville, KLS, 8/ 2/07; Winnebago: Beaver Cr., Lake Mills, CTH, 7/22/03; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08; Woodbury: Wolf Cr., Correctionville, TMM, 9/22/14; Worth: Elk Cr., Kensett, CTH, 8/26/08; Wright: Otter Cr., Goldfield, MWB, 10/13/04. Distribution: Transcontinental in the USA, although there are three subspecies. Habitat preference is sunwarmed shallows of permanent pools and ponds, although most Iowa records were from flowing streams.</p> <p>Laccophilus proximus Say. Allamakee: Clark Cr., New Albin, CTH, 9/22/14; Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Benton: Bear Cr., Shellsburg, CTH, 10/1/01; Black Hawk: Dry Run Cr., Cedar Falls, CTH, 9/14/10;</p> <p>Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 9/20/10; Clinton: Brophy Cr., McCausland, CTH, 8/11/04; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 8/12/10; Jackson: Bear Cr., Baldwin, CTH, 8/28/03; Jasper: Walnut Cr., Prairie City, CTH, 10/3/ 03; Johnson: State Hygienic Lab parking lot puddle, CTH, 7/13/15; Kossuth: East Fk. Des Moines R., St. Joseph, TMM, 9/10/13; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Polk: Four Mile Cr., Ankeny, CTH, 9/30/99; Pottawattamie: Mosquito Cr., Underwood, JML, 10/11/00; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/17/08; Shelby: East Branch West Nishnabotna R., Avoca, CTH, 9/21/94; Tama: South Branch Salt Cr., Clutier, CTH, 8/7/14; Van Buren: Coppers Cr., Keosauqua, CTH, 9/15/10; Winneshiek: Yellow R., Castalia, CTH, 9/15/04. Distribution: Eastern USA, extending west to the Rocky Mountains. Laccophilus proximus is characterized as a pioneer species as it commonly occurs in temporary habitats, although the Iowa records were all from flowing streams.</p> <p>Liodessus affinis (Say). Allamakee: Upper Iowa R., Dorchester, MWB, 10/6/05; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Black Hawk: West Fk.Cedar R., Finchford, MWB, 9/15/08; Buchanan: Pine Cr., Quasqeton, CTH, 9/1/15; Butler: Boylan Cr., Aredale, MWB, 11/11/02; Cedar: Sugar Cr., Tipton, CTH, 8/23/01; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/ 12; Cherokee: Mill Cr., Larrabee, CTH, 10/7/98; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 10/14/10; Clay: Little Sioux R., Spencer, KLS, 9/10/13; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Tetes Des Morts Cr., St. Donatus, CTH, 8/1/05; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Cedar R., Floyd, CTH, 9/23/08; Franklin: Beaver Cr., Ackley, MWB, 8/12/05; Grundy: South Beaver Cr., Parkersburg, CTH9/10/14; Hamilton: South Skunk R., Ellsworth, CTH, 9/23/97; Hancock: East Branch Iowa River, Goodell, TMM, 10/7/10; Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/14/01; Howard: Crane Cr., Riceville, CTH, 7/10/06; Iowa: Old Mans Cr., Ladora, CTH, 10/2/13; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Kossuth: East Fk. Des Moines R., Irvington, MWB, 8/25/03; Linn: Indian Cr., Cedar Rapids, CTH, 9/19/00; Mahaska: South Skunk R., Oskaloosa, CTH, 10/7/13; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Muscatine: Cedar R., Conesville, CTH, 8/29/ 12; O’ Brien: Floyd R., Sheldon, CTH, 9/14/99; Page: East Nishnabotna R., Shenandoah, TMM, 9/18/08; Plymouth: Floyd R., James, CTH, 10/ 26/99; Pottawattamie: Jordan Cr., Cumberland, CTH, 9/20/94; Sac: North Raccoon R., Sac City, CTH, 10/8/98; Story: South Skunk R., Story City, CTH, 9/23/97; Tama: Little Wolf Cr., Gladbrook, CTH, 7/1/04; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Van Buren: Coppers Cr., Keosauqua, CTH, 10/6/09; Warren: North R., Norwalk, CTH, 10/15/99; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08. Distribution: Transcontinental in the USA. Its habitat preference is widely varied, ranging from temporary ponds to slow streams.</p> <p>Liodessus flavicollis (LeConte). Henry: Skunk R., Oakland Mills, MWB, 9/24/08; Johnson: Iowa R., Iowa City, CTH, 8/8/05. Distribution: Eastern USA. Iowa is on the western edge of its distribution. Its habitat preference is relatively deep water, with many records from clear water pools or ponds with sandy substrates. Within Iowa, collection records were from larger, predominantly finer substrate rivers. New state record.</p> <p>Liodessus obscurellus (LeConte). (Larson et al. 2000). Distribution: Western USA, extending east to Wisconsin in the north, Indiana, and Louisiana in the south.</p> <p>Lioporeus triangularis (Fall). Black Hawk: West Fk. Cedar R., Finchford, MWB, 8/6/12. Distribution: Eastern USA. Specimens have only been collected in the northeastern quarter of the state from two locations in the same county. Iowa forms part of the northwestern edge of its distribution. Its habitat preference is the margin and undercuts of small, clear streams, although the Iowa collection locations are sandy, mediumsized rivers. New state record.</p> <p>Matus bicarinatus (Say). Clinton: Elk R., Andover, CTH, 8/7/07; Jackson: Bear Cr., Eden Valley County Park, CTH, 8/31/11; Johnson: (Wickham 1911); Muscatine: Pike Run, Nichols, CTH, 8/23/04; Scott: (Wickham 1911); Story: (Wickham 1911). Distribution: Eastern USA, with Iowa forming part of the western border in the Midwest. Its habitat preference is permanent ponds and marshes near, or associated with, larger streams.</p> <p>Matus ovatus Leech. Johnson: Iowa City, HFW, 7/ 28/1898. Distribution: Eastern USA, extending west to Wisconsin in the north, Iowa, and Texas in the south.</p> <p>Neoporus clypealis (Sharp). Black Hawk: Beaver Cr., Cedar Falls, MWB, 8/6/12; Buchanan: Wapsipinicon R., Independence, MWB, 8/6/12; Butler: West Fk. Cedar R., Lake Considine County Park, CTH, 10/2/12; Cedar: UNT Bennett Cr., Tipton, CTH, 7/16/12; Clinton: Brophy Cr., McCausland, CTH, 8/11/04; Louisa: Honey Cr., Conesville, CTH, 8/21/12; Muscatine: Honey Cr., Conesville, CTH, 7/18/ 06; Scott: Lost Cr., Princeton, CTH, 8/30/04; Worth: Beaver Cr., Fertile, KLS, 9/19/06. Distribution: Eastern USA, with Iowa forming the western border. Within Iowa, collections were primarily in the northeastern quarter. Its habitat preference is among emergent vegetation along the margins of slow, marshy streams, or small lakes.</p> <p>Neoporus dimidiatus (Gemminger and Harold). Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: French Cr., Churchtown, CTH, 7/1/96; Appanoose: Chariton R., Rathbun, CTH, 8/17/04; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Buchanan: Bear Cr., Brandon, CTH, 8/26/13; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cass: East Nishnabotna R., Lewis, KLS, 9/19/02; Cedar: Rock Cr., Tipton, CTH, 8/20/08; Cerro Gordo: Winnebago R., Mason City, CTH, 9/4/13; Cherokee: Mill Cr., Larrabee, TMM, 9/18/12; Clarke: South Whitebreast Cr., Weldon, TMM, 8/17/15; Clay: Henry Cr., Peterson, TMM, 8/23/10; Clayton: Hewitt Cr., Volga, MWB, 9/1/04; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Crawford: Otter Cr., Deloit, TMM, 10/11/10; Dallas: Mosquito Cr., Panora, JML, 8/8/02; Davis: Soap Cr., Floris, CTH, 7/24/02; Decatur: Long Cr., Van Wert, JML, 10/9/01; Delaware: Fenchel Cr., Strawberry Point, CTH, 9/17/07; Dubuque: Cloie Branch, Durango, CTH, 8/5/ 14; Fayette: Bell Cr., Clermont, CTH, 7/20/04; Floyd: Cedar R., Carrville, CTH, 9/4/13; Franklin: Beaver Cr., Ackley, CTH, 8/22/13; Grundy: South Beaver Cr., Parkersburg, CTH, 8/13/01; Guthrie: Middle Raccoon R., Bagley, KLS, 9/20/05; Hardin: Iowa R., Eldora, CTH, 8/16/05; Harrison: Soldier R., Pisgah, KLS, 10/ 22/02; Henry: Big Cr., New London, CTH, 8/ 30/05; Howard: Crane Cr., Riceville, CTH, 8/ 16/10; Jackson: North Fk. Maquoketa R., Maquoketa, CTH, 10/10/02; Jasper: Mud Cr., Baxter, TMM, 9/14/04; Johnson: Iowa R., Iowa City, CTH, 9/21/05; Jones: Bear Cr., Wyoming, CTH, 7/13/04; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Skunk R., Augusta, CTH, 10/8/12; Linn: Indian Cr., Cedar Rapids, CTH, 9/18/00; Louisa: Long Cr., Columbus Jct., CTH, 10/1/13; Lucas: Whitebreast Cr., Lacona, CTH, 8/1/97; Madison: Middle R., Winterset, CTH, 9/16/98; Mahaska: Muchakinock Cr., Eddyville, CTH, 8/30/00; Marion: English Cr., Knoxville, TMM, 8/14/14; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mills: Keg Cr., Mineola, CTH, 7/30/97; Mitchell: Wapsipinicon R., McIntire, CTH, 8/17/10; Monona: Soldier R., Moorhead, KLS, 10/6/ 04; Montgomery: Walnut Cr., Red Oak, CTH, 10/14/96; Muscatine: Mud Cr., Wilton, CTH, 9/ 15/03; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Plymouth: Beaver Cr., Akron, KLS, 8/12/13; Polk: Camp Cr., Mitchellville, TMM, 7/20/05; Pottawattamie: East Nishnabotna R., Griswold, JML, 9/19/02; Poweshiek: West Fk.trib. to Arbor Lake, Grinnell, KLS, 10/10/03; Ringgold: Lotts Cr., Lamoni, TMM, 7/21/14; Shelby: Mosquito Cr., Panama, JML, 9/5/02; Story: College Cr., Ames, TMM, 9/24/07; Taylor: Honey Cr., Bedford, KBK, 8/18/04; Union: Unnamed trib. to Thompson R., Lorimor, TMM, 9/17/15; Van Buren: Chequest Cr., Pittsburg, CTH, 7/17/97; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, TFW, 8/20/02; Wayne: Unnamed trib. to Steel Cr., Humeston, TMM, 6/17/14; Winneshiek: Coon Cr., Decorah, CTH, 8/30/99; Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Transcontinental in the USA, except for extreme western states. Its habitat preference is spring-fed pools, small ponds, and shorelines of lakes and larger rivers, although nearly all Iowa records are wadeable streams. New state record.</p> <p>Neoporus hybridus (Aubé). Des Moines: Brush Cr., Burlington, CTH, 9/29/00; Louisa: Iowa R., Wapello, CTH, 9/24/08; Scott: (Wickham 1911). Distribution: Eastern USA, with Iowa forming the northwestern border.</p> <p>Neoporus striatopunctatus (Melsheimer). Appanoose: Packard Cr., Exline, CTH, 9/15/15; Davis: North Fabius R., Moulton, CTH, 7/17/ 06; Des Moines: Flint Cr., Danville, CTH, 7/15/ 03; Johnson: (Wickham 1911); Lee: Lick Cr., Farmington, CTH, 7/14/03; Marion: English Cr., Knoxville, MWB, 7/18/05; Van Buren: Coppers Cr., Keosauqua, CTH, 10/6/09; Warren: South R, Ackworth, TMM, 8/19/13. Distribution: Eastern USA with Iowa forming the northwestern edge of its range.</p> <p>Neoporus superioris (Balfour-Browne). Allamakee: Yellow R., Ion, CTH, 9/14/04; Winnebago: Beaver Cr., Lake Mills, CTH, 7/22/03. Distribution: Northcentral USA, with Iowa forming part of the southern edge of its distribution. Its habitat preference is permanent warm water in slow moving streams and impoundments on streams. New state record.</p> <p>Neoporus tennetum (Wolfe). Howard: Little Turkey R., Protivin, CTH, 8/19/02. Distribution: Northeastern USA, extending west to Iowa, Missouri, and Kansas. Its habitat preference is small to medium creeks in silty, backwater areas. New state record.</p> <p>Neoporus undulatus (Say). Allamakee: Dousman Cr., Ion, CTH, 8/12/15; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: West Fk. Cedar R., Finchford, CTH, 9/9/14; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/23/11; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 7/21/10; Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Clinton: Brophy Cr., McCausland, CTH, 8/11/ 04; Fayette: Little Volga R., Maynard, CTH, 7/ 23/14; Guthrie: Unnamed trib. Middle Raccoon R., Panora, TMM, 6/16/14; Harrison: Potato Cr., Persia, KLS, 7/12/05; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Jackson: Tetes Des Morts,Cr., St. Donatus, CTH, 9/6/11; Johnson: Ciha Fen, Sutliff, CTH, 4/17/17; Jones: Prairie Cr., Scotch Grove, CTH, 7/23/ 03; Lee: Lick Cr., Shimek State Forest, CTH, 7/ 28/14; Linn: Cedar R., Cedar Rapids, CTH, 9/ 19/13; Louisa: Honey Cr., Conesville, CTH, 8/ 21/12; Lucas: (Wickham 1911); Madison: Howerdon Cr., Winterset, JML, 7/12/01; Muscatine: Pike Run, Nichols, CTH, 9/1/98; Pottawattamie: Lapworth Cr., Crescent, KLS, 7/1/03; Poweshiek: Walnut Cr., Hartwick, CTH, 7/27/09; Story: Dye Cr., Colo, TMM, 6/19/13; Tama: Otter Cr., Chelsea, DPH, 4/16/04; Van Buren: Town Branch, Pittsburg, KJS, 7/29/14; Wayne: Unnamed trib. to Steel Cr., Humeston, TMM, 6/20/13; Winnebago: Winnebago R., Lake Mills, TMM, 9/28/15; Winneshiek: UV light, KR, 11/3/01; Worth: Elk Cr., Kensett, TMM, 9/29/15. Distribution: Transcontinental in the USA. Its habitat preference is permanent habitats where there is some flowing water.</p> <p>Neoporus venustus (LeConte). (Downie and Arnett 1996). Distribution: Eastern USA, with Iowa forming part of the western border.</p> <p>Neoporus vittatus (LeConte). Boone: Unnamed trib. Little Beaver Cr., Bouton, JML, 10/4/06; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Calhoun: Unnamed trib. Purgatory Cr., Lohrville, KLS, 8/14/07; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 7/21/10; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Dubuque: Hickory Cr., New Vienna, CTH, 9/18/ 08; Fayette: Dibble Cr., Clermont, CTH, 8/14/ 13; Franklin: Maynes Cr., Hampton, CTH, 8/18/ 08; Guthrie: Unnamed trib. to Middle Raccoon R., Panora, TMM, 6/16/14; Hamilton: White Fox Cr., Webster City, CTH, 8/25/97; Hancock: Unnamed trib. to Drainage Ditch 114, Gladfelter Marsh, TMM, 8/29/11; Hardin: Unnamed trib. to South Fk. Iowa R., Buckeye, TMM, 5/24/12; Howard: Crane Cr., Riceville, CTH, 7/10/06; Ida: Silver Cr., Ida Grove, CTH, 8/5/97; Jasper: North Skunk R., Kellogg, CTH, 8/7/01; Johnson: Old Womans Cr., Coralville, CTH, 9/16/15; Mitchell: Deer Cr., Carpenter, CTH, 8/25/08; O’ Brien: Little Waterman Cr., Hartley, CTH, 7/9/96; Story: Unnamed trib. to Willow Cr., Collins, TMM, 8/25/14; Webster: Prairie Cr., Lehigh, TMM, 8/2/11; Winnebago: Beaver Cr., Lake Mills, CTH, 7/22/03; Winneshiek: Silver Cr., Burr Oak, CTH, 8/16/11; Wright: Buttermilk Cr., Goldfield, KJS, 9/6/06. Distribution: Northcentral USA, with Iowa forming part of the southern edge of its distribution. Its habitat preference is small prairie streams. New state record.</p> <p>Platambus semivittatus (LeConte). Allamakee: French Cr., Churchtown, CTH, 9/10/02; Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Buchanan: Lime Cr., Brandon, CTH, 8/3/10; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/27/13; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 9/22/15; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 5/18/10; Clayton: Hewitt Cr., Volga, MWB, 9/1/04; Davis: Burr Oak Cr., Troy, CTH, 6/5/13; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/7; Dickinson: Private Wetland, Spirit Lake, MTF, 7/24/07; Dubuque: Hickory Cr., New Vienna, CTH, 8/11/99; Emmett: Drainage Ditch 40, Dolliver, TMM, 7/30/15; Fayette: Unnamed trib. Dry Branch Turkey R., Fort Atkinson, MWB, 6/27/ 06; Guthrie: Springbrook Cr., Springbrook State Park, TMM, 6/24/14; Hancock: Unnamed trib. to Drainage Ditch 114, Gladfelter Marsh, TMM, 8/29/11; Harrison: Boyer R., Missouri Valley, TMM, 9/24/14; Howard: Crane Cr., Riceville, CTH, 7/10/06; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: Old Womans Cr., Coralville, CTH, 9/16/15; Lee: Bonnell Cr., Franklin, CTH, 7/16/02; Marshall: Unnamed trib. to Linn Cr., State Center, TMM, 8/26/15; Pottawattamie: Lapworth Cr., Crescent, KLS, 7/1/03; Sioux: Dry Cr., Hawarden, KLS, 6/28/ 04; Story: Ballard Cr., Cambridge, KJS, 7/18/ 07; Tama: South Branch Salt Cr., Clutier, CTH, 8/7/14; Van Buren: Town Branch, Pittsburg, CTH, 7/29/14; Wayne: Unnamed trib. to Steel Cr., Humeston, TMM, 6/20/13; Winnebago: Private Wetland, Forest City, MTF, 7/18/07; Winneshiek: Middle Bear Cr., Highlandville, CTH, 7/27/03; Woodbury: Unnamed trib. to Little Sioux R., Correctionville, TMM, 9/22/14; Wright: Buttermilk Cr., Goldfield, TMM, 9/4/07. Distribution: Transcontinental in the USA. Its habitat preference is lotic systems.</p> <p>Platambus obtusatus (Say). Johnson: Iowa City (Wickham 1911); (Downie and Arnett 1996). Distribution: Eastern USA, with Iowa forming part of the western border.</p> <p>Rhantus calidus (Fabricius). Johnson: North Liberty, MHB, 7/14/15. Distribution: Southeastern USA, with Iowa forming the northwestern border. New state record.</p> <p>Rhantus consimilis Motschulsky. Cerro Gordo: Private Wetland, Clear Lake, MTF, 7/2/08. Distribution: Northern transcontinental in the USA. Iowa forms part of the southern border in the Midwest. Its habitat preference is warm, weedy, temporary or permanent ponds, generally in open grassland areas. New state record.</p> <p>Rhantus gutticollis (Say). Allamakee: French Cr, Churchtown, CTH, 9/16/04; Cerro Gordo: Private Wetland, Clear Lake, MTF, 7/2/08; Clayton: Mill Cr., Osterdock, CTH, 9/6/11; Jackson: South Big Mill Cr., Bellevue, CTH, 8/28/07; Winneshiek: UV light, KR, 11/3/01; Woodbury: Dutch Cr., Danbury, TMM, 6/20/16. Distribution: Western USA, extending to the Midwest. Iowa forms part of the eastern border of its range. Its habitat preference is quiet areas of small, clear streams. New state record.</p> <p>Rhantus sericans Sharp. Cerro Gordo: Union Hills West Wetland, Thornton, MTF, 7/20/07; Dickinson: Private Wetland, Spirit Lake, MTF, 7/24/07; Hancock: Eagle Lake, Duncan, MTF, 7/ 20/07. Distribution: Western USA, extending to the Midwest. Wisconsin, Iowa, and Missouri form part of the eastern border. Its habitat preference is lentic systems, but with a preference for rich, open, sun-warmed ponds.</p> <p>Rhantus wallisi Hatch. (Larson et al. 2000). Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border in the Midwest.</p> <p>Sanfilippodytes pseudovillis (Young). Allamakee: French Cr., Churchtown, CTH, 9/10/02; Clayton: Bear Cr., Edgewood, CTH, 9/9/04; Fayette: Duttons Cr., West Union, CTH, 9/11/07; Jackson: South Big Mill Cr., Bellevue, CTH, 8/6/14;</p> <p>Kossuth: Stork Ditch-Union Slough NWR, Bancroft, CTH, 7/22/97; Winneshiek: Trout R., Decorah, CTH, 7/18/07; Wright: Buttermilk Cr., Goldfield, JML, 9/5/07. Distribution: Scattered throughout the USA, with records from Indiana, Michigan, Utah, and Wisconsin. Its habitat preference is cold stenothermal springs. In Iowa, it has been collected from predicted habitat, as well as waterways supplied by tile-drained land. New state record.</p> <p>Thermonectus basillaris (Harris). Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Clinton: Brophy Cr., McCausland, CTH, 8/14/ 12; Montgomery: East Nishnabotna R., Red Oak, TMM, 10/12/07. Scott: Brophy Cr., McCausalnd, MWB, 8/14/12. Distribution: Eastern, extending west to Wisconsin in the north, Iowa, Missouri, and Texas in the south. Its habitat preference is clear, temporary ponds, although the records from Iowa include a large, sandy river, a sandy drainage ditch, and an intermittent gravel/sand stream. New state record.</p> <p>Thermonectus nigrofasciatus ornaticollis (Aubé). Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Davis: Soap Cr., Floris, CTH, 7/23/02; Emmet: Private Wetland, Wallingford, MTF, 7/ 27/07; Fremont: Hamburg, Bailey and Davis, 7/ 19/36; Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Ida: Little Maple R., Galva, CTH, 8/6/ 97; Johnson: (Wickham 1911); Wright: Lake Cornelia, Clarion, MTF, 7/27/07. Distribution: Transcontinental in the USA except for the extreme western and northwestern states. Its habitat preference is open areas within ponds and lakes, although it has been collected in small to medium-sized streams in Iowa.</p> <p>Uvarus granarius (Aubé). Davis: North Fabius R., Moulton, CTH, 7/17/06; Johnson: (Wickham 1911); Muscatine: Pike Run, Nichols, CTH, 8/1/ 16. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Texas in the south. Iowa forms part of the western border of the range. Its habitat preference is shallow, sunwarmed shorelines of small pools, although in Iowa the Davis County collection record is from a small intermittent, sandy-bottomed stream.</p> <p>Uvarus lacustris (Say). Allamakee: West Branch French Cr., Waukon, CTH, 9/5/13; Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Crawford: Big Cr., Denison, CTH, 8/12/98; Fayette: Volga R., Maynard, CTH, 8/30/10; Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/14/01; Story: (Wickham 1911); Van Buren: Town Branch, Pittsburg, CTH, 9/14/15. Distribution: Eastern USA, extending to the western Great Plains in Colorado. Its habitat</p> <p>preference is small, muddy/clay bottomed pools and shallow, sun-warmed margins of larger water bodies. In Iowa, as reported for Wisconsin, records were from margins of streams.</p></div> 	https://treatment.plazi.org/id/03F487A3FFC8FFC2FEB4FBB39267FE84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFDDFFC7FEB2FEAE9388FC6C.text	03F487A3FFDDFFC7FEB2FEAE9388FC6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Elmidae Curtis 1830	<div><p>ELMIDAE</p> <p>Ancyronyx variegatus (Germar). Allamakee: Yellow R., Postville, CTH, 8/17/11; Black Hawk: Black Hawk Cr., Hudson, MWB, 8/8/02; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/25/14; Buchanan: Lime Cr., Brandon, DPH, 9/4/07; Butler: Beaver Cr., New Hartford, CTH, 8/14/03; Cedar: Rock Cr., Tipton, CTH, 7/30/01; Chickasaw: Crane Cr., Jerico, CTH, 9/7/94; Clayton: Volga R., Elkport, CTH, 10/20/00; Clinton: Barber Cr., Grand Mound, CTH, 9/9/98; Delaware: Plum Cr., Earlville, CTH, 8/16/06; Dubuque: North Fk. Maquoketa R., Dyersville, MWB, 7/25/05; Fayette: Little Turkey R., Gouldsburg County Park, CTH, 9/10/03; Floyd: Little Cedar R., Colwell County Park, MWB, 8/20/02; Grundy: South Beaver Cr., Parkersburg, MWB, 9/3/08; Howard: Crane Cr., Riceville, CTH, 7/10/06; Jackson: Prairie Cr., Maquoketa, CTH, 8/14/01;; Jones: Maquoketa R., Monticello, MWB, 9/6/ 05; Linn: Buffalo Cr., Central City, MWB, 9/2/ 08; Louisa: Iowa R., Columbus Junction, CTH, 9/10/08; Mitchell: Little Cedar R., New Haven, CTH, 8/17/05, Muscatine: Mud Cr., Wilton, DPH, 9/15/03; Scott: Lost Cr., Princeton, CTH, 8/30/04; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05. Distribution: Eastern USA, extending west to Kansas and Texas in the south. Iowa forms part of the western edge of its distribution with collections occurring in the eastern half of the state. New state record.</p> <p>Dubiraphia bivittata (LeConte). Clayton: Roberts Cr., Postville, CTH, 8/14/08; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08. Distribution: Eastern USA, extending west to North Dakota in the north, Missouri, and Iowa. Habitat preferences are coarse substrate, including wood in streams and rivers. New state record.</p> <p>Dubiraphia minima Hilsenhoff. Allamakee: Yellow R., Ion, CTH, 9/14/04; Appanoose: Cooper Cr., Centerville, CTH, 8/11/05; Benton: Bear Cr., Shellsburg, CTH, 10/1/01; Black Hawk: Wapsipinicon R., Dunkerton, CTH, 8/2/ 06; Bremer: East Fk. Wapsipinicon R., Sweet Marsh SWMA, CTH, 8/23/11; Buchanan: Lime Cr., Brandon, CTH, 9/4/07; Buena Vista: Brooke Cr., Rembrandt, JML, 10/5/05; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/13/07; Cass: Sevenmile Cr., Cumberland, TMM, 6/ 28/06; Cedar: Rock Cr., Tipton, CTH, 8/20/ 08; Cherokee: Silver Cr., Cherokee, KLS, 9/20/02; Chickasaw: Wapsipinicon R., New Hampton, CTH, 8/18/04; Clay: Stony Cr., Everly, TMM, 8/30/05; Clayton: Hewitt Cr., Volga, MWB, 9/1/04; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Dallas: Little Beaver Cr., Woodward, TMM, 7/24/07; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Long Cr., Van Wert, CTH, 9/14/95; Delaware: South Fk. Maquoketa R., Dundee, CTH, 7/12/05; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dickinson: Little Sioux R., Milford, CTH, 9/4/ 96; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/ 07; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Little Cedar R., Colwell, CTH, 8/20/02; Franklin: Maynes Cr., Hampton, CTH, 7/18/95; Greene: West Buttrick Cr., Grand Junction, TMM, 9/4/08; Guthrie: Deer Cr., Stuart, CTH, 7/30/98; Hamilton: White Fox Cr., Webster City, CTH, 11/10/97; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/1/07; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Ida: Battle Cr., Holstein, KLS, 6/29/04; Jackson: Prairie Cr., Maquoketa, CTH, 10/1/08; Jasper: North Skunk R., Newton, CTH, 8/8/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Bear Cr., Wyoming, MWB, 7/13/04; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Linn: West Otter Cr., Toddville, CTH, 10/11/01; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Wolf Cr., Chariton, CTH, 8/10/05 Lyon: Mud Cr., Lester, JML, 7/21/04; Madison: North Branch North R., Winterset, TMM, 8/21/07; Mahaska: North Skunk R., Rose Hill, CTH, 8/ 14/02; Marion: English Cr., Knoxville, MWB, 7/ 18/05; Marshall: South Minerva Cr., St. Anthony, TMM, 8/21/08; Mitchell: Rock Cr., Rock Cr., CTH, 8/15/96; Montgomery: Unnamed trib. to Tarkio R., Stanton, TMM, 8/6/13; Muscatine: Pike Run, Nichols, CTH, 7/22/05; O’ Brien: Little Waterman Cr., Hartley, CTH, 7/9/96; Page: West Tarkio Cr., Essex, KLS, 7/7/04; Pocahontas: Big Cedar Cr., Fonda, KBK, 9/19/01; Polk: Four Mile Cr., Ankeny, TMM, 8/9/06; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Sac: Boyer R., Schaller, JML, 8/4/04; Scott: Lost Cr., Princeton, CTH, 8/30/04; Shelby: East Branch West Nishnabotna R., Avoca, TMM, 8/16/07; Sioux: Deep Cr., Granville, KLS, 9/26/ 06; Story: South Skunk R., Ames, CTH, 9/19/97; Tama: Richland Cr., Haven, CTH, 9/22/99; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05;</p> <p>Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Webster: Lyons Cr., Webster City, TMM, 9/16/08; Winnebago: Winnebago R., Lake Mills, CTH, 7/30/12; Winneshiek: Yellow R., Postville, CTH, 8/1/ 06; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Worth: Deer Cr., Carpenter, CTH, 8/21/01; Wright: Otter Cr., Goldfield, JML, 8/11/01. Distribution: Northeastern USA, with Iowa forming part of the western border of its range.</p> <p>Dubiraphia quadrinotata (Say). Allamakee: Yellow R., Waukon, CTH, 8/8/06; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Quarter Section Run, Waverly, CTH, 8/9/04; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/ 27/13; Butler: Boylan Cr., Aredale, MWB, 11/ 11/02; Calhoun: Unnamed trib. to Purgatory Cr., Lohrville, KLS, 8/14/07; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/13/07; Cedar: Rock Cr., Tipton, CTH, 8/20/08; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Cherokee: Bear Cr., Cleghorn, CTH, 9/4/01; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 10/2/ 95; Clay: Prairie Cr., Spencer, JML, 9/4/01; Clayton: Roberts Cr., Postville, CTH, 8/13/08; Clinton: Brophy Cr., McCausland, CTH, 10/7/04; Davis: Soap Cr., Peach Ave., CTH, 7/17/13; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Lake Park, TMM, 8/11/14; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Emmet: Ditch 151, Wallingford, TMM, 6/17/03; Fayette: Dibble Cr., Clermont, CTH, 8/5/04; Floyd: Cedar R., Charles City, MWB, 9/9/08; Franklin: Maynes Cr., Hampton, CTH, 8/23/01; Grundy: South Beaver Cr., Parkersburg, CTH, 7/30/96; Hamilton: Long Dick Cr., Ellsworth, TMM, 10/6/08; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 7/23/12; Hardin: Iowa R., Eldora, CTH, 8/16/05; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/2/11; Howard: Crane Cr., Riceville, 8/ 16/10; Ida: Ashton Cr., Washta, JML, 7/6/06; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Jasper: North Skunk R., Newton, CTH, 8/8/ 01; Johnson: Iowa R., Iowa City, CTH, 8/8/ 05; Jones: Maquoketa R., Monticello, CTH, 9/6/ 05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Four Mile Cr., Algona, TMM, 7/11/06; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Buffington Cr., Columbus City, CTH, 8/23/10; Lyon: Little Rock R., George, CTH, 9/ 5/96; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mitchell: Deer Cr., Carpenter, CTH, 8/25/08; Monona: Jordan Cr., Soldier, TMM, 7/5/06; Muscatine: Mad Cr., Muscatine, CTH, 6/30/04; O’ Brien: Little Waterman Cr., Hartley, CTH, 7/9/96; PaloAlto: Prairie Cr., Whittemore, CTH, 10/2/01; Plymouth: Beaver Cr., Akron, TMM, 8/12/13; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Polk: Four Mile Cr., Ankeny, CTH, 9/29/99; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/17/08; Sac: North Raccoon R., Sac City, TMM, 8/12/13; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: Walnut Cr., Ames, CTH, 7/15/99; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/04; Van Buren: Chequest Cr., Pittsburgh, CTH, 7/17/97; Webster: Deer Cr., Clare, TMM, 7/28/04; Winnebago: Little Buffalo Cr., Thompson, DPH, 7/27/04; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Worth: Elk Cr., Kensett, CTH, 8/26/08; Wright: Otter Cr., Holmes, KLS, 8/11/00. Distribution: Primarily northeastern USA, extending from New York in the north to South Carolina in the south and west to the Great Plains.</p> <p>Dubiraphia vittata (Melsheimer). Allamakee: Yellow R., Ion, CTH, 9/29/98; Benton: Mud Cr., Vinton, CTH, 8/29/05; Black Hawk: Cedar R., Waterloo, CTH, 9/3/13; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Buena Vista: Brooke Cr., Rembrandt, JML, 10/5/05; Cedar: Clear Cr., Lisbon, CTH, 7/26/01; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Cherokee: Maple Cr., Aurelia, CTH, 8/6/97; Clay: Ocheyedan River, Spencer, TMM, 8/26/03; Delaware: Plum Cr., Earlville, CTH, 8/16/06; Dickinson: Little Sioux R., Lake Park, CTH, 9/3/96; Floyd: Cedar R., Charles City, MWB, 9/5/00; Franklin: Maynes Cr., Hampton, CTH, 8/18/08; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Greene: East Buttrick Cr., Dana, TMM, 7/30/09; Guthrie: Deer Cr., Stuart, CTH, 7/30/98; Hamilton: Boone R., Stratford, CTH, 9/17/98; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 7/ 23/12; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Henry: Heather Branch, Mt. Pleasant, CTH, 10/14/98; Howard: Little Turkey R., Protivin, CTH, 8/19/02; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Johnson: (Wickham 1911); Jones: Maquoketa R., Monticello, CTH, 9/6/05; Kossuth: Plum Cr., Wesley, TMM, 8/15/06; Linn: Crabapple Cr., Springville, CTH, 7/17/01; Lyon: Mud Cr., Alvord, KLS, 8/29/05; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Monona: Little Sioux R., Turin, TMM, 8/7/12; O’ Brien: Waterman Cr., Sutherland, TMM, 8/26/08; Page: East Nishnabotna R., Shenandoah, KLS, 10/17/02; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Pottawattamie: Pony Cr., Council Bluffs, KJS, 9/27/05; Sac: North Raccoon R., Sac City, TMM, 9/17/08; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Floyd R., Alton, TMM, 8/6/03; Story: Long Dick Cr., Roland, CTH, 9/22/97; Webster: Lizard Cr., Clare, CTH, 9/11/96; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Worth: Willow Cr., Hanlontown, DPH, 9/4/01; Wright: East Branch Iowa R., Belmond, KLS, 8/25/05. Distribution: Primarily eastern USA, extending from New Hampshire in the north to Florida in the south and west to the Great Plains.</p> <p>Macronychus glabratus Say. Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/01; Appanoose: Chariton R., Centerville, JML, 9/19/01; Benton: Bear Cr., Shellsburg, CTH, 8/2/01; Black Hawk: Black Hawk Cr., Hudson, CTH, 8/29/96; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: East Branch Wapsipinicon R., Tripoli, CTH, 9/3/03; Buchanan: Lime Cr., Brandon, CTH, 8/26/97; Butler: Johnson Cr., Aplington, MWB, 9/11/02; Calhoun: Cedar Cr., Lohrville, JML, 7/31/02; Carroll: Marrowbone Cr., Lanesboro, TMM, 9/ 13/07; Cass: West Nodaway R., Grant, CTH, 8/ 19/98; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Willow Cr., Clear Lake, CTH, 7/ 28/04; Cherokee: Willow Cr., Quimby, JML, 8/14/01; Chickasaw: Wapsipinicon R., New Hampton, CTH, 10/6/04; Clarke: South Whitebreast Cr., Weldon, TMM, 9/29/10; Clay: Little Sioux R., Spencer, TMM, 9/6/05; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Silver Cr., DeWitt, CTH, 7/21/98; Crawford: Trinkle Cr., Kiron, TMM, 10/5/ 15; Dallas: Mosquito Cr., Panora, CTH, 7/15/ 96; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Weldon R., Woodland, KLS, 9/25/03; Delaware: Plum Cr., Hopkinton, CTH, 9/2/03; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Lake Park, TMM, 7/23/03; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Emmett: Ditch 151, Wallingford, TMM, 6/17/03; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Cedar R., Floyd, CTH, 9/4/ 02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/ 03; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Greene: Buttrick Cr., Grand Junction, CTH, 8/ 30/95; Grundy: South Beaver Cr., Parkersburg, CTH, 9/3/08; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 11/794; Hancock: East Branch Iowa R., Goodell, TMM, 7/24/12; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Harrison: Stowe Cr., Pisgah, TMM, 8/26/15; Henry: Big Cr., New London, CTH, 8/30/05; Howard: Crane Cr., Riceville, CTH, 7/10/06; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 8/2/12; Iowa: Big Bear Cr., Victor, CTH, 9/20/99; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: Mud Cr., Baxter, CTH, 9/18/98; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 9/20/05; Jones: Maquoketa R., Monticello, 9/6/05; Keokuk: Cedar Cr., Delta, CTH, 8/18/03; Kossuth: Little Buffalo Cr., Titonka, JML, 8/ 21/00; Linn: Buffalo Cr., Central City, CTH, 8/2/96; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Whitebreast Cr., Lacona, TMM, 7/29/14; Lyon: Little Rock R., George, TMM, 10/1/08; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mahaska: Buck Cr., Barnes City, CTH, 8/3/94; Marion: English Cr., Knoxville, MWB, 7/18/05; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mills: Keg Cr., Mineola, CTH, 7/30/97; Mitchell: Rock Cr., Rock Cr., CTH, 8/15/96; Monona: Soldier R., Moorhead, TMM, 10/6/04; Montgomery: East Nishnabotna R., TMM, Red Oak, 10/12/07; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Little Floyd R., Sheldon, JML, 9/12/ 01; Page: West Tarkio Cr., Shenandoah, CTH, 10/12/95; Plymouth: Floyd R., LeMars, TMM, 8/24/04; Pocahontas: Des Moines R., West Bend, TMM, 8/17/05; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Pidgeon Cr., Neola, CTH, 7/29/97; Poweshiek: Bear Cr., Brooklyn, CTH, 9/25/02; Ringgold: Middle Fk. Grand R., Mount Ayr, TMM, 8/24/11; Sac: North Raccoon R., Sac City, CTH, 10/26/99; Scott: Lost Cr., Princeton, CTH, 9/10/98; Shelby: West Nishnabotna R., Kirkman, CTH, 8/15/94; Sioux: Floyd R., Alton, TMM, 8/6/03; Story: South Skunk R., Ames, CTH, 9/15/95; Tama: Richland Cr., Haven, CTH, 9/8/95; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05; Warren: North R., Norwalk, CTH, 10/15/98; Washington: English R., Riverside, CTH, 6/9/00; Wayne: Jackson Cr., Corydon, TMM, 9/14/11; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winnebago: Winnebago R., Lake Mills, CTH, 9/6/00; Winneshiek: Canoe Cr., Decorah, CTH, 9/9/03; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Wright: Boone R., Renwick, TMM, 8/20/03. Distribution: Eastern USA, extending west to Texas in the south, Oklahoma, Nebraska, and Minnesota in the north.</p> <p>Microcylloepus pusillus (LeConte). (Downie and Arnett 1996). Distribution: Eastern USA, extending west to Utah, Colorado, and Wyoming.</p> <p>Optioservus fastiditus (LeConte). Allamakee: Yellow R., Waukon, CTH, 8/8/06; Buchanan: Pine Cr., Quasqeton, CTH, 8/28/97; Cedar: Clear Cr., Lisbon, CTH, 7/26/01; Cerro Gordo: Calmus Cr., Mason City, MWB, 9/24/01; Clayton: Silver Cr., Gunder, CTH, 8/2/00; Clinton: Silver Cr., DeWitt, CTH, 7/14/04; Delaware: Fenchel Cr., Strawberry Point, CTH, 9/17/07; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Dibble Cr., Clermont, CTH, 8/4/98; Floyd: Little Cedar R., Colwell County Park, CTH, 9/9/09; Franklin: Bailey Cr., Sheffield, CTH, 7/24/03; Howard: Unnamed trib. To Bohemian Cr., Protivin, MWB, 6/17/03; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Jones: Unnamed trib. To North Fk. Walnut Cr., Morley, CTH, 8/15/02; Linn: Indian Cr., Cedar Rapids, CTH, 9/25/00; Mitchell: Burr Oak Cr., Osage, CTH, 8/14/95; Winneshiek: Coldwater Cr., Bluffton, CTH, 7/10/95; Worth: Willow Cr., Hanlontown, CTH, 8/17/95. Distribution: Northeastern USA, extending west to North Dakota and south to Iowa, Illinois, and Indiana. Habitat preference is gravel and rocky substrate in medium to fast flowing streams.</p> <p>Stenelmis bicarinata LeConte. Buchanan: (Wickham 1911); Johnson: (Wickham 1911). Distribution: Eastern United States, extending west to New Mexico in the south and Minnesota in the north. Iowa, along with Minnesota, forms the northwestern margin of its range.</p> <p>Stenelmis cheryl Brown. Allamakee: Upper Iowa R., Hanover, MWB, 7/21/07; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Bremer: Cedar R., Janesville, CTH, 9/9/14; Butler: Shell Rock R., Shell Rock, CTH, 9/8/11; Cedar: Cedar R., West Branch, CTH, 9/12/06; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Clayton: Turkey R., Elkader, JOK, 9/13/88; Davis: Soap Cr., Eldon, CTH, 7/23/02; Fayette: Turkey R., Eldorado, CTH, 10/1/02; Floyd: Cedar R., Floyd, CTH, 9/4/02; Franklin: Beaver Cr., Ackley, CTH, 8/22/13; Hamilton: Boone R., Stratford, TMM, 8/16/12; Hardin: Iowa R., Eldora, CTH, 8/16/05; Howard: Upper Iowa R., Florenceville, CTH, 10/1/02; Jackson: Maquoketa R., Maquoketa, CTH, 9/13/12; Johnson: Iowa R., Iowa City, CTH, 9/22/05; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Cedar R., Bertram, CTH, 10/ 10/05; Louisa: Iowa R., Wapello, CTH, 10/8/ 13; Mahaska: South Skunk R., Oskaloosa, CTH, 10/7/13; Mitchell: Cedar R., Osage, CTH, 7/6/06; Monona: Little Sioux R., Rodney, TMM, 7/18/06; Muscatine: Cedar</p> <p>R., Conesville, CTH, 8/29/12; Poweshiek: North Skunk R., Millgrove, CTH, 8/6/01; Scott: Wapsipinicon R., DeWitt, CTH, 9/16/13; Van Buren: Des Moines R., Farmington, CTH, 9/6/ 06; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Winneshiek: Upper Iowa R., Decorah, CTH, 7/25/06. Distribution: Northeastern USA, extending southwest to New Mexico in the south and Minnesota in the north, with Iowa forming part of the western limit of its range.</p> <p>Stenelmis crenata (Say). Allamakee: Yellow R., Ion CTH, 9/28/98; Appanoose: Chariton R., Centerville, JML, 9/19/01; Benton: Bear Cr., Shellsburg, CTH, 8/8/95; Black Hawk: Black Hawk Cr., Hudson, CTH, 7/31/96; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Horton Cr., Horton, CTH, 7/24/01; Buchanan: Lime Cr., Brandon, CTH, 6/12/95; Butler: Coldwater Cr., Greene, CTH, 8/29/96; Calhoun: Cedar Cr., Lohrville, JML, 7/31/02; Carroll: Brushy Cr., Dedham, TMM, 9/16/09; Cass: West Nodaway R., Grant, CTH, 8/19/98; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Calmus Cr., Mason City, MWB, 9/24/01; Cherokee: Maple R., Aurelia, CTH, 8/6/97; Chickasaw: Wapsipinicon R. New Hampton, CTH, 10/6/04; Clay: Willow Cr., Royal, CTH, 8/26/02; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Silver Cr., DeWitt, CTH, 7/21/98; Crawford: Otter Cr., Deloit, KLS, 8/11/04; Dallas: Mosquito Cr., Panora, CTH, 7/15/96; Davis: Soap Cr., Peach Ave., CTH, 7/17/13; Decatur: Little R., Leon, CTH, 9/23/02; Delaware: Plum Cr., Hopkinton, CTH, 9/2/03; Dickinson: Little Sioux R., Lake Park, CTH, 9/3/ 96; Dubuque: Lytle Cr., Zwingle, CTH, 8/16/99; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Little Cedar R., Colwell, CTH, 10/3/ 95; Franklin: Maynes Cr., Hampton, CTH, 7/ 18/95; Greene: Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hancock: East Branch Iowa R., Goodell, JML, 10/17/00; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Harrison: Soldier R., Pisgah, CTH, 10/27/99; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/2/99; Howard: Little Turkey R., Protivin, CTH, 8/ 19/02; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/10/13; Ida: Maple R., Ida Grove, CTH, 8/5/97; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: Indian Cr., Colfax, TMM, 10/1/12; Johnson: (Wickham 1911); Jones: Bear Cr., Wyoming, CTH, 7/13/04; Kossuth: Black Cat Cr., Algona, CTH, 8/13/97; Lee: Lick Cr., Farmington, CTH, 7/14/03; Linn: Buffalo Cr., Central City, CTH, 8/2/96; Louisa: Long Cr., Columbus Junction, CTH, 9/26/95; Lucas: Wolf Cr., Chariton, TMM, 8/26/11; Lyon: Little Rock R., George, TMM, 10/1/08; Madison: North Branch North R., Winterset, CTH, 6/20/95; Marion: Whitebreast Cr., Knoxville, TMM, 8/ 5/13; Marshall: Minerva Cr., St. Anthony, TMM, 8/8/06; Mills: Keg Cr., Mineola, JML, 8/2/01; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Monona: Monona-Harrison Ditch, Blencoe, TMM, 7/19/06; Montgomery: East Nishnabotna R., Red Oak, TMM, 10/12/07; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Floyd R., Sheldon, CTH, 9/6/95; Plymouth: Floyd R., James, JML, 9/18/08 Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/98; Pottawattamie: Mosquito Cr., Underwood, TMM, 8/17/04; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Sac: North Raccoon R., Sac City, KBK, 9/20/01; Scott: Lost Cr., Princeton, CTH, 8/9/12; Shelby: East Branch West Nishnabotna R., Avoca, JML, 7/18/00; Sioux: Sixmile Cr., Hawarden, JML, 8/2/ 00; Story: South Skunk R., Ames, CTH, 9/15/95; Tama: Little Wolf Cr., Gladbrook, CTH, 7/1/04; Taylor: Honey Cr., Bedford, CTH, 8/18/98; Van Buren: Chequest Cr., Pittsburgh, CTH, 7/17/97; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, CTH, 10/ 15/99; Washington: English R., Riverside, CTH, 6/9/00; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Winneshiek: Middle Bear Cr., Highlandville, CTH, 7/11/95; Woodbury: West Fk. Little Sioux R., Bronson, TMM, 8/9/05; Worth: Willow Cr., Hanlontown, CTH, 9/4/01; Wright: Otter Cr., Goldfield, JML, 8/11/01. Distribution: Eastern USA, exclusive of Florida, extending west to Oklahoma, Colorado, and North Dakota in the north.</p> <p>Stenelmis decorata Sanderson. Allamakee: Upper Iowa R., Dorchester, CTH, 8/24/05; Appanoose: Chariton R., Centerville, CTH, 10/15/99; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Bremer: Cedar R., Janesville, CTH, 9/8/11; Buchanan: Bear Cr., Brandon, CTH, 9/8/09; Butler: Shell Rock R., Shell Rock, CTH, 9/8/ 14; Carroll: Elk Run Cr., Lanesboro, TMM, 8/ 4/11; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 10/14/10; Clayton: Volga R., Elkport, CTH, 9/27/02; Clinton: Silver Cr., DeWitt, CTH, 7/14/04; Davis: Soap Cr., Floris, CTH, 7/24/02; Delaware: Buck Cr., Delhi, MWB, 8/16/01; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dubuque: North Fk. Maquoketa R., New Vienna, CTH, 8/21/01; Fayette: Little Turkey R., West Union, CTH, 9/ 10/03; Floyd: Cedar R., Floyd, CTH, 9/4/02; Grundy: South Beaver Cr., Parkersburg, CTH, 9/10/14; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Howard: Upper Iowa R., Florenceville, CTH, 10/1/02; Jackson: Bear Cr., Baldwin, CTH, 8/28/03; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Maquoketa R., Monticello, CTH, 9/6/05; Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Wapsipinicon R., Central City, CTH, 9/27/06; Louisa: Iowa R., Wapello, CTH, 9/26/02; Mahaska: North Skunk R., Rose Hill, CTH, 8/14/02; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Muscatine: Pike Run, Nichols, CTH, 7/26/04; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Scott: Wapsipinicon R., DeWitt, CTH, 9/16/13; Van Buren: Chequest Cr., Pittsburg, CTH, 7/15/03; Wapello: Des Moines R., Eldon, CTH, 9/30/03; Warren: North R., Norwalk, CTH, 10/15/99; Washington: English R., Riverside, MWB, 9/10/08; Wayne: South Fk. Chariton R., Corydon, TMM, 8/8/16; Winneshiek: Ten Mile Cr., Ridgeway, CTH, 8/ 16/11. Distribution: Eastern USA, extending west to the southern Great Plains, with Iowa forming part of the western border.</p> <p>Stenelmis grossa Sanderson. Adair: Middle Nodaway R., Bridgewater, CTH, 10/21/98; Allamakee: Upper Iowa R., Dorchester, CTH, 9/27/02; Benton: Pratt Cr., Mt. Auburn, CTH, 9/ 5/00; Black Hawk: West Fk. Cedar R., Finchford, CTH, 9/14/01; Boone: Squaw Cr., Zenorsville, KBK, 7/14/00; Bremer: Cedar R., Janesville, CTH, 9/8/11; Buchanan: Lime Cr., Brandon, CTH, 8/9/00; Butler: Beaver Cr., New Hartford, CTH, 8/14/03; Cass: Sevenmile Cr., Grant, KLS, 7/26/06; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Cerro Gordo: Winnebago R., Mason City, CTH, 9/8/14; Cherokee: Mill Cr., Larrabee, CTH, 10/7/98; Chickasaw: Wapsipinicon R., New Hampton, CTH, 8/18/04; Clay: Prairie Cr., Spencer, JML, 9/4/01; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Barber Cr., Grand Mound, CTH, 8/30/04; Crawford: Big Cr., Denison, TMM, 8/10/04; Dallas: Mosquito Cr., Panora, TMM, 9/18/09; Davis: Soap Cr., Eldon, CTH, 7/23/02; Delaware: Buck Cr., Delhi, MWB, 8/16/01; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dickinson: Little Sioux R., Milford, TMM, 7/23/03; Fayette: Dibble Cr., Clermont, CTH, 8/4/98; Floyd: Little Cedar R., Colwell, CTH, 8/20/02; Franklin: Bailey Cr., Sheffield, CTH, 7/24/03; Greene: West Buttrick Cr., Grand Junction, CTH, 8/30/95; Grundy: South Beaver Cr., Parkersburg, CTH, 8/13/01; Hamilton: White Fox Cr., Webster City, CTH, 8/22/95; Hancock: Winnebago R., Forest City, TMM, 9/18/06; Hardin: South Fk. Iowa R., Iowa Falls, CTH, 7/19/95; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/12/00; Howard: Little Turkey R., Protivin, CTH, 8/19/02; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/9/14; Ida: Maple R., Galva, CTH, 8/6/97; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: North Skunk R., Newton, CTH, 8/8/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Bear Cr., Wyoming, CTH, 7/13/04; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Kossuth: Little Buffalo Cr., Titonka, JML, 8/21/00; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Buffalo Cr., Central City, CTH, 9/2/08; Louisa: Buffington Cr., Columbus City, CTH, 9/25//95; Lucas: Wolf Cr., Chariton, TMM, 8/26/11; Lyon: Little Rock R., George, KLS, 8/5/03; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mahaska: North Skunk R., Rose Hill, CTH, 8/14/02; Marshall: South Minerva Cr., Clemons, TMM, 8/8/07; Mitchell: Deer Cr., Carpenter, CTH, 8/15/ 95; Monona: Maple R., Turin, TMM, 8/26/03; Muscatine: Mud Cr., Durant, CTH, 9/16/96; O’ Brien: Little Floyd R., Sheldon, CTH, 9/13/ 99; Page: West Tarkio Cr., Essex, KLS, 7/7/04; Plymouth: Floyd R., James, CTH, 10/26/99; Pocahontas: North Branch Lizard Cr., Havelock, TMM,7/27/04; Pottawattamie: Boyer R., Missouri Valley, TMM, 8/13/03; Poweshiek: Little Bear Cr., Brooklyn, CTH, 8/18/05; Sac: Boyer R., Early, TMM, 8/19/03; Scott: Wapsipinicon R., DeWitt, CTH, 8/30/12; Sioux: Sixmile Cr., Hawarden, JML, 8/2/00; Story: South Skunk R., Ames, CTH, 9/19/97; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/04; Van Buren: Chequest Cr., Pittsburg, CTH, 7/15/03; Warren: Middle R., Indianola, TMM, 9/18/13; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05; Webster: Prairie Cr., Lehigh, CTH, 8/22/96; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Woodbury: Little Sioux R., Smithland, TMM, 8/18/14; Worth: Deer Cr., Carpenter, CTH, 8/21/01. Distribution: Eastern USA, extending west to North Dakota in the north, Nebraska, and Texas in the south.</p> <p>Stenelmis lignicola (Schmude). Des Moines: Burlington (Schmude et al. 1992). Distribution: Southeastern USA, with Iowa forming part of the northwestern border.</p> <p>Stenelmis musgravei Sanderson. Allamakee: Upper Iowa R., Dorchester, MWB, 8/24/05. Distribution: Eastern USA, extending west to Wisconsin in the north and Texas in the south. Iowa forms part of the western border of its distribution.</p> <p>Stenelmis sexlineata Sanderson. Cerro Gordo: Calmus Cr., Mason City, MWB, 9/24/01; Cherokee: Little Sioux R., Larrabee, TMM, 8/ 21/13; Clay: Willow Cr., Royal, CTH, 9/13/99; Clayton: Nutting Cr., Clermont, CTH, 9/21/09; Dallas: South Raccoon R., Redfield, TMM, 9/6/ 13; Des Moines: Brush Cr., Spring Grove, CTH, 9/28/00; Dickinson: Little Sioux R., Milford, CTH, 9/4/96; Fayette: Nutting Cr., Clermont, CTH, 9/21/09; Guthrie: Deer Cr., Stuart, TMM, 7/29/04; Henry: Big Cr., New London, CTH, 8/ 30/05; Lee: Lick Cr., Farmington, CTH, 7/14/03; Louisa: Buffington Cr., Columbus City, CTH, 9/ 25/95; Madison: North Branch North R., Winterset, CTH, 8/12/94; Mitchell: Deer Carpenter, CTH, 8/15/95; O’ Brien: Little Waterman Cr., Hartley, JML, 8/27/02; Sioux: Floyd R., Hospers, TMM, 8/16/06; Van Buren: Honey Cr., Bonaparte, CTH, 6/24/03; Warren: North R., Norwalk, CTH, 10/15/98; Washington: Crooked Cr., Crawfordsville, CTH, 7/14/05. Distribution: Central USA, west to South Dakota and Kansas, east to Indiana and Kentucky.</p> </div>	https://treatment.plazi.org/id/03F487A3FFDDFFC7FEB2FEAE9388FC6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFD8FFC5FC90FBC894FCFB4C.text	03F487A3FFD8FFC5FC90FBC894FCFB4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gyrinidae	<div><p>GYRINIDAE</p> <p>Dineutus assimilis (Kirby). Allamakee: Yellow R., Postville, CTH, 8/14/00; Benton: Cedar R., Vinton, MWB, 9/14/07; Black Hawk: West Fk. Cedar R., Finchford, CTH, 9/15/08; Bremer: Cedar R., Janesville, CTH, 8/6/12; Buchanan: Lime Cr. Brandon, CTH, 8/31/94, 9/4/07; Butler: West Fk. Cedar R., Lake Considine County Park, CTH, 10/2/12; Cass: Indian Cr., Lewis, TMM, 9/22/11; Cedar: Cedar R., West Branch, CTH, 9/12/06; Cerro Gordo: Winnebago R., Mason City, CTH, 8/7/12; Clay: Little Sioux R., Spencer, TMM, 8/14/12; Clayton: Miners Cr., Guttenberg, CTH, 10/8/07; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Crawford: Otter Cr., Deloit, TMM, 9/30/13; Dallas: Raccoon R., Des Moines, TMM, 9/1/11; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Long Cr., Decatur, CTH, 9/14/95; Des Moines: Skunk R., Burlington, CTH, 9/7/06; Dickinson: Little Sioux R., Lake Park, TMM, 9/29/11; Floyd: Cedar R., Charles City, MWB, 9/9/08; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Grundy: South Beaver Cr., Parkersburg, CTH, 9/10/14; Guthrie: South Raccoon R., Stuart, TMM, 7/25/12; Hardin: Hardin City Woodlands (UV light), ELF, 7/16/05; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/9/14; Jackson: Big Mill Cr., Bellevue, CTH, 8/28/07; Johnson: Old Mans Cr., Williamstown, CTH, 10/11/10; Kossuth: Buffalo Cr., Titonka, JML, 8/23/00; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/ 03; Louisa: Iowa R., Columbus Junction, CTH, 9/10/08; Mahaska: South Skunk R., Oskaloosa, CTH, 10/8/12; Marion: English Cr., Knoxville, TMM, 9/11/13; Marshall: Iowa R., Marshalltown, TMM, 9/11/14; Mills: West Nishnabotna R., Malvern, TMM, 10/2/13; Muscatine: Pike Run, West Liberty, CTH, 7/19/06; Polk: Des Moines R., Des Moines, TMM, 9/24/13; Pottawattamie: Jordan Cr., Macedonia, TMM, 9/17/13; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/17/08; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Scott: Wapsipinicon R., DeWitt, CTH, 8/30/12; Story: South Skunk R., Cambridge, TMM, 10/7/14; Van Buren: Des Moines R., Farmington, CTH, 10/10/06; Warren: Middle R., Indianola, TMM, 7/23/12; Wayne: Unnamed trib. to Steel Cr., Humeston, TMM, 6/17/ 14; Webster: Lizard Cr., Clare, TMM, 8/22/11; Winnebago: Pilot Knob Wetland, Forest City, MTF, 7/18/07; Winneshiek: UV light, KJL, 10/ 7/93; Worth: Elk Cr., Kensett, CTH, 8/26/08; Wright: East Branch Iowa R., Belmond, TMM, 7/23/12. Distribution: Eastern USA, extending west to the Great Plains.</p> <p>Dineutus discolor Aubé. Allamakee: Upper Iowa R., Dorchester, MWB, 10/6/05; Black Hawk: Wapsipinicon R., Dunkerton, CTH, 8/2/06; Buchanan: Wapsipinicon R., Independence, CTH,9/3/13; Johnson: (Wickham 1911); Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Texas in the south.</p> <p>Dineutus hornii Roberts. Cherokee: Cherokee, FJM, 6/10/56. Distribution: Eastern USA, extending west to Minnesota in the north, Kansas and Arizona in the south.</p> <p>Dineutus nigrior Roberts. Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05; Bremer: Quarter Section Run, Waverly, CTH, 8/9/04; Floyd: Cedar R., Carrville, CTH, 9/4/13. Distribution: Eastern USA, extending west to North Dakota in the north, Iowa, and Louisiana in the south. New state record.</p> <p>Gyrinus aeneolus LeConte. Appanoose: Packard Cr., Exline, CTH, 9/15/15; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Black Hawk: Indian Cr., Gilbertville, CTH, 6/28/06; Boone:</p> <p>Montgomery Cr., Gilbert, TMM, 8/9/13; Buchanan: Pine Cr., Quaqeton, CTH, 9/1/15; Clarke: South Whitebreast Cr., Weldon, CTH, 9/28/95; Clayton: Unnamed trib. to Turkey R., Elgin, CTH, 9/24/14; Dubuque: Lytle Cr., Zwingle, CTH, 7/31/07; Fayette: Volga R., Randalia, CTH, 8/31/10; Jackson: Bear Cr., Baldwin, CTH, 8/ 31/11; Keokuk: Twin Grove Cr., Ollie, CTH, 6/2/ 15; Scott: Lost Cr., Princeton, CTH, 8/30/04; Winneshiek: Canoe Cr., Canoe Creek SWMA, CTH, 8/18/15. Distribution: Eastern USA, extending west to the Great Plains. Its habitat preference is both lentic and lotic environments. New state record.</p> <p>Gyrinus affinis Aubé. Cedar: (Wickham 1911); Johnson: (Wickham 1911). Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border in the Midwest.</p> <p>Gyrinus analis Say. Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05; Black Hawk: West Fk. Cedar R., Finchford, MWB, 9/15/08; Butler: Shell Rock R., Shell Rock, CTH, 8/6/12; Clayton: Turkey R., Garber, CTH, 10/7/14; Clinton: Brophy Cr., McCausland, CTH, 10/7/ 04; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/12/08; Johnson: Pechman Cr., Hills, CTH, 4/ 17/17; Kossuth: Winnebago R., Lake Mills, TMM, 7/30/12; Lee: Des Moines R., Keokuk, CTH, 10/1/03; Linn: Cedar R., Palo, CTH, 10/7/ 11; Louisa: Iowa R., Columbus Junction, CTH, 9/10/08; Mitchell: Burr Oak Cr., Osage, CTH, 8/3/11; Scott: Wapsipinicon R., DeWitt, CTH, 8/ 30/12; Story: South Skunk R., Cambridge, TMM, 10/7/14; Van Buren: Des Moines R., Keosauqua, CTH, 10/9/12; Wapello: Des Moines R., Ottumwa, CTH, 10/8/12; Winnebago: Winnebago R., Lake Mills, JML, 7/22/03. Distribution: eastern USA, extending to the Great Plains and south to the Gulf Coast. Its habitat preference is slow moving portions of rivers and backwaters areas.</p> <p>Gyrinus bifarius Fall. Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05; Mahaska: Buck Cr., Barnes City, KLS, 9/18/07; Winnebago: Beaver Cr., Lake Mills, CTH, 7/22/03. Distribution: Widely distributed and transcontinental in the USA. Its habitat preference is lentic and lotic environments. New state record.</p> <p>Gyrinus dichrous LeConte. Johnson: (Wickham 1911). Distribution: Northeastern USA, with Iowa forming the southwestern border.</p> <p>Gyrinus gibber LeConte. (Oygur and Wolfe 1991). Distribution: Eastern USA, extending west to the Midwest. Iowa forms part of the western border.</p> <p>Gyrinus latilimbus Fall. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/10/11. Distribution: Northeastern USA, extending west to North Dakota in the north. Iowa forms part of the southwestern border. New state record.</p> <p>Gyrinus lecontei Fall. Allamakee: Unnamed trib. to Village Cr., Lansing, CTH, 8/26/10; Clayton: Bluebell Cr., Millville, CTH, 5/28/15; Fayette: Unnamed trib. to Volga Lake outlet, Fayette, CTH, 8/4/15; Hamilton: Boone R., Tunnel Mill access, TMM, 10/5/10; Mitchell: Poor Cr., St. Ansgar, CTH, 3/30/15; Winneshiek: Coldwater Cr., Bluffton, CTH, 9/22/04. Distribution: Northeastern USA with Iowa forming part of the southwestern border.</p> <p>Gyrinus maculiventris LeConte. Black Hawk: Beaver Cr., Cedar Falls, CTH, 10/9/14; Butler: West Fk. Cedar R., Dumont, CTH, 10/11/06; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 10/14/10; Clayton: Volga R., Elkport, CTH, 10/7/14; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 8/23/11; Guthrie: Middle Raccoon R., Coon Rapids, TMM, 10/6/11; Hamilton: Boone R., Albright access, TMM, 10/8/10; Jackson: Farmers Cr., Fulton, CTH, 10/20/11; Johnson: (Wickham 1911); Linn: (Wickham 1911); Marshall: Iowa R., Marshalltown, TMM, 9/11/14; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/06; Muscatine: (Wickham 1911); Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Winneshiek: UV light, ED, 10/6/93. Distribution: Northern transcontinental in the USA with Iowa forming part of the southern border in the Midwest.</p> <p>Gyrinus marginellus Fall. Allamakee: Upper Iowa R., Hanover, CTH, 5/25/07; Black Hawk: Dry Run Cr., Cedar Falls, CTH, 9/14/10; Buchanan: Bear Cr., Brandon, CTH, 8/3/10; Cerro Gordo: Bailey Cr., Thornton, CTH, 7/23/03; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Fayette: Brush Cr., Wadena, CTH, 9/27/05; Floyd: Cedar R., Floyd, CTH, 9/23/08; Hamilton: East Br. Iowa R., Belmond, TMM, 7/23/12; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Mitchell: Cedar R., Floyd, MWB, 9/23/08; Tama: Little Wolf Cr., Gladbrook, CTH, 8/12/04; Winnebago: Little Buffalo Cr., Thompson, DPH, 7/27/04; Winneshiek: Yellow R., Castalia, CTH, 9/15/ 04; Wright: East Branch Iowa R., Belmond, TMM, 7/23/12. Distribution: Eastern USA, extending to the eastern Great Plains, with Iowa forming part of the western border. Its habitat preference is clear rivers with gravel or clay bottoms, particularly along the stream margin. New state record.</p> <p>Gyrinus minutus (Fabricius). Black Hawk: (Wickham 1911); Johnson: (Wickham 1911); (Ferkinhoff and Gundersen 1983). Distribution: Northern transcontinental in the USA, extending south to Illinois, Iowa, and Kansas in the Midwest.</p> <p>Gyrinus parcus Say. (Oygur and Wolfe 1991). Distribution: Eastern USA, extending west to the Great Plains.</p> <p>Gyrinus pectoralis LeConte. A single specimen is on record at the Illinois Natural History Survey (INHS), but has no collection/location data associated with it, other than it was collected in Iowa (Ed DeWalt, INHS, personal communication). Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border in the Midwest.</p> <p>Gyrinus sayi Aubé. Howard: (Wickham 1911). Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Gyrinus ventralis Kirby. Allamakee: Paint Cr., Yellow River State Forest, CTH, 8/13/10; Black Hawk: Dry Run Cr., Cedar Falls, CTH, 8/29/11; Buchanan: Buffalo Cr., Winthrop, CTH, 9/28/ 10; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 9/20/10; Clayton: Peck Cr., Osterdock, CTH, 9/6/11; Clinton: Silver Cr., DeWitt, CTH, 7/17/12; Delaware: Bear Cr., Dyersville, CTH, 7/27/11; Fayette: Dibble Cr., Clermont, CTH, 9/24/14; Howard: North Branch Turkey R., Lime Springs, CTH, 8/5/15; Jackson: Farmers Cr., Fulton, CTH, 8/14/12; Johnson: (Wickham 1911); Linn: West Otter Cr., Toddville, CTH, 10/2/13; Mitchell: Burr Oak Cr., Osage, CTH, 7/30/12. Distribution: Northeastern USA, extending west to Missouri in the south, Iowa, and Minnesota in the north.</p> </div>	https://treatment.plazi.org/id/03F487A3FFD8FFC5FC90FBC894FCFB4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFDAFFC8FC9BFAEC9113FA9A.text	03F487A3FFDAFFC8FC9BFAEC9113FA9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haliplidae	<div><p>HALIPLIDAE</p> <p>Haliplus borealis LeConte. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 9/11/07; Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Benton: Cedar R., Vinton, MWB, 9/14/ 07; Black Hawk: West Fk. Cedar R., Finchford, CTH, 9/9/14; Bremer: Horton Cr., Horton, CTH, 7/24/01; Buchanan: Pine Cr., Quasqeton, CTH, 9/1/15; Cass: West Nodaway R., Grant, TMM, 8/19/04; Cerro Gordo: Winnebago R., Mason City, CTH, 9/4/13; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 5/15/12; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 8/26/14; Clarke: South Whitebreast Cr., Weldon, TMM, 8/17/15; Clay: Willow Cr., Rossie, JML, 8/26/02; Clayton: Buck Cr., Garnavillo, CTH, 8/11/11; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Decatur: Redmans Cr., Dekalb WMA, TMM, 9/15/15; Des Moines: Flint Cr., Danville, CTH, 7/15/ 03; Dickinson: Milford Cr., Milford, JML, 9/ 5/01; Dubuque: North Fk. Maquoketa R., Dyersville, CTH, 7/25/05; Fayette: Floyd: Franklin: Guthrie: South Raccoon R., Stuart, KLS, 9/13/00; Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/14/01; Harrison: Boyer R., Missouri Valley, TMM, 9/24/14; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Ida: Maple R. R., Ida Grove, CTH, 8/5/97; Jasper: Walnut Cr., Prairie City, CTH, 5/6/97; Johnson: Iowa R., Iowa City, CTH, 8/8/05; Jones: Silver Cr., Monticello, CTH, 7/31/01; Kossuth: Four Mile Cr., Algona, TMM, 7/11/06; Louisa: Iowa R., Columbus Junction, CTH, 9/10/08; Lucas: Wolf R., Chariton, TMM, 8/3/15; Madison: Middle R., Winterset, TMM, 10/13/15; Marion: English Cr., Knoxville, TMM, 8/14/14; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Montgomery: East Nishnabotna R., Red Oak, TMM, 9/6/07; Muscatine: Honey Cr., Conesville, CTH, 7/18/ 06; Plymouth: Beaver Cr., Akron, 8/12/13, KLS; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/ 06; Poweshiek: Walnut Cr., Hartwick, CTH, 7/ 27/09; Story: Drainage Ditch #81, Nevada, TMM, 8/29/07; Taylor: Honey Cr., Bedford, TMM, 7/22/14; Union: Unnamed trib. to Thompson R., Lorimor, TMM, 9/17/15; Van Buren: Chequest Cr., Pittsburg, CTH, 7/15/03; Wapello: Bear Cr., Ottumwa, CTH, 6/28/05; Warren: South R., Ackworth, TMM, 9/23/15; Washington: English R., Riverside, MWB, 9/ 10/08; Winnebago: Pike Run, Leland, MWB, 9/ 19/06; Winneshiek: South Pine Cr., Sattre, CTH, 8/ 14/07; Woodbury: Little Sioux R., Smithland, TMM, 9/6/11; Worth: Elk Cr., Kensett, CTH, 8/26/ 08; Wright: East Branch Iowa R., Belmond, KLS, 8/25/05. Distribution: The Great Lakes area, east to Maine, west to Montana, and south to Missouri.</p> <p>Haliplus connexus Matheson. Allamakee: French Cr., Churchtown, CTH, 7/1/96; Wayne: Unnamed trib. Steel Cr., Humeston, TMM, 5/23/12. Distribution: Northeastern USA, extending west to North Dakota, Minnesota, and Iowa.</p> <p>Haliplus fasciatus Aubé. Clinton: Barber Cr., Grand Mound, CTH, 8/9/12; Jackson: Farmers Cr., Fulton, MWB, 8/14/12. Distribution: Eastern USA, extending west to Wisconsin in the north, Iowa, Kansas, and Texas in the south. Iowa forms part of the western border.</p> <p>Haliplus immaculicollis Harris. Allamakee: Clear Cr., Lansing, MWB, 8/16/07; Butler:</p> <p>Palmer Cr., Greene, CTH, 5/29/07; Clayton: Peck Cr., Osterdock, CTH, 7/26/11; Delaware: Little Turkey R., Colesburg, MWB, 9/8/06; Dickinson: Little Sioux R., Lake Park, TMM, 7/23/03; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 9/5/12; Fayette: Mink Cr., Wadena, MWB, 7/13/06; Johnson: (Wickham 1911); Kossuth: Black Cat Cr., Algona, MWB, 8/25/03; Mitchell: Rock Cr., Rock Cr., CTH, 8/ 15/96; O’ Brien: Mill Cr., Paullina, JML, 8/21/ 02; Plymouth: Beaver Cr., Akron, 8/12/13, KLS; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: (Wickham 1911); Winnebago: Beaver Cr., Lake Mills, CTH, 7/22/03; Winneshiek: South Pine Cr., Sattre, CTH, 8/14/07; Worth: Willow Cr., Hanlontown, TMM, 9/28/10; Wright: Buttermilk Cr., Goldfield, KJS, 9/6/ 06. Distribution: Primarily northern transcontinental in the USA, with limited collections south of Iowa.</p> <p>Haliplus longulus LeConte. Kossuth: Kettle Wetland-Union Slough NWR, Bancroft, CTH, 5/ 6/97. Distribution: Northern transcontinental in the USA, with Iowa forming part of the southern border in the Midwest.</p> <p>Haliplus triopsis Say. Cerro Gordo: Winnebago R., Mason City, CTH, 9/8/14; Clinton: Silver Cr., DeWitt, CTH, 7/17/12; Davis: Fox R., Bloomfield, CTH, 7/15/13; Dickinson: Milford Cr., Milford, JML, 9/5/01; Hancock: Winnebago R., Forest City, MWB, 9/18/06; Harrison: Boyer R., Missouri Valley, TMM, 9/24/14; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/12/08; Jackson: Tetes Des Morts, St. Donatus, CTH, 8/27/07; Johnson: (Wickham 1911); Linn: Cedar R., Cedar Rapids, KJS, 10/9/14; Madison: North Branch North R., Winterset, CTH, 6/14/ 96; Marion: Cedar Cr., Marysville, JML, 8/7/06; Story: (Wickham 1911); Winneshiek: UV light, KR, 9/29/01; Worth: Elk Cr., Kensett, TMM, 9/ 29/12. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, Colorado, and New Mexico in the south.</p> <p>Peltodytes duodecimpunctatus (Say). Clinton: Elk R., Teeds Grove, CTH, 8/17/99; Davis: Fox R., Bloomfield, CTH, 7/15/13; Des Moines: Skunk R., Burlington, MWB, 9/7/06; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Harrison: Potato Cr., Persia, KLS, 7/12/05; Jackson: Pleasant Cr., Springbrook, CTH, 9/12/02; Johnson: (Wickham 1911); Lee: Lick Cr., Shimek State Forest, 7/28/ 14; Louisa: Honey Cr., Conesville, CTH, 8/ 21/12; Mills: Silver Cr., Malvern, JML, 10/11/ 06; Montgomery: Little Indian Cr., Red Oak, TMM, 9/26/13; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Scott: (Wickham 1911); Story:</p> <p>(Wickham 1911); Van Buren: Chequest Cr., Pittsburg, CTH, 9/14/15. Distribution: Northeastern USA, extending west to Iowa and Missouri.</p> <p>Peltodytes edentulus (LeConte). Adair: North R., Stuart, TMM 9/21/05; Allamakee: Yellow R., Ion, CTH, 8/29/02; Appanoose: Chariton R., Centerville, CTH, 10/15/99; Benton: Bear Cr., Shellsburg, CTH, 10/1/01; Black Hawk: Black Hawk Cr., Hudson, CTH, 8/8/02; Boone: Montgomery Cr., Gilbert, TMM, 8/9/13; Bremer: East Branch Wapsipinicon R., Tripoli, CTH, 8/27/97; Buchanan: Pine Cr., Quasqeton, CTH, 8/28/97; Buena Vista: Brooke Cr., Rembrandt, JML, 10/5/05; Butler: Johnson Cr., Aplington, MWB, 9/11/02; Carroll: Marrowbone Cr., Lanesboro, JML, 10/9/02; Cass: West Nodaway R., Grant, TMM, 8/19/04; Cedar: Clear Cr., Lisbon, CTH, 7/26/01; Cerro Gordo: Bailey Cr., Thornton, TMM, 10/7/03; Cherokee: Mill Cr., Larrabee, CTH, 10/7/98; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 10/2/95; Clarke: South Whitebreast Cr., Weldon, TMM, 8/17/15; Clay: Willow Cr., Royal, CTH, 9/13/99; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Elk R., Teeds Grove, CTH, 8/17/99; Crawford: Big Cr., Denison, JML, 8/12/98; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Long Cr., Van Wert, JML, 10/9/01; Delaware: Maquoketa R., Manchester, CTH, 9/4/02; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dickinson: Little Sioux R., Lake Park, CTH, 9/3/96; Dubuque: North Fk. Maquoketa R., New Vienna, CTH, 8/21/01; Emmett: Ditch 151, Wallingford, JML, 6/17/03; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Cedar R., Charles City, CTH, 9/26/02; Franklin: Beaver Cr., Ackley, CTH, 8/15/05; Fremont: Plum Cr., Thurman, KLS, 8/3/05; Greene: Buttrick Cr., Grand Junction, TMM, 7/30/14; Grundy: Middle Fk. South Beaver Cr., Wellsburg, CTH, 8/14/01; Guthrie: South Raccoon R., Guthrie Center, KLS, 6/12/03; Hamilton: Long Dick Cr., Ellsworth, TMM, 10/6/08; Hardin: Middle Fk. South Beaver Cr., Ackley, CTH, 8/14/01; Harrison: Soldier R., Pisgah, CTH, 10/27/99; Henry: Big Cr., New London, CTH, 8/30/05; Howard: North Branch Turkey R., Lime Springs, CTH, 8/5/15; Humboldt: West Fk. Des Moines R., Humboldt, TMM, 9/9/14; Ida: Maple R., Ida Grove, CTH, 8/5/97; Iowa: Old Mans Cr., Ladora, CTH, 10/2/13; Jackson: Bear Cr., Baldwin, CTH, 9/3/97; Jasper: Walnut Cr., Prairie City, CTH, 8/20/99; Johnson: Muddy Cr., North Liberty, CTH, 10/12/09; Jones: Prairie Cr., Scotch Grove, CTH, 7/23/03; Kossuth: Little Buffalo Cr., Titonka, JML, 8/ 22/00; Lee: Skunk R., Augusta, CTH, 10/8/12; Linn: Buffalo Cr., Central City, CTH, 8/29/01; Louisa: Honey Cr., Conesville, CTH, 9/2/98; Lucas: Wolf Cr., Chariton, CTH, 8/10/05; Lyon: Little Rock R., George, CTH, 9/5/96; Madison: Howerdon Cr., Winterset, CTH, 6/19/95; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Marion: English Cr., Knoxville, MWB, 7/18/05; Marshall: Burnett Cr., Marshalltown, CTH, 8/22/00; Mills: Keg Cr., Mineola, TMM, 8/15/12; Mitchell: Burr Oak Cr., Osage, CTH, 8/14/95; Monona: Soldier R., Moorhead, TMM, 10/6/04; Montgomery: Walnut Cr., Red Oak, CTH, 10/14/96; Muscatine: Mud Cr., Wilton, CTH, 9/20/96; O’ Brien: Waterman Cr., Sutherland, JML, 8/15/01; Page: East Nishnabotna R., Shenandoah, TMM, 9/18/08; Palo Alto: Prairie Cr., Whittemore, CTH, 10/2/01; Plymouth: West Branch Floyd R., Struble, TMM, 8/1/06; Pocahontas: Big Cedar Cr., Fonda, JML, 9/27/06; Polk: Jordan Cr., West Des Moines, TMM, 7/29/05; Pottawattamie: Mosquito Cr., Manawa, JML, 10/5/00; Poweshiek: Walnut Cr., Belle Plaine, CTH, 7/29/03; Ringgold: Lotts Cr., Lamoni, TMM, 7/21/ 14; Sac: Boyer R., Schaller, JML, 8/4/04; Scott: Wapsipinicon R., DeWitt, CTH, 10/2/ 14; Shelby: East Branch West Nishnabotna R., Avoca, CTH, 9/21/94; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: South Skunk R., Ames, CTH, 9/15/95; Tama: Richland Cr., Haven, CTH, 9/8/95; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Union: Unnamed trib. to Thompson R., Lorimor, TMM, 9/ 17/15; Van Buren: Rock Cr., Keosauqua, CTH, 7/29/14; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Warren: North R., Norwalk, CTH, 10/15/99; Washington: English R., Riverside, MWB, 9/10/08; Wayne: Dick Cr., Corydon, TMM, 8/12/09; Webster: Deer Cr., Clare, TMM, 7/28/04; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Winneshiek: Coldwater Cr., Bluffton, CTH, 7/10/95; Woodbury: Little Sioux R., Smithland, TMM, 9/6/11; Worth: Willow Cr., Hanlontown, CTH, 8/ 17/95; Wright: Otter Cr., Goldfield, MWB, 10/ 13/04. Distribution: Northern transcontinental in the USA, extending south to Kansas and Missouri in the Midwest.</p> <p>Peltodytes lengi Roberts. (Downie and Arnett 1996). Distribution: Eastern USA, extending west to the Midwest, with Iowa forming part of the western border.</p> <p>Peltodytes litoralis Matheson. Des Moines: Skunk R., Burlington, MWB, 9/7/06. Distribution:</p> <p>Southcentral USA with Iowa forming part of the northern border with records extending south to Texas, east to Indiana, and west to Colorado. New state record.</p> <p>Peltodytes sexmaculatus Roberts. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/27/14; Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Cass: West Nodaway R., Grant, TMM, 8/20/12; Davis: Soap Cr., Drakesville, CTH, 8/ 16/04; Decatur: Long Cr., Van Wert, TMM, 7/ 15/14; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dubuque: Little Maquoketa R., Dubuque, CTH, 9/25/12; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Johnson: Iowa R., Iowa City, CTH, 9/14/12; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03; Louisa: Honey Cr., Conesville, CTH, 7/26/04; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Marion: English Cr., Knoxville, MWB, 7/18/ 05; Mills: Douglas Cr., Emerson, TMM, 9/25/13; Muscatine: Mad Cr., Muscatine, CTH, 6/30/04; Poweshiek: Walnut Cr., Belle Plaine, CTH, 9/17/ 08; Ringgold: Lotts Cr., Lamoni, TMM, 7/18/03; Sioux: Deep Cr., Granville, KLS, 9/26/06; Taylor: Honey Cr., Bedford, TMM, 7/22/14; Van Buren: Coppers Cr., Keosauqua, CTH, 10/6/09; Warren: South R., Ackworth, TMM, 9/ 23/15; Wayne: Dick Cr., Corydon, TMM, 8/12/ 09; Winneshiek: Yellow R., Postville, CTH, 7/ 25/07. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, Kansas, and Texas in the south. New state record.</p> <p>Peltodytes tortulosus Roberts. Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Henry: Big Cedar Cr., Oakland Mills, MWB, 9/12/08; Louisa: Honey Cr., Conesville, CTH, 7/26/04; Polk: Walnut Cr., Windsor Heights, CTH, 8/26/ 98; Poweshiek: Walnut Cr., Holiday Lake, CTH, 7/27/09; Wayne: South Fk. Chariton R., Corydon, TMM, 8/24/12. Distribution: Northeastern USA, with Iowa forming part of the southwestern border. New state record.</p> </div>	https://treatment.plazi.org/id/03F487A3FFDAFFC8FC9BFAEC9113FA9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFD7FFC8FEE4FAAC9392FCCB.text	03F487A3FFD7FFC8FEE4FAAC9392FCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helophoridae Leach 1815	<div><p>HELOPHORIDAE</p> <p>Helophorus lacustris LeConte. Franklin: Boylan Cr., Aredale, CTH, 8/11/14; Howard: North Branch Turkey R., Lime Springs, CTH, 8/5/15; Mitchell: Deer Cr., Carpenter, CTH, 8/15/95; Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Northern transcontinental in the USA, with Iowa forming part of southern border in the Midwest.</p> <p>Helophorus linearis LeConte. (Downie and Arnett 1996). Distribution: Transcontinental in the USA.</p> <p>Helophorus lineatus Say. Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/29/11; Dubuque: White Pine Hollow Cr., Luxemburg, MWB, 3/20/05; Johnson: Private wetland, Sutliff, JAG, 7/25/13; Story: (Wickham 1911); Winneshiek: Martha Cr., Cresco, CTH, 9/23/14. Distribution: Eastern USA, extending west to North Dakota in the north, Iowa, and Missouri in the south.</p> <p>Helophorus marginicollis Smetana. Appanoose: Sharon Bluffs State Park (UV light), ELF, 7/21/ 07. Distribution: Eastern USA, extending west to Louisiana in the south, Missouri, and Iowa in the north.</p> <p>Helophorus nitiduloides Orchymont. (Downie and Arnett 1996). Distribution: Transcontinental in the USA.</p> <p>Helophorus orientalis Motschulsky. Black Hawk: Indian Cr., Gilbertville, CTH, 6/28/06; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cedar: Baldwin Mason Cr., Mechanicsville, CTH, 6/23/11; Fayette: Unnamed trib. to Mink Cr., Wadena, CTH, 7/17/03. Distribution: Transcontinental in the USA.</p> </div>	https://treatment.plazi.org/id/03F487A3FFD7FFC8FEE4FAAC9392FCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFD7FFC9FCC3FC629157FD87.text	03F487A3FFD7FFC9FCC3FC629157FD87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrochidae Thomson 1859	<div><p>HYDROCHIDAE</p> <p>Hydrochus excavatus LeConte. Johnson: (Wickham 1911); Winnebago: (Wickham 1911). Distribution: Eastern USA, extending west to the Midwest.</p> <p>Hydrochus neosquamifer Smetana. Winnebago: Winnebago R., Lake Mills, TMM, 7/30/12. Distribution: Eastern USA, extending west to North Dakota in the north and Mississippi in the south. New state record.</p> <p>Hydrochus pseudosquamifer Miller. Jackson: South Big Mill Cr., Bellevue, CTH, 9/3/10; Kossuth: Black Cat Cr., Algona, MWB, 8/25/ 03. Distribution: Northern transcontinental in the USA.</p> <p>Hydrochus rufipes Melsheimer. Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Linn: Wapsipinicon R., Troy Mills, CTH, 9/20/04. Distribution: Eastern USA, extending west to Wyoming in the north and Texas in the south.</p> <p>Hydrochus scabratus Mulsant. Buchanan: Pine Cr., Quasqueton, CTH, 9/1/15; Hardin: Tipton Cr., Buckeye, CTH, 7/30/99; Johnson: (Wickham 1911); Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08. Distribution: Transcontinental in the USA.</p> <p>Hydrochus squamifer LeConte. Johnson: (Wickham 1911); Linn: Wickiup Nature Center (UV light), ELF, 7/28/04; Story: (Wickham 1911). Distribution: northern Transcontinental in the USA, extending south to Missouri and Kansas in the Midwest.</p> <p>Hydrochus subcupreus Randall. Black Hawk: Beaver Cr., Cedar Falls, MWB, 8/6/12; Cedar: Sugar Cr., Wilton, CTH, 8/23/01; Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Muscatine: Pike Run, Nichols, CTH, 8/20/12. Distribution: Northeastern USA, extending west to Minnesota and Iowa.</p> </div>	https://treatment.plazi.org/id/03F487A3FFD7FFC9FCC3FC629157FD87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFD6FFF3FED4FD9594D5FA4E.text	03F487A3FFD6FFF3FED4FD9594D5FA4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrophilidae Latreille 1802	<div><p>HYDROPHILIDAE</p> <p>Anacaena lutescens (Stephens). Allamakee: Hecker Cr., Postville, TMM, 7/24/07; Buchanan: Pine Cr., Quasqueton, CTH, 8/9/11; Clayton: Unnamed trib. Silver Cr., Luana, CTH, 8/7/06; Delaware: Bear Cr., Dyersville, CTH, 7/27/ 11; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 8/12/10; Floyd: Cedar R., Charles City, Mark Johnston, 9/9/08; Franklin: Bailey Cr., Sheffield, CTH, 7/20/11; Kossuth: Buffalo Cr., Titonka, KLS, 8/23/00; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/06; Winneshiek: South Pine Cr., Sattre, CTH, 8/4/10. Distribution: Western USA, extending east to the upper Midwest.</p> <p>Berosus aculeatus LeConte. Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/2006. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Texas in the south.</p> <p>Berosus fraternus LeConte. Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Butler: Palmer Cr., Greene, CTH, 5/29/2007; Dubuque: North Fk. Maquoketa R., Dyersville, CTH, 7/25/05; Hancock: East Branch Iowa R., Goodell, JML, 10/17/00; Henry: Skunk R., Oakland Mills, CTH, 9/24/08; Jackson: North Fk. Maquoketa R., Maquoketa, CTH, 9/13/12; Linn: Cedar R., Cedar Rapids, CTH, 8/24/15; Muscatine: Mud Cr., Wilton, CTH, 9/15/03. Distribution: Transcontinental in the USA.</p> <p>Berosus infuscatus LeConte. Clinton: Brophy Cr., McCausland, CTH, 8/14/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 8/29/12; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 8/20/14; Johnson: Iowa R., Iowa City, CTH, 9/18/13; Linn: Cedar R., Cedar Rapids, CTH, 9/19/13; Louisa: Honey Cr., Conesville, CTH, 8/24/04; Mahaska: Muchakinock Cr., Eddyville, CTH, 9/ 2/11; Muscatine: Pike Run, West Liberty, CTH, 7/21/05; Polk: Des Moines R., Des Moines, KLS, 9/24/13; Washington: English R., Riverside, CTH, 9/24/02. Distribution: Transcontinental in the USA.</p> <p>Berosus pantherinus LeConte. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/27/14; Cedar: Cedar R., West Branch, CTH, 9/12/06; Clinton: Brophy Cr., McCausland, CTH, 8/14/12; Johnson: Old Mans Cr., Hills, CTH, 9/18/13; Linn: Cedar R., Cedar Rapids, CTH, 9/19/13; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Muscatine: Cedar R., Conesville, CTH, 8/29/ 12; Polk: Des Moines R., Des Moines, CTH, 8/ 27/12; Wayne: Jordan Cr., Millerton, TMM, 8/ 24/12. Distribution: Eastern USA, extending west to Minnesota in the north, Nebraska, Missouri, and Texas in the south.</p> <p>Berosus peregrinus (Herbst). Appanoose: Chariton R., Rathbun, CTH, 8/17/04; Buchanan: Pine Cr., Quasqueton, CTH, 8/9/11; Butler: Shell Rock R., Shell Rock, CTH, 9/8/14; Cedar: Sugar Cr., Wilton, CTH, 8/23/01; Clayton: Silver Cr., Monona, CTH, 8/7/06; Clinton: Brophy Cr., McCausland, CTH, 10/2/14; Dallas: South Raccoon R., Redfield, TMM, 8/8/12; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Weldon R., Woodland, TMM, 8/8/ 13; Delaware: Bear Cr., Dyersville, CTH, 7/ 27/11; Des Moines: Flint Cr., Danville, CTH, 7/15/03; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08; Floyd: Cedar R., Charles City, CTH, 9/9/08; Franklin: Beaver Cr., Ackley, CTH, 8/22/13; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hamilton: Boone R., Stratford, TMM, 8/28/ 14; Henry: Big Cr., New London, CTH, 8/30/ 05; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Johnson: (Wickham 1911); Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Linn: Indian Cr., Cedar Rapids, CTH, 8/27/13; Louisa: Honey Cr., Conesville, CTH, 8/21/12; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Madison: Middle R., Wintereset, TMM, 7/16/12; Marion: English Cr., Knoxville, CTH, 7/18/05; Mitchell: Deer Cr., Carpenter, CTH, 8/21/13; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Polk: Des Moines R., Des Moines, TMM, 8/27/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Taylor: Honey Cr., Bedford, TMM, 8/18/04; Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13; Wayne: Jordan Cr., Millerton, TMM, 9/12/12; Webster: Des Moines R., Ft. Dodge, TMM, 9/29/14; Winnebago: Buffalo Cr., Forest City, MWB, 9/16/06; Woodbury: West Fk. Little Sioux R.,</p> <p>Bronson, TMM, 8/9/05. Distribution: Transcontinental in the USA.</p> <p>Berosus sayi Hansen. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Appanoose: Chariton R., Centerville, TMM, 7/20/12; Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Clay: Big Muddy Cr., Spencer, TMM, 8/27/13; Clayton: Hewitt Cr., Volga, CTH, 8/4/ 14; Clinton: Brophy Cr., McCausland, CTH, 9/ 30/13; Davis: North Fabius R., Moulton, CTH, 7/17/06; Dickinson: Little Sioux R., Lake Park, TMM, 8/11/14; Dubuque: Middle Fk. Maquoketa R., Rickardsville, CTH, 8/10/04; Fremont: Waubonsie State Park (UV light), ELF, 8/15/12; Kossuth: Lotts Cr., West Bend, MWB, 9/25/ 02; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/ 06; Story: Dye Cr., Colo, TMM, 5/27/14; Wayne: Jordan Cr., Millerton, TMM, 9/12/ 12; Winneshiek: South Pine Cr., Sattre, CTH, 8/14/07; Wright: East Br. Iowa R., Belmond, TMM, 7/23/12. Distribution: Transcontinental in the USA.</p> <p>Chaetarthria atra (LeConte). Delaware: Maquoketa R., Dundee, MWB, 6/24/01. Distribution: Eastern USA, extending west to Missouri and Iowa.</p> <p>Chaetarthria pallida (LeConte). Johnson: (Wickham 1911). Distribution: Primarily eastern USA, with isolated records in the west.</p> <p>Cymbiodyta blanchardi Horn. Allamakee: French Cr., Churchtown, CTH, 9/16/04; Clayton: North Cedar Cr., Clayton, CTH, 9/2/15; Clinton: Elk R., Teeds Grove, CTH, 8/12/13; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 9/9/ 04; Fayette: Unnamed trib. to Volga Lake outlet, Fayette, CTH, 8/4/15; Mitchell: Burr Oak Cr., Osage, CTH, 8/3/11; Winneshiek: UV light, KR, 11/3/01. Distribution: Eastern USA, extending west to Missouri and Iowa.</p> <p>Cymbiodyta chamberlaini Smetana. Allamakee: Suttle Cr., Monona, CTH, 8/17/11; Clayton: Unnamed trib. to Volga R., Volga, CTH, 9/21/15; Delaware: Twin Springs Cr., Colesburg, CTH, 9/ 17/07; Dubuque: Catfish Cr., Dubuque, CTH, 7/ 28/95; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 9/28/11; Jackson: Storybook Hollow Cr., Bellevue, CTH, 8/29/07; Winneshiek: Middle Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Wisconsin in the north, Iowa, and Kansas in the south.</p> <p>Cymbiodyta minima Notman. (Downie and Arnett 1996). Distribution: Northeastern USA, with Iowa forming part of the western border.</p> <p>Cymbiodyta semistriata (Zimmermann). Davis: Burr Oak Cr., Troy, CTH, 6/1/15; Van Buren: Town Branch, Pittsburg, CTH, 7/29/14. Distribution: Eastern USA, extending west to Wisconsin in the north and Colorado.</p> <p>Cymbiodyta toddi Spangler. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/27/13; Butler: South Beaver Cr., Parkersburg, CTH, 7/30/96; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 9/29/ 14; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 8/13/13; Clayton: Bixby Springs, Edgewood, MWB, 9/9/04; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Fayette: Brush Cr., Arlington, CTH, 9/10/04; Franklin: Boylan Cr., Aredale, CTH, 8/11/14; Grundy: South Beaver Cr., Parkersburg, CTH, 7/30/96, 9/ 3/08; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/2/12; Howard: North Branch Turkey R., Lime Springs, CTH, 8/5/15; Jasper: Walnut Cr., Prairie City, CTH, 8/21/95; Johnson: Muddy Cr., North Liberty, CTH, 10/2/ 06; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Lick Cr., Farmington, KJS, 7/28/ 14; Linn: West Otter Cr., Toddville, CTH, 10/11/ 01; Madison: Howerdon Cr., Winterset, JML, 7/ 12/01; Mitchell: Rock Cr., Rock, CTH, 9/10/09; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/ 14; O’ Brien: Little Floyd R., Sheldon, JML, 8/ 22/02; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Sioux: Sixmile Cr., Hawarden, JML, 8/2/00; Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13; Winneshiek: Trout R., Decorah, CTH, 7/18/07, South Pine Cr., Sattre, CTH, 8/14/ 07. Distribution: Eastern USA, extending west to North Dakota in the north and Oklahoma in the south.</p> <p>Cymbiodyta vindicata Fall. Benton: Mud Cr., Van Horne, CTH, 9/30/15; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Clayton: Roberts Cr., Gunder, CTH, 8/11/08; Clinton: Silver Cr., DeWitt, CTH, 7/17/12; Dubuque: Hickory Cr., New Vienna, CTH, 9/18/08; Fayette: Deep Cr., Wadena, CTH, 9/28/05; Ida: Odebolt Cr., Ida Grove, CTH, 8/5/97; Jackson: South Big Mill Cr., Bellevue, CTH, 9/17/14; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06; Sioux: Deep Cr., Granville, KLS, 8/2/06; Woodbury: Dutch Cr., Danbury, TMM, 6/20/16. Distribution: transcontinental in the USA, exclusive of the Southwest.</p> <p>Enochrus cinctus (Say). Davis: Unnamed trib. to North Fabius R., Bloomfield, CTH, 7/15/13; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: (Wickham 1911); Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13. Distribution: Eastern USA, extending west to South Dakota in the north, Kansas, and Texas in the south.</p> <p>Enochrus collinus Brown. Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Henry: Little Potomac Cr., Mt. Pleasant, CTH, 8/7/15; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06. Distribution: Northeastern USA, extending west to Minnesota in the north. Iowa forms the southwestern border of its range. New state record.</p> <p>Enochrus consortus Green. Davis: Soap Cr., Peach Ave., CTH, 7/17/13; Linn: Wapsipinicon R., Viola, DPH, 6/9/01; Louisa: Iowa R., Wapello, MWB, 10/8/13; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Van Buren: Shimek State Forest (UV light), ELF, 5/18/12. Distribution: eastern USA, extending west to Minnesota in the north, Iowa, Missouri, and Louisiana in the south.</p> <p>Enochrus diffusus (LeConte). Howard: (Wickham 1911). Distribution: Northwestern USA, extending east to Michigan.</p> <p>Enochrus fimbriatus (Melsheimer). Johnson: (Wickham 1911); Scott: (Wickham 1911); Story: (Wickham 1911; Gunderson 1967). Distribution: Transcontinental in the USA.</p> <p>Enochrus hamiltoni (Horn). Allamakee: French Cr., Churchtown, CTH, 7/1/02; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Clay: Elk Cr., Gillett Grove, TMM, 9/24/13; Dickinson: (Wickham 1911); Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/4/12; Fayette: Volga R., Maynard, CTH, 8/30/10; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 6/26/13; Jackson: South Big Mill R., Bellevue, CTH, 9/3/10; Jasper: Walnut Cr., Prairie City, CTH, 8/21/95; Linn: East Otter Cr., Alburnett, CTH, 7/10/02; Poweshiek: Walnut Cr., Malcom, CTH, 7/28/09; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06; Story: Dye Cr., Colo, TMM, 6/19/13; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Woodbury: Dutch Cr., Danbury, TMM, 6/20/16. Distribution: Transcontinental in the USA.</p> <p>Enochrus ochraceus (Melsheimer). Allamakee: Yellow R., Postville, CTH, 8/14/00; Black</p> <p>Hawk: Cedar R., Waterloo, CTH, 8/6/12; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Crawford: Otter Cr., Deloit, TMM, 9/30/13; Decatur: Thompson Fk. Grand R., Davis City, TMM, 9/3/13; Dickinson: Little Sioux R., Lake Park, TMM, 7/23/03; Dubuque: Lytle Cr., Zwingle, CTH, 7/17/12; Fayette: Bell Cr., Clermont, CTH, 9/2/04; Floyd: Cedar R., Charles City, MWB, 9/14/01; Franklin: Boylan Cr., Aredale, TMM, 8/11/14; Fremont: Nishnabotna R., Hamburg, TMM, 10/1/13; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/ 07; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/2/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 8/29/12; Jackson: Unnamed trib. to Big Mill Cr., La Motte, CTH, 9/13/11; Jasper: Walnut Cr., Prairie City, CTH, 5/1/96; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Kossuth: East Fk. Des Moines R., Irvington, MWB, 8/25/ 03; Mahaska: Buck Cr., Barnes City, CTH, 7/26/ 12; Mitchell: Wapsipinicon R., McIntyre, CTH, 8/27/08; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Montgomery: Pilot Branch, Stennett, TMM, 9/16/13; Muscatine: Honey Cr., Conesville, CTH, 7/18/06; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/ 14; Polk: Camp Cr., Mitchellville, KBK, 9/26/ 01; Pottawattamie: Jordan Cr., Macedonia, TMM, 9/17/13; Poweshiek: Walnut Cr., Hartwick, CTH, 7/27/09; Sac: North Raccoon R., Auburn, TMM, 10/1/13; Scott: Lost Cr., Princeton, CTH, 8/ 30/04; Van Buren: Town Branch, Pittsburg, CTH, 9/14/15; Wapello: Des Moines R., Ottumwa, CTH, 10/8/12; Wayne: South Fk. Chariton R., Corydon, TMM, 8/24/12; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Woodbury: Little Sioux R., Smithland, TMM, 10/3/12. Distribution: Eastern USA, extending west to the Midwest, although West Coast collections exist.</p> <p>Enochrus pygmaeus (Fabricius). Allamakee: Waterloo Cr., Dorchester, CTH, 8/1/12; Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Boone: Beaver Cr., Beaver, KBK, 7/25/01; Butler: West Fk. Cedar R., Lake Considine Park, CTH, 10/2/12; Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Dallas: Beaver Cr., Perry, JML, 7/25/01; Davis: North Fabius R., Moulton, CTH, 7/17/06; Decatur: Long Cr., Van Wert, CTH, 9/14/95; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 10/2/03; Fayette: Brush Cr., Wadena, CTH, 9/27/05; Floyd: Cedar R., Charles City, MWB, 9/14/01; Franklin: Beaver Cr., Ackley, MWB, 8/15/05; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Harrison: Soldier R., Pisgah, JML, 10/25/00; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: Old Mans Cr., Hill, CTH, 9/18/13; Keokuk: North Skunk R., Sigourney, CTH, 10/7/ 13; Linn: Cedar R., Bertram, CTH, 10/10/05; Louisa: Long Cr., Columbus Junction, CTH, 9/ 14/95; Mahaska: North Skunk R., Rose Hill, CTH, 8/22/11; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Page: West Tarkio Cr., Coin, TMM, 9/7/04; Pottawattamie: Mosquito Cr., Manawa, TMM, 9/ 24/12; Shelby: East Branch West Nishnabotna R., Harlan, KLS, 8/4/05; Van Buren: Town Branch, Pittsburg, CTH, 9/14/15; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Webster: Allen Cr., Stratford, JML, 8/15/07; Woodbury: West Fk.Little Sioux R., Hornick, TMM, 8/18/14. Distribution: Transcontinental in the USA, except the West Coast.</p> <p>Enochrus sayi Gundersen. Des Moines: Flint Cr., Danville, CTH, 7/15/03; Floyd: Cedar R., Carrville, CTH, 8/7/12; Jackson: Maquoketa R., Maquoketa, CTH, 9/13/12. Distribution: Eastern USA, extending west to Iowa in the north, Missouri, and Texas in the south. Iowa forms the northwestern edge of the range.</p> <p>Helochares maculicollis Mulsant. Buchanan: Wapsipinicon R., Independence, CTH, 8/6/12; Linn: Wapsipinicon R., Central City, CTH, 9/27/ 06; Louisa: Iowa R., Wapello, CTH, 10/15/03; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Scott: Wapsipinicon R., DeWitt, CTH, 8/30/ 12. Distribution: Northeastern USA, extending west to Missouri and Iowa.</p> <p>Helocombus bifidus (LeConte). Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/ 12; Howard: (Wickham 1911). Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Missouri in the south.</p> <p>Hydrobius fuscipes (Linnaeus). Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/ 12; Jackson: South Big Mill Cr., Bellevue, CTH, 9/17/14. Distribution: Northern transcontinental in the USA, Iowa forms part of the southern border in the Midwest.</p> <p>Hydrobius melaenus (Germar). Allamakee: Clear Cr., Lansing, MWB, 8/16/07; Clayton: North Cedar Cr., Clayton, CTH, 9/24/07; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Fayette: Unnamed trib. to Volga Lake outlet, Fayette, CTH, 8/4/15; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Wisconsin and Iowa.</p> <p>Hydrochara leechi Smetana. Winneshiek: Decorah, HFW, 7/20/33; (Downie and Arnett 1996). Distribution: Central USA, from North Dakota to New Mexico in the west, and Pennsylvania, Indiana, and Texas in the east.</p> <p>Hydrochara obtusata (Say). Hancock: Pilot Knob Wetland, Forest City, MTF, 7/18/07; Linn: Cedar R., Cedar Rapids, CTH, 8/24/15. Distribution: Northern transcontinental in the USA, Iowa forms part of the southern border in the Midwest.</p> <p>Hydrochara simula Hilsenhoff and Tracy. Jackson: South Big Mill Cr., Bellevue, CTH, 8/ 28/07. Distribution: Great Lakes region, with Iowa forming the southwestern border.</p> <p>H. soror Smetana. Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/27/14; Appanoose: Sharon Bluffs State Park (UV light), ELF, 7/26/ 11; Black Hawk: Beaver Cr., Cedar Falls, MWB, 8/6/12; Butler: West Fk. Cedar R., Dumont, CTH, 10/11/99; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: UV lighting, ELF, 5/19/12; Jackson: South Big Mill Cr., Bellevue, CTH, 8/6/14. Distribution: Eastern USA, extending west to Texas in the south, Oklahoma, Iowa, and Wisconsin in the north.</p> <p>H. spangleri Smetana. Allamakee: Dousman Cr., Ion, CTH, 8/12/15; Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Fremont: Waubonsie State Park (UV light), ELF, 6/22/12; Guthrie: Springbrook Cr., Guthrie Center, TMM, 7/16/13; Johnson: Pechman Cr., Hills, CTH, 4/17/17; Kossuth: Private Wetland, Algona, MTF, 7/14/08; Story: Bear Cr., Ames, TMM, 7/7/13. Distribution: Eastern, exclusive of extreme northeastern USA, extending west to Nebraska in the north and Texas in the south. Iowa forms part of the northern border in the Midwest.</p> <p>Hydrophilus ovatus Gemminger and Harold. Johnson: (Wickham 1911). Distribution: Eastern USA, extending west to Iowa in the north, Kansas, and Texas in the south. Iowa forms part of the northwestern border.</p> <p>Hydrophilus triangularis Say. Cerro Gordo: Private Wetland, Ventura, MTF, 7/20/07; Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Franklin: Hampton, FHS, 9/5/ 1896; Fremont: Sapp Bros. Truck stop (UV light), ELF, 4/17/04; Greene: Snake Cr., Rippey, MTF, 8/14/07; Johnson: State Hygienic Laboratory, Coralville, KJS, 5/20/13; Winnebago: Private Wetland, Thompson, MTF, 7/18/07; Worth: Shell Rock R., Kensett, CTH, 10/15/02. Distribution: Transcontinental in the USA.</p> <p>Laccobius agilis (Randall). Madison: North Branch North R., Winterset, CTH, 6/14/96. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.</p> <p>Laccobius minutoides (Orchymont). Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/13; Wayne: South Fk. Chariton R., Corydon, TMM, 8/24/12. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.</p> <p>Laccobius reflexipenis Cheary. Allamakee: Hecker Cr., Postville, CTH, 7/24/07; Appanoose: Packard Cr., Exline, CTH, 9/15/15; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Dallas: Puckerbush Access (UV light), ELF, 11/ 1/12; Davis: North Fabius R., Moulton, CTH, 7/ 17/06; Delaware: Maquoketa R., Dundee, MWB, 6/24/01; Des Moines: Flint Cr., Danville, CTH, 7/15/03; Dubuque: Hickory Cr., New Vienna, 9/18/08; Fayette: Brush Cr., Wadena, CTH, 9/27/05; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 6/26/13; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Jones: Maquoketa R., Monticello, CTH, 9/6/05; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Shimek State Forest (UV light), ELF, 4/3/05; Linn: Cedar R., Bertram, MWB, 10/10/05; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Winnebago: Buffalo Cr., Forest City, MWB, 9/ 16/06; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.</p> <p>Laccobius spangleri Cheary. Cherokee: Ashton Cr., Washta, TMM, 7/6/06; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 8/26/14; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 6/29/15; Dubuque: Cloie Branch, Durango, CTH, 8/5/14; Fayette: North Branch Volga R., Randalia, CTH, 9/18/01; Ida: Ashton Cr., Washta, JML, 7/6/06; Jackson: Prairie Cr., Maquoketa, CTH, 8/30/06; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Sioux: Floyd R., Alton, KLS, 8/6/03; Van Buren: Town Branch, Pittsburg, CTH, 9/14/16; Wayne: Jordan Cr., Millerton, TMM, 9/12/12. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.</p> <p>Laccobius truncatipenis Miller. Clayton: Bear Cr., Edgewood, MWB, 9/9/04; Fayette: Bell Cr., Clermont, CTH, 7/20/04; Harrison: Soldier R., Pisgah, JML, 7/11/00; Jackson: North Fk. Maquoketa R., Maquoketa, CTH, 9/16/13; Shelby: Unnamed trib. to West Nishnabotna R., Irwin, JML, 7/6/05; Winneshiek: South Pine Cr., Sattre, CTH, 8/27/14. Distribution: Northwestern USA, extending east to Michigan. Iowa forms part of the southern border in the Midwest.</p> <p>Paracymus despectus (LeConte). Davis: Fox R., Moulton, CTH, 6/29/06. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.</p> <p>Paracymus subcupreus (Say). Allamakee: Yellow R., Postville, CTH, 8/14/00; Black Hawk: Beaver Cr., Cedar Falls, CTH, 10/9/14; Buchanan: Bear Cr., Brandon, CTH, 8/3/10; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/27/13; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cedar: Unnamed trib. to Bennett Cr., Tipton, CTH, 7/16/12; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 9/20/10; Clay: Little Sioux R., Spencer, TMM, 9/6/05; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Davis: Fox R., Moulton, CTH, 6/29/06; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Tetes Des Morts Cr., St. Donatus, CTH, 8/1/05; Fayette: Little Volga R., Maynard, CTH, 7/23/14; Floyd: Cedar R., Charles City, CTH, 9/14/01; Franklin: Bailey Cr., Sheffield, CTH, 7/20/11; Guthrie: Middle Raccoon R., Bagley, KLS, 9/20/05; Henry: Big Cedar Cr., Oakland Mills, 9/2/11; Howard: North Branch Turkey R., Lime Springs, CTH, 8/ 5/15; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: Muddy Cr., North Liberty, CTH, 10/2/ 06; Jones: Helmer Cr., Prairieburg, CTH, 8/31/ 05; Lee: Lick Cr., Farmington, CTH, 7/28/14; Mills: Mud Cr., Hastings, JML, 8/2/05; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Montgomery: Pilot Branch, Stennett, CTH, 9/ 9/96; Muscatine: Mud Cr., Wilton, CTH, 9/15/ 03; Pocahontas: Big Cedar Cr., Fonda, JML,9/ 27/06; Poweshiek: Walnut Cr., Brooklyn, CTH, 7/28/09; Story: Ballard Cr., Cambridge, KJS, 7/ 18/07; Van Buren: Town Branch, Pittsburg, CTH, 8/3/15; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08. Distribution: Transcontinental in the USA, except extreme western states.</p> <p>Sperchopsis tessellata (Ziegler). Allamakee: Yellow R., Ion, CTH, 8/29/02; Buchanan: Pine</p> <p>Cr., Quasqeton, CTH, 8/28/97; Butler: Coldwater Cr., Greene, CTH, 7/22/96; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Chickasaw: Wapsipinicon R., New Hampton, CTH, 10/6/04; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Delaware: Plum Cr., Hopkinton, CTH, 8/13/ 15; Dubuque: Little Maquoketa R., Dubuque, CTH, 9/21/95; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Little Cedar R., Colwell, CTH, 10/3/95; Franklin: Bailey Cr., Sheffield, CTH, 7/22/96; Howard: Crane Cr., Lourdes, CTH, 7/19/94; Jackson: Prairie Cr., Maquoketa, CTH, 8/14/01; Jones: Silver Cr., Monticello, MWB, 7/31/01; Linn: East Big Cr., Springville, CTH, 7/18/01; Mitchell: Burr Oak Cr., Osage, CTH, 8/14/95; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Poweshiek: North Skunk R., Millgrove County access, CTH, 7/26/12; Scott: Lost Cr., Princeton, CTH, 8/9/12; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Wright: Otter Cr., Goldfield, KLS, 8/11/00. Distribution: Eastern USA, extending west to Iowa and Arkansas.</p> <p>Tropisternus affinis Motschulsky. Allamakee: Upper Iowa R., Dorchester, MWB, 9/27/02; Butler: West Fk. Cedar R., Lake Considine County park, CTH, 10/2/12; Clayton: Silver Cr., Monona, CTH, 8/7/06; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Deep Cr., Wadena, CTH, 9/28/ 05; Guthrie: Unnamed trib. Middle Raccoon R., Panora, TMM, 9/30/14; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Kossuth: Plum Cr., Algona, MWB, 8/24/05; Linn: Cedar R., Cedar Rapids, CTH, 9/28/12; Mitchell: Deer Cr., Carpenter, CTH, 8/28/08; Muscatine: Pike Run, West Liberty, CTH, 7/19/06; Winnebago: Buffalo Cr., Forest City, JML, 9/16/06; Winneshiek: Canoe Cr., Decorah, CTH, 9/14/11; Worth: Beaver Cr., Fertile, KLS, 9/19/06. Distribution: Western USA, extending east to Wisconsin in the north and Texas in the south.</p> <p>Tropisternus blatchleyi Orchymont. Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Wapello: Des Moines R., Ottumwa, CTH, 10/7/ 13. Distribution: Eastern USA, with Iowa forming the northwestern border.</p> <p>Tropisternus collaris (Fabricius). Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Henry: Big Cr., New London, CTH, 8/30/05; Sac: North Raccoon R., Auburn, TMM, 10/1/13;</p> <p>Van Buren: Des Moines R., Farmington, CTH, 9/6/06; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05. Distribution: Southeastern USA, with Iowa forming the northwestern border.</p> <p>Tropisternus columbianus Brown. Allamakee: Unnamed trib. Yellow R., Postvile, CTH, 7/24/ 07; Clayton: Brownfield Cr., Colesburg, CTH, 9/18/07; Muscatine: Pike Run, West Liberty, CTH, 7/19/06; Winneshiek: South Pine Cr., Sattre, CTH, 10/27/10. Distribution: Western USA, extending east to Ohio in the north and Texas in the south.</p> <p>T. glaber (Herbst). Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05; Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/ 15; Buchanan: Lime Cr., Brandon, CTH, 9/4/07; Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Clinton: Elk R., Teeds Grove, CTH, 8/13/12; Delaware: Little Turkey R., Colesburg, MWB, 9/8/06; Fayette: Brush Cr., Wadena, CTH, 9/27/ 05; Floyd: Little Cedar R., Colwell County Park, CTH, 9/9/09; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Muscatine: Cedar R., Conesville, CTH, 8/29/12; Winneshiek: Coldwater Cr., Bluffton, CTH, 9/22/04. Distribution: Northeastern USA, extending west to South Dakota and Iowa.</p> <p>Tropisternus lateralis (Fabricius). Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Appanoose: Cooper Cr., Centerville, CTH, 8/11/ 05; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Black Hawk: Black Hawk Cr., Hudson, CTH, 8/ 8/02; Bremer: Cedar R., Janesville, CTH, 9/9/ 14; Buchanan: Bear Cr., Brandon, CTH, 8/26/ 13; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 5/15/12; Chickasaw: Wapsipinicon R., New Hampton, CTH, 9/6/94; Clay: Little Sioux R., Spencer, TMM, 9/9/14; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Clinton: Elk R., Andover, CTH, 8/7/07; Dallas: North Raccoon R., Adel, TMM, 9/29/05; Davis: Fox R., Bloomfield, CTH, 7/15/13; Decatur: Little R., Leon, CTH, 9/23/02; Delaware: Maquoketa R., Manchester, CTH, 9/25/13; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dubuque: North Fk. Maquoketa R., Dyersville, CTH, 7/25/05; Emmet: Drainage Ditch 40, Dolliver, TMM, 7/30/15; Fayette: Volga R., Randalia, CTH, 8/31/10; Floyd: Cedar R., Charles City, CTH, 9/26/02; Franklin: Beaver Cr., Ackley, CTH, 9/10/14; Greene: West Buttrick Cr., Spring Lake Park, TMM, 7/31/14; Hancock: Pilot Knob Wetland, Forest City, MTF, 7/5/07; Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Harrison: Soldier R., Pisgah, TMM, 9/5/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/2/11; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 8/27/07; Jasper: Walnut Cr., Prairie City, CTH, 8/15/01; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Bear Cr., Wyoming, CTH, 7/13/04; Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Lee: Skunk R., Augusta, CTH, 10/8/13; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03; Louisa: Long Cr., Columbus Junction, CTH, 9/14/95; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Lyon: Little Rock R., George, TMM, 9/28/11; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Mitchell: Cedar R., Osage, CTH, 7/6/06; Montgomery: Pilot Branch, Stennett, TMM, 9/16/13; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/14; Plymouth: Big Sioux R., Akron, TMM, 10/10/ 12; Polk: Des Moines R., Des Moines, TMM, 9/ 24/13; Pottawattamie: Mosquito Cr., Manawa, TMM, 9/24/12; Sac: North Raccoon R., Sac City, TMM, 7/31/12; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/ 00; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Union: Unnamed trib. to Thompson R., Lorimor, TMM, 9/17/15; Van Buren: Town Branch, Pittsburg, CTH, 10/1/14; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Winnebago: Buffalo Cr., UWW10, JML, 9/16/06; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Woodbury: Wolf Cr., Correctionville, TMM, 9/22/14; Worth: Elk Cr., Kensett, CTH, 8/26/ 08; Wright: Lower Morse Lake Wetland Complex, Belmond, MTF, 7/15/09. Distribution: Eastern USA, extending west to the Rocky Mountains.</p> <p>Tropisternus mixtus (LeConte). Allamakee: Dousman Cr., Ion, CTH, 8/12/15; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Fayette: Dibble Cr., Clermont, CTH, 9/24/14; Floyd: Cedar R., Floyd, CTH, 9/23/08; Howard: Crane Cr., Lourdes, CTH, 9/28/09; Johnson: Pechman Cr., Hills, CTH, 4/17/17; Linn: Cedar R., Cedar Rapids, CTH, 9/28/12; Muscatine: Muscatine Slough, Muscatine, CTH, 9/17/02; Winnebago: Private Wetland, Thompson, MTF, 7/18/07; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08; Worth: Beaver Cr., Fertile, KLS, 9/19/06. Distribution: Transcontinental in the USA.</p> <p>Tropisternus natator Orchymont. Allamakee: Clear Cr., Lansing, MWB, 8/16/07; Appanoose: Chariton R., Rathbun, CTH, 8/17/04; Black Hawk: West Fk. Cedar R., Finchford, MWB, 9/ 15/08; Bremer: Cedar R., Janesville, CTH, 8/6/ 12; Buchanan: Lime Cr., Brandon, CTH, 8/21/ 08; Butler: West Fk. Cedar R., Dumont, CTH,</p> <p>10/11/06; Clarke: West Long Cr., Grand River, TMM, 8/25/15; Clay: Little Sioux R., Spencer, TMM, 9/8/14; Clayton: Turkey R., Garber, MWB, 10/3/03; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Dallas: South Raccoon R., Redfield, TMM, 8/8/12; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: Little R., Leon, CTH, 9/ 23/02; Delaware: Little Turkey R., Colesburg, MWB, 9/8/06; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08; Fayette: Little Turkey R., West Union, CTH, 9/10/03; Floyd: Cedar R., Charles City, CTH, 9/14/01; Greene: West Buttrick Cr., Spring Lake Park; Guthrie: South Raccoon R., Stuart: Hamilton: Boone R., Stratford, TMM, 8/16/12; Hancock: Winnebago R., Forest City, KLS, 9/18/06; Henry: Skunk R., Oakland Mills, CTH, 9/24/ 08; Howard: Crane Cr., Lourdes, CTH, 9/24/ 13; Jackson: Maquoketa R., Canton, CTH, 9/ 7/05; Jasper: Walnut Cr., Prairie City, CTH, 8/15/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 9/ 22/05; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Keokuk: Twin Grove Cr., Ollie, CTH, 5/22/14; Kossuth: East Fk. Des Moines R., Irvington, 8/25/03; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03; Louisa: Long Cr., Columbus Junction, CTH, 10/1/13; Lucas: Whitebreast Cr., Woodburn, JML, 9/24/02; Madison: North Branch North R., Winterset, CTH, 8/25/95; Mahaska: Muchakinock Cr., Eddyville, CTH, 9/2/11; Marion: English Cr., Knoxville, CTH, 8/1/13; Mills: West Nishnabotna R., Malvern, TMM, 9/4/12; Mitchell: Wapsipinicon R., McIntyre, CTH, 9/10/07; Muscatine: Pike Run, Nichols, CTH, 8/23/04; Polk: Four Mile Cr., Ankeny, TMM, 8/9/06; Poweshiek: Walnut Cr., Holiday Lake, CTH, 8/25/10; Scott: Lost Cr., Princeton, CTH, 8/9/12; Tama: Richland Cr., Haven, CTH, 10/6/10; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Warren: South R., Ackworth, TMM, 7/23/12; Washington: English R., Riverside, CTH, 10/9/12; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05; Winneshiek: Upper Iowa R., Decorah, CTH, 7/25/06; Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Eastern USA, extending west to Texas in the south, Colorado, and Minnesota in the north.</p></div> 	https://treatment.plazi.org/id/03F487A3FFD6FFF3FED4FD9594D5FA4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
03F487A3FFECFFF0FC9CF9DD921CFE19.text	03F487A3FFECFFF0FC9CF9DD921CFE19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noteridae	<div><p>NOTERIDAE</p> <p>Hydrocanthus iricolor Say. Cass: West Nodaway R., Grant, TMM, 8/19/04; Jackson: Pleasant Cr., Springbrook, CTH, 8/31/11; Johnson: Ciha Fen, Sutliff, CTH, 4/17/17; Louisa: Iowa R., Lone Tree, CTH, 8/25/15; Muscatine: Pike Run, West</p> <p>Liberty, CTH, 7/19/06; Scott: (Wickham 1911). Distribution: Eastern USA, extending west to Wisconsin in the north, Kansas, and New Mexico in the south. Iowa forms part of the western border.</p> <p>Suphisellus puncticollis (Crotch). Johnson: Ciha Fen, Sutliff, CTH, 4/17/17. Distribution: Eastern USA, extending west to Wisconsin in the north, Missouri, and Louisiana in the south. Iowa forms part of the western border. New state record.</p> </div>	https://treatment.plazi.org/id/03F487A3FFECFFF0FC9CF9DD921CFE19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hubbard, Todd	Hubbard, Todd (2017): Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes. The Coleopterists Bulletin 71 (3): 471-505, DOI: 10.1649/0010-065X-71.3.471, URL: http://dx.doi.org/10.1649/0010-065x-71.3.471
