taxonID	type	description	language	source
03F687E7301AFF88FF5B7D36FDE99095.taxon	materials_examined	Type species. Feronia molopina Chaudoir, 1868 by original designation.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301AFF88FF5B7D36FDE99095.taxon	description	Redescription. Medium-sized pterostichine (BL 11.2 – 16.2 mm), piceous brown to black, smooth, shiny though some females matt. Habitus very similar to Molops s. str., mainly representatives of the “ piceus ” species group, which also represented in NW Turkey by a single polytypic species. Head relatively large and robust, smooth, with prominent eyes. Frons with two supraorbital setigerous punctures. Clypeus with two long setae, labrum anteriorly with six marginal setae. Mentum with pair of setigerous punctures; tooth prominent and bifide at apex. Submentum with two pairs of setae, outer setae much shorter than inner ones. Glossal sclerite rectangular with two setigerous punctures on anterior margin; paraglossae apparently longer than glossal sclerite. Antennae relative short, reaching anterior angles of elytra, Antennomere I twice as long as II and much thicker then remaining, antennomeres IV – XI covered with dense pubescence. Pronotum almost as wide as long, or slightly wider than long, with rounded lateral sides. Anterior angles slightly prominent, rounded at apex. Posterior angles distinct, acute (T. molopinus, T. wiedemanni, T. phrygius) to rectangular (T. molopiformis), without setigerous punctures. Base of pronotum on each side with single modestly deep, smooth (sometimes with fine transversal wrinkles), linear posterolateral impression, spreading over basal third. Median longitudinal impression distinct, not reaching basal or anterior margins. Lateral groove evenly narrow along sides, with single setigerous punctures in anterior third. Elytra oval or more or less cylindrical, with distinct, finely punctate striae, with rudimentary scutellar stria, scutellar setigerous punctures missing. Third elytral interval with two setigerous punctures (normal 2 nd and 3 rd dorsal elytral pores in carabids) situated in apical half of elytra; exceptionally with one or three punctures. Umbilicate series consisting of 13 – 17 pores in all but one species (10 pores in T. relegatus sp. nov., see below); one setigerous punctures at the end of 7 th elytral stria. Mesepisternum without punctation. Last abdominal segment (sternum VII) with two pores (T. machardi, some specimens of T. phrygius) or four pores (other species). Legs relatively short, robust, tarsomeres 1 – 4 glabrous above, pubescent ventrally. Onychium without setae underneath (except T. machardi - with one or two pairs of setae). Two setigerous punctures on middle coxa, only one on middle trochanter. Three setigerous punctures on hind coxa, including medial one situated below meeting point of coxae; single seta on hind trochanter. Profemur with two setigerous punctures on anterior ventral margin; mesofemur with three medio-ventral seta on anterior margin and 3 (rarely 4) on posterior ventral margin; metafemur with two setae on posterior ventral margin. Mesotibial ctenidium clear but poorly differentiated (as in Molops spp. and Myas). Males with first three protarsomeres dilated, ventrally with densely arranged adhesive setae. Male genitalia: Aedeagus relatively large, median lobe rectangulary arcuate and apex variable (ventral view), short and subtriangular (dorsal view) in T. molopiformis to very long, narrow, and hooked at extremity in T. wiedemanni, apical orifice deflected to left lateral position. Left paramere with transverse apophysis; right paramere elongate apically. Female genitalia: Tergum VIII with convex or subtriangular distal margin and ± short proximal “ legs ”. Sternum VIII consisting, in most species, of two chitinised plates connected by a membrane which Connected by membrane which is disappeared in T. wiedemanni. Syntergum IX + X long, narrow, bearing a pair of small, articulated ovipositors. Ovipositor consisting of large valvifer and one-segmented stylus (in the most species) or two-segmented stylus (in T. molopiformis). Spermatheca with seminal canal and receptaculum discrete; receptaculum shorter than seminal canal; spermathecal canal long or short, inserted at junction of receptaculum and seminal canal. Larvae. Unknown.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301AFF88FF5B7D36FDE99095.taxon	distribution	Distribution. The presence of T. (M.) molopinus (cf. Chaudoir, 1868) in Europe is doubtful. In Asia, Molopsis inhabit the western half of the Anatolian Peninsula but the eastern limits of the range remain poorly documented at present. The species occur in forested regions, elevated alpine meadows, near wet places, streams, thawing snow, under stones, leaves, and pieces of wood.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301CFF8AFF5B7AF4FAC89548.taxon	materials_examined	Type locality. Anatolie, Bosch-Dagh. Taxonomic decisions. Feronia (Haptoderus) rectangula Chaudoir, 1868: 246. Platysma (Crisimus) molopiforme Lutshnik, 1922: 77, replacement name for Feronia rectangula Chaudoir, 1868: 246, nec Fairmaire, 1866: 252.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301CFF8AFF5B7AF4FAC89548.taxon	materials_examined	Material examined. 29 specimens. Type material. Lectotype Ƥ of Feronia rectangula here designated, relatively well preserved, “ rectangula Chaud. Anatolie. Lederer. ” (h label pinned on the bottom of box) / LECTOTYPE Feronia (Haptoderus) rectangula Chaudoir G. Dubault désign. 2003 (red p) / Tapinopterus (Molopsis) molopiformis G. Dubault détermin. (p) (Fig. 3) (MNHN, “ Collection Chaudoir ”, box no 216 “ Argutor ”). Other material examined. Boz daglari: 2 33 2 ƤƤ, “ Caucas ” / “ Coll. Türk 188 ” (NMW); 1 Ƥ, “ Boszdagh Coll. Javet ” (MNHN); 1 Ƥ, “ Bosdagh ” (MNHN); 4 33 5 ƤƤ, “ Tmolos-Gbg., Lydien, West-Kleinasien, Weirather, Innsbruck ” (MCSNM, MCSNT, MNHN; ZMAN); 1 3, “ Asia Minor Bos-Dagh. ” / “ Coll. Türk 188 ” / “ Pt. rectangulus Chd. Coll. Reitter ” (HNHM); 1 3, “ Bos dagh As. min. ” / “ rectangulus ” / “ Coll. Apfelbeck ” / “ Tapinopterus rectangulus Chd. det. Ing. Jedlička ” (HNHM); 1 3, Caucase? (h) (ZISP); 1 Ƥ, Klein Asien, Bosz-Dagh, Lederer (h) (DEI); 1 3 1 Ƥ, TR (Izmir) Boz-daġ, ob. Bozdagköy, 1500 – 1700 m, 2. – 3.5.1996, Heinz lgt. (cWH); 2 33, TURKEY, vil. Izmir, Boz daġlari, Boz dag köy env., 1200 – 1500 m, 29.5.2003, R. Lohaj & I. Smatana lgt. (cRL) (Fig. 4); 3 33 2 ƤƤ, NW Turkey, prov. Izmir, Boz dag, 1520 m, NE Ödemish, 6.5.0 7 lgt. E. Hajdaj (cEH, cRL).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301CFF8AFF5B7AF4FAC89548.taxon	diagnosis	Diagnosis. BL 11.2 – 13.8 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301CFF8AFF5B7AF4FAC89548.taxon	description	Male genitalia. Median lobe of aedeagus short, with subtriangular apex, apical orifice deflected to left lateral position (Figs. 26 – 27); basal bulb short, with concave basal orifice, apical part longer, copulatory sclerite straight, obliquely situated in medial part, apex slighlty bent downwards (Figs. 46, 56). Right paramere relatively short, slightly curved medially, with thicker basal part and thinner apical one (Figs. 81, 96). Left paramere with long, thin transverse apophysis and oblique paramedial apophysis (Figs. 111, 125). Female genitalia. Tergum VIII with convex distal margin and short proximal “ legs ” (Fig. 139). Sternum VIII consisting of two chitinised plates connected by membrane, with well-developed “ legs ” (Fig. 145). Syntergum IX + X long, narrow, bearing pair of small, articulated ovipositors (Fig. 151). Ovipositor consisting of a large valvifer and smaller two-segmented stylus (Figs. 155 – 156); valvifer with both internal and external margins each with pointed tip and well-developed process across; basal stylomere relatively larger than apical one, latter with single minute (probably ensiform) seta. Spermatheca with seminal canal and receptaculum discrete; seminal canal long; receptaculum slighty shorter than seminal canal, coiled apically (Fig. 163); spermathecal gland prolonged, spermathecal canal long, inserted at junction of receptaculum and seminal canal (Fig. 167). Taxonomic notes. This taxon was first recorded as Molops wiedemanni Chaudoir from Boz dag by Fairmaire (1866: 252). Chaudoir (1866) noted that Faimaire's specimens were different from his specimens of F. wiedemanni and described the taxon as Feronia rectangula. The name Feronia rectangula was preoccupied by Feronia (Orthomus) rectangula Fairmaire, 1859, currently a synonym of Orthomus (Orthomus) barbarus barbarus (Dejean, 1828) (see Bousquet, 2003: 477). Lutshnik (1922: 77 – 78) first noted this homonymy and proposed a new name, Platysma molopiforme, for Feronia rectangula Chaudoir, 1868. Müller (1937: 132) studied a series of T. molopiformis from the type locality and compared it with specimens of his T. phrygius s. str. and T. phrygius pisidicus and stated that the former is a distinct species.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301CFF8AFF5B7AF4FAC89548.taxon	distribution	Distribution. Boz daġlari (Tmolos Mountains by the ancient authors) near Izmir (“ Lydia ”), W Turkey.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301FFF84FF5B78ECFE529563.taxon	materials_examined	Type locality. Turkey? Material examined. One specimen. Type material. Holotype 3, with left hind leg and three right metatarsomeres missing, Armenie (h) / Fer. (Tapinopt.) rectangula Chd. (pale orange h) * / Zool. Inst St. Petersburg (yellow p) / HOLOTYPE Tapinopterus (sg. Molopsis) aenigmaticus n. sp. Lohaj, Guéorguiev, Dubault & Lassalle, 2004 (red p) / (Fig. 5) (ZISP). * Note: One male of Molopsis molopiformis in ZISP bears label “ Caucase? ” written with the same handwriting and on same pale orange paper as the first two labels of the new species. We assume these labels were later additions.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301FFF84FF5B78ECFE529563.taxon	description	Description. BL 12.1 mm, maximum width 4.4 mm. Head, pronotum and elytra black, shiny, antennae, legs and abdomen piceous brown. Head large, robust, almost as long as wide, shiny, smooth, with very fine microsculpture. Antennae relatively short, reaching hind angles of pronotum, with dense, decumbent pubescence on antennomeres 4 – 11. Pronotum wider than long, ratio PL / PW 0.67, with rounded lateral sides, shiny, smooth, with very fine transversal wringles along the midline. Anterior edge strongly sinuate, with prominent, rounded, apical angles. Posterior angles small, almost rectangular. Base of pronotum with single deep linear posterolateral impression on each side of midline, reaching basal third, smooth, with very fine transversal wrinkles. Midline distinct, not reaching basal or anterior margins. Elytra parallel-sided, slightly flattened, with distinct, finely punctate striae. Third elytral interval with two setigerous pores situated in apical half of elytra, 2 nd in middle and 3 rd in apical fourth. Umbilicate series consists of 14 setigerous punctures on each side, in the middle widely interrupted, with distribution: left side 5 + 1 + 8, right side 6 + 8. Male genitalia. Median lobe of aedeagus relatively short, straight, with short and fine apex in dorsal view and a small twisted tooth in ventral position (Figs. 28 – 29); apical orifice deflected to left laterad position, ventral margin forming slight acute angle at medial part, lobe with distinct lateral process in left lateral view (Figs. 47, 57). Right paramere strongly curved apically, with thicker basal part and thin apical part truncate towards tip (Fig. 82, 97). Left paramere with subangular plate, long, thin transverse apophysis, slightly bent inwards, and oblique paramedial apophysis (Figs. 112, 126). Female genitalia. Unknown. PLATE 1. Habitus of Tapinopterus species (Figs. 1 – 12). Fig. 1. Tapinopterus (Pseudomolopsis) rebellis (Greece, Taygetos); Fig. 2. T. (Pseudomolopsis) rebellis kumanensis (holotype); Fig. 3. T. (Molopsis) molopiformis (lectotype of Feronia rectangula); Fig. 4. T. (Molopsis) molopiformis (Turkey, Boz daġ); Fig. 5. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 6. T. (Molopsis) phrygius (syntype); Fig. 7. T. (Molopsis) phrygius (Turkey, Davraz daġ); Fig. 8. T. (Molopsis) phrygius (Turkey, Kumalar daġlari); Fig. 9. T. (Molopsis) phrygius (holotype of Tapinopterus dipojranus); Fig. 10. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 11. T. (Molopsis) phrygius (Turkey, Beyşehir); Fig. 12. T. (Molopsis) phrygius (Turkey, Sultan daġlari).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301FFF84FF5B78ECFE529563.taxon	etymology	Etymology. From the Latin adjective aenigmaticus, meaning “ enigmatic ”. The name reflects the uncertainty about the type locality of this species.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301FFF84FF5B78ECFE529563.taxon	diagnosis	Diagnosis. This species is easily recognized from all other species of the group by the shape of the median lobe of aedeagus and right paramere; median lobe dorsally with short, fine apex, with distinct lateral process at left lateral position which forms a small tooth ventrally; ventral margin of forming slight acute angle at medial part (Figs. 28 – 29, 47, 57). Right paramere strongly curved apically, with thin apical part not pointed towards tip (Fig. 82, 97).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7301FFF84FF5B78ECFE529563.taxon	distribution	Distribution. The type locality of this new species remains uncertain. If the locality label “ Armenie ” is correct (see also Note above), the new species originated either from Armenia (former republic of USSR) or from “ Armenian Taurus ” of ancient authors. This region, also called “ Small Armenia ” is located in Kurdistan, south of Lake Van (see Casale & Vigna Taglianti, 1999: 401, note 21). Correct type locality needs to be confirmed by the examination of additional material.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73011FF86FF5B780FFB9E952B.taxon	materials_examined	Type locality. Kleinasien, Ak-Dagh im Phrygischen Taurus. Taxonomic decisions.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73011FF86FF5B780FFB9E952B.taxon	materials_examined	Material examined. 72 specimens. Type material. One syntype (3) of Tapinopterus (Crisimus) phrygius, Ak dagh bei Tschiwril Asm. (h) / phry- gius Müll. (h) / Paratypus (p) (red label) / 322 (h) (red label) / (Fig. 6) (MCSNM). Two syntypes (33) of Tapinopterus (Crisimus) phrygius pisidicus, “ Klein Asien Pisidischer Taurus Barla Gruppe N. O. von Isparta ” (green p / h) / “ vend. Weirather Tapinopterus (Crisimus) phrygius pisidicus J. Müll. ” (p / h) / “ Tapinopterus phrygius pisidicus J. Mü. ” (pink h) (NMP). Holotype 3 of Tapinopterus (Molopsis) dipojranus, Dipojras Gbg. Pisid. Taurus, Klein-Asien, Weirather, Innsbruck. (p) / Holotypus (p) Tapinopt. dipojranus (h) (red label) / Holotypus Tapinopterus dipojranus (h), det. S. L. Straneo 1985 (p) / TAPINOPTERUS (sg. MOLOPSIS) phrygius Müller 1932 R. LOHAJ det. 2003 (p) / (Fig. 9) (MCSNM). Four paratypes (33) of Tapinopterus (Molopsis) dipojranus ssp. cilicius, TURCHIA — vil. Mersin, Namrun (Çamliyayla) 5. VI. 1983, G. Sama leg. (p) /, Bolkar Daġ, altipiano m 2400 - (h) / PARATYPUS (p) Tap. dipojr. s. cilicius (h) (red label) / Paratypus T. dipojranus s. cilicius (h), det. S. L. Straneo 1985 (p) / TAPINOPTERUS (sg. MOL- OPSIS) phrygius Műller 1932 R. LOHAJ det. 2003 (p) / (Fig. 10) (2 33 MCSNM, 1 3 cBL, 1 3 cWH). Other material examined. Barla daglari: 1 3, Barla-Gbg. Pisid. Taurus (MCSNM); 1 Ƥ, Gelindschik Ana- Gbg. nö Isparta, Pisidia, Asm. (MCSNM); 1 3, Turcia, 8.5.1991, Barla dagi, 200 – 2500 m, M. Mikát lgt. (cVS); 1 Ƥ, “ Barla-Gbg. Pisid. Taurus ” (MNHN). Kumalar daglari: 2 33 2 ƤƤ, Turkey occ., 21. vi. 1999, Kumalar Daglari mts., alp., Basören S, 1900 – 2200 m (50 km S-of Afyon) S. Benedikt leg. (Fig. 8) (cVS, cRL). Sultan daglari: 1 Ƥ, Turkey, 23. – 26.5.96, Sultan Daglari, Cankurtaran, 1800 m, Smatana lgt. (cRL) (Fig. 12); 1 3 2 ƤƤ, “ Asia minor Sultan Dagh v. Bodemeyer ” (MNHN, ZMAN); 1 Ƥ, Asia minor, Sultan Dagh, v. Bodemeyer, O. Leonhard (DEI); 1 Ƥ, Asia minor, Sultan Dagh, v. Bodemeyer, ex-coll. Dr. Melichar (BMNH); 1 Ƥ, Anatolien, Ak-Chehir, 1900, Korb. (DEI); 1 3 1 Ƥ, Anatolia centr, Sultan Dagh b. Çay, 1900 – 2200 m, 18.7.1965, Heinz lgt. (cWH); 4 33 2 ƤƤ, Asia minor, Sultan Dagh, v. Bodemeyer (HNHM, NMP, ZISP). Davraz dag: 9 33 2 ƤƤ, TURKEY W, 1900 m, 22.5.2001, Isparta, Davras dag, lgt. M. Snížek (Fig. 7) (cVS, cDW, cRL); 1 3, TR. Vil. Isparta, 24.5. 2001, 10 km W of Yukarigökdre, Kasnak, J. Mertlik lgt. (cVS); 9 33 2 ƤƤ, Davraz, Isparta Turquie, VI. 96, G. Dubault, B. Lassalle (sGD, cBL); 1 Ƥ, Davraz Daġ, w. Anatolien, 6.5.72 (cWH); 4 33 2 ƤƤ, “ Turquie (Isparta) Mt. Davras 2.000 / 2.200 H. Coiffait. 20. V. 54 ” (MNHN). Dedegöl daġlari: 1 3, Dedegöl-daġ, Nordsei, alpine zone, s üdl. Egirdir, 28. VII. 1971, 1800 – 2300, Anatolia mer, Heinz lgt. (cWH); 1 3 1 Ƥ, TR vill. Isparta, Yenisar Bademli env., Dipoyraz daġi, Josef Mertlik lgt, 1.6.1995 (cVS). Bolkar daglari: 1 Ƥ, Dumbelek, Taurus Cil, Anatolien (MNHUB); 2 33 1 Ƥ, Namrun, Anat. m., 10.5. – 3.6.63, leg. F. Schubert (MNHUB, NMP); 3 33 2 ƤƤ, Turkei, Mersin, Findik pinari, Arslanköi, 2.5.0 1, leg. Megger (cDW, cRL). Konya Villayet: 131 Ƥ, “ Turkey, 90 IV 18. Konya. Beysehir leg. Podlussány A. ” (Fig. 11) (HNHM).! unspecified localities: 1 3, “ 76 ” (green h) / “ T. phrygius pisidicus ” (yellow h) (MCSNT); 1 3, “ 89 ” (green h) / “ Tapinopt. phrygius m det. J. Müller ” (h / p) (MCSNT); 1 Ƥ, “ 89 ” (green h); 1 3, “ H 406 ” (h) (MCSNT).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73011FF86FF5B780FFB9E952B.taxon	diagnosis	Diagnosis. BL 11.8 – 15.9 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73011FF86FF5B780FFB9E952B.taxon	description	Male genitalia. Median lobe of aedeagus short to medium-sized, apical lamella long, thin, more or less regularly curved to left (Figs. 32 – 34) or rarely slightly sinuous before left-orientated tip (Figs. 30 – 31); apical orifice deflected to left lateral position; basal bulb relatively long, with basal orifice concave (Figs. 48, 50 – 52), apical part longer than basal one (Figs. 50 – 52, 60 – 62), rarely almost equal in size (Figs. 48, 58), curved copulatory sclerite in subapical position (Figs. 49, 51, 59, 61), apex becoming thinner towards tip, hardly deflexed (Figs. 48, 58) or straight (Figs. 50 – 52, 60 – 62). Aedeagus of syntype of T. phrygius pisidicus from Barla-daġlari (Fig. 31) shows slight deformation very probably caused by aging of this old specimen. Examination of another male from this locality confirms that it is identical with other specimens of T. phrygius s. l. Right paramere relatively long, more or less curved medially, basal part thicker than apical one, with thickened marginal part in internal view, apical part pointed towards apex (Figs. 83 – 84, 98 – 99). Left paramere with shortened transverse apophysis, and long, oblique paramedial apophysis (Figs. 113 – 114, 127 – 128). Female genitalia. Sternum VIII very long, consisting of two chitinised plates characteristic in shape, connected by membrane, proximal part with reduced, round “ legs ” (Fig. 146). Ovipositor with equal in size valvifer and stylus, as longer axis of former ± perpendicular to longer axis of latter (Figs. 157); valvifer with long, cylindrical proximal part, plate-like distal one, and small process across at distal part; stylus consisting of one, basal stylomere of subcylindrical shape (apical stylomere disappeared). Taxonomic notes. In the course of studying the types of T. dipojranus s. str. and T. dipojranus cilicius, one paratype of phrygius as well as a numerous specimens from the type localities of phrygius s. str., phrygius pisidicus (Sultan daġlari, Kumalar daġlari, Barla daġi, Davraz daġ, Dipojraz daġlari), and additional material from Bolkar daġlari (type locality of dipojranus cilicius), we concluded that members of all above mentioned species and subspecies are conspecific with the typical phrygius since no significant morphological differencies were found. T. (Molopsis) phrygius is, just like the other species of this subgenus, a very variable species. Specimens from various populations vary in size (significant size discrepancies were discovered even in the populations of different years from Davraz dag; specimens from 2001 were obviously smaller and more slender than those collected in 1996), and in shape of pronotum and median lobe of aedeagus. Understanding this, it was not possible to establish subspecies that would have more constant features. The population from Bolkar daglari (described as diporjanus ssp. cilicius), which lives the farthest from the phrygius type-locality, is the most different from those of typical phrygius, but in fact there are no distinct features to fullfil the criteria for establishing a valid subspecies.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73011FF86FF5B780FFB9E952B.taxon	distribution	Distribution. Taurus mountain ranges about lakes Beyşehir, Eġirdir and Burdur: Ak daġ, Sultan daġlari, Kumalar daġlari, (“ Phrygic Taurus ”), Barla daġi, Davraz daġ, Dipojraz daġlari (“ Pisidic Taurus ”), SW Turkey, Bolkar daġlari (Bulghar Dagh by the ancient authors, “ Cilician Taurus ”), S Turkey.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73013FF80FF5B78D7FC6C93B4.taxon	materials_examined	Type locality. Turquie, Antalya, Col Karaovabeli, 1560 m. Taxonomic decisions. Molopsis machardi Jeanne, 2005: 383.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73013FF80FF5B78D7FC6C93B4.taxon	materials_examined	Material examined. Four specimens. Type material. One paratype 3 of Tapinopterus (Molopsis) machardi, Col. KARAOVABELI 1400 Antalya Turquie, 0 3.05.96 B. Lassalle (p) // PARATYPE (red p) / Molopsis machardi m. Jeanne det. 2002 (p) (Fig. 13) (cBL). Other material examined. 3 33, Col. KARAOVABELI 1400 Antalya Turquie, 0 3.05.96 G. Dubault & D. Echaroux (cGD, cDE).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73013FF80FF5B78D7FC6C93B4.taxon	diagnosis	Diagnosis. BL 14.5 – 16.2 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73013FF80FF5B78D7FC6C93B4.taxon	description	Male genitalia. Median lobe of aedeagus long in dorsal view, apical lamella long, straight, and thin, apical orifice deflected to left lateral position (Figs. 35); basal bulb relatively long, with concave basal orifice, apical part longer than basal one, curved copulatory sclerite in subapical position, apex becoming thinner towards tip, hardly bent downwards (Figs. 53, 63). Right paramere long, regularly curved medially, thicker basally than apically, with well-differentiated, curved subbasal part, apical part shortly acute towards apex (Figs. 85, 100). Left paramere with short transverse apophysis and long oblique paramedial apophysis (Figs. 115, 129). Female genitalia. Unknown. This species differs from all other species of Molopsis by the presence of very fine setae underneath the onychium and by the specific shape of the median lobe of aedeagus and right paramere (Figs. 35, 53, 63, 85, 100). Taxonomic notes. The single paratype studied possesses different numbers of setae underneath the onychium, respectivelly: 1 + 1 (left anterior leg), 1 + 1 (right anterior leg), 1 + 1 (left middle leg), 1 + 0 (right midlle leg), 2 + 1 (left posterior leg), 1 + 0 (right posterior leg). The setae vary in size and thickness, and generally are shorter and finer than typical setae in the most species of Tapinopterus s. l. We consider that the presence / absence of ventral setae on onychium may have different phyletic value to specify monophyletic lines in Pterostichini. This process reflects morphological regressions appeared in different, unrelated clades of the tribe as a result of an adaptation to different environments. The two states of this character occur either in species from different subgenera of the same genus (Ancholeus Dejean, 1828 and Poecilus Bonelli, 1810 of Poecilus s. l.) or among species from one and same subgenus — Phonias Gozis, 1886 (of the genus Pterostichus Bonelli, 1810; cfr. Hůrka, 1996), Molopsis Schatzmayr (this paper).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73013FF80FF5B78D7FC6C93B4.taxon	distribution	Distribution. South Taurus, Karaovabeli pass between Kaş and Elmali, W of Antalya (“ Lycian Taurus ”). PLATE 2. Habitus of Tapinopterus species (Figs. 13 – 24). Fig. 13. Tapinopterus (Molopsis) machardi (paratype); Fig. 14. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 15. T. (Molopsis) molopinus (holotype of Feronia molopina); Fig. 16. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Fig. 17. T. (Molopsis) molopinus (Turkey, Uludaġ); Fig. 18. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 19. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 20. T. (Molopsis) chaudoiri sp. nov. (paratype, Gök daġlari); Fig. 21. T. (Molopsis) wiedemanni (lectotype); Fig. 22. T. (Molopsis) wiedemanni (Turkey, Abant); Fig. 23. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari); Fig. 24. T. (Molopsis) relegatus sp. nov. (holotype). PLATE 3. Median lobe of aedeagus (Figs. 25 – 36), dorsal view (Figs. 25 – 28, 30 – 36) and ventral view (Fig. 29) of Tapinopterus species. Fig. 25. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kalamata); Fig. 26. T. (Molopsis) molopiformis (Tmolos- Gbg., Lydien, West-Kleinasien, Weirather); Fig. 27. T. (Molopsis) molopiformis (Coll Türk 1888); Figs. 28 – 29. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 30. T. (Molopsis) phrygius (syntype); Fig. 31. T. (Molopsis) phrygius (syntype of T. phrygius pisidicus); Fig. 32. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 33. T. (Molopsis) phrygius (Turkey, Beyşehir); Fig. 34. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 35. T. (Molopsis) machardi (paratype); Fig. 36. T. (Molopsis) oyukluensis sp. nov. (holotype). Scale bar: 1 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73015FF81FF5B7F73FB739460.taxon	materials_examined	Type locality. Faşikan pass, 1600 m, Konya, Turkey. Material examined. One specimen. Type material. Holotype 3, Faşikan pass, 1600 m, Konya Turquie, VII. 86 B. Lassalle (p) / HOLOTYPE Tapinopterus (sg. Molopsis) oyukluensis n. sp. Lohaj, Guéorguiev, Dubault & Lassalle, 2007 (red p) / (Fig. 14). The holotype is deposited in cBL.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73015FF81FF5B7F73FB739460.taxon	description	Description. BL 13 mm, maximum width 4.1 mm. Head, pronotum and elytra dark brown, shiny, antennae, legs and abdomen piceous brown. Head large, robust, almost as long as wide, shiny, smooth, with very fine microsculpture. Antennae relatively short, reaching hind angles of pronotum, with dense, decumbent pubescence from antennomere 4. Pronotum weakly transverse, wider than long, ratio PL / PW 0.73, with rounded lateral sides, shiny, smooth, with very fine transversal wringles along the midline. Anterior edge strongly sinuated, with prominent, at apex rounded angles. Posterior angles small, almost rectangular. Base of pronotum with single deep linear posterolateral impression on each side of midline, reaching basal third, smooth, disc without transversal wrinkles. Midline distinct, not reaching basal or anterior margins. Elytra oval, slightly flattened, with maximum width in posterior third, with distinct, impunctate striae. Third elytral interval with two setigerous pores situated in apical half of elytra, 2 nd behind the middle and 3 rd behind apical third, situated in second elytral striae. One irregular setigerous pore is also present in the middle of first left elytral stria. Umbilicate series consists of 17, respective 16 setigerous punctures, in the first fourth interrupted, with distribution: left side 6 + 1 + 10, right side 6 + 10. Male genitalia. Median lobe of aedeagus in dorsal view relatively short, apical lamella wide, directly curved to left (Figs. 36); apical orifice deflected to left lateral position, ventral margin forming right angle at medial part, apex regularly deflexed (Figs. 54, 64). Right paramere with basal part thicker than apical part, in middle constricted and curved, apex pointed towards tip (Fig. 86, 101). Left paramere subelongate, transverse apophysis reduced, paramedial apophysis, oblique, well-developed (Figs. 116, 130). Female genitalia. Unknown.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73015FF81FF5B7F73FB739460.taxon	etymology	Etymology. Topotypic, referring to the type locality, the mountain range Oyuklu daġi, northern part of Taurus Mountains, ca 100 km south of Konya.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73015FF81FF5B7F73FB739460.taxon	diagnosis	Diagnosis. T. oyukluensis sp. nov. is closely related to T. phrygius, a species widely distributed in various mountain ranges of the Taurus (see also Check list). This species differs from all other species of Molopsis by the shape of the median lobe of aedeagus and left paramere; median lobe in dorsal view (Fig. 36) with apical lamella wide, directly curved to left (this state occurs also in T. phrygius, but here the lamella is thinner and either regularly curved to left (see Figs. 32 – 34) or slightly sinuous before apex (see Figs. 30 – 31); left paramere subelongate, with reduced transverse apophysis (Figs. 116, 130).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73015FF81FF5B7F73FB739460.taxon	distribution	Distribution. Type specimen was collected in Faşikan geçidi (pass) between the villages of Taşkent and Sariveliler, ca 30 km southwest of ridge of mountain range Oyuklu daġi (Central Turkey, villayet Konya).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73014FF9EFF5B793CFD4A9153.taxon	materials_examined	Type locality. Constantinople. Taxonomic decisions. Feronia (Haptoderus) molopina Chaudoir, 1868: 245.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73014FF9EFF5B793CFD4A9153.taxon	materials_examined	Material examined. 97 specimens. Type material. Holotype 3 of Feronia molopina, designated by monotypy, well preserved, “ molopinus Chaud. Constantinople Sturm. ” (h label pinned on the bottom of box) / HOLOTYPE Feronia (Haptoderus) molopina Chaudoir 1868 G. Dubault désign. 2003 (red p) / Tapinopterus (Molopsis) molopinus Chaudoir G. Dubault détermin. (p) (Fig. 15) (MNHN, “ Collection Chaudoir ”, box no 216 “ Argutor ”). Holotype 3 of Tapinopterus (Molopsis) dipojranus brussanus, “ Brussa Anatolien G. Frey IV. 32. ” (p) / “ Tapinopterus molopinus Chd. ” (h) / “ Holotypus T. dipojran. s. brussan. (red p / h) / “ Tapinopterus dipojranus s. brussanus (h), det. S. L. Straneo 1985 Holotypus ” (p / h) / TAPINOPTERUS (sg. MOLOPSIS) molopinus (Chaudoir, 1868) R. LOHAJ det. 2003 (p) (Fig. 16) (MCSNM). Other material examined. Uludag: 1 3, Turkei 1848 (NMP); 13, Asia minor (HNHM); 1 3, BALKAN FRIV. (HNHM); 3 33, Olympe (ZISP); 2 33, Bithynischen Olymp, leg. Merkl, coll. Leonhard (DEI); 1 Ƥ, Olymp, coll. O. Leonhard (DEI); 1 3, Olympos Bithyn. Asia min. Dr. Jureček 1931 (NMP); 4 33, BRUSSA, 1870, leg. Pável (HNHM); 5 33 ƤƤ, “ Uludag Bursa 1.500 West. Turkey J. y S. Klapperich. ” (MNHN); 6 33 ƤƤ, “ Ulu Dagh Brousse 1400 - VI 66 ” (MNHN); 6 33 ƤƤ, “ Ulu Dagh Turquie 1700 m VI 68 ” (MNHN); 2 33, Merkl. 92, Türkei (NMNHS); 1 3, Turcia Merkl 98. (MCSNT); 1 3, “ rebellis Turcia ” (MNHN); 1 3, “ Museum Paris 1906 Coll Léon Fairmaire ” / “ Pter. olympicus Turquie ” (MNHN); 1 3, “ olympicus Kr. Turkei Merkl. ” (MNHN); 1 Ƥ, Türkei, Ulu dagh 2000 m, leg. Kenyery, VI. 1963 (NMP); 3 33 1 Ƥ, Anatolia occ, Uludagh bei Bursa (gipfelzone) 1500 m, 28. V. 1966, Blumental leg. (NMP); 8 33 4 ƤƤ, Turchia, Bursa Uludag, 29.5.1974, lg. Sama, Coll. Gudenzi (cMP); 3 33, Anatol. Bursa Uludag 25.5.82 Drioli (MCSNT); 1 3 1 Ƥ, Turkei, Uludag, 13. IV. 1983, H. + L. Freude (cMP); 8 33 7 ƤƤ, Uludag 1500, Bursa Turquie, VII 87 G. Dubault & D. Echaroux lgt. (cGD, cDE); 1 Ƥ, Uludaġ 1500 m, (Bursa) 15. V. 89, leg. R. Pittino (cWH); 2 ƤƤ, Uludag 1500 Bursa VI 90 ” / “ Lassale ” (cBL); 7 33 3 ƤƤ, TR (Bursa) Uludaġ b. Soġurpinar, 1200 – 1400 m, 26. IV. 1992 Heinz leg. (cRL); 4 33, Tr. occ., Bursa env. Uludag, 27.5.1994, Skoupý leg. (cVS); 1 3, “ NW Turkey — Bursa Uludag pineto — abietum, 1500 m 3.10.97, D. Čatlos lgt. (cRL) (Fig. 17); 1 3, Tr. occ, Bursa Uludag 1650 m, 15.5.2000, Skoupý leg. (cVS); 1 3, Tr. occ, Bursa Uludag 2100 m, 15.5.2000, Skoupý leg. (cVS); 1 Ƥ, Turkey, Cumalikizik, Bursa, Ulu daġ, 24.5.2004, 5 – 900 m, T. Tichý (cEH).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73014FF9EFF5B793CFD4A9153.taxon	diagnosis	Diagnosis. BL 10.8 – 15 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73014FF9EFF5B793CFD4A9153.taxon	description	Male genitalia. Median lobe of aedeagus long in dorsal view, with long, thin, almost straight apical lamella, apical orifice deflected to left lateral position (Figs. 37 – 38); basal bulb relatively long, with concave basal orifice, apical part longer, copulatory sclerite curved, situated in medial part, apex slighlty bent downwards (Figs. 65 – 67, 70 – 72). Right paramere relatively long, more or less strongly curved and constricted medially, with thicker basal and thinner apical parts (Figs. 87 – 88, 102 – 103). Left paramere with short, more or less reduced transverse apophysis and fairly long, oblique paramedial apophysis (Figs. 117 – 118, 131 – 132). Female genitalia. Tergum VIII with convex distal margin and short, thin proximal “ legs ” (Fig. 140). Sternum VIII consisting of two chitinised parts connected by membrane, without “ legs ” on proximal margin (Fig. 147). Syntergum IX + X long, narrow, bearing pair of small, articulated, distant each other ovipositors (Fig. 152). Ovipositor with large valvifer and smaller stylus, as longer axis of former ± perpendicular to longer axis of latter (Figs. 158); valvifer with short cylindrical part, more acute internal margin, less acute external margin, and with developed, slightly curved externally, process at distal part; stylus consist of one, basal, elongate stylomere (apical stylomere disappeared), having incised outer margin. Spermatheca with seminal canal and receptaculum discrete; seminal canal long; receptaculum shorter than seminal canal, slightly curved apically (Fig. 164). This species differs from its closest relatives, T. chaudoiri sp. nov. and T. relegatus sp. nov. in the structure of the male and female genitalia. T. molopinus has a smaller aedeagus, with relatively shorter and thinner apical lamella, about 1 / 3 entire length of lobe (in dorsal view). T. chaudoiri sp. nov. has a larger aedeagus, with apical lamella about 2 / 5 that of median lobe (compare Figs. 37 – 39). In lateral view, the aedeagus of T. molopinus is smaller, with narrower medial part and shortened apical lamella (Figs. 65 – 74). The transverse apophysis on left paramere in T. molopinus is more or less reduced while in T. chaudoiri possesses a longer, well-developed transverse apophysis (Figs. 117 – 120, 131 – 134). The females of this species are easily distinguished from the females of T. chaudoiri in the absence of incision at the apex of elytra and apex of sternum VII (Figs. 168 – 171). Besides, sternum VII has finer posterior border. For the distintions between T. molopinus and T. relegatus, see paragraph “ Diagnosis ” under the latter species. Taxonomic notes. The first mention of this taxon has been made by Sturm (1843: 25), but the species was described by Chaudoir in 1868 as Feronia (Haptoderus) molopina based on single male specimen. The last author wrote “ C’est le Steropus brevis du catalogue de Sturm, indiqué comme venant de Constantinople, et que j’ai acheté à la vente de sa collection. ”. In view of the relativeness of the geographical designation in the old-days, it should be noted that the type locality of T. molopinus is doubtful. We are unaware of any subsequent record or material of Molopsis in collections from Constantinople (= Istanbul). Regarding the present bionomy of the species, we suppose that the type material was collected in the area of the Uludag Mt.; the same has already been supposed (Apflelbeck, 1904: 244, note 1). PLATE 5. Median lobe of aedeagus (Figs. 45 – 54) in left lateral view of Tapinopterus species. Fig. 45. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kalamata); Fig. 46. T. (Molopsis) molopiformis (Tmolos-Gbg., Lydien, West-Kleinasien, Weirather); Fig. 47. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 48. T. (Molopsis) phrygius (syntype); Fig. 49. T. (Molopsis) phrygius (syntype of T. phrygius pisidicus); Fig. 50. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 51. T. (Molopsis) phrygius (Turkey, Beyşehir); Fig. 52. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 53. T. (Molopsis) machardi (paratype); Fig. 54. T. (Molopsis) oyukluensis sp. nov. (holotype). Scale bar: 1 mm. The holotype of Feronia molopina was found in MNHN. Meantime, two of us (BG, RL) tried to find the holotype of Pterostichus olympicus in DEI and MNHUB. According to Dr. Lothar Zerche (personal communication), no type of this species exists in DEI. The search of BG in NMHUB led nowhere too. The material of Tapinopterus from DEI, very probably including also the type of Pterostichus olympicus, was many years ago loaned by Dr. Jozef Mařan (Praha, Czech Republic) and maybe have never been returned (Dr. L. Zerche, personal communication). According to the information provided by Dr. Josef Jelinek, the former curator of Coleoptera in NMP, the Tapinopterus loan was in possession of Dr. J. Mařan at the time of his death. The type material was, however, not located after his demise. Kraatz (1875) did not indicate clearly the exact massif from which the type series of Pterostichus olympicus comes, e. g. the Olymp Mt. in Central Greece or the Uludag Mt. (Bythinic Olymp near Bursa by the ancient authors) in Northwestern Anatolia. The title of the Kraatz paper has points that the author dealt with species from Europe (“ Drei neue europäische Pterostichus Bon. “) and author noted: „ Mein Ex. stammt aus der Schaum´schen Sammlung und wurde wahrscheinlich von Dr. Krüper aufgefunden “. Dr. Theobald Johaness Krüper (1829 – 1921) a German ornithologist and entomologist and long-time curator and director of University Museum in Athens. He collected in Greece and also often in Turkey. Authors (RL, BG) could not find the type sample in the studied collections which may have successfully associated with the specimen / s from the type series. For this reason, until the type material is located and examined, we prefer to treat this taxon as synonym of T. molopinus.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73014FF9EFF5B793CFD4A9153.taxon	distribution	Distribution. This species inhabits the massif of Uludaġ near Bursa, NW Anatolia. PLATE 6. Median lobe of aedeagus Figs. 55 – 64) in right lateral view of Tapinopterus species. Fig. 55. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kalamata); Fig. 56. T. (Molopsis) molopiformis (Tmolos-Gbg., Lydien, West-Kleinasien, Weirather); Fig. 57. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 58. T. (Molopsis) phrygius (syntype); Fig. 59. T. (Molopsis) phrygius (syntype of T. phrygius pisidicus); Fig. 60. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 61. T. (Molopsis) phrygius (Turkey, Beyşehir); Fig. 62. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 63. T. (Molopsis) machardi (paratype); Fig. 64. T. (Molopsis) oyukluensis sp. nov. (holotype). Scale bar: 1 mm. PLATE 7. Median lobe of aedeagus (Figs. 65 – 79), left lateral view (Figs. 65 – 69, 75) and right lateral view (70 – 74, 76 – 79) of Tapinopterus species. Figs. 65, 70. T. (Molopsis) molopinus (holotype of Feronia molopina); Figs. 66, 71. T. (Molopsis) molopinus (“ olympicus ”, HNHM); Figs. 67, 72. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Figs. 68, 73. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Figs. 69, 74. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Figs. 75 – 76. T. (Molopsis) wiedemanni (lectotype of Feronia wiedemanni); Fig. 77. T. (Molopsis) wiedemanni (Turkey, Abant Lake); Fig. 78. T. (Molopsis) wiedemanni (Turkey, Aladaġlari); Fig. 79. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari). Scale bars: 1 mm (Figs. 65 – 74); 0.5 mm (Figs. 75 – 79).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300BFF9BFF5B7C1FFA0692AC.taxon	materials_examined	Type locality. Forest between Domanic and Tahtaköprü, 1000 – 1500 m, NW Turkey. Material examined. 65 specimens. Type material. Holotype 3, TR Wld zw. Domanic u. Tahtaköprü, IV 1973, 1000 – 1500 m, Heinz leg. (p) / HOLOTYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) / (cWH). Paratypes: 24 33 12 ƤƤ, TR Wld zw. Domanic u. Tahtaköprü, IV 1973, 1000 – 1500 m, Heinz leg. (Fig. 18) (p) / PARATYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) /, (MCSNM, NMNHS, cBL, cGD, cRL, cWH), 9 33 5 ƤƤ, Turkey, Domanic — Bursa, 0 3.05.98, G. Dubault & D. Echaroux lgt. (p) / PARATYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) / (cDE, cGD), 3 33, Oylat 1200 Bursa Turquie, 0 7.05.98 G. Dubault (Fig. 19) (p) / PARATYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) (cBL, cDE, cGD), 2 33 8 ƤƤ, Anatolia occ. Heinz leg. (p) / Gök-daġ: Keltepe, ca. 1100 – 1400 m, 14. IV. 1982 (h) / PARATYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) (Fig. 20) (MCSNM, NMNHS, cRL, cWH), 1 3, Cangaz, Anat 1962 (h) / Tapinopterus olympicus Kr. (h) det. Ing. Jedlička (p) / PARATYPE Tapinopterus (Molopsis) chaudoiri n. sp., Lohaj, Guéorguiev, Dubault & Lassalle des. 2007 (red p) (NMP).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300BFF9BFF5B7C1FFA0692AC.taxon	description	Description. BL 11.3 – 15 mm (Holotype 12.6 mm), maximum width up to 5.5 mm. Head, pronotum and elytra black, shiny, some females matt, antennae, legs and abdomen in some specimens paler, piceous brown. Head large, robust, almost as long as wide, shiny, smooth, with very fine microsculpture. Antennae relatively short, reaching hind angles of pronotum. Pronotum weakly transverse, slightly wider than long, ratio PL / PW 0.65 – 0.76 (HT 0.67), sides rounded, shiny, smooth, with very fine transversal wrinkles along the midline. Anterior margin concave, anterior angles prominent, rounded at apex .. Posterior angles strongly prominent, sharp, acute-angled. Base of pronotum with single deep linear posterolateral impression on each side of midline, reaching basal third, smooth. Midline distinct, not reaching basal or anterior margins. Elytra elongate oval, with maximum width in apical third, slightly convex, with distinct, finely punctate striae. Apex of elytra with deep and symmetric incision, which is wide in females, and very narrow, but visible in males (Fig. 169). Third elytral interval with two setigerous punctures situated at apical half of elytra, 2 nd behind middle and 3 rd in apical fifth-sixth of elytral length. Umbilicate series consists of 14 – 17 setigerous punctures at each side, in the middle widely interrupted. Distal margin of last visible abdominal sternum (sternum VII) with thick posterior border, surface smooth, not punctate, with two pairs of SP; apex of sternum VII in females with very deep circular incision at tip (Fig. 171). Male genitalia. Median lobe of aedeagus in dorsal view long, with long, wide, almost straight apical lamella, apical orifice deflected to left lateral position (Figs. 39); basal bulb short, with basal orifice slightly concave, apical part longer, copulatory sclerite curved, situated in medial part, apex more or less bent downwards (Figs. 68 – 69, 73 – 74). Right paramere relatively long, more or less strongly curved and constricted medially, with thicker basal part and thinner apical one (Figs. 89 – 90, 104 – 105). Left paramere with long transverse apophysis and fairly long, oblique paramedial apophysis (Figs. 119 – 120, 133 – 134). Female genitalia. Tergum VIII with regularly convex distal margin and very short proximal “ legs ” (Fig. 141). Sternum VIII consists of two round, chitinised parts connected with membrane, without “ legs ” on proximal margin (Fig. 148). Ovipositor with valvifer and stylus, as longer axis of former more or less perpendicular to longer axis of latter (Figs. 159); valvifer with short cylindrical part, internal margin strongly one-humped, external margin regularly convex, and with clear process across at distal part; stylus consist of one, basal, elongate stylomere (apical stylomere disappeared), with slightly incised internal margin.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300BFF9BFF5B7C1FFA0692AC.taxon	etymology	Etymology. Patronymic, dedicated to baron Maximilien Stanislavovitch de Chaudoir (1816 – 1881), famous Russian entomologist and outstanding specialist on the carabid beetles.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300BFF9BFF5B7C1FFA0692AC.taxon	diagnosis	Diagnosis. This species is distinct from its closest relatives, T. molopinus and T. relegatus sp. nov. in the structure of the male and female genitalia. T. chaudoiri sp. nov. has a larger aedeagus, with longer and wider apical lamella, with length about 2 / 5 of entire length of lobe (in dorsal view); T. molopinus has smaller aedeagus, with relatively shorter and thinner apical lamella only 1 / 3 of length of median lobe (compare Figs. 37 – 39). In lateral view, the aedeagus of the new species is larger, with wider medial part and prolonged apical lamella (Figs. 65 – 74). The transverse apophysis on left paramere in T. chaudoiri sp. nov. well-developed while in T. molopinus it is more or less reduced (Figs. 117 – 120, 131 – 134). T. chaudoiri sp. nov. is easily distinguished from the closely distributed T. molopinus from the Uludaġ Mt. by the presence of deep, symmetric incision on the apex of elytra (in both sexes) and by the presence of deep circular incision on the apex of sternum VII in females, which is missing in T. molopinus (Figs. 168 – 171). Besides, the sternum VII in T. chaudoiri sp. nov. has thicker posterior border. T. chaudoiri sp. nov. differs also morphologically from T. relegatus sp. nov. in the different shape of tergum VIII (Figs. 1410142), sternum VIII (Figs. 148 – 149), and valvifer and stylus of ovipositor (Figs. 159 – 160). PLATE 8. Right paramere (internal face) (Figs. 80 – 94) of Tapinopterus species. Fig. 80. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kumani); Fig. 81. T. (Molopsis) molopiformis (Coll Türk 1888); Fig. 82. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 83. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 84. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 85. T. (Molopsis) machardi (paratype); Fig. 86. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 87. T. (Molopsis) molopinus (holotype of Feronia molopina); Fig. 88. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Fig. 89. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 90. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 91. T. (Molopsis) wiedemanni (lectotype of Feronia wiedemanni); Fig. 92. T. (Molopsis) wiedemanni (Turkey, Abant Lake); Fig. 93. T. (Molopsis) wiedemanni (Turkey, Aladaġlari); Fig. 94. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari). Scale bar: 0.5 mm. PLATE 9. Right paramere (external face) (Figs. 95 – 109) of Tapinopterus species. Fig. 95. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kumani); Fig. 96. T. (Molopsis) molopiformis (Coll Türk 1888); Fig. 97. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 98. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 99. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 100. T. (Molopsis) machardi (paratype); Fig. 101. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 102. T. (Molopsis) molopinus (holotype of Feronia molopina); Fig. 103. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Fig. 104. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 105. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 106. T. (Molopsis) wiedemanni (lectotype of Feronia wiedemanni); Fig. 107. T. (Molopsis) wiedemanni (Turkey, Abant Lake); Fig. 108. T. (Molopsis) wiedemanni (Turkey, Aladaġlari); Fig. 109. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari). Scale bar: 0.5 mm. PLATE 10. Left paramere (internal face) (Figs. 110 – 123) of Tapinopterus species. Fig. 110. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kumani); Fig. 111. T. (Molopsis) molopiformis (Coll Türk 1888); Fig. 112. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 113. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 114. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 115. T. (Molopsis) machardi (paratype); Fig. 116. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 117. T. (Molopsis) molopinus (holotype of Feronia molopina); Fig. 118. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Fig. 119. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 120. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 121. T. (Molopsis) wiedemanni (lectotype of Feronia wiedemanni); Fig. 122. T. (Molopsis) wiedemanni (Turkey, Abant Lake); Fig. 123. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari). Scale bar: 0.5 mm. The several unique morphological differences between the populations from the massifs of Gök Daglari and Uludaġ as well as the lack of any records from the intermediate area between them suppose presence of two different and apparently isolated (allopatric) species as the different aedeagus manifests the isolation.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300BFF9BFF5B7C1FFA0692AC.taxon	distribution	Distribution. This species inhabits southern and eastern slopes of Uludaġ mountain range (pass between Domanic and Tahtaköprü, surrounding of Oylat) and mountain of Gök Daglari, situated south of the Sapanca Lake, (villayet Sakarya), NW Anatolia. Authors tried to designate the position of the locality “ Cangaz ”, by which is one male paratype specimen labelled, but without success. PLATE 11. Left paramere (external face) (Figs. 124 – 137) of Tapinopterus species. Fig. 124. Tapinopterus (Pseudomolopsis) rebellis (Greece, Kumani); Fig. 125. T. (Molopsis) molopiformis (Coll Türk 1888); Fig. 126. T. (Molopsis) aenigmaticus sp. nov. (holotype); Fig. 127. T. (Molopsis) phrygius (holotype of T. dipojranus); Fig. 128. T. (Molopsis) phrygius (paratype of Tapinopterus dipojranus cilicius); Fig. 129. T. (Molopsis) machardi (paratype); Fig. 130. T. (Molopsis) oyukluensis sp. nov. (holotype); Fig. 131. T. (Molopsis) molopinus (holotype of Feronia molopina); Fig. 132. T. (Molopsis) molopinus (holotype of Tapinopterus dipojranus brussanus); Fig. 133. T. (Molopsis) chaudoiri sp. nov. (paratype, Domanic); Fig. 134. T. (Molopsis) chaudoiri sp. nov. (paratype, Oylat); Fig. 135. T. (Molopsis) wiedemanni (lectotype of Feronia wiedemanni); Fig. 136. T. (Molopsis) wiedemanni (Turkey, Abant Lake); Fig. 137. T. (Molopsis) wiedemanni (Turkey, Sundiren daġlari). Scale bar: 0.5 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300EFF95FF5B7C40FA079475.taxon	materials_examined	Type locality. Dorukhan Pass near Zonguldak, NE Turkey. Material examined. One specimen. Type material. Holotype Ƥ, relatively well preserved, only apical borders of elytra bitten, Turkey, col Dorukhan, Zonguldak, VII. 1990 (p) / Holotype Tapinopterus (Molopsis) relegatus sp. n. Lohaj, Guéorguiev, Dubault & Lassalle, 2007 ” (red p) (Fig. 24) (cBL).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300EFF95FF5B7C40FA079475.taxon	description	Description. BL 12.2 mm; maximum width 4 mm. Body smooth and glabrous (excluding antennae), head and pronotum without microsculpture, elytra with distinct fine isodiametric microsculpture. Head, pronotum and elytra black, shiny, antennae, palpi, legs and body ventrally piceous brown. Head large, robust, subconvex, 1.33 times longer than wide; disc smooth. Mandibles stout and protruding. Antennae relatively short, not reaching hind angles of pronotum, with dense, decumbent pubescence from antennomere 4, antennomere 2 nearly two times shorter than scapus. Length of eyes almost equal to length of scapus; temporae not protruding, shorter than length of eye. Frontal furrows short, very smooth. Mentum with short bifid tooth. Pronotum 1.48 times wider as long and 1.53 times wider than head, subcordate and subconvex, smooth. Lateral sides more convergent posteriorly than anteriorly; ratio anterior margin / basal margin 1.09. Anterior margin moderately sinuate, with rounded and slightly prominent angles. Posterior margin almost straight; angles sharply pointed, prominent outside. Basal surface with single impunctate basal fovea on each side of midline, fovea with single linear posterolateral impression slightly divergent anteriorly and not reaching basal margin. Midline distinct and deeper in basal third, not reaching anterior, and hardly reaching basal margin. Elytra suboval, coalesced along suture, 1.32 times longer than wide and 1.41 times wider than pronotum; disc convex. Shoulders obtusely rounded. Scutellar stria absent; basal border not reaching scutellum; striae fine, barely punctate. Intervals slightly convex; interval 3 of left elytron with single setigerous puncture at apical third, same one of left elytron without puncture. Umbilicate series of each elytron consist of 10 setigerous punctures (left elytron 5 + 0 + 5, right elytron 5 + 0 + 5), in the middle largely interrupted. Prosternum, mesosternum, metasternum and abdominal sterna smooth, glabrous, impunctate; anterior margin of metaepisterna rather longer than internal margin; abdominal sterna II – VI with two setigerous punctures, sternum VII with four marginal setigerous punctures. Legs moderately long, femora stout; protibia with two clip setae; mesotibial ctenidium well differentiated; hind coxa with medial setigerous puncture below meeting point of coxae; both median and hind trochanter with setigerous punctures; tarsomere 5 without ventral setae. Female genitalia. Tergum VIII with convex, slighlty angled at top, distal margin and very short proximal “ legs ” (Fig. 142). Sternum VIII consists of two oblong chitinised parts connected with membrane, without “ legs ” on proximal margin (Fig. 149). Ovipositor very small, with relativelly bigger valvifer and smaller stylus, as longer axis of former more or less perpendicular to longer axis of latter (Figs. 160); valvifer having two-humped internal margin, smoother external margin, and well-developed process across; stylus consisting of one basal degenerate stylomere (apical stylomere disappeared). Spermatheca with seminal canal and receptaculum discrete, seminal canal longer, receptaculum shorter than seminal canal, strongly curved apically (Fig. 165). Male genitalia. Unknown.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300EFF95FF5B7C40FA079475.taxon	etymology	Etymology. The specific epithet derives from the Latin adjective relegatus, meaning “ exile ” or “ ejected ” (in English), reflecting its isolation from the ranges its known to be inhabited by its congeners.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300EFF95FF5B7C40FA079475.taxon	diagnosis	Diagnosis. This species is distinct from its congeners by: (i) the reduction of setigerous punctures in interval 3 of elytra, i. e presence of single puncture on left elytron, and none on rigth elytron; (ii) umbilicate series on each elytron consisting of 10 setigerous punctures; (iii) peculiarities in the structure of the female genital armature (see above paragraph “ Female genitalia ”). The new species possess receptaculum strongly curved apically that differs quite clearly from the receptaculum in T. molopinus, which is only slightly curved apically. In the structure of the female genital components, i. e. tergum VIII, sternum VIII, stylus of ovipositor, and spermatheca (Figs. 140 – 142, 147 – 149, 158 – 160, 164 – 165), T. relegatus sp. nov. looks like T. molopinus and T. chaudoiri sp. nov. Most probably, three aforementioned species of Molopsis form a monophyletic complex. It worth noting that the species inhabits an area situated very closely to the range of T. wiedemanni, but beyond the ranges of T. molopinus and T. chaudoiri sp. nov. On the other hand, T. relegatus sp. nov. and T. wiedemanni seem to be not closely related phyletically. There are clear distinctions between the last two species, as well as between T. wiedemanni and the pair T. molopinus - T. chaudoiri sp. nov., in the structure of the female genitalia (see Figs. 140 – 143, 147 – 150, 158 – 162, 164 – 166). From a zoogeographic perspective, we may postulate the following two hypotheses. In the past the lineage “ T. molopinus - T. chaudoiri sp. nov. - T. relegatus sp. nov. ” had a wider range in NW Anatolia and occurred east of the Sakarya River. After that, probably as a result of climatic changes, the spreading of this group was reduced as the central and most of the eastern parts of former range were occupied by T. wiedemanni. Today, two taxa of the complex live in the western part of the former range, and a third taxon survived in a restricted area in the east. Or, alternatively, a single migration of the same lineage, started from the area of west of the Sakarya River, reached the region of Dorukhan Pass. The migration took place along the north parts of the Bolu Range. These assumptions should be tested by phylogenetic and molecular analyses.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E7300EFF95FF5B7C40FA079475.taxon	distribution	Distribution. The type specimen was found in the Dorukhan geçidi (pass), between Mengen and Devrek. The point lies ca 20 km N of Mengen (NW Turkey, villayet Zonguldak). PLATE 12. Terga VIII (dorsal view) (Figs. 138 – 143) of Tapinopterus species. Fig. 138. Tapinopterus (Pseudomolopsis) rebellis (Greece, Taygetos, Anogia); Fig. 139. T. (Molopsis) molopiformis (Tmolos-Gbg., Lydien, West-Kleinasien, Weirather); Fig. 140. T. (Molopsis) molopinus (Turkey, Uludaġ); Fig. 141. T. (Molopsis) chaudoiri sp. nov. (paratype, Gök-daġ); Fig. 142. T. (Molopsis) relegatus sp. nov. (holotype); Fig. 143. T. (Molopsis) wiedemanni (Turkey, Bolu, Kartalkaya). Scale bar: 0.5 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73000FF96FF5B792FFDC993EB.taxon	materials_examined	Type locality. Nord de l´Anatolie. Taxonomic decision. Tapinopterus (Molopsis) wiedemanni (Chaudoir, 1850), combinatio novum of Feronia (Molops) wiedemanni Chaudoir, 1850: 145 (nec Fairmaire, 1866: 252).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73000FF96FF5B792FFDC993EB.taxon	materials_examined	Material examined. 103 specimens. Type material. Two syntypes (3 Ƥ) of Feronia wiedemanni (lectotype male and paralectotype female by present designation), well preserved, “ Wiedemanni Chaud. Anatolie C. Falderm. ” (h label pinned on the bottom of box) / LECTOTYPE and PARALECTOTYPE Feronia (Molops) wiedemanni Chaudoir D. Dubault désign. 2003 (red p) / Tapinopterus (Molopsis) wiedemanni G. Dubault détermin. (p) (Fig. 21) (MNHN, “ Collection Chaudoir ”, box no 216 “ Argutor ”). Other material examined. Abant: 29 33 ƤƤ, “ Aband 1.300 m. Turkey J. y S. Klapperich. ” (MNHN); 1 3, TR. bor. occ. ABANT lake, 5.6.1992, V. Skoupý lgt. (cVS); 1 3 1 Ƥ, TURKEY occ. Abant Gölü env. (35 km SW of BOLU), 1993 - 06 - 21, Kuboň lgt. (cGD); 2 33, Türkie bor., Abant, 9. – 10.5.1994, A. Trmal lgt. (cRL); 1 3 3 ƤƤ, TURKEY (Bolu), Elmacik dagi, Golu lake, 10. V. 1994, Major (cDW); 3 33 1 Ƥ, N TURKEY, Abant env., 5.1995, F. Moravec lgt. (cRL) (Fig. 22); 1 3, TR-prov. Bolu, Abant gölü, 1200 m, 4.6.1996, Z. Malinka lgt. (cVS); 1 3 1 Ƥ, TURKEY, Abant Golu, 30 km W Bolu, 4.5.1997, leg J. Bašta (cDW); 1 3, TURKEY, Abant Golu, W Bolu, 8.6.1998, lgt. E. & P. Hajdaj (cDW); 1 Ƥ, TR vill. Bolu, Abant Gölü, 23. – 24.5.2000, Josef Mertlik lgt. (cVS). Aladaglari: 15 33 8 ƤƤ, Anatolia bor., Aladaġ ca 20 km südl. Bolu, 1400 – 1600 m, 27. IV. 1981, Heinz leg. (MCSNM, cWH, cBL, cRL). Köroġlu daġlari: 1 Ƥ, Anatolia bor, Tasliyayla, ca 21 km ne Seben (Bolu), 1500 – 1600 m, 27. IV. 1984, W. Heinz lgt. (cWH); 13 33 9 ƤƤ, Kartalkaya, 1800 Bolu Turquie, 19.05.94, G. Dubault (cGD, cRL, cBL). Sündiren daġlari: 7 33 1 Ƥ, Anatolia centr., Wald n. Mihaliççik, 1500 – 1600 m, Heinz leg. (cBL, cGD) (Fig. 23).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73000FF96FF5B792FFDC993EB.taxon	diagnosis	Diagnosis. BL 11.3 – 15.8 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73000FF96FF5B792FFDC993EB.taxon	description	Male genitalia. Median lobe of aedeagus in dorsal view long, with long, thin, and slightly curved to the left apical lamella, which formed more or less distinct transverse disc in lateral position (Figs. 40 – 44, 75 – 79); apical orifice deflected to left lateral position; basal bulb shorter than apical, with basal orifice concave, apical part long, copulatory sclerite curved, with widened medial part, situated in paramedial position, apex of lobe bent downwards, but reflexed before end. Right paramere relatively short, strongly curved and slightly constricted medially, with thinner apical and thicker basal part (Figs. 91 – 94, 106 – 109). Left paramere with long transverse apophysis and long, oblique paramedial apophysis (Figs. 121 – 123, 135 – 137). Female genitalia. Tergum VIII relatively long, with convex, subpyramidal distal margin, straight margin and short “ legs ” in proximal position (Fig. 143). Sternum VIII unified, in proximal position with two closed “ holes ” and without “ legs ” (Fig. 150). Syntergum IX + X long, narrow, bearing pair of small, articulated, distant each other ovipositors (Fig. 153). Ovipositor with valvifer and stylus (Figs. 161 – 162); valvifer with twin-peaked internal margin, convex external margin, and process across at distal part; stylus consist of one, basal, subelongate stylomere (apical stylomere disappeared), having strongly incised internal margin. Spermatheca with seminal canal and receptaculum discrete; receptaculum shorter than seminal canal, slightly curved apically (Fig. 166); spermathecal gland elongate, spermathecal canal short, inserted at junction of receptaculum and seminal canal. Taxonomic notes. This species was described from three males and one female as Feronia (Molops) wiedemanni from the north of Anatolia (Chaudoir, 1850: 145 – 146). Later, Gemminger & Harold (1868: 333) placed it under Molops, but Chaudoir (1876: 348) specified that it did not belongs to that genus. Kraatz (1875: 419) is first to indicate that F. molopina and F. wiedemanni were close relatives. Marseul (1882) put F. wiedemanni Chaudoir among the Molops species. For the lack of basolateral setigerous punctures and presence of single basal impression of pronotum, Ganglbauer (1889: 52) supposed that this taxon belonged to Tapinopterus. It seems that since the work of Jakobson (1907) this taxon disappeared from the literature. It appears again only recently, when F. wiedemanni Chaudoir, 1850 was considered either as a representative of Molops (Mlynář, 1977) or was placed among the species “ incertae sedis ” of the same genus (Bousquet, 2003: 475). In fact, no real explanation about the systematic position of this species has been taken after the above note of Chaudoir (1876). After prolonged search for the Chaudoirś types in MNHN by one of us (GD), two syntypes of Feronia wiedemanni were eventually found. Their study revealed that both belong to a valid species of Tapinopterus, and not to Molops.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73000FF96FF5B792FFDC993EB.taxon	distribution	Distribution. Mountain ranges around Bolu: Abant, surrounding of the Abant Lake, Aladaġlari, Köroġlu daġlari, Sündiren daġlari, NW Turkey.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73003FF97FF5B7E8FFE1493D9.taxon	materials_examined	Material examined. 24 specimens. Type material. Holotype 3 of Pterostichus rebellis v. Kumanensis, Morea Cumani Brenske (p) / rebellis, v. Kumanensis m. (h) / coll. Reitter (p) / Holotypus (p) 1884 Pterostichus rebellis v. Kumanensis Reitter (h) (red label) / (Fig. 2) (HNHM). Other material examined. Peloponnesos: 3 33 1 Ƥ, GREECE (Peloponissos) (Arkadia) Taygetos Geb., Langadapaβ 1000 m, bei Abzweigung Kokkinilakka (Gesiebe an Platanen in schmalem Bachtal), 28. IV. 1999 Ingo Wolf (Fig. 1) (cDW); 3 33 2 Ƥ, Mt. TAYGETE 1300, Péloponnése, Gréce V. 94, G. Dubault, B. Lassalle (cGD, cBL); 1 3, Mt. TAYGETE 1300, Péloponnése, Gréce 17.05.93, G. Dubault (cGD); 1 3, “ Rte KALAMATA-Sparte km 28 1000, Péloponnése, Gréce, 20.10.99, G. Dubault ” (cGD); 1 Ƥ, GRECE Péloponnése, Taygéte Forét, Au dessus d´Anogia, 7. V. 2001, G. Ledoux (cGD); 1 3, TAYGETE, Péloponnése Gréce, X. 00 G. Dubault (cGD); 4 33 1 Ƥ, KOSMAS, Péloponnése Gréce, X. 00 G. Dubault, B. Lassalle (cGD, cBL); 1 Ƥ, Kalamata GR, Oxihori, Taygétes, 1300 m, 26 / IV / 2000, Péloponnése, Gréce, B. Lassalle (cBL); 1 3, Morea Cumani Brenske (HNHM); 2 33, Grece (ZISP); 2 33, N 37 ° 47´EO 21 ° 44´, Griechenland Pelop Umg. Koumani 630 m Lompe 1.4.1997 (cDW). Taxonomic notes. In the course of this work, we came to the conclusion that Tapinopterus rebellis s. l. is not closely related to the known species of Molopsis, which form a distinct monophyletic complex. Diagnostic features that distinguish these groups of Tapinopterus are: 1 / general shape of the pronotum; 2 / length of the antennae; 3 / puncturation of the abdominal sterna; 4 / setation of the hind coxa; 5 / structure of the aedeagus; 6 / structure of the ovipositor. Our results were confirmed by the subsequent study of Italian authors and species of Tapinopterus rebellis s. l. will be placed to the new genus Pseudomolopsis (Giachino et al, in press).	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
03F687E73003FF97FF5B7E8FFE1493D9.taxon	distribution	Distribution. Greece: Peloponnesos. PLATE 13. Sterna VIII (ventral view) (Figs. 144 – 150) of Tapinopterus species. Fig. 144. Tapinopterus (Pseudomolopsis) rebellis (Greece, Taygetos, Anogia); Fig. 145. T. (Molopsis) molopiformis (Turkey, Boz daġ); Fig. 146. T. (Molopsis) phrygius (Turkey, Davraz daġ); Fig. 147. T. (Molopsis) molopinus (Turkey, Uludaġ); Fig. 148. T. (Molopsis) chaudoiri sp. nov. (paratype Gök daġ); Fig. 149. T. (Molopsis) relegatus sp. nov. (holotype); Fig. 150. T. (Molopsis) wiedemanni (Turkey, Bolu, Kartalkaya). Scale bar: 0.5 mm.	en	Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard, Lassalle, Bernard (2012): Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species. Zootaxa 3185: 1-35, DOI: 10.5281/zenodo.279988
