taxonID	type	description	language	source
03F687A7FFF3F80CFF01BB4A43E2F8DC.taxon	materials_examined	Type species: Mortoniella bilineata Ulmer, 1906. As a subgenus, distinct from the newly recognized subgenus Nanotrichia, this taxon is restricted to include taxa formerly referred to as members of the bilineata and leroda species groups (Blahnik and Holzenthal 2008, 2011), as well as all of the taxa previously unplaced to species group, except for M. rodmani Blahnik and Holzenthal, 2008. A formal assignment of all species in the genus to subgenus and species group, including those described in this paper, can be found in Table 1. A generic synonymy for the genus as wholde and a more complete listing of literature citations and distributional records for the individual species can be found in the Catalog of Neotropical Trichoptera (Holzenthal and Calor 2017). We have continued to designate species of M. (Mortoniella) within the two recognized species groups (the bilineata and leroda groups), since they are diagnostically distinct and represent about 85 % of the species. A third group of species is also considered under the category of “ unplaced species, ” but is probably not a natural group. It includes species retaining some plesiomorphic characters, but lacking the apomorphic characters used to define either the bilineata or leroda species groups. The individual species may either be basal to one or the other of these two species groups, or basal to both groups combined. Characters suggestive of their phylogenetic placement are discussed under the subgroup headings or species descriptions and in the accompanying phylogeny. The subgenus Mortoniella can be recognized by a combination of characters. About 75 % of the species have more than 1 fork in the hind wing (forks III and / or fork V present, in addition to fork II). Species with the hind wing venation reduced to fork II, as in M. (Nanotrichia), typically have the costal margin of the hind wing more abruptly angulate (Fig. 101 B). These species belong to the leroda group and are also characterized by a relatively short ventral process on segment VI and males with the anterior margin of segment IX broadly rounded (diagnostic characters of the leroda group). Hind wing configuration for other species in the subgenus Mortoniella includes having forks II, III, and V present (present in most species of the bilineata group, as well as in the species unplaced to species group) (Fig. 97 B, 99 B); forks II and III present (various species subgroups of the leroda group) (Fig. 100 B); or forks II and V present (the flinti subgroup of the bilineata group) (Fig. 98 B). Overall character similarities are difficult to define, due to the variability in genitalia, but there is a tendency for tergum X of males to have the apicolateral lobes distinctly sclerotized and defined, whereas tergum X in members of M. (Nanotrichia) usually have the apicolateral lobes broadly rounded or simple in form.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFF7F817FF01BBC64215FD8F.taxon	description	Fig. 1, 106	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFF7F817FF01BBC64215FD8F.taxon	description	Adult — Length of forewing: male (pharate adult) ca. 4.0 mm; female (not developed enough to measure). Wing venation not determined. Apex of forewing angulate. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish brown. Legs yellowish, tibial spurs dark brown, contrasting with legs. Wing without evident bar at anastamosis. Male genitalia — Ventral process of segment VI posteriorly projecting, very short, narrow basally, length only slightly greater than width at base. Tergum VIII distinctly elongate dorsally, membranous connection to tergum IX only moderately developed. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with rounded (subangular) projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, bulbous, lateral margins rounded, laterally with subacute lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, subtruncate, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection (mesal notch absent); tergum ventromesally with paired, rounded, lightly sclerotized, ventromesal lobes at about midlength, each with short setae. Inferior appendages with short rounded dorsolateral lobes and paired, apically rounded, ventromesal lobes. Mesal pockets of inferior appendage with relatively short, spine-like, posteriorly-directed, apicoventral projections. Paramere appendage short, linear, slightly widened preapically, apex acute. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with undulate contour, strongly curved basally, ventral margin only slightly produced and rounded in middle (not angulate), nearly rectilinearly upturned in apical third, apex rounded (flattened and compressed as viewed dorsally). Phallicata with distinctly sclerotized basodorsal projection and paired, lightly sclerotized, ventral lobes; ventral lobes, as viewed ventrally, relatively short, broad, rounded apically. Endophallic membrane simple in structure, without evident lateral lobes or spines; phallotremal spines absent. Material examined — ECUADOR: 11 mi. W of Pujili, 12500 ft., 15. iii. 1958, RW Hodges, male Holotype (pharate adult, USNM type # 66019) – 3 males, 1 female Paratypes (pharate adults, in alcohol) (NMNH). Distribution — Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE8F816FF01BAE64578FCCF.taxon	description	Fig. 2	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE8F816FF01BAE64578FCCF.taxon	description	Adult — Length of forewing: male 5.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color medium golden or tawny brown. Tibial spurs darker, contrasting with legs. Wing without distinct bar at anastamosis. Male genitalia — Ventral process of segment VI posteriorly projecting, very short, narrow basally, length only slightly greater than width at base. Tergum VIII distinctly elongate dorsally, membranous connection to tergum IX only moderately developed. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with rounded (subangular) projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins rounded, laterally with subacute lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, subtruncate, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection (mesal notch absent); tergum ventromesally with paired rounded, lightly sclerotized ventromesal lobes in basal half, each with short setae. Inferior appendages with short rounded dorsolateral lobes and paired linear, apically tapering, ventromesal lobes. Mesal pockets of inferior appendage with relatively short, spine-like, posteriorly-directed, apicoventral projections. Paramere appendage short, linear, nearly uniform in width, apex acute. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with undulate contour, strongly curved basally, articulating with fused basal segments of paramere appendages, ventral margin only slightly produced and rounded in middle (not angulate), nearly rectilinearly upturned in apical third, apex distinctly enlarged and rounded, slightly recurved (shape somewhat variable, flattened and compressed as viewed dorsally). Phallicata without apparent basodorsal projection, ventrally with paired, lightly sclerotized, ventral lobes; ventral lobes, as viewed ventrally, moderately elongate, broad, rounded apically. Endophallic membrane simple in structure, with only weakly developed membranous lateral lobes; phallotremal spines absent. Material examine d — ECUADOR: Cañar: Río Chauchas, 3 km N Zhud, 2910 m, 17. ix. 1990, OS Flint, Jr – 1 male (pinned) (NMNH); Pichincha: 7 km E Pifo, 2950 m, 26 - 28. ix. 1990, OS Flint, Jr – 1 male (pinned) (NMNH). Distribution — Ecuador. — bilineata subgroup Included species: Mortoniella bilineata Ulmer; M. bulbosa, n. sp.; M. chicana Sykora; M. hamata, n. sp.; M. monopodis, n. sp.; M. paralineata Sykora; and M. roldani Flint. The species in this subgroup are characterized by having 2 white forewing bands, 1 at the anastamosis, as found in many species of Mortoniella, and 1 on the proximal part of the wing. Outside the bilineata subgroup, this character is found only in two species of the flinti subgroup, which are otherwise very different in morphology. Mortoniella iridescens Flint, which is here placed in its own species group, also has two wing bands, but these are an iridescent turquoise, rather than white, and other characters do not indicate an obvious close relationship to the taxa included in this subgroup (although it may belong here). The general coloration of species in the bilineata subgroup ranges from a golden brown to brownish-black, but generally not as dark in color as species in the flinti or foersteri subgroups. The species are also characterized by the structure of tergum X, which has the ventrolateral margins curled inward in its apical part, converging on the median plane to form a “ linear seam. ” The resulting apex of the tergum is more or less truncate, without a pronounced apicomesal notch. Species in the apiculata subgroup are similarly developed, but differ from species of the bilineata subgroup in lacking an angulate ventral projection on the dorsal phallic spine, as well as in lacking wing bands. Species in the enchrysa, foersteri, and wygodzinskii subgroups have tergum X with the lateral margins curled inward, as in species in the bilineata subgroup, but the ventral margins of the tergum do not completely converge on the median line, leaving the apex with a distinct apicomesal notch. The loss of this notch in the bilineata and apiculata subgroups, accompanied by the distinct ventromesal seam, is considered a derived apomorphic character, and a probable indication of their close relationship.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE9F815FF01BBA64293FC0F.taxon	description	Fig. 3, 105	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE9F815FF01BBA64293FC0F.taxon	description	Adult — Length of forewing: male 4.5 - 4.8 mm; female 5.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown. Legs same color, tibial spurs darker, contrasting with legs. Palps and basal segments of antenna blackish-brown, base of antenna contrasting with subsequent whitish or light brown segments, apex of antenna dark brown (like wings). Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow basally, length about 2 ½ times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins subparallel, laterally with acute, finger-like, lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, subtruncate, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection near apex (mesal notch nearly absent); tergum ventromesally with paired, rounded, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes, and single tapering ventromesal lobe; mesal lobe, as viewed laterally, short and strongly flexed at base, apex widened and subtruncate. Mesal pockets of inferior appendage with moderately elongate, sinuous, posteriorly-directed, spinelike, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, extending about same length as dorsal phallic spine; basal segment of appendage articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin strongly curved and arched from base, sinuously and nearly rectilinearly upturned in apical 1 / 4 or 1 / 5, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly and strongly widened on ventral margin at about basal 1 / 3, forming acute ventral projection, narrowing apically from projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, basodorsal projection with lateral margins forming short rounded lobes; phallicata ventrally with very elongate, narrow, projecting, sclerotized lobes, extending about same length as paramere appendages. Endophallic membrane simple in structure, with only weakly developed membranous lateral lobes; phallotremal spines absent. Material examined — COLOMBIA: Antioquia: Quebrada El Aguelo, 2 km E El Retiro, 8. ii. 1983, OS Flint, Jr – 1 male (pinned) (NMNH); Km 50 Río Aurra, E San Jeronimo, 22. ii. 1984, OS Flint, Jr – 1 female (pinned) (NMNH); ECUADOR: El Oro: Pinas / Zaruma, Río La Calera, 19 - 20. viii. 1977, LE Peña G – 1 male (pinned) (NMNH). Distribution — Colombia, Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEAF814FF01BC664497FC6F.taxon	description	Fig. 4	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEAF814FF01BC664497FC6F.taxon	materials_examined	This species can be distinguished from others in the bilineata subgroup by the distinctive form of the inferior appendages, which have a bulbous, microsetose, ventromesal projection, which forks apically into 2 narrow, hooked processes, each with 1 or 2 branches. The females listed as additional examined material closely resemble the associated female of M. bulbosa, but are not included in the paratype material, since they were collected from different localities. The female genitalia generally resemble other examined females in the bilineata subgroup, except that the anterior extension of tergum IX is somewhat shorter and pheromone sacks are absent on both terga VI and VII. Adult — Length of forewing: male 3.4 mm; females 5.7 - 6.8. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color (in alcohol) yellowishbrown. Tibial spurs darker, contrasting with legs. Forewing of male without evident wing bars (in alcohol), but anastomosis evident due to unpigmented crossveins. Pinned females dark brown with 2 evident white wing bars. Male genitalia — Ventral process of segment VI prominent, posteriorly projecting, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, margins rounded laterally, subparallel in demarcated dorsal region, laterally with acute, finger-like, lateral lobes, each with prominent apical seta and one or more preapical setae; apex of tergum distinctly sclerotized, bluntly rounded apically, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection near apex (mesal notch very narrow and shallow); tergum ventromesally with paired, rounded, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short, upright, dorsolateral lobes and bulbous ventromesal lobe with minute microsetae, lobe divided apically to form pair of narrow processes (forked or unforked). Mesal pockets of inferior appendage with moderately elongate, sinuous, posteriorly-directed, spinelike apicoventral projections. Paramere appendage short, narrow, nearly uniform in width, apex acute, extending slightly more than ½ length of dorsal phallic spine; basal segment of appendage articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin strongly curved and arched from base, sinuously and nearly rectilinearly upturned in apical 1 / 4, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly and strongly widened on ventral margin at midlength, forming acute ventral projection; spine, as viewed dorsally, nearly uniformly narrow in width (very slightly widened at apical inflection). Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, phallicata ventrally with paired sclerotized lobes, extending about same length as apical inflection of dorsal phallic spine, lobes very broadly rounded basally; as viewed ventrally, narrowed and rounded apically. Endophallic membrane simple in structure, with only weakly developed, membranous, lateral lobes; phallotremal spines absent. Holotype male (alcohol) — PERU: Madre de Dios: Manu, Erika (near Salvación), 550 m, 4 - 6. ix. 1988, O Flint and N Adams (UMSP 000097104) (MJP). Paratypes — PERU: Madre de Dios: same data as Holotype – 1 female (pinned) (NMNH). Additional material examined — PERU: Cuzco: Paucartambo, Puente San Pedro, ca. 50 km NS Pilcopata, 1600 m, 2 - 3. ix. 1988, O Flint and N Adams – 1 female (pinned) (NMNH); Paucartambo, Quinta Calzon, ca. 30 km NW Pilcopata, Km 164, 13.15000 ° S, 71.36667 ° W, 1030 m, 1 - 2. ix. 1989, N Adams, et al. – 1 female (pinned) (NMNH). Etymology — This species is named M. bulbosa for the bulbous ventromesal lobe of the inferior appendage, which character easily distinguishes it from closely related species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEBF813FF01BC464218FBAF.taxon	description	Fig. 5	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEBF813FF01BC464218FBAF.taxon	description	Adult — Length of forewing: male 4.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown. Legs same color as wings, tibial spurs darker, contrasting with legs. Palps and basal segments of antenna blackish-brown, base of antenna contrasting with subsequent whitish or light brown segments, apex of antenna dark brown (like wings). Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX (apparently) elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, margins rounded laterally, subparallel in demarcated dorsal region, laterally with short acute, finger-like, lateral lobes, each with prominent apical seta; apex of tergum very distinctly sclerotized, lateral margins rounded and converging mesally (not truncate), with ventrolateral margins incurved and converging mesally to form linear “ seam, ” mesal notch very narrow; tergum ventromesally with paired, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short angular, retrorse, dorsolateral lobes and distinctive and prominent paired, upward-curved, lateral lobes; lobes widely separated, as viewed ventrally, with short angular, closely apposed, mesal projections on posteromesal margin. Mesal pockets of inferior appendage with elongate, narrow, strongly arched, spine-like, apicoventral projections, conforming to arched ventral margin of phallicata. Paramere appendage moderately elongate, narrow, nearly uniform in width, apex acute, extending more than ½ length of dorsal phallic spine; basal segment of paramere articulating near base of dorsal phallic spine. Phallobase without evident dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin very strongly and distinctively arched, sinuously and nearly rectilinearly upturned in about apical 1 / 6 th, with slight dip at point of inflection, apex of spine rounded; base of spine narrow and stalk-like, abruptly and strongly widened on ventral margin in basal ½, forming acute ventral projection, spine narrowing apically from projection; spine, as viewed dorsally, narrow in width throughout length. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, basodorsal projection with lateral margins forming short rounded lobes; phallicata ventrally with pair of arched and rather weakly sclerotized lobes, extending about same length as apex of paramere appendages. Endophallic membrane simple in structure, with only weakly developed membranous lateral lobes, dorsally with sclerotized mesal “ pocket, ” to accommodate apical inflection of dorsal phallic spine; phallotremal spines absent. Material examined — ECUADOR: Napo: Río Jondachi, 30 km N Tena, 950 m, 10. ix. 1990, OS Flint, Jr – 1 male Paratype (pinned) (NMNH); Zamora-Chinchipe: 6 km E Zumbi, 980 m, 21. ix. 1990, OS Flint, Jr – 1 male Paratype (pinned) (NMNH). Distribution — Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFECF812FF01BC0644E6FB0F.taxon	description	Fig. 6	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFECF812FF01BC0644E6FB0F.taxon	description	Adult — Length of forewing: male 4.2 - 4.6 mm; female 5.2 - 5.4 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color medium brown or goldenbrown. Legs same color as wings, tibial spurs only slightly darker, somewhat contrasting with legs. Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis; apex of wing with fringe of whitish setae. Male genitalia — Ventral process of segment VI prominent, posteriorly projecting, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins subparallel, laterally with acute, finger-like, lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, subtruncate, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection near apex (mesal notch nearly absent); tergum ventromesally with paired, rounded, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with subtriangular lateral lobes and short narrow, paired, ventromesal lobes, each forked near apex. Mesal pockets of inferior appendage with moderately elongate, posteriorlydirected, sinuous, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, extending about same length as dorsal phallic spine; basal segment of appendage articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin strongly curved and arched from base, sinuously and nearly rectilinearly upturned in apical 1 / 5, apex of spine rounded; base of spine narrow, curved and stalk-like, moderately widened on ventral margin at about basal 1 / 3, forming angular ventral projection, narrowing apically from projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, basodorsal projection with lateral margins forming short rounded lobes; phallicata ventrally with elongate, narrow, projecting, sclerotized lobes, extending about same length as paramere appendages; apices of lobes, as viewed ventrally, nearly straight on lateral margins and forming rounded lobes on mesal margins. Endophallic membrane simple in structure, with only weakly developed membranous lateral lobes; dorsal margin with weakly sclerotized area, apparently to accommodate flexed part of dorsal phallic spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFECF812FF01BC0644E6FB0F.taxon	materials_examined	Holotype male (pinned) — COLOMBIA: Cauca: Municipio de Inzá, Quebrada San Andrés, ca. 500 m W Restaurante “ La Portada ”, San Andrés de Pisimbalá, 2.58222 ° N, 76.04333 ° W, 1750 m, 21. xii. 1997, F Muñoz-Q et al. (UMSP 000209440) (UMSP). Paratypes — COLOMBIA: Cauca: same data as holotype – 2 males, 2 females (pinned) (UMSP), 1 male (NMNH); Municipio de Inzá, Quebrada San Andrés, 1 km S del centro de San Andrés de Pisimbalá, 2.57667 ° N, 76.03639 ° W, 1730 m, 20. xii. 1997, F Muñoz-Q et al. – 1 female (pinned) (UMSP). Etymology — This species is named M. hamata from the Latin work hamus, a hook or barb or angle, and referring to the pair of hooked mesal processes on the inferior appendages.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEDF811FF01BD6644A8FBEF.taxon	description	Fig. 7 This species is easily diagnosed by the very elongate, narrow, mesal projection of the inferior appendages. It is unlikely to be confused with any other described species. The left side of segment IX is somewhat deformed in the holotype specimen and the illustration shows the contour of the opposite side. The posterolateral margins of segment IX in this species are rounded and less angular than most other species in the bilineata subgroup, and the ventral lobe or lobes of the phallicata somewhat less developed, but otherwise the species conforms well to the generalized features of the group. Adult — Length of forewing: male 4.8 - 5.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown. Legs same color, tibial spurs slightly darker, not greatly contrasting with legs. Palps and basal segments of antenna blackishbrown, base of antenna contrasting with subsequent light brown segments, apex of antenna dark brown (like wings). Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis. Male genitalia — Ventral process of segment VI prominent, posteriorly projecting, relatively wide basally, length about 1 to 1 ½ times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with broadly rounded projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, lateral margins rounded, laterally with acute, finger-like, lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, rounded laterally, weakly truncate at extreme apex, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection near apex (mesal notch nearly absent); tergum ventromesally with paired, lightly sclerotized, rounded and compressed, ventromesal lobes in basal half, each with short setae. Inferior appendages with very short rounded dorsolateral lobes, and with single elongate, narrow ventromesal lobe; mesal lobe, as viewed laterally, upcurved apically, very uniformly tapering and acute apically, as viewed ventrally. Mesal pockets of inferior appendage with short, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, slightly widened preapically, apex acute, appendage extending about same length as dorsal phallic spine; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, curved and arched basally, linearly extended in middle, and sinuously upturned in about apical 1 / 5, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly widened on ventral margin in basal ½, forming very acute ventral projection; spine, as viewed dorsally, narrow throughout, only slightly widened in middle, rounded apically. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with sclerotized lobes, lateral margins compressed and slightly widened near base, narrowed apically, forming single apicomesal projection, mesal projection with several small spines (more extensively developed in paratype specimen). Endophallic membrane with well- developed, pleated and membranous, lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEDF811FF01BD6644A8FBEF.taxon	materials_examined	Holotype male (pinned) — COLOMBIA: Chocó: km 130, 86 km E Quibdo, 17. ii. 1983, OS Flint, Jr (UMSP 000157314) (NMNH). Paratype — ECUADOR: Imbabura: Reserva Los Cedros, Río los Cedros, 00.30359 ° N, 78.78233 ° W, 1312 m, 18 - 19. x. 2011, Holzenthal, Rios, Encalada, Acosta – 1 male (pinned) (UMSP). Etymology — The name monopodis is taken from the Greek words mono for one and podos for foot and referring to the single elongate mesal projection of the inferior appendages.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEEF810FF01BCC64215FA0F.taxon	description	Fig. 8	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEEF810FF01BCC64215FA0F.taxon	materials_examined	Mortoniella paralineata is a close sister species to M. bilineata Ulmer. Character similarities include the elongate, paired, ventral sclerites of the phallicata and short unbranched mesal projection of the inferior appendages. The ventral sclerites of the phallicata in M. bilineata are distinctly longer than in M. paralineata and the mesal process of the inferior appendages is strongly bent and trianguloid in shape. It also has distinctly longer paramere appendages, subequal in length to the dorsal phallic spine and ventral sclerites of the phallicata. Although we suggested the possible placement of M. paralineata in the enchrysa subgroup (Blahnik and Holzenthal 2011), based on color characters given in the type description, the species was correctly placed in the bilineata subgroup by Sykora. The species is somewhat lighter brown than other species in the bilineata subgroup, but has two obvious white wing bands on the forewing, a defining character of the subgroup. The genitalia are also very similar to other members of the subgroup. A single, nonparatype specimen was examined, from a site not too distant from the paratype specimens. It was much smaller in size (accounting for most of the size variation listed below), but the only significant difference in the genitalia was that the paramere appendages seemed to be more distinctly bent in the middle. We are considering this intraspecific variation. The question may bear reinvestigation when more material is available. Adult — Length of forewing: male 4.0 - 5.7 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color medium brown, head and thorax golden brown. Legs same color as wings, tibial spurs darker, weakly contrasting with legs. Palps blackishbrown. Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, relatively narrow basally, subacute apically, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins subparallel, laterally with narrow, apically acute, finger-like, lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, subtruncate, with ventrolateral margins incurved and converging mesally to form linear “ seam, ” apicodorsally with lightly sclerotized connection near apex (mesal notch nearly absent); tergum ventromesally with paired, rounded, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes and single short ventromesal lobe; mesal lobe bluntly rounded apically, as viewed laterally. Mesal pockets of inferior appendage with moderately elongate, posteriorlydirected, spine-like, apicoventral projections. Paramere appendage only moderately elongate (much shorter than dorsal phallic spine), narrow, nearly uniform in width, nearly straight or very weakly bent in middle; basal segments of appendage fused mesally and articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin strongly curved and arched from base, sinuously and nearly rectilinearly upturned in about apical 1 / 5, apex of spine rounded; base of spine narrow, curved, and stalk-like, abruptly widened on ventral margin in basal ½, forming acute ventral projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, rounded apically. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, basodorsal projection with lateral margins forming short rounded lobes; phallicata ventrally with moderately elongate, narrow, lightly sclerotized lobes; lobes only slightly projecting apically, extending slightly beyond paramere appendages. Endophallic membrane with evident membranous lateral lobes and distinct sclerotized dorsomesal pocket, apparently to accommodate apical inflection of dorsal phallic spine; phallotremal spines absent. Material examined — ECUADOR: Morona-Santiago: Río Salado, Hwy E 46 (via Riobamba Macas), 2.242530 ° S, 78.277910 ° W, 1646 m, 26. i. 2015, Holzenthall, Huisman, Rios-Touma, Amigo – 26 males, 58 females (pinned) (UMSP); Zamora-Chinchipe: Río Jamboe, 21 km S Zamora, 1340 m, 22. ix. 1990, OS Flint, Jr – 2 male Paratypes (pinned) (NMNH); Zamora, 4. xii. 1978, JJ Anderson – 1 male (alcohol) (NMNH). Distribution — Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEFF81EFF01BE664361FBEF.taxon	description	Fig. 9, 97	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFEFF81EFF01BE664361FBEF.taxon	description	Adult — Length of forewing: male 3.5 - 4.7 mm; female 4.8 - 5.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown. Legs same color as wings, tibial spurs slightly darker, not distinctly contrasting with legs. Antennae with several segments after basal 4 or 5 whitish, contrasting with basal and apical segments. Forewing with 2 distinct white wing bars, 1 at anastomosis and 1 on proximal part of wing, approximately midway between base and anastomosis, setae of bands with turquoise iridescence at some light angles; apex of forewing with fringe of whitish setae. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, lateral margins rounded, laterally with acute, finger-like lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, truncate, shorter than basal projection, with ventrolateral margins incurved and nearly converging mesally (typically separated by distinct gap), apicodorsally with lightly sclerotized connection near apex (mesal notch nearly absent); tergum ventromesally with paired, lightly sclerotized, rounded, ventromesal lobes in basal half, each with short setae. Inferior appendage with short rounded lateral lobe, usually with acute posterior projection, and paired narrow ventromesal lobes, each with several small papillae or short setae. Mesal pockets of inferior appendage with moderately elongate, sinuous, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, usually with distinct curve near apex, apex acute, extending nearly as far as dorsal phallic spine, subequal in length to ventral lobes of phallicata; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin sinuous, strongly curved and arched at base, sinuously and nearly rectilinearly upturned in apical 1 / 4, apex of spine rounded; base of spine narrow, curved and stalk-like, moderately widened on ventral margin at about basal ½, forming angular ventral projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, basodorsal projection with lateral margins forming rounded and distinctly projecting lobes; phallicata ventrally with elongate, narrow, sclerotized lobes, extending about same length as paramere appendages; apices of lobes, as viewed ventrally, distinctly rounded or spatulate on lateral margins, nearly straight on mesal margins. Endophallic membrane simple in structure, with short membranous lateral lobes; phallotremal spines absent. Material examined — COLOMBIA: Antioquia: Km 50, Río Aurra San Jeronimo, 14. ii. 1983, OS Flint, Jr, 1 male Paratype (pinned) (NMNH); Valle: Municipio de Buenaventura, Río Escalerete, 1 km E casa de “ AquaValle ” (ca. 16 km SE Cordoba), 3.82722 ° N, 76.87083 ° W, el 210 m, 2. xii. 1997, F Muñoz-Q et al., 1 male (pinned) (UMSP); ECUADOR: Pichincha: Santo Domingo de los Colorados, 14 km E, 5. vii. 1975, Langley and Cohen, 1 male Paratype (M. similis) (NMNH); VENEZUELA: Zulia: Parque Nacional Perijá, Río Negro in Toromo, 10.051 ° N, 72.712 ° W, el 360 m, 15. i. 1994, Holzenthal, Cressa, Rincón, 24 males, 6 females (pinned) (UMSP). Distribution — Colombia, Ecuador, Venezuela. — catherinae subgroup Included species: Mortoniella catherinae, n. sp. As defined here, this group includes only a single unusual species. We initially placed the species with the “ unplaced species ” in the subgenus Mortoniella, because of the apparent absence of a mesal invagination of segment VIII in the female genitalia and the presence of other characters inferred to be primitive and which also characterize a number of the “ unplaced species, ” including a segment IX with a broadly rounded anterior margin, inferior appendage with an elongate, retrorsely curved, dorsal appendage, and elongate spine-like, projections from the mesal pockets of the inferior appendages. However, close examination shows that there is a very slight invagination in the dorsal margin of segment VIII of the female, although the posteromarginal setae remain evenly spaced and not clustered on either side of the invagination. The character is only suggestively developed, but its presence would allow its inclusion in the bilineata group, as it is strictly defined here. Other characters clearly show a relationship with the bilineata group. Characters indicating this include the very angular ventral margin of the dorsal phallic spine, which is also very abruptly upturned apically; the presence of a distinct mesal apodeme on the dorsal margin of the phallicata, which articulates with the dorsal phallic spine; and an elongate, narrow tergum X, with strongly sclerotized apicolateral lobes and also angular ventrolateral lobes. It seems likely that this is the most basal member of the bilineata group recognized to date. However, the absence of some of the synapomorphies listed above in the flinti and quinuas subgroups makes its definitive placement uncertain.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE1F81DFF01BCC6427CF8CF.taxon	description	Fig. 10, 108	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE1F81DFF01BCC6427CF8CF.taxon	description	Adult — Length of forewing (pharate adults): about 4 mm. Wing venation not determinable (in pinned specimen of unassociated female, with forks I, II, and III present in forewing, forks II, III, and V present in hind wing). Spur formula 0: 4: 4. Overall color (in alcohol) yellowish-brown, wings slightly darker. Wing markings not evident (in pinned specimen of unassociated female, uniformly fuscous, without evident wing bar). Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX slightly elongated. Segment IX with anterolateral margin broadly rounded, with greatest width at about middle of segment, posterolateral margin with distinct, irregularly rounded, projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X moderately elongate, with short inflated basomesal projection, lateral margins subparallel, with elongate setae (setae absent on mesal part of tergum), apicolateral margins of tergum distinctly sclerotized, forming narrow, declivous lobes; tergum laterally with acutely angled ventrolateral lobes, ventromesal lobes absent. Inferior appendages with very elongate, narrow, retrorsely curved, dorsolateral lobes, appendages ventrally without mesal invagination or projection. Mesal pockets of inferior appendage with elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage moderately elongate, narrow, nearly uniform in width, apex acute, extending about same length as apical bend of dorsal phallic spine; fused basal segments of appendage articulating near base of dorsal phallic spine. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin very slightly arched, sharply and nearly rectilinearly upturned in apical 1 / 3 or 1 / 4, apical part with numerous small spines, apex acute; base of spine narrow, stalk-like, nearly straight, abruptly and strongly widened on ventral margin in basal 1 / 2, forming very prominent acute ventral projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, apex acute. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with lightly sclerotized, rounded and projecting, basal lobes, and much smaller, angular, apicolateral lobes. Endophallic membrane simple in structure, without membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE1F81DFF01BCC6427CF8CF.taxon	materials_examined	Holotype male (pharate adult, alcohol) — PERU: Cuzco: Paucartambo to Pilcopata rd., Puente Morro Leguia, 13.12400 ° S, 71.72283 ° W, el 2200 m, 20 - 21. vi. 1993, R Blahnik and M Pescador (UMSP 000097161) (MJP) Paratypes — PERU: Cuzco: same data as holotype, 4 males, 1 female (NMNH). Additional material examined — PERU: Cuzco: Paucartambo, E Buenos Aires, km 135, 13.13333 ° S, 71.55000 ° W, 28 - 29. viii. 1989, NE Adams – 1 female (pinned) (NMNH). Etymology — The first author takes great pleasure in naming this unique and interesting species M. catherinae for his mother, Catherine Blahnik, now deceased, without whose support this paper would probably never have been completed. — enchrysa subgroup Included species: Mortoniella adamsae, n. sp.; Mortoniella denticulata Sykora; M. enchrysa Flint; M. langleyae, n. sp.; M. paraenchrysa Sykora; M. silacea, n. sp.; and M. squamata Sykora. The species in this subgroup are characterized by a uniformly golden orange forewing coloration, without a wing bar, and with hind wings and setation on the ventral surface of the forewings fuscous. In a few species, the wing membrane is also infuscated, accentuating the overall golden coloration. Exceptions are M. denticulata Sykora, which has forewings a uniform light brown in color, and possibly M. langleyae n. sp., which is only known from alcohol. Both of these species are placed in the enchrysa subgroup based on structural features of the male genitalia. In general, most species have the ventrolateral margins of the apex of tergum X curved mesally, as in members of the bilineata and M. apiculata subgroups, but the ventral margins do not quite converge mesally. As a result, there is a distinct apicomesal notch (shallow to deep) on tergum X, rather than the structure being truncate or nearly truncate apically. Nevertheless, the general appearance is of a distinctly sclerotized apical “ cap. ” This is less evident in species with a deep mesal invagination. As in members of the bilineata subgroup, the posterolateral margin of segment IX has a more or less angular and very distinctly produced projection. All of the species have a very short ventral process on segment VI, but the general form of the process is typical of species of the bilineata group (narrow basally and posteriorly directed). A very short ventral process is also found in species of the apiculata subgroup.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE3F81BFF01B8A6433CFECF.taxon	description	Fig. 11	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE3F81BFF01B8A6433CFECF.taxon	description	Adult — Length of forewing: male 6.3 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Dorsal side of forewings, head, basal segments of antennae, and legs, except distal parts of tibiae and tarsi, golden-orange; ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, palps, and distal part of tibiae and tarsi dark brownish-black. Wing membranes (apparently) somewhat infuscated. Tibial spurs black, contrasting with legs. Wing bars absent. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, lateral margins rounded, ventrolaterally with acute, tapering, lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, emarginate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with broad U-shaped connection near apex (mesal notch distinct); tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half each with short setae. Inferior appendages with short narrow setose dorsolateral lobes and paired elongate, narrow, ventromesal lobes; ventromesal lobes curved, each with distinct preapical spine on ventral margin. Mesal pockets of inferior appendage with very elongate, posteriorly-directed, spine-like, apicoventral projections. Parameres with paired appendages on either side, 1 moderately elongate, with small spines on apical ½, the other short, about ½ length of longer spine, without spines, both appendages slightly widened preapically, apices acute; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin arched, sinuously and nearly rectilinearly upturned in apical ¼, apex of spine rounded; base of spine narrow, undulately curved, and stalk-like, abruptly and very strongly widened on ventral margin in basal ½, forming acute ventral projection, narrowing apically from projection; spine, as viewed dorsally, somewhat widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with moderately elongate sclerotized lobes, extending about same length as longer paramere appendages, lateral margins of lobes subparallel, apices enlarged and mesally curved. Endophallic membrane simple in structure, with membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE3F81BFF01B8A6433CFECF.taxon	materials_examined	Holotype male (pinned) — PERU: Cuzco: Paucartambo; Puente San Pedro, ca. 50 km NW Pilcopata, 13.15000 ° S, 71.43333 ° W, 1600 m, 2 - 3. ix. 1988, O Flint and N Adams (UMSP 000157307) (MJP). Etymology — We take great pleasure in naming this species M. adamsae for Nancy Adams, now deceased, who was co-collector of the type specimen, and spent many years as assistant curator of the Trichoptera collection at the Smithsonian.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE4F81AFF01B9A6424CFDAF.taxon	description	Fig. 12	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE4F81AFF01B9A6424CFDAF.taxon	description	Adult — Length of forewing: male 4.9 - 6.0 mm; female 5.8 - 6.0. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color of male, including dorsal side of forewings, head, legs, and base of antennae, golden-brown, female slightly darker. Wing membrane of fore- and hind wings distinctly infuscated, more distinctly evident on ventral side of forewings and hind wings, due to short scant setae (of same color as dorsal side of wings). Palps, apices of antennae, and marginal setae of wings dark brown. Tibial spurs dark brown, contrasting with legs. Wings of male without wing bars, of female with indistinct or interrupted white wing bar at anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, moderate in length, narrow basally, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, dorsal margin somewhat widened, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half, projection with minute microsetae in addition to usual lateral setae; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, base of segment not inflated, lateral margins constricted at base, subparallel laterally, apex of tergum with deep V-shaped mesal incision, apex sclerotized, but not forming evident “ cap, ” tergum ventrolaterally with elongate narrow, curved, apically acute, lateral lobes, each with several preapical setae, lobe widely separated from dorsal part of tergum by rounded notch; tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half, each with short setae (not readily visible in lateral view). Inferior appendages with moderately elongate narrow, posteriorly recurved, dorsolateral lobes and short acute, paired, apicoventral lobes. Mesal pockets of inferior appendage with elongate, posteriorly directed, spine-like, apicoventral projections. Fused basal segments of parameres articulating near middle of stalklike basal part of dorsal phallic spine, paramere appendages absent. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin sinuously undulate, gradually upturned in about apical 1 / 3, base of spine narrow, stalk-like and sinuously curved, distinctly widened on ventral margin in basal ½, forming acute ventral projection, apical part of spine rather uniformly broad, apex rounded, spine with many small lateral spines in about apical 1 / 3; spine, as viewed dorsally, slightly widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with indistinct, lightly sclerotized, lobes, extending about same length as recurved dorsal lobe of inferior appendages. Endophallic membrane with subdivided, membranous lateral lobes, basal part distinctly projecting, with minute spines; phallotremal spines absent. Material examined — VENEZUELA: Merida: Río Albarregas, ca. 1 km NW Univ. de los Andes, 8.634 ° N, 71.158 ° W, el 1980 m, 24. iv. 1995, Holzenthal, Gulic, Segnini – 15 males, 3 females (pinned) (UMSP); Parque Nacional Sierra Nevada, Quebrada La Mucuy, 7 km E Tabay, 8.637 ° N, 71.034 ° W, 2200 m, 18. i. 1994, Holzenthal, Cressa, Rincón – 1 male (pinned) (UMSP). Distribution — Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE5F819FF01BA064229FD6F.taxon	description	Fig. 13	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE5F819FF01BA064229FD6F.taxon	description	Adult — Length of forewing: male 6.0 mm; female 6.8. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Dorsal side of forewings, head, basal segments of antennae, and legs golden-orange; wing membrane of fore- and hind wings distinctly infuscated, ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, and palps dark brown. Tibial spurs brownish-black, strongly contrasting with legs. Wing bars absent. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 1 ½ to 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with nearly rectilinearly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins rounded, ventrolaterally with subangular lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, emarginate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with lightly sclerotized connection near apex (mesal notch shallow, but distinct); tergum ventromesally with paired, sclerotized, rounded (short paddle-like), ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes and short paired ventromesal lobes, ventromesal lobes strongly curved, each with distinct preapical spine on ventral margin; basodorsal lobes and base of ventromesal lobes with numerous microsetae. Mesal pockets of inferior appendage with moderately elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, distinctly dorsally curved and slightly widened near apex, apex acute; ventral margin of appendage with linear row of sensilla; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin arched in middle, sinuously and nearly rectilinearly upturned in apical ¼, apex of spine rounded; base of spine narrow and stalk-like, abruptly and strongly widened on ventral margin in basal ½, forming subangular ventral projection, spine narrowing apically from projection; spine, as viewed dorsally, somewhat widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with 2 pairs of diverging, sclerotized lobes, basal ones retrorsely oriented, rounded laterally, posterior pair diverging from posterior margin of basal lobes, apices bluntly rounded, posteriorly projecting; ventromesal margin of phallicata extending beyond paired lobes, very lightly sclerotized, extending about same length as paramere appendages. Endophallic membrane simple in structure, with very small membranous preapical lateral lobes; phallotremal spines absent. Material examined — COLOMBIA: Valle: Municipio El Cerrito, Río Cerrito, 7.1 km E Hacienda “ El Paraiso ”, 3.64972 ° N, 76.17278 ° W, 1950 m, 3. xii. 1997, F Muñoz et al. – 1 male, 1 female (pinned) (UMSP). Distribution — Colombia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE6F818FF01BB46426BFD0F.taxon	description	Fig. 14 This is a very distinctive species, probably most closely related to M. denticulata Sykora, though superficially very different. Both species have a dorsal phallic spine with small apical spines (also present in M. silacea, n. sp.) and an endophallic membrane with conspicuous membranous lateral lobes with minute spines. Both species also lack paramere appendages. As compared to either M. denticulata or M. silacea, the apex of the dorsal phallic spine is much narrower, as viewed laterally. Especially distinctive aspects of M. langleyae include the very elongate and apically curved tergum X, with a very deep mesal invagination and very elongate, pencil-like, ventrolateral lobes. Additionally, the tergum has very conspicuously developed, enlarged and projecting, ventromesal lobes. Like M. denticulata, the tergum lacks either obvious apicoventral sclerotization or a distinctly inflated and demarcated basal portion. As compared to other species in the bilineata group, the anteroventral margin of segment IX is only weakly produced. Adult — Length of forewing: male 3.5 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Color (in alcohol) yellowish-brown; setae of forewing (apparently) golden or golden-brown. Wing membranes not obviously infuscated. Wing bars not evident. Male genitalia — Ventral process of segment VI posteriorly projecting, relatively short, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX; membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and weakly produced in ventral half, posterolateral margin with nearly rectilinearly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X very elongate and curved apically, base of segment not inflated, lateral margins slightly constricted at base, subparallel laterally, apex of tergum with very deep V-shaped mesal incision, apex sclerotized, but not forming evident “ cap, ” tergum ventrolaterally with very elongate narrow, pencil-like, lateral lobes, each with elongate apical seta, lobe narrowly separated from and subparallel to apicolateral lobes; tergum ventromesally with very prominent, paired, sclerotized and projecting, ventromesal lobes in basal half, each with short setae. Inferior appendages with very short (apparently vestigial) dorsolateral lobes and short paired, curved, apicoventral lobes, each with a distinct apicoventral spine-like projection. Mesal pockets of inferior appendage with elongate, posteriorly-directed, spine-like, apicoventral projections. Fused basal segments of parameres articulating near middle of stalk-like basal part of dorsal phallic spine, paramere appendages absent. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin nearly straight, obtusely upturned in about apical 1 / 3, base of spine narrow, stalk-like and sinuously curved, distinctly widened on ventral margin in basal ½, forming acute ventral projection; upturned apex of spine very distinctly narrowed, with scattered small lateral spines, apex rounded; spine, as viewed dorsally, narrow basally and apically, slightly widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with indistinct, paired, lightly sclerotized lobes, extending beyond lateral lobes of endophallic membrane. Endophallic membrane with distinct projecting membranous lateral lobes at midlength, each with minute spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE6F818FF01BB46426BFD0F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Zamora-Chinchipe: Cumbaratza, 12. vi. 1976, A Langley et al. (NMNH) (UMSP 000097033). Etymology — This species is named in honor of Andrea Langley, who collected the type specimen as part of a Peace Corps project in the 1970 ’ s, in recognition of the value of this endeavor to the study of Neotropical caddisflies.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE7F827FF01BB664244FDEF.taxon	description	Fig. 15	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFE7F827FF01BB664244FDEF.taxon	description	Adult — Length of forewing: male 6.4 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Dorsal side of forewings, head, basal segments of antennae, and legs golden-orange; ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, and palps dark brownish-black (fuscous). Wing membrane not apparently infuscated. Tibial spurs brownish-black, contrasting with legs. Wing bars absent. Male genitalia — Ventral process of segment VI posteriorly projecting, very short, narrow basally, length about 1 ½ times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X with basal part slightly inflated, distinctly set off from apical part, tergum moderately elongate, lateral margins rounded, ventrolaterally with short acute lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, emarginate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with broad U-shaped connection near apex (mesal notch distinct); tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short narrow setose dorsolateral lobes and paired ventromesal lobes, each with narrow, strongly hemispherically curved, dorsal process and short acute apicomesal process. Mesal pockets of inferior appendage with very elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendages elongate, narrow, slightly widened preapically, apices acute, subequal in length to dorsal phallic spine; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin curved basally, slightly undulate in middle, and nearly rectilinearly upturned in apical ¼, apex of spine rounded; base of spine narrow, undulately curved and stalk-like, abruptly and very strongly widened on ventral margin in basal ½, forming obtusely angular ventral projection, narrowing apically from projection; spine, as viewed dorsally, somewhat widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with elongate, lightly sclerotized, lobes, extending about same length as paramere appendage, lateral margins of lobes subparallel, apices mesally curved. Endophallic membrane with conspicuous, membranously pleated, lateral lobes; phallotremal spines absent. Material examined — PERU: Cuzco: Paucartambo, E Buenos Aires, km 135, 13.13333 ° S, 71.55000 ° W, 2150 m, 28 - 29. viii. 1989, N Adams – 1 male (NMNH). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD8F826FF01BAC643DCFD0F.taxon	description	Fig. 16 This species probably bears the greatest overall resemblance to M. denticulata Sykora, especially because of the minute spines near the apex of the dorsal phallic spine, which is also distinctly broadened and club-shaped apically (as viewed laterally). Mortoniella langleyae, n. sp., also has small spines on the apex of the dorsal phallic spine, but the spine is greatly narrowed apically, in lateral view, and its general resemblance to M silacea is not very strong. Mortoniella silacea is more distinctly golden in coloration and differs in a number of other details from M. denticulata. Very distinctive is the nearly rectilinear angle on the posterior margin of segment IX. Other differences from M. denticulata include the structure of the inferior appendage, which has a ventral apex that is posteriorly-directed and nearly straight, rather than strongly arched; ventrolateral projections on tergum X that are only very narrowly separated from the dorsal part of the tergum; a ventral margin of the dorsal phallic spine that is obtusely, rather than acutely, angled; and the absence of spines on the membranous lateral projections of the endophallic membrane. Adult — Length of forewing: male 6.1 mm; female 7.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color of male, including dorsal side of forewings, head, legs, and base of antennae, golden-orange, female lightly darker. Wing membrane of fore- and hind wings somewhat infuscated, setae on ventral side of forewings and hind wings dark brown. Palps and apices of antennae dark brown. Tibial spurs brownish-black, contrasting with legs. Wing bars absent. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, dorsal margin somewhat widened, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with nearly rectilinearly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, base of segment inflated, lateral margins subparallel, apex of tergum with deep V-shaped mesal incision, apex truncate, sclerotized, but not forming evident “ cap ” (only suggestively developed), tergum ventrolaterally with tapering, apically acute, lateral lobes, each with elongate apical seta, lobes narrowly separated from dorsal part of tergum; tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with narrow, lightly sclerotized, apically recurved, dorsolateral lobes and paired narrow, apically acute, apicoventral lobes, each with short acute projection on ventral surface at about midlength. Mesal pockets of inferior appendage with very elongate, posteriorly-directed, spine-like, apicoventral projections. Fused basal segments of parameres articulating near middle of stalk-like basal part of dorsal phallic spine, paramere appendages relatively short, narrow, acute apically. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin strongly curved basally, then straight, and strongly, nearly rectilinearly, upturned in about apical 1 / 3; base of spine narrow, stalk-like, strongly curved, spine distinctly widened on ventral margin in basal ½, forming obtuse, subangular, ventral projection, apical part of spine relatively wide, apex rounded, slightly widened and knob-like, with small lateral spines in about apical 1 / 3; spine, as viewed dorsally, slightly widened in middle, basal and apical parts narrow, apex compressed. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with moderately elongate, subparallel, lightly sclerotized, lobes, extending slightly past elongate spine-like projections of mesal pockets of inferior appendages. Endophallic membrane with prominent, membranously pleated, lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD8F826FF01BAC643DCFD0F.taxon	materials_examined	Holotype male (pinned) — COLOMBIA: Cauca: Municipio de Silvi, Río Piendamó, ca. 5 km NE Silvia, 2.63194 ° N, 76.33806 ° W, 2610 m, 30. xii. 1997, F Muñoz-Q et al., (UMSP 000209642) (UMSP). Paratypes — COLOMBIA: Cauca: same data as holotype, 2 females (pinned) (UMSP); ECUADOR: Bolivar: W Pilalo, 1800 m, 9 - 10. x. 1977, LE Peña G, 1 male (pinned) (NMNH). Etymology — This species is named M. silacea from the Latin word silaceus, the color of yellow ochre, and referring to the color of the forewings of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD9F824FF01BB664594FF4F.taxon	description	Fig. 17	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD9F824FF01BB664594FF4F.taxon	description	Adult — Length of forewing: male 5.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Dorsal side of forewings, head, basal segments of antennae, and legs, except distal parts of tibiae and tarsi, golden (ochraceous); ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, and palps, dark brownish-black; legs with scattered dark setae, especially at apices of tibiae and tarsal segments. Forewings (apparently) somewhat infuscated. Tibial spurs brownish-black, contrasting with legs. Wing bars absent Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X with basal part inflated and distinctly set off from apical part, tergum moderately elongate, lateral margins weakly rounded, ventrolaterally with short acute lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized and emarginated mesally, lateral lobes truncate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with short U-shaped connection near apex (mesal notch distinct); tergum ventromesally with paired, rounded and sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short setose dorsolateral lobes and single short ventromesal lobe, as viewed ventrally. Mesal pockets of inferior appendage with only moderately elongate, posteriorly-directed, spine-like, apicoventral projections, projections apparently fused to ventral margin of phallicata. Parameres with paired appendages on either side, both relatively elongate, narrow, with numerous small adpressed, spine-like projections, dorsal pair relatively straight, ventral pair strongly bowed or curved in apical ½; fused basal segments of parameres articulating near base of dorsal phallic spine, ventral margins with numerous microsetae and (apparently) sclerously fused to and more or less continuous with base of phallicata. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin curved basally, nearly straight in middle part and strongly, nearly rectilinearly, upturned in apical ¼, apex of spine rounded; base of spine narrow, undulately curved and stalk-like, widened on ventral margin in basal ½, forming rounded, non-angular, ventral projection, narrowing apically from projection; spine, as viewed dorsally, somewhat widened at ventral enlargement, basal and apical parts narrow, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with rounded ventral projection of dorsal phallic spine; phallicata ventrally with 2 pairs of diverging sclerotized lobes (butterfly-like), basal ones retrorsely oriented, rounded laterally, posterior pair diverging from posterior margin of basal lobes, apices bluntly rounded, posteriorly projecting; ventromesal margin of phallicata extending somewhat beyond paired lobes, very lightly sclerotized, extending about same length as paramere appendages. Endophallic membrane simple in structure, without apparent membranous lateral lobes; phallotremal spines absent. Material examined — ECUADOR: Napo: 5 km S Baeza, 1900 m, 10. ix. 1990, OS Flint, Jr – 1 male Paratype (pinned) (NMNH). Distribution — Ecuador. — flinti subgroup Included species: Mortoniella bifurcata Sykora; M. flinti Sykora; M. tanyrhabdos, n. sp.; and M. tusci, n. sp. This subgroup of the bilineata group was originally proposed by Sykora (1999), but included what is probably a heterogeneous assemblage of species. We are restricting the definition of the group to include two species listed by Sykora, M. flinti and M. bifurcata, and two new species. A distinct, but unassociated, female specimen reveals that at least 1 additional species belongs to the group. As redefined, the group is very homogeneous and all of the species are obviously closely related. Known species in the group are restricted to Venezuela. All of the species are relatively small for species in the bilineata group and very dark in color, with the mesotarsi white or whitish, except at the very apex. A white or whitish band is also evident in the basal part of the antennae in most specimens; the band occurs in the apical part of the antennae in the undescribed female specimen. Similar white markings are found on the legs and antennae of at least some species of the bilineata subgroup, but this is probably a homoplasious similarity, since the overall differences between the two subgroups are considerable. The flinti subgroup shares basic features that characterize the bilineata group as a whole, including an anteroventrally produced segment IX, elongate narrow ventral process on segment VI, and females with a strongly invaginated tergum VIII. However, unlike most other species in the bilineata group, the posterior margin of segment IX of males in the flinti subgroup is not angular and only slightly produced, the ventral margin of the of the dorsal phallic spine is not angularly articulated with the phallicata, and the apex of the dorsal phallic spine is neither abruptly upturned nor rounded apically in lateral view. In all of the species of this group the inferior appendages have very fine, elongate setae on the dorsal margin of the dorsal lobe, and the assemblage of the inferior appendages with the phallicata seems to be more or less fused, so that the elongate spines from the mesal pockets of the inferior appendages are separated from and project below the ventral margin of the phallicata, which is characteristically arched. Tergum X is elongate and more or less entire (notched apically in M. flinti), without the paired sclerotized apicolateral lobes that characterize species in the M bilineata, apiculata, and enchrysa subgroups. The species in this subgroup also lack the projecting ventromesal lobes on tergum X that characterize the species of the bilineata, apiculata, enchrysa, foersteri, iridescens, and wygodzinskii subgroups. The flinti subgroup species are unusual for species of the bilineata group in having only 3 tibial spurs on the mesotibiae and in lacking fork III in the hind wing (only forks II and V present). The latter venational combination is unique with the genus Mortoniella.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDBF824FF01B9264226F8CF.taxon	description	Fig. 18	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDBF824FF01B9264226F8CF.taxon	description	Adult — Length of forewing: male 3.8 mm. Forewing with forks I, II, and III present, hind wing with forks II and V. Spur formula 0: 3: 4. Overall color dark brownish-black (fuscous). Mesotarsi whitish, except at very apex. Tibial spurs slightly darker than legs, not strongly contrasting with legs. Forewing with distinct white wing bar at anastomosis, and evidence (on one wing of somewhat rubbed specimen) of second white wing bar on proximal part of wing, closer to base than anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow basally, length about 3 ½ times width at base. Tergum VIII narrow, subtending ventral margin of segment IX, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin nearly straight, without distinct projection; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X elongate, apex rounded, with only slight mesal invagination, lateral margins subparallel, with paired longitudinal ridges extending from basolateral margins to past midlength, ridges somewhat converging posteriorly; tergum with bluntly rounded ventrolateral lobes, ventromesal lobes absent. Inferior appendages with short rounded dorsolateral lobes, each with fringing row of very elongate setae, and short, bluntly rounded, ventromesal projection, subtending apical spine-like projections of mesal pockets of inferior appendages. Mesal pockets of inferior appendage with moderately elongate, posterodorsally curved, spine-like, apicoventral projections, projecting distinctly below ventral margin of phallicata. Paramere appendage relatively elongate, not extending to apex of dorsal phallic spine, narrow, nearly uniform in width, apex acute, strongly bent in about apical 1 / 3; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow throughout, tapering apically, slightly dorsally curved in about apical 1 / 4, apex acutely bifid in holotype, unilaterally asymmetric in paratype (Fig. 18 D); base of spine with short, curved stalk and distinct, rounded, ventral deflection in basal ¼; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, apex bifid or acute and asymmetric. Phallicata with elongate sclerotized basodorsal projection, articulating with rounded ventral deflection of dorsal phallic spine, basolaterally with small rounded projection, ventral margin sclerotized and strongly arched. Endophallic membrane simple in structure, without membranous lateral lobes; phallotremal spines absent. Material examined — VENEZUELA: Barinas: 22 km NW Barinitas, 19. ii. 1976, CM and OS Flint, Jr – 1 male Paratype (pinned) (NMNH). Distribution — Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDCF823FF01B8A64226F92F.taxon	description	Fig. 19	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDCF823FF01B8A64226F92F.taxon	description	Adult — Length of forewing: male 3.1 - 3.4 mm; female 3.5 - 3.7 mm. Forewing with forks I, II, and III present, hind wing with forks II and V. Spur formula 0: 3: 4. Overall color dark brownish-black (fuscous). Mesotarsi whitish, except at very apex; antennae with indistinct whitish band or annulus, from about segments 4 - 6. Tibial spurs only slightly darker than legs, not strongly contrasting with legs. Forewing with 2 distinct white wing bars, 1 at anastomosis, and 1 on proximal part of wing, closer to base than anastomosis. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow basally, acute apically, length about 3 ½ times width at base. Tergum VIII narrow, subtending ventral margin of segment IX, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin nearly straight, without distinct projection; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X elongate, apex with distinct shallow mesal invagination, extending less than ¼ length of tergum, lateral margins slightly converging apically, with paired longitudinal ridges, extending from basolateral margins to past midlength, ridges somewhat converging posteriorly; tergum with bluntly rounded ventrolateral lobes, ventromesal lobes absent. Inferior appendages with short rounded dorsolateral lobes, each with fringing row of very elongate setae, and short ventromesal projection, subtending apical spine-like projections of mesal pockets of inferior appendages. Mesal pockets of inferior appendage with relatively short and strongly curved, spine-like, apicoventral projections, projecting below ventral margin of phallicata. Paramere appendage relatively elongate, not extending to apex of dorsal phallic spine, narrow, tapering from base to apex, apex acute, appendage ventrally curved (falcate); fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow throughout, tapering apically, slightly dorsally curved in about apical 1 / 4, apex acutely trifid; base of spine with short curved stalk and distinct rounded ventral deflection in basal 1 / 3; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, apex trifid (Fig. 19 D). Phallicata with sclerotized basodorsal projection, articulating with rounded ventral deflection of dorsal phallic spine, basolaterally with small rounded projection, ventral margin sclerotized and very strongly arched, with protruding rounded lateral lobes near base and in apical 1 / 2. Endophallic membrane simple in structure, with slightly produced membranous lateral lobes; phallotremal spines absent. Material examined — VENEZUELA: Aragua: Est. Exp. Cataurito, 1. ii. 1983, OS Flint, Jr – male Holotype, 1 male, 2 female Paratypes (pinned) (NMNH). Distribution — Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDCF822FF01BE864360FA4F.taxon	description	Fig. 20 This species is most similar to M. bifurcata Sykora, especially in the shape of tergum X, which is only weakly notched apicomesally, and in the relative length of the spine from the mesal pockets of the inferior appendages. The apex of the dorsal phallic spine is symmetrically curved, without preapical projections. Characters considered diagnostic for the species include the elongate paramere appendages and the sclerotized apical lobes of the endophallic membrane. Adult — Length of forewing: male 4.0 - 4.8 mm; female 5.0 mm. Forewing with forks I, II, and III present, hind wing with forks II and V. Spur formula 0: 3: 4. Overall color dark brownish-black (fuscous). Mesotarsi whitish, except at very apex; antennae of female with indistinct whitish band or annulus, from about segments 4 - 6, males without annulus. Tibial spurs slightly darker than legs, not strongly contrasting with legs. Forewing without wing bars. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow, length about 4 times width at base. Tergum VIII narrow, subtending ventral margin of segment IX, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin nearly straight, without distinct projection; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X elongate, apex rounded, with slight mesal invagination, lateral margins subparallel, with paired longitudinal ridges extending from basolateral margins to past midlength, ridges somewhat converging posteriorly; tergum with very short, apically rounded, ventrolateral lobes, ventromesal lobes absent. Inferior appendages with short rounded dorsolateral lobes, each with fringing row of very elongate setae, and short ventromesal projection, apparently fused to apical spine-like projections of mesal pockets of inferior appendages. Mesal pockets of inferior appendage with moderately elongate, posterodorsally curved, spine-like, apicoventral projections, projecting distinctly below ventral margin of phallicata. Paramere appendage elongate, extending nearly to apex of dorsal phallic spine, narrow, nearly uniform in width, bowed or curved mesally at apex, apex acute; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow throughout, distinctly undulate in contour, nearly straight (not upturned) apically, apex rounded in both lateral and dorsal views; base of spine with short, curved stalk and distinct rounded ventral deflection in basal 1 / 3; spine, as viewed dorsally, nearly uniformly narrow in width throughout length. Phallicata with elongate sclerotized basodorsal projection, articulating with rounded ventral deflection of dorsal phallic spine, basolaterally with small rounded projection, ventrally with rounded lateral lobes near base and paired rounded, ventrally deflexed, apical lobes (apparently somewhat separated from sclerotized basal part of phallicata). Endophallic membrane simple in structure, without prominent membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDCF822FF01BE864360FA4F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Tachira: Quebrada Mesa del Palmar, 5 km S El Cobre, 7.99750 ° N, 72.06325 ° W, 2370 m, 18 - 20. iv. 1995, Holzenthal, Cressa, Gulic (UMSP 000001394) (UMSP). Paratypes — VENEZUELA: Tachira: same data as holotype — 1 male, 1 female (pinned) (UMSP); Quebrada La Honda, 10 km E La Grita, 8.14695 ° N, 71.93378 ° W, 2300 m, 23. iv. 1995, Holzenthal, Cressa, Gulic — 3 males (pinned) (UMSP), 1 male (pinned) (NMNH), 1 male (pinned) (MIZA). Etymology – This species is named M. tanyrhabdos from the Greek words tany, meaning long, and rhabdos, a rod, and referring to the elongate paramere appendages of this species, which help to distinguish it from other species in the flinti subgroup.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDDF821FF01BE26438AF8CF.taxon	description	Fig. 21, 98, 107 Among species in the flinti subgroup, this species is most readily diagnosed by the very elongate spines from the mesal pockets of the inferior appendages. The phallicata is also distinctive in having elongate microsetae from its ventral margin and a widened and extended dorsal margin with minute spines. Like M. tanyrhabdos, n. sp., it is black in color, without wing bars. In addition to the characters mentioned, it differs from this species in having much shorter paramere appendages and in having the apex of the dorsal phallic spine more narrowed apically, as viewed both laterally and dorsally. Adult — Length of forewing: male 3.0 - 3.4 mm; female 3.2 - 3.4 mm. Forewing with forks I, II, and III present, hind wing with forks II and V. Spur formula 0: 3: 4. Overall color dark brownish-black (fuscous). Mesotarsi whitish, except at very apex; antennae of female with indistinct whitish band or annulus, from about segments 4 - 6, males apparently without annulus. Tibial spurs slightly darker than legs, not strongly contrasting with legs. Forewing without wing bars. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, narrow, length about 3 ½ times width at base. Tergum VIII narrow, subtending ventral margin of segment IX, membranous connection to tergum IX moderately elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with very slight rounded projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X elongate, apex rounded, with slight mesal invagination, lateral margins subparallel, with paired longitudinal ridges extending from basolateral margins to past midlength, ridges somewhat converging posteriorly; tergum without apparent ventrolateral lobes, ventromesal lobes absent. Inferior appendages with short rounded dorsolateral lobes, each with fringing row of very elongate setae, and ventral projection, apparently fused to apical spine-like projections of mesal pockets of inferior appendages. Mesal pockets of inferior appendage with very elongate, posteriorly directed, spine-like (or tusk-like), apicoventral projections, projecting distinctly below ventral margin of phallicata. Paramere appendage moderately elongate, shorter than dorsal phallic spine, narrow, nearly uniform in width, ventrally curved, apex acute; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow throughout, somewhat undulate in contour, but overall nearly straight, only weakly upturned at extreme apex, apex acute in both lateral and dorsal views; base of spine with short curved stalk and distinct rounded ventral deflection in basal 1 / 4; spine, as viewed dorsally, nearly uniformly narrow in width throughout length. Phallicata very elongate, with dorsal margin sclerotized and flattened, lateral margins forming projecting longitudinal ridges, with minute spines in basal part; phallicata laterally with small rounded projection, ventral margin strongly arched, with distinct microsetae in basal part; as viewed ventrally, with basal part subparallel, apically with small subangular lateral lobes. Endophallic membrane very reduced and simple in structure, without membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDDF821FF01BE26438AF8CF.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Merida: La Campana, 12 km SE Santo Domingo, 24. ii. 1976, CM and OS Flint, Jr (UMSP 000157406) (NMNH). Paratypes — VENEZUELA: Merida: same data as Holotype – 3 males, 3 females (pinned) (NMNH). Etymology — This species is named M. tusci, from the Anglo-Saxon word tusk (or tusc), in reference to the elongate spines from the mesal pockets of the inferior appendages, which appear somewhat like an elephant’s tusk. — foersteri subgroup Included species: Mortoniella foersteri (Schmid); and M. longiterga, n. sp. Mortoniella foersteri was considered a member of the bilineata subgroup by Sykora (1999), but is removed and treated separately here, largely because of its lack of a proximal white forewing band. The second species in this group, M. longiterga n. sp., is obviously closely related. Wing color in both species is a dark brownish-black or fuscous, somewhat darker than in members of the bilineata subgroup. Both species have the posterior margin of segment IX broadly rounded, as opposed to the distinctly angular margin in the M. bilineata and M. enchrysa subgroups. Like (most) members of both the bilineata and enchrysa subgroups, the ventral margin of the dorsal phallic spine is angular and articulates with a protuberance on the dorsal margin of the phallicata; the angular development in both M. foersteri and M. longiterga is notably exaggerated and very acute. Both species of this subgroup also have elongate paramere appendages and inferior appendages without a distinct ventromesal projection.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDFF820FF01B8A64229F94F.taxon	description	Fig. 22	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDFF820FF01B8A64229F94F.taxon	description	Adult — Length of forewing: male 6.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color brownish-black (fuscous). Tibial spurs slightly darker than legs, but not greatly contrasting in color. Forewing without white wing bars, but anastomosis distinct because of unpigmented veins. Male genitalia — Ventral process of segment VI posteriorly projecting, moderately prominent, narrow basally, length about 2 - 2 ½ times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX (apparently) elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin broadly rounded; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, basal part distinctly inflated and set off from short sclerotized apical part, laterally with very short rounded ventrolateral lobes, each with 1 or 2 prominent setae; apex of tergum distinctly sclerotized, with ventrolateral margins incurved and converging mesally, mesal notch (apparently) either small or nearly absent; tergum ventromesally with paired, lightly sclerotized ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes, ventromesal lobe either very short or absent. Mesal pockets of inferior appendage with very short curved, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, slightly widened preapically, apex acute, extending about same length as dorsal phallic spine; fused basal segments of appendage articulating near base of dorsal phallic spine. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow, with dorsal margin curved basally, very strongly upturned in apical 1 / 3, slightly deflected before apical inflection, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly and strongly widened on ventral margin at about basal 1 / 2, forming very acute, spine-like, ventral projection, narrowing apically from projection. Phallicata with sclerotized basodorsal projection, articulating with spine-like ventral projection of dorsal phallic spine, sclerotization extending dorsally from spine and conforming to ventral margin of dorsal phallic spine; phallicata ventrally with pair of sclerotized and projecting basal lobes, ventral sclerotization not extending much beyond lobes. Endophallic membrane simple in structure, without apparent membranous lateral lobes; small phallotremal sclerite present, but without distinct spines. Material examined — COLOMBIA: Cundinamarca: Monterredonda, 10. xii. 1958, JE Foerster – 1 male Paratype (pinned) (NMNH). Distribution — Colombia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDFF82EFF01BF26418EFE4F.taxon	description	Fig. 23 This species is most closely related to M. foersteri (Schmid) and like that species has the posterior margin of segment IX broadly rounded and the ventral margin of the dorsal phallic spine very angularly developed. It is distinguished from M. foersteri by its distinctly longer tergum X and also by its smaller size. Adult — Length of forewing: male 4.7 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color brownish-black (fuscous). Tibial spurs slightly darker than legs, but not greatly contrasting in color. Forewing without white wing bars. Male genitalia — Ventral process of segment VI posteriorly projecting, moderately prominent, narrow basally, length about 2 - 2 ½ times width at base, apex acute. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin broadly rounded; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, basal part slightly inflated and set off from elongate sclerotized apical part, laterally with very short rounded ventrolateral lobes, each with 1 or 2 prominent setae; tergum, as viewed dorsally, very elongate, narrow, lateral margins subparallel, apices truncate; apex of tergum distinctly sclerotized, with ventrolateral margins incurved and nearly converging mesally, mesal notch small, but distinct, dorsally, with short U-shaped connection; tergum ventromesally with paired, lightly sclerotized, ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes, ventromesal lobes absent. Mesal pockets of inferior appendage with short, posteriorly-curved, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, extending about same length as dorsal phallic spine; fused basal segments of appendage articulating near base of dorsal phallic spine. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively narrow, dorsal margin nearly straight from base, sinuously deflected and then strongly, nearly rectilinearly, upturned in apical 1 / 4, apex of spine rounded; base of spine narrow and stalk-like, abruptly and strongly widened on ventral margin at about basal half, forming acute spine-like ventral projection, narrowing apically from projection; as viewed dorsally, nearly uniformly narrow throughout length. Phallicata with sclerotized basodorsal projection, articulating with spine-like ventral projection of dorsal phallic spine, sclerotization extending dorsally from spine and conforming to ventral margin of dorsal phallic spine; phallicata ventrally with pair of projecting sclerotized basal lobes, sclerotization not extending beyond lobes. Endophallic membrane with pleated membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFDFF82EFF01BF26418EFE4F.taxon	materials_examined	Holotype male (pinned) — ECUADOR: Pichincha: 2.3 km S Tandayapa, 1800 m, 6. ix. 1990, OS Flint, Jr (UMSP 000157312) (NMNH). Etymology — This species is named M. longiterga for its very elongate tergum X, which helps to distinguish it from M. foersteri. — hodgesi subgroup Included species: Mortoniella hodgesi Flint. This species was included in the bilineata subgroup by Sykora, but is removed and placed in its own subgroup here because of its unusual combination of characters. The only specimens examined were from alcohol, but the color is apparently brownish black (fuscous), without forewing wing bars. Males of this species have tergum X very deeply invaginated mesally, without the evident enrolling of the lateral margins typically found in the apiculata, bilineata, and enchrysa subgroups. Like species in the M. apiculata and M. enchrysa groups, the ventral process of segment IV is very small. Otherwise, the angular posterolateral margin of segment IX and the dorsal phallic spine, with a very distinctly produced ventral margin (although rounded and not angular), and with the apical part bent at nearly a right angle and with the apex rounded, as viewed laterally, is very similar to species of several subgroups, including the bilineata, apiculata, and enchrysa subgroups. The primary feature distinguishing this species group and suggesting a relatively basal position in the entire bilineata group clade, is the structure of the female genitalia (Fig. 109), which is rather minimally modified, with the usual mesal notch in tergum VIII indicated only by being lightly sclerotized, and without an anterior projection of tergum IX, which normally extends into the mesal notch of tergum VIII in other members of the bilineata group. Male characters that are unusual include a basal part of tergum X that is not inflated or distinctly separated from the apical part; elongate, curved apicolateral processes of tergum X with very elongate setae; and very minimally developed ventromesal processes of tergum X, which lack a spatulate mesal projection and are retracted so that they are not directly visible in lateral view.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD1F82DFF01BA264570FBAF.taxon	description	Fig. 24, 109	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD1F82DFF01BA264570FBAF.taxon	description	Adult — Length of forewing: male 4.7 mm; female 5.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color (in alcohol) dark brown (fuscous), tibiae and tarsi slightly paler. Tibial spurs darker than tibiae and tarsi, contrasting in color. Forewing without apparent wing bands. Head noticeably small. Male genitalia — Ventral process of segment VI posteriorly projecting, short and very narrow, length about 3 times width at base, process without sclerotized posterolateral line (in specimens examined). Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with distinctly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins broadly curved in dorsal view, ventrolateral lobes acute, tapering ,,, each with prominent apical seta; apex of tergum, in dorsal view with deep U-shaped mesal invagination, extending nearly ½ length of segment, apicolaterally with distinctly sclerotized and ventromesally curved projections, dorsum of tergum with prominent elongate, curved setae; tergum ventromesally with paired, lightly sclerotized, ventromesal lobes with short setae, lobes retracted and not directly visible in lateral view. Inferior appendages with very short rounded dorsolateral lobes and single tapering, deltoid ventromesal lobe. Mesal pockets of inferior appendage with short, posteriorly-directed, spine-like, apicoventral projections, projecting somewhat below ventral margin of phallicata. Paramere appendage relatively short, nearly uniform in width, apex acute, extending about same length as ventral lobes of phallicata; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase with small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin curved and arched from base, sinuously and nearly rectilinearly upturned in apical 1 / 3, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly and strongly widened on ventral margin in basal ½, forming rounded ventral projection, narrowing apically from projection; spine, as viewed dorsally, nearly uniformly narrow in width throughout length. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine, and with lightly sclerotized dorsal area extending into endophallic membrane; phallicata ventrally with moderately elongate, lightly sclerotized, apically rounded, lobes, extending about same length as paramere appendages. Endophallic membrane inflated, but without distinct lateral lobes, ventromesally with lightly sclerotized projection; phallotremal spines absent. Material examined — ECUADOR: Napo: 5 mi S Antisana, 13500 ft., 28. iv. 1958, RW Hodges, male Holotype (U. S. N. type 66020, pharate adult, alcohol) (NMNH); Reserva Ecologica Antisana, streams draining Crespo Glacier, 9.6 mi SE Secas, 0.53472 ° S, 78.225560 ° W, 18. i. 2012, B Kondratieff – 1 male, 1 female (alcohol) (NMNH). Distribution — Ecuador. — iridescens subgroup Included species: Mortoniella iridescens Flint. This species was originally placed in the bilineata subgroup by Sykora (1999) and was also speculatively placed in that subgroup by Blahnik and Holzenthal (2011), mostly based on the presence of 2 wing bars. However, as noted by Flint in its original description, the wing bars are unusual in that they more or less disappear when the light source is directly overhead, and only appear when the light strikes the wing at an angle. The color of the wing bars is a brilliant iridescent turquoise. Species in the bilineata subgroup with white wing bars may also have a slight turquoise iridescence at some light angles, but direct examination of the genitalia of M. iridescens makes its placement in the bilineata subgroup problematic, since it lacks some of the defining characters of that subgroup and has some unusual features of its own, notably the shape of tergum X, which is short and peculiarly truncate apically, as viewed laterally, and also has a distinct mesal notch. Other features of the genitalia, such as the rounded posterior margin of segment IX, simple inferior appendages, with absence of a mesal process, and the very acute ventral projection on the dorsal phallic spine suggest a relationship with the foersteri subgroup, which Sykora also included in the bilineata subgroup. We are presently placing M. iridescens in its own species group to draw attention to its unusual combination of characters.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD2F82CFF01BC06426BF92F.taxon	description	Fig. 25	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD2F82CFF01BC06426BF92F.taxon	description	Adult — Length of forewing: male 4.8 - 5.1 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown (fuscous). Legs with basal part of mesotibiae and apices of mesotarsi somewhat paler. Tibial spurs slightly darker than legs, not greatly contrasting in color. Forewing with 2 iridescent turquoise wing bands, 1 at anastomosis and 1 on proximal part of wing, bands only visible at oblique angle, disappearing with light directly overhead. Male genitalia — Ventral process of segment VI posteriorly projecting, prominent, length about 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin broadly rounded; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X short, basal part inflated and set off from apical part; overall shape, as viewed dorsally, subquadrate, with distinct small U-shaped apicomesal invagination, tergum laterally with short rounded ventrolateral lobes, each with prominent apical setae; apex of tergum, in lateral view, distinctly sclerotized, subtruncate, with ventrolateral margins incurved and nearly converging mesally, mesal notch distinct; tergum ventromesally with paired, lightly sclerotized, ventromesal lobes at about midlength, each with short setae. Inferior appendages with short rounded setose dorsolateral lobes, appendages constricted ventromesally, without mesal projection. Mesal pockets of inferior appendage with short, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage short, narrow, nearly uniform in width, apex acute; fused basal segments of appendage articulating near middle of basal stem of dorsal phallic spine. Phallobase with evident rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin curved basally, then extending nearly straight and obtusely angled in apical ¼, apex of spine rounded; base of spine narrow, curved and stalk-like, abruptly and strongly widened on ventral margin at about midlength, forming very acute ventral projection, narrowing apically from projection; spine, as viewed dorsally, distinctly widened at ventral projection, narrowing basally and apically, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata lightly sclerotized ventrally, sclerotization narrowly extending mesally. Endophallic membrane with welldeveloped membranous dorsolateral lobes, and smaller ventrolateral lobes with minute spines; phallotremal spines absent. Material examined — COLOMBIA: Antioquia: 12 km N Fredonia, 2000 m, 22. ii. 1983, OS Flint, Jr – 2 male Paratypes (pinned) (NMNH). Distribution — Colombia. — quinuas subgroup Included species: Mortoniella gilli, n. sp. and M. quinuas Harper and Turcotte. Mortoniella quinuas Harper and Turcotte was placed in the flinti subgroup by Sykora (1999), possibly because of the contour of the dorsal phallic spine, which lacks an angular ventral projection. However, the species is distinctly different from species in the flinti subgroup. We have placed the two species considered here in their own subgroup, even though it is conceivable that the two subgroups may be related. Mortoniella quinuas does seem to be correctly placed in the bilineata group, based on overall structural considerations. Female genitalia, which are distinctive for the group, would help to confirm this, but are not known or not available for confirmation. The second species described in this subgroup, M. gilli, n. sp., has a similarly developed dorsal phallic spine, with the apex very strongly recurved and covered with minute spines, and apparently also has a similarly shaped tergum X, with projecting apicolateral lobes. Both species also have elongate spine-like projections from the mesal pockets of the inferior appendages. The structure of the inferior appendages seems to be very different in the two species. Nevertheless, their relationship seems likely. Unfortunately, the holotype of M. quinuas, which was reillustrated by Sykora, could not be located for direct comparison. Mortoniella gilli lacks ventromesal processes on tergum X and also lacks an elongate membranous extension between segments VIII and IX. It is likely that the same is true for M. quinuas.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD3F82BFF01BE8643E9FA4F.taxon	description	Fig. 26 This species is most closely related to M. quinuas Harper and Turcotte. Both species have a dorsal phallic spine with a strongly dorsally curved apex covered with minute spines, but lack the acute or spine-like ventrolateral lobes of tergum X, typical of most species in the bilineata group. Mortoniella gilli differs significantly from M. quinuas in having paramere appendages with numerous small spines in the apical half, rather than a single apical spine, and in having very differently formed inferior appendages, which lack spiniform processes. Also, it possesses a very prominent, posteroventrally-oriented, ventral process on segment VI, rather than a small weakly formed one. Adult — Length of forewing: male 5.0 - 5.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color (in alcohol) brown. Legs paler than body, tibial spurs very dark, contrasting with legs. Forewing largely denuded, apparently without wing bars. Head noticeably small. Male genitalia — Ventral process of segment VI posteroventrally projecting, prominent, length about 2 times width at base, apex acute. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX normal (not elongate). Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin weakly produced and rounded in dorsal ½; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X, as viewed dorsally, moderately elongate, basal part very slightly inflated and set off from sclerotized apical part, lateral margins subparallel, apex with deep V-shaped mesal invagination, extending almost ½ length of segment; as viewed laterally, with rounded setose ventrolateral lobes at about midlength, apex of tergum distinctly sclerotized and forming bluntly rounded and slightly ventrally curved apical projections, ventromesal lobes absent. Inferior appendages prominent, setose, moderately elongate, without distinct dorsolateral lobe; as viewed ventrally, without mesal projection (lateral appendages separated by deep mesal invagination). Mesal pockets of inferior appendage with elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, bowed outward in apical ½, apical part with numerous spines on lateral margin; fused basal segments of appendages articulating near base of dorsal phallic spine. Phallobase without dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, nearly uniform in width, slightly ventrally bowed in basal 2 / 3, conforming to sclerotized dorsal margin of phallicata, apical 1 / 3 with sinuous deflection and strong apical dorsal inflection, apex acutely narrowed and with numerous small spines; spine, as viewed dorsally, nearly uniformly narrow in width throughout length, apex acute. Phallicata without basodorsal projection, sclerotized dorsal margin extending into endophallic membrane, with nipple-like projection apically, conforming to sinuous apical inflection of dorsal phallic spine; phallicata lightly sclerotized ventrally, extending about as far as paramere appendages, lateral margins bowed outward and converging apically. Endophallic membrane with projecting and pleated membranous lateral lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD3F82BFF01BE8643E9FA4F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Napo: PAP 8 unnamed trib., Papallacta River, Hwy E- 28, ca. 1.7 mi SW Papallacta, 0.385893 ° S, 78.143530 ° W, 25. i. 2012, B Gill (NMNH) (UMSP 000097165). Paratypes — ECUADOR: Napo: same data as holotype, 2 males (alcohol) (NMNH). Etymology — This species is named M. gilli after its collector.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD4F82AFF01BE26442AFDEF.taxon	distribution	Distribution — Ecuador. — wygodzinskii subgroup Included species: Mortoniella wygodzinskii (Schmid, 1958). This species subgroup, with a single contained species, was recognized by Sykora (1999) and is retained here because of its unusual morphology and lack of characters that would clearly place it in one of the other species groups. Mortoniella wygodzinskii is a uniform brownish in color, without wing bars. The species has a very distinct ventromesal process of tergum X, as in species of the apiculata, bilineata, enchrysa, foersteri, and iridescens subgroups. The other modifications of tergum X, with the apicolateral angles distinctly sclerotized and enrolled ventrally to form an apical cap, but with a distinct mesal notch, resembles species in the enchrysa, foersteri, and iridescens subgroups. The posterior margin of segment IX is slightly produced, but this is neither distinctly angular nor broadly rounded, as in members of the other species groups mentioned above.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD5F82AFF01BAC64392FB6F.taxon	description	Fig. 27	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD5F82AFF01BAC64392FB6F.taxon	materials_examined	Material examined — VENEZUELA: Trujillo: Quebrada Potrerito, 7.5 km NE Boconó, 9.27392 ° N, 70.21837 ° W, 1530 m, 29 - 30. iv. 1995, Holzenthal, Cressa, Gulic – 1 male (pinned) (UMSP). Distribution — Argentina, Bolivia, Ecuador, Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD7F837FF01B8A64244FECF.taxon	description	Fig. 28, 110	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFD7F837FF01B8A64244FECF.taxon	description	Adult — Length of forewing: male 2.9 - 3.6 mm; female 3.5 - 4.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Tibial spurs somewhat darker than legs, not strongly contrasting in color. Wing bar absent in male, forewing sometimes marked with whitish setae at arculus in female. Male with both surfaces of fore- and hind wings densely covered with short prostrate scale-like setae (sometimes lost in alcohol); female with unmodified setation. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Tergum VIII relatively wide (nearly as wide as previous segment), subtending ventral margin of segment IX, anterior margin of tergum with evident apodeme, posterior margin densely setose; membranous connection to tergum IX elongate, ballooned when expanded, surface slightly textured. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin very slightly projecting, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short and strongly sclerotized, with deep U-shaped mesal excision, extending more than ½ length of segment; mesally with short sclerotized, apically rounded, ventral projection, apparently articulating with dorsal phallic spine. Inferior appendages with tapering setose dorsolateral lobes and elongate, narrow, tapering, ventral lobes. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages relatively short, widely forked near base, forming 2 narrow, apically acute projections, ventral projection longer than dorsal one; fused basal segments of paramere articulating with dorsal phallic spine before sinuous middle flexure. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, more or less uniform in width, broadly S-shaped over its length, apex acutely narrowed and upturned; in dorsal view, nearly uniform in width throughout length. Phallicata very short, with short sclerotized dorsal projection and short rounded ventral lobe. Endophallic membrane short, with prominent pair of strongly sclerotized, spine-like sclerites on basodorsal margin (possibly phallotremal sclerites), and very small mesal spine distal to basal sclerites; ventrally with short curved, lightly sclerotized, ventromesal spine. Material examined — BOLIVIA: La Paz: AMNI Madidi, Río Tuichi and tributary at entrance to Chalalan lodge, 14.41695 ° S, 67.90630 ° W, 300 m, 27. vii. 2003, Robertson and Blahnik – 1 male (pinned) (UMSP); Rayo Mayo river at Wabacuro trail, Chalalan Ecolodge, 14.44257 ° S, 67.91095 ° S, 351 m, 28. vii. 2003, Robertson and Blahnik – 1 male, 1 female (pinned) (NMNH); Santa Cruz: Saaveda Exp. Station, 1. iv. 1960, R Cumming – 1 male, 2 females (alcohol) (NMNH); PN and ANMI Amboró, Guardia Parque Mataracú, Río Yapacaní, 17.52072 ° S, 63.86795 ° W, 329 m, 26. xi. 2004, Robertson, Garcia, Vidaurre – 8 males, 8 females (alcohol) (UMSP); PERU: Yurac, 67 mi E Tingo Maria, EI Schlinger and ES Ross, 28. ix. 1954 – 2 males, 1 female (alcohol) (CAS); Avispas, -. x. 1962, LE Peña G – 1 male (alcohol) (NMNH); Cuzco: Pilcopata, premontane moist forest, 600 m, 8 - 10. xii. 1979, JB Heppner – 2 males, 3 females (alcohol) (NMNH); Madre de Dios: Manu, Pakitza Biological Station, Quebrada Paujil-Picoflor, 11.94417 ° S, 71.28300 ° W, 350 m, 2. vii. 1993 Blahnik and Pescador – 1 male (pinned) (NMNH); Manu, Erika (near Salvacion), 12.88333 ° S, 71.23333 ° W, 550 m, 4 - 6. ix. 1988, O Flint and N Adams – 6 males, 15 females (alcohol) (NMNH); Hostel Erica (near Salvacion), 12.88333 ° S, 71.23333 ° W, 3 - 5. ix. 1989, RA Faitoute, et al. – 8 males, 6 females (alcohol) (NMNH); same locality and date, N Adams et al. – 8 males, 8 females (alcohol) (NMNH); Amazonia Lodge, Río Alto Madre de Dios, 12.87033 ° S, 71.37600 ° W, 500 m, 30. vi. 1993, R Blahnik and M Pescador – 5 males, 4 females (alcohol) (NMNH); Pasco: Puerto Bermudez, Río Pichis, 10.2964 ° S, 74.9364 ° W, 15. vii. 1920 – 3 males (alcohol) (NMNH). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC8F836FF01B9A64183FD2F.taxon	description	Fig. 29, 112 Mortoniella brevis, n. sp. is perhaps most similar overall to M. atenuata Flint, especially because of its similarly shaped paramere appendages, which are broadly forked, with the ventral projection of the fork more elongate. Mortoniella brevis is the only species in the atenuata subgroup to have inferior appendages very short, without elongate ventral projections and thus is easily diagnosed based on this character. As an additional character difference, the apicolateral lobes of tergum X are also more rounded in M. brevis than in M. atenuata. Adult — Length of forewing: male 3.0 - 3.5 mm, female 3.4 - 4.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color (in alcohol) medium brown. Tibial spurs somewhat darker than legs, not strongly contrasting in color. Wing bar absent in male, forewing sometimes marked with whitish setae at arculus in female. Male with both surfaces of fore- and hind wings densely covered with short prostrate scale-like setae; female with unmodified setation. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Tergum VIII relatively wide (nearly as wide as previous segment), subtending ventral margin of segment IX, anterior margin of tergum with evident apodeme, posterior margin densely setose; membranous connection to tergum IX elongate, ballooned when expanded, surface distinctly textured. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin very slightly projecting, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short and strongly sclerotized, with deep U-shaped mesal excision, extending more than ½ length of segment; mesally with short sclerotized, apically rounded, ventral projection, apparently articulating with dorsal phallic spine. Inferior appendages with tapering setose dorsolateral lobes and very short acute ventral lobes. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages relatively short, widely forked near base, forming 2 narrow, apically acute projections, ventral projection longer than dorsal one; fused basal segments of paramere articulating with dorsal phallic spine before sinuous middle flexure. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, more or less uniform in width, broadly S-shaped over its length, apex acutely narrowed and upturned; in dorsal view, nearly uniform in width throughout length. Phallicata very short, with short sclerotized dorsal projection and short rounded ventral lobe. Endophallic membrane short, with prominent pair of strongly sclerotized, spine-like sclerites on basodorsal margin (possibly phallotremal sclerites), and very small mesal spine distal to basal sclerites; ventrally with short curved, lightly sclerotized, ventromesal spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC8F836FF01B9A64183FD2F.taxon	materials_examined	Holotype male (alcohol) — VENEZUELA: Barinas: Río Singüis in Caño Grande, 8.40000 ° N, 70.77417 ° W, 520 m, 22. iii. 1997, Holzenthal (UMSP 00092465) (UMSP). Paratypes — ECUADOR: Napo: Puyo, 6. v. 1977, PJ Spangler and DR Givens – 1 male, 2 females (alcohol) (NMNH); Puerto Nuevo, 8. vii. 1976, J Cohen – 1 male, 1 female (alcohol) (NMNH); Tena (17 km SW), 28. v. 1977, PJ Spangler and DR Givens – 3 males, 21 females (alcohol) (NMNH); Río Misahuallí, Archidona, 650 m, 11. ix. 1990, OS Flint, Jr – 1 male, 1 female (pinned) (NMNH); Pastaza: Puyo, 5. v. 1977, PJ Spangler and DR Spangler – 4 males, 11 females (alcohol) (NMNH); same locality and collectors, 6. v. 1977 – 1 male (alcohol) (NMNH); same locality and collectors 10. v. 1977 – 2 males (alcohol) (NMNH); same locality and collectors, 13. v. 1977 – 9 males, 16 females (alcohol) (NMNH); same locality and collectors, 14. v. 1977 – 1 male, 2 females (alcohol) (NMNH); same locality and collectors, 16. v. 1977 – 1 male, 3 females (alcohol) (NMNH); same locality and collectors, 21. v. 1977 – 5 males, 7 females (alcohol) (NMNH); Puyo, 30. i. 1976, Spangler et al. – 15 males, 3 females (alcohol) (NMNH); same locality and collectors, malaise trap, 1 - 7. ii. 1976 – 1 male (alcohol) (NMNH); Puyo (27 km N) Est. Fluvia Metrica, 4. ii. 1976, Spangler et al. – 2 males (alcohol) (NMNH); VENEZUELA: Barinas: same data as Holotype – 2 males (pinned), 8 males (alcohol) (UMSP), 4 males, 6 females (alcohol) (MIZA); Río Santo Domingo, 17. ii. 1976, CM and OS Flint, Jr – 40 males, 29 females (alcohol) (NMNH); Zulia: El Tucuco, Sierra de Perija, montane forest, 28 - 29. i. 1978, JB Heppner – 1 male (alcohol) (NMNH); Perijo El Tucuco, Missíon El Tucuco, Río El Tucucu, 1 / 2 km from church, 1 - 5. x. 1979, HM Savage – 4 males (alcohol) (NMNH); El Tucuco (45 km SW of Machiques), 5 - 6. vi. 1976, AS Menke and D Vincent – 1 male, 10 females (alcohol) (NMNH). Etymology — This species is named M. brevis, from the Latin word brevis, meaning short, in reference to the short inferior appendages of this species, which helps distinguish it from closely related species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC9F835FF01BA86412BFECF.taxon	description	Fig. 30 Mortoniella dinotes n. sp. is most similar to M. atenuata Flint in having inferior appendages with elongate ventral projections and noticeably forked paramere appendages. However, the ventral projections of the inferior appendages are more distinctly curved and rounded basally in M. dinotes and have a less prominent, upright, dorsal processes. Additionally, the paramere appendages in M. dinotes are more narrowly forked and the dorsal branch is very short. Adult — Length of forewing: male 3.2 - 3.4 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Tibial spurs somewhat darker than legs, not strongly contrasting in color. Wing bar absent in male. Male with both surfaces of fore- and hind wings densely covered with short prostrate scale-like setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Tergum VIII relatively wide (nearly as wide as previous segment), subtending ventral margin of segment IX, anterior margin of tergum with evident apodeme, posterior margin densely setose; membranous connection to tergum IX elongate, ballooned when expanded, sur-face noticeably textured. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin very slightly projecting, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short and strongly sclerotized, with deep U-shaped mesal excision, extending more than ½ length of segment; mesally with short sclerotized, apically rounded, ventral projection, apparently articulating with dorsal phallic spine. Inferior appendages with very short, nearly obsolete, dorsolateral lobes and very strongly curved, elongate, tapering, ventral lobes. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages moderately elongate, narrowly forked near base, forming 2 narrow, apically acute projections, dorsal projection very short, ventral projection elongate; fused basal segments of paramere articulating with dorsal phallic spine before sinuous middle flexure. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, more or less uniform in width, broadly S-shaped over its length, apex acutely narrowed and upturned; in dorsal view, nearly uniform in width throughout length. Phallicata very short, with short sclerotized dorsal projection and short rounded ventral lobe. Endophallic membrane short, with prominent pair of strongly sclerotized, spine-like, sclerites on basodorsal margin (possibly phallotremal sclerites), and very small mesal spine distal to basal sclerites; ventrally with short curved, lightly sclerotized, ventromesal spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC9F835FF01BA86412BFECF.taxon	materials_examined	Holotype male (pinned) — PERU: Huanuco: Tingo Maria, premontane rain forest, 672 m, 1 - 6. ii. 1980, JB Heppner (UMSP 000118533) (MJP). Paratypes — PERU: Huanuco: same data as Holotype – 2 males (alcohol) (NMNH). Etymology — This species is named M. dinotes, from the Greek word dinotos, meaning turned or rounded, and referring to the strongly rounded ventral projections of the inferior appendages in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCAF834FF01B9A64214FAEF.taxon	description	Fig. 31	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCAF834FF01B9A64214FAEF.taxon	materials_examined	Mortoniella leei is perhaps most similar overall to M. atenuata Flint, differing most distinctly in the shape of its paramere appendages. In its original description, it was differentiated from M. atenuata by having a short curved, unbranched, paramere appendage. However, in material examined (which did not include the holotype) the appendage seems to be consistently branched, with the ventral branch very narrow and hyaline or very lightly sclerotized (and thus easily overlooked). The shape and length of this ventral branch varies, and it is possible that it is absent in some specimens. The short, thick, curved, dorsal branch is distinctive for this species. The inferior appendages are more elongate than in M. brevis, n. sp., but the apices of the appendages are only lightly sclerotized, and thus may not be readily noticed. This is in contrast to the distinctly sclerotized apices of the inferior appendages in M. atenuata and M. dinotes, n. sp. The characteristic difference in the structure of the paramere appendage is diagnostic, and generally evident even in uncleared specimens. Adult — Length of forewing: male 2.9 - 3.4 mm, female 3.4 - 3.8 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color medium brown. Tibial spurs somewhat darker than legs, not strongly contrasting in color. Wing bar absent in male, forewing sometimes marked with whitish setae at arculus in female. Male with both surfaces of fore- and hind wings densely covered with short prostrate scale-like setae; female with unmodified setation. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Tergum VIII relatively wide (nearly as wide as previous segment), subtending ventral margin of segment IX, anterior margin of tergum with evident apodeme, posterior margin densely setose; membranous connection to tergum IX elongate, ballooned when expanded, surface slightly textured. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin very slightly projecting, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short and strongly sclerotized, with deep U-shaped mesal excision, extending more than ½ length of segment; mesally with short sclerotized, apically rounded, ventral projection, apparently articulating with dorsal phallic spine. Inferior appendages with setose, apically rounded, dorsolateral lobes and elongate, narrow, tapering, lightly sclerotized, ventral lobes. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages short, forked near base, forming 2 apically acute projections, ventral projection longer than dorsal one, very narrow and lightly sclerotized (easily overlooked), dorsal projection short, ventrally curved, and strongly sclerotized; fused basal segments of paramere articulating with dorsal phallic spine before sinuous middle flexure. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, more or less uniform in width, broadly S-shaped over its length, apex acutely narrowed and upturned; in dorsal view, nearly uniform in width throughout length. Phallicata very short, with short sclerotized dorsal projection and very short rounded ventral lobe. Endophallic membrane short, with prominent pair of strongly sclerotized, spine-like, sclerites on basodorsal margin (possibly phallotremal sclerites), and very small mesal spine distal to basal sclerites; ventrally with short curved, lightly sclerotized, ventromesal spine. Material examined — COLOMBIA: Antioquia: Río Claro, 5.59000 ° N, 75.86720 ° W, 3. v. 1984, U Matthias – 1 male (alcohol) (NMNH); Chocó: Río Atrato, Yuto, 18. ii. 1983, OS Flint, Jr – 2 males, 1 female (pinned), 13 males, 2 females (alcohol) (NMNH); Valle de Cauca: Río Raposo, - iii. 1965, 3.8933 ° N, 77.0697 ° W, VH Lee – 40 males, 10 females (Paratypes-alcohol) (NMNH); ECUADOR: Cotopaxi: Latacunga, 133 km W, 1080 m, 2. vii. 1975, Langley and Cohen – 28 males, 1 female (alcohol) (NMNH); Quevedo (36 km NE), 21. vii. 1976, 335 m, J Cohen – 3 males, 5 females (alcohol) (NMNH); El Oro: 9 mi S Santa Rosa, 3.45000 ° S, 79.96667 ° W, 23. i. 1955, EI Schlinger and ES Ross, 23. i. 1955 – 8 males, 24 females (alcohol) (CAS); Canton de Arenillas, Las Lajas, 600 m, 30. v. 1979, JJ Anderson – 1 male (alcohol) (NMNH); Pasaje (6 km E), 13. i. 1978, PJ Spangler and J Anderson – 2 males, 1 female (pinned), 7 males, 5 females (alcohol) (NMNH); Loja: Río Puyango, 300 m, 17 - 18. viii. 1977, LE Peña G – 32 males, 15 females (alcohol) (NMNH); Los Rios: Quevedo (56 km N), Río Palenque Biological Station, 1.03306 ° S, 79.45000 ° W, 250 m, 28 - 29. vii. 1976, J Cohen – 93 males, 1 female (alcohol) (NMNH); Quevedo (11 km S), 3. vii. 1975, Langley and Cohen – 1 male, 2 females (alcohol) (NMNH); Manabi: Santo Domingo (29 km SW), Rancho Ronald, 20. vii. 1976, JJ Anderson – 1 male (alcohol) (NMNH); Pichincha: Santo Domingo (47 km S), Río Palenque Biol. Station, 750 m, 29. vii. 1976, J Cohen – 254 males (alcohol) (NMNH); Tungurahua: Yanayacu, 300 m, 29 - 30. viii. 1977, LE Peña G – 3 males (alcohol) (NMNH). Distribution — Colombia, Ecuador. — bolivica subgroup Included species: Mortoniella bolivica (Schmid); M. flexuosa, n. sp.; and M. spatulata, n. sp. Only three closely related species are included in this subgroup. The species have retained fork III in the hind wing and have inferior appendages with an elongate and more or less recurved dorsal process, both probably plesiomorphic characters. They also lack basodorsal processes on the phallicata. Paired ventrolateral spines are present on the endophallic membrane, but a ventromesal spine is absent. It seems likely that these are alternate character states representing the same character development. The species have a general similarity to members of the punensis subgroup in having a dorsal phallic spine that is somewhat expanded and compressed (dorsoventrally flattened) apically and in the form of tergum X, which has the apical lobes separated by a nearly horizontal mesal invagination. The shape of the ventral process of segment VI, which is short, but posteriorly directed, is also similar. Among species in the leroda group, only M. guyanensis, n. sp., in the limona subgroup, has similarly shaped inferior appendages; reasons for assigning M. guyanensis to the limona subgroup are given in the discussion of that subgroup. It is easily distinguished from species in the bolivica subgroup by the presence of small apical spines on the dorsal phallic spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCBF833FF01BDC64257FC0F.taxon	description	Fig. 32	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCBF833FF01BDC64257FC0F.taxon	description	Adult — Length of forewing: male 4.4 - 5.0 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color light brown. Tibial spurs slightly darker in color, contrasting with legs. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae. Male genitalia — Ventral process of segment VI laterally compressed, very large, subtriangular, posteroventrally projecting, width at base nearly as long as segment, subequal to length. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X with excision between lateral lobes nearly linear; apicolateral lobes distinctly sclerotized, acute, curved inward from lateral margins, mesal margins subparallel. Inferior appendages relatively large and prominent, setose, nearly completely divided ventromesally, apicolateral angles acute, dorsolateral lobes narrow and posteriorly recurved, apices acute. Mesal pockets of inferior appendage with apical processes relatively short, dorsally curved. Paramere appendages moderately elongate, narrow, extending about as far as apical inflection of dorsal phallic spine. Dorsal phallic spine, as viewed laterally, nearly uniform in width, narrowing apically, base distinctly curved, apex weakly and obtusely upturned in about apical ¼ or 1 / 5; in dorsal view, with apical inflection distinctly widened (dorsoventrally flattened), apex irregularly serrate or bifurcate (Fig. 32 D and inset). Phallicata with dorsal margin weakly sclerotized, apparently to accommodate dorsal phallic spine. Endophallic membrane moderately elongate and relatively simple, with small membranous apical and preapical lobes, ventrally with pair of distinctly sclerotized, curved spines; phallotremal spines absent. Material examined — BOLIVIA: Coroico, 3 - 13. xii. 1955, LE Peña G – 2 male Paratypes (pinned) (NMNH); PERU: Cuzco: Paucartambo to Pilcopata rd., river at Puente Unión, 13.07033 ° S, 71.56667 ° W, 1670 m, 21 - 23. vi. 1993, Blahnik and Pescador – 2 males (pinned) (NMNH); Paucartambo to Pilcopata rd., Río San Pedro at Puente San Pedro, 13.05500 ° S, 71.64633 ° W, 1445 m, 24. vi. 1993, Blahnik and Pescador – 1 male (pinned) (UMSP). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCCF832FF01BC66418BFCAF.taxon	description	Fig. 33, 111 This species is very similar to M. bolivic a (Flint), as discussed in the diagnosis for that species, and the two form a closely related species pair. The species are unusual for members of the M leroda species group in having an elongate, reflexed, dorsolateral lobe on each of the inferior appendages. That of M. flexuosa is diagnostic in being more elongate and sinuously bent, and the apicoventral projection of the inferior appendage is also much more acute. Mortoniella flexuosa also has a small, sclerotized projection or tubercle on the phallicata that is apparently absent in M. bolivica. There is some size variation in the material examined. The holotype appears as in the illustration; the paratypes, which include the female specimens, are smaller in size and have the ventral projection of the inferior appendages slightly shorter. There are no other significant structural differences and we infer the differences to be populational variation. Adult — Length of forewing: male 3.8 - 4.8 mm; female 4.1 - 4.6 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color light brown. Tibial spurs slightly darker in color, contrasting with legs. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae. Male genitalia — Ventral process of segment VI laterally compressed, very large, subtriangular, posteroventrally projecting, width at base nearly as long as segment, subequal to length. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X with excision between lateral lobes nearly linear; apicolateral lobes distinctly sclerotized, acute, curved inward from lateral margins, mesal margins subparallel. Inferior appendages relatively large and prominent, setose, nearly completely divided ventromesally, apicolateral angles projecting and very acutely narrowed, dorsolateral lobes elongate, narrow, sinuous, posteriorly recurved, apices acute. Mesal pockets of inferior appendage with apical processes relatively short, dorsally curved. Paramere appendages moderately elongate, narrow, extending about as far as apical inflection of dorsal phallic spine. Dorsal phallic spine, as viewed laterally, only slightly widened in basal ½, narrowing apically, base distinctly curved, apex weakly and obtusely upturned in about apical 1 / 3 or 1 / 4; in dorsal view, with apical inflection distinctly widened (dorsoventrally flattened), apex asymmetrical and irregularly serrate. Phallicata with dorsal margin weakly sclerotized, apparently to accommodate dorsal phallic spine, laterally with rounded protrusion, apparently to accommodate flexed dorsal lobe of inferior appendages. Endophallic membrane moderately elongate and relatively simple, with small membranous apical and preapical lobes, ventrally with pair of distinctly sclerotized, curved spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCCF832FF01BC66418BFCAF.taxon	materials_examined	Holotype male (pinned) — COLOMBIA: Quindió: Río Quindió, Retén “ La Playa ”, ca. 2 km NE Salento, 4.64028 ° N, 75.55667 ° W, 1890 m, 2. i. 1998, F Muñoz-Q et al. (UMSP 000209628) (UMSP). Paratypes — COLOMBIA: Cauca: Municipio de Belalcazar, Quebrada Tálaga, ca. 14 km N Páez (Belalcazar), 2.70667 ° N, 76.01806 ° W, 1680 m, 22. xii. 1997, F Muñoz-Q et al. – 2 males, 2 females (pinned) (UMSP), 1 male, 1 female (pinned) (NMNH). Etymology — This species is named M. flexuosa from the Latin flexuosus, meaning winding or with many bends, and referring to the sinuously bent dorsolateral process of the inferior appendage in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCDF830FF01BB0641D7FC0F.taxon	description	Fig. 34 Mortoniella spatulata bears a general similarity to the other species of the bolivica subgroup, but the dorsolateral lobes of the inferior appendages are shorter and not as strongly flexed. Also, the dorsal phallic spine, while being widened and spatulate apically, is more nearly symmetrical, the ventrolateral lobes of tergum X are acutely developed (separated from the apicolateral lobes by a distinct notch), and the ventral spines of the endophallic membrane are very weakly developed. The holotype specimen is over cleared and only the genital capsule (segments IX-X and phallic apparatus) is present. Adult — Length of forewing: male 3.8 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color light brown, apices of tarsi and basal segments of antennae slightly paler. Tibial spurs about same color as legs, not contrasting in color. Wing bar at anastamosis relatively indistinct, interrupted, marked with whitish setae. Male genitalia — Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X with excision between lateral lobes slightly concave; apicolateral lobes acute, curved inward from lateral margins, mesal margins subparallel. Inferior appendages only moderately large, setose, incompletely divided ventromesally, apicolateral angles short and acute, dorsolateral lobes narrow and slightly posteriorly recurved, apices acute. Mesal pockets of inferior appendage with apical processes relatively short, dorsally curved. Paramere appendages elongate, narrow, extending about as far as dorsal phallic spine. Dorsal phallic spine, as viewed laterally, distinctly undulate in contour, nearly uniform in width, narrowing apically, base distinctly curved, apex very weakly and obtusely upturned in about apical ¼; in dorsal view, with apex slightly widened and spatulate in shape (dorsoventrally flattened). Phallicata with dorsal margin weakly sclerotized, apparently to accommodate dorsal phallic spine. Endophallic membrane moderately elongate and relatively simple, ventrally with pair of very short, weakly sclerotized spines; phallotremal spines (apparently) present, very small.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCDF830FF01BB0641D7FC0F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Barinas: Parque Nacional Sierra Nevada, Quebrada San Juan in Santa Rosa, 8.46450 ° N, 70.84867 ° W, 1000 m, 21. iii. 1997, Holzenthal (UMSP 000041337) (UMSP). Etymology — This species is named M. spatulata for the dorsal phallic spine of the male, which has its apex flattened and distinctly spatulate in shape. — florica / leroda subgroups included species (South America): Mortoniella bothrops, n. sp.; M. cressae, n. sp.; M. curtispina, n. sp.; M. draconis, n. sp.; M. elongata (Flint); M. furcula, n. sp.; M. grandiloba n. sp., M. ruedae, n. sp.; M. schlingeri, n. sp.; and M. simla (Flint). Blahnik and Holzenthal (2008) defined the leroda and florica subgroups based on species from Central America and commented that the two groups were probably related. As in the majority of the subgroups of the leroda group treated in the current work, the hind wings in both subgroups have both forks II and III present. Also, the dorsolateral projections of the inferior appendages are either absent or greatly reduced in both subgroups, usually appearing as short rounded projections. The latter is considered an apomorphy for the combined subgroups, since a projecting dorsal lobe is probably the plesiomorphic state for the subgenus Mortoniella. Many of the species, especially of the leroda subgroup, have a rather characteristically formed ventral process on segment VI, which is short, ventrally projecting, and with the apex more or less blunt or bluntly rounded. The anterobasal margin of the ventral process is characteristically somewhat retracted (Fig. 38 E). However, many species in the florica subgroup from Central America have the ventral process more angular apically, and not as evidently retracted anteriorly, though always distinctly ventrally deflected. A bluntly rounded ventral process otherwise occurs only in the atenuata subgroup, which is apomorphically distinct, and also in M. guyanensis, n. sp., in the limona subgroup. An additional, although more difficult to define, character uniting the two subgroups is the general structure of the dorsal phallic spine, which has a characteristic undulate or sinuous inflection, probably due to the tendency for the ventral margin of the spine to have a rounded deflection that articulates with a corresponding contour in the dorsal margin of the phallicata. However, this may be a plesiomorphic character for the leroda group as a whole, since a similar character state also occurs in the akantha and bolivica subgroups. Another character, which seems to characterize most of the species of the two subgroups, is that the paramere appendages are displaced ventrally on elongate membranous lobes, with the base of the appendages curved inward. This is already evident in M. leroda, which seems to be a basal, or near basal, species in the combined subgroups. The character is more developed in species with paired dorsolateral processes on the phallicata, especially those of the florica subgroup (eg. Fig. 35 C, 36 C). An additional character similarity of the two subgroups is the structure tergum X, which has apicolateral lobes that are distinctly sclerotized and projecting and have a very reduced or nearly absent ventrolateral lobes. There is also some tendency for the lobes to be flattened and converge mesally, and for the dorsomesal margin to be projecting. Although the majority of species from Central America seemed to fall into 2 groups, based on a combination of characters, in treating the species from South America the distinctness of the two groups is less evident. It therefore seems prudent to combine them here. The primary character difference used to define the two groups was the structure of the inferior appendages; members of florica subgroup have paired symmetrical apical processes on the inferior appendages (long or short), and those of the leroda subgroup have a single asymmetrical apicomesal process. Because the names were used previously, and the primary defining character is diagnostically useful, we have continued to group the species from South America in the two subgroups, while recognizing that they may not be mutually monophyletic. Many of the species of the leroda subgroup from South America have upright dorsal processes on the phallicata, a ventromesal spine on the endophallic membrane, and scabrous apices of the paramere appendages (characters generally found in the florica subgroup of Central America, but absent in the species assigned to the leroda subgroup). These characters occur in addition to the presence of a prominent asymmetric ventromesal process on the inferior appendages, used to characterize the leroda subgroup. Most of the South American species of the leroda subgroup also have the apex of this mesal process scabrous (or sensillate), as in M. meralda (Mosely) and M. panamensis Blahnik and Holzenthal from Central America. It seems likely that an asymmetric ventromesal process on the inferior appendages is a plesiomorphic character that has been subsequently lost in the florica subgroup. This may also explain the presence (retention) of this character in a number of species of the albolineata subgroup from Brazil, and an indication of the relationship of these subgroups. The most likely explanation for the presence of paired rounded or upright basodorsal processes on the phallicata in the majority of species of the florica subgroup is that they represent a lineage from within the leroda subgroup that derived this apomorphy, followed by the subsequent loss of the ventromesal process and gain of apicolateral projections on the inferior appendages. The new species described below as M. cressae (Fig. 39) possibly represents a transitional species (or one that has converged on similar character modifications), with apicolateral projections on the inferior appendages present, but with the ventromesal process much reduced. Absence of a ventromesal spine on the endophallic membrane in various species of the leroda subgroup, especially those from Central America, probably represents a character loss, judging by its general presence (retention) in the florica subgroup and in a number of other subgroups. However, there seems to be no parsimonious solution for its loss, which must have occurred repeatedly. Possibly, the plesiomorphic state was for the spine to be very small, as in M. leroda, or in the species of the atenuata subgroup, making its loss more probable. If redefined in a combined sense to include both members of the leroda and florica subgroups, the composite subgroup would include all of the species from Central America, in addition to the species described below, with the exception of the three species of the akantha subgroup and M. pacuara (Flint) and M. rodmani Blahnik and Holzenthal of the subgenus Nanotrichia. This entire group constitutes a revised definition of the leroda subgroup, and will be referred to subsequently as the leroda subgroup, sensu lato. — “ florica ” subgroup	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCFF83FFF01BC664229FD6F.taxon	description	Fig. 35	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFCFF83FFF01BC664229FD6F.taxon	description	Adult — Length of forewing: male 3.4 - 4.0 mm; female 4.0 - 4.2 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish. Tibial spurs slightly darker than legs, not distinctly contrasting in color. Wing bar at anastamosis relatively indistinct, marked with light brown setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderately elongate, lateral margins slightly converging from base, with acute, trianguloid apicomesal projection, apicolateral lobes moderately elongate, sclerotized, slightly mesally curved; ventrolateral lobe nearly obsolete; ventromesal lobes with few short setae, extending from ventral margin of apicolateral lobes. Inferior appendages setose, without distinct dorsolateral projections, apically with pair of narrow, attenuate, ventrally curved projections. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages elongate, narrow, extending beyond dorsal phallic spine, slightly widened preapically, apex very acutely narrowed, awllike; base of appendage emerging from projecting membranous lobe. Dorsal phallic spine, as viewed laterally, undulate in contour, slightly widened on ventral margin in basal ½, apex acute, weakly and obtusely upturned in about apical ¼; in dorsal view, nearly uniform in width, apex very acutely narrowed. Phallicata relatively elongate, with upright, sclerotized, more or less rounded projections on dorsal margin; basolaterally with slightly produced, rounded and compressed projection. Endophallic membrane relatively simple, with single prominent curved ventromesal spine; phallotremal spines absent. Material examined — COLOMBIA: Antioquia: Quebrada Honda, 12 km SW Fredonia, 1450 m, 22. ii. 1983, OS Flint, Jr – 1 male (pinned), 1 male, 1 female (alcohol) (NMNH); Quebrada La Ayura, Municipio Envigado, 1750 m, 8. viii. 1983, U Matthias – 1 male, 1 female (alcohol) (NMNH); same locality and collector, 20 - 28. iv. 1984 – 1 male, 1 female (alcohol) (NMNH). Distribution — Colombia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC0F83EFF01BB46414EFF4F.taxon	description	Fig. 36 Mortoniella grandiloba very closely resembles M. florica (Flint) and M. propinqua Blahnik and Holzenthal in having upright dorsal projections on the phallicata and short inferior appendages, without elongate mesal or apicolateral projections. It can be distinguished from either of those species in that the dorsal projections of the phallicata are very prominent and enlarged, and also in the distinctive shape of the dorsal phallic spine, which is relatively thick through the middle, as viewed laterally, and has the apex narrowed and slightly ventrally recurved. Adult — Length of forewing: male 3.8 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color dark brown, apices of tarsal segments and basal segments of antennae whitish. Tibial spurs slightly darker than legs, not distinctly contrasting in color. Wing bar at anastamosis relatively indistinct, marked with light brown setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apically with subacute, trianguloid apicomesal projection, apicolateral lobes bluntly rounded, slightly mesally curved; ventrolateral lobe nearly obsolete; ventromesal lobes with few short setae, extending from ventral margin of apicolateral lobes. Inferior appendages setose, with short rounded dorsolateral projections, and very short acute mesal projection. Mesal pockets of inferior appendage with apical processes short, posteroventrally curved. Paramere appendages elongate, narrow, extending beyond dorsal phallic spine, slightly widened preapically, apex acute; base of appendage emerging from projecting membranous lobe. Dorsal phallic spine, as viewed laterally, with dorsal margin broadly curved, ventral margin slightly undulate in contour, spine distinctly widened in middle, apex narrow, acute, slightly ventrally recurved; spine in dorsal view, widened through middle, gradually narrowed to acute apex. Phallicata relatively elongate, with very prominent, upright, sclerotized, rounded and inflated projections on dorsal margin; laterally with slightly produced, crease-like projection. Endophallic membrane simple, with single prominent curved, asymmetrically positioned, ventral spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC0F83EFF01BB46414EFF4F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Barinas: 22 km NW Barinitas, 19. ii. 1976, CM and OS Flint, Jr, (UMSP 000157413) (NMNH). Etymology — This species is named M. grandiloba for the enlarged, upright basodorsal lobes of the phallicata.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC1F83EFF01B92641D1F98F.taxon	description	Fig. 37 This species probably most closely resembles M. munozi Blahnik and Holzenthal, especially in the shape of the dorsal phallic spine, which has a pronounced rounded ventral bulge that articulates with the base of the phallicata. It can be distinguished in that the acute apicolateral projections of the inferior appendages are not down-turned apically. It also lacks the pair of acute scabrous sclerites that are associated with the phallicata in M. munozi; instead, it has a pair of apically acute, wing-like, dorsolateral processes. The endophallic membrane has a very small curved spine. The combination of characters possesses by these two species make it possible that they are not closely related to the core of species included in the florica subgroup, and thus the placement of M. schlingeri in this subgroup is partly a matter of diagnostic convenience. Adult — Length of forewing: male 3.6 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish brown (specimen faded and largely denuded). Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apically with very weakly developed mesal projection, apicolateral lobes elongate, slightly mesally curved; ventrolateral lobes very small, rounded; ventromesal lobes not evident. Inferior appendages with short rounded dorsolateral lobes and elongate, narrow, paired, apicoventral projections, apices of projections acute. Mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, distinctly widened and somewhat cupped preapically, apex acute. Dorsal phallic spine, as viewed laterally, undulate in contour, spine distinctly widened on ventral margin at about middle, ventral bulge articulating with dorsal margin of phallicata, apex of spine narrow, acute, obtusely upturned in about apical 1 / 5; spine, in dorsal view, widened in middle, apex acute. Phallicata moderately elongate, basodorsal margin with upright, apically acute, dorsolateral projections. Endophallic membrane simple, with single small curved ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC1F83EFF01B92641D1F98F.taxon	materials_examined	Holotype male (alcohol) — COLOMBIA: 3 mi W Villavicencio, 11. iii. 1955, EI Schlinger and ES Ross (UMSP 000129555) (CAS). Etymology — The species is named M. schlingeri in honor of Evert I. Schlinger, one of the co-collectors of this species. — “ leroda ” subgroup	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC1F83DFF01BEE645ABF92F.taxon	description	Fig. 38	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC1F83DFF01BEE645ABF92F.taxon	description	Adult — Length of forewing: male 2.3 - 2.8 mm; female 2.9 - 3.2 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis relatively indistinct, marked with light brown setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection either suggestively developed or absent, apicolateral lobes relatively elongate, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral projections, and very elongate, asymmetric, mesal projection, apex of mesal projection distinctly inflated and dorsally curved, with numerous sensilla. Mesal pockets of inferior appendage with apical processes short, posteroventrally curved. Paramere appendages short (much shorter than dorsal phallic spine), narrow, apex scabrous and slightly enlarged; base of appendage emerging from projecting membranous lobe. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin widened in about middle, apex narrow, acute, only slightly dorsally curved; spine, in dorsal view, widened in middle, apex very acutely narrowed. Phallicata with small rounded and sclerotized projections on dorsal margin, ventrally with microsetae at base. Endophallic membrane elongate, simple in structure, without ventromesal spine; phallotremal spines distinct, small, spine-like.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC1F83DFF01BEE645ABF92F.taxon	materials_examined	Holotype male (pinned) — PERU: Madre de Dios: Manu, Pakitza Biol. Station, 11.94417 ° S, 71.28300 ° W, Quebrada Trompetero, 350 m, 6. vii. 1993, Blahnik and Pescador (MJP) (UMSP 000157376). Paratypes — PERU: Madre de Dios: same data as Holotype, 7 males, 6 females (pinned) (NMNH), 3 males, 3 females (pinned) (UMSP); same locality and collectors, 3. vii. 1993 – 3 females (pinned), 38 females (alcohol) (NMNH); Manu, Pakitza Biol. Station, Quebrada Paujil-Picoflor, 11.94417 ° S, 71.28300 ° W, 350 m, 2. vii. 1993, Blahnik and Pescador – 4 females (pinned) (NMNH), 2 females (pinned) (UMSP); same locality and collectors, 4 - 6. vii. 1993 – 3 females (pinned) (NMNH); Manu, Pakitza Biol. Station, Quebrada Pachija near intersection with Río Manu, 350 m, 4. vii. 1993, Blahnik and Pescador – 1 female (pinned) (NMNH); Manu, Pakitza, 11.93333 ° S, 71.30000 ° W, 250 m, 12 - 23. ix. 1989, N Adams et al. – 1 female (pinned), 4 males, 16 females (alcohol) (NMNH); Manu, Pakitza, 11.93333 ° S, 71.30000 ° W, 250 m, 9 - 14. ix. 1988, O Flint and N Adams – 1 female (alcohol) (NMNH); Manu, Pakitza, 12.11667 ° S, 70.96667 ° W, malaise trap (day collection), 250 m, 14 - 23. ix. 1988, O Flint and N Adams – 1 male, 2 females (pinned), 9 males, 22 females (alcohol) (NMNH); same locality and collectors, 18. ix. 1988 – 1 male, 14 females (alcohol) (NMNH); same locality and collectors, 17 - 20. ix. 1988, – 1 female (pinned), 8 males, 27 females (alcohol) (NMNH); same locality and collectors 18. ix. 1988 – 2 females (pinned) (NMNH). Etymology — This species is named M. bothrops, from the generic name for a rattlesnake, used as a noun in apposition, for the scabrous apex of the mesal process of the inferior appendages, which is characteristically curled upward in this species and somewhat resembles the rattle of a rattlesnake.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC2F83CFF01BE86422EFA0F.taxon	description	Fig. 39 This is perhaps the most distinctive of the South American species placed in the leroda subgroup. It has a very narrow and lightly sclerotized ventromesal process on the inferior appendages, without an enlarged scabrous apex. Because the process is very reduced and lightly sclerotized, it may not always be readily evident. The species is easily diagnosed by the distinctive form of the dorsal phallic spine, which has its apex very much narrowed and recurved downward. Adult — Length of forewing: male 2.8 - 3.2 mm; female 3.0 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish. Tibial spurs slightly darker than legs, contrasting in color. Wing bar at anastamosis relatively indistinct, marked with light brown setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection only weakly developed, apicolateral lobes short, acute, slightly mesally curved, with small acute lateral point at flexure with ventrolateral lobe; ventrolateral lobes rounded, weakly developed. Inferior appendages without dorsal lobes, apicoventral lobes relatively broad, acute apically, ventromesal projection short, very narrow, lightly sclerotized, and asymmetrically curved to one side. Mesal pockets of inferior appendage with apical processes short, posteroventrally curved. Paramere appendage moderate in length (shorter than dorsal phallic spine), relatively broad, apex scabrous; base of appendage emerging from projecting membranous lobe. Dorsal phallic spine, as viewed laterally, strongly curved at base, dorsal margin nearly straight, ventral margin widened in basal 1 / 2, apex very narrow, acute, and ventrally recurved; spine in dorsal view, somewhat widened in middle, apex very acutely narrowed. Phallicata with rounded sclerotized projections on dorsal margin; laterally with crease-like projection, ventrolateral margins with weakly developed longitudinal sclerites. Endophallic membrane simple in structure, without ventromesal spine; phallotremal spines very lightly sclerotized, indistinct, appearing as acutely tapering membranous projections.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC2F83CFF01BE86422EFA0F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Lara: Parque Nacional Terepaima, Río Auro, near Sabana Alta, 9.74567 ° N, 69.27690 ° W, 480 m, 16. vi. 2001, Holzenthal, Blahnik, Paprocki, Cressa (UMSP 000074409) (UMSP). Paratypes — VENEZUELA: Aragua: Est. Exp. Cataurito, ca. 32 km E Villa de Cura, 1100 m, OS Flint, Jr – 1 male, 1 female (alcohol) (NMNH); Guárico: Parque Nacional Guatopo, Quebrada Guatopo, 0.5 km N Estacion L Colina, 10.014 ° N, 66.363 ° W, 600 m, Holzenthal, Cressa, Rincón – 3 males, 10 females (alcohol) (UMSP), 1 male, 5 females (alcohol) (MIZA). Etymology — We take pleasure in naming this species M. cressae for Dra. Claudia Cressa, (Central University of Venezuela, Caracas), who helped in collecting the type specimen, in recognition of our numerous collaborations and also gratitude for organizing the collecting expedition during which the type specimen was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC3F83BFF01BE6645ABFC4F.taxon	description	Fig. 40 As discussed in the diagnosis for M. simla (Flint), both this species and M. ruedae, n. sp., are very similar to M. simla. Unlike other species with an elongate, asymmetric mesal process on the inferior appendages, and with a scabrous apex, all of these species have prominent upright lateral projections on the phallicata. The species differ in the relative lengths of the paramere appendages: very short in M. curtispina, n. sp., of intermediate length (generally shorter than the dorsal phallic spine or mesal process of the inferior appendages) in M. simla, and elongate (longer than either the dorsal phallic spine or mesal process of the inferior appendages) in M. ruedae. Additionally, this species lacks a ventromesal endophallic spine, although it does have paired sclerotized phallotremal spines. In contrast, Mortoniella simla has a prominent ventromesal endophallic spine and M. ruedae has a relatively small one. The very short paramere appendages of M. curtispina are especially distinctive and should easily distinguish this species. Adult — Length of forewing: male 2.5 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish brown (specimen largely denuded and faded). Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection obtusely angular, weakly projecting, apicolateral lobes relatively elongate, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral projections, and very elongate, asymmetric, mesal projection, apex of mesal projection somewhat inflated, with numerous sensilla. Mesal pockets of inferior appendage with apical processes very short. Paramere appendage very short and spine-like, apex acute. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin distinctly widened in basal ½, apex narrow, acute, only slightly dorsally curved; spine, in dorsal view, relatively narrow throughout, apex very acutely narrowed. Phallicata with rounded, sclerotized, projections on dorsal margin. Endophallic membrane simple in structure, without ventromesal spine; phallotremal spines distinct, small, spine-like.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC3F83BFF01BE6645ABFC4F.taxon	materials_examined	Holotype male (alcohol) — VENEZUELA: Zulia: El Tucuco, Sierra de Perija, montane forest, 28 - 29. i. 1978, JB Heppner (UMSP 000124881) (NMNH). Etymology — This species is named M. curtispina, Latin for short-spined, in reference to the very short paramere appendages of this species, which helps to distinguish it from closely related species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC4F83AFF01BC26451DFCCF.taxon	description	Fig. 41 This species is easily diagnosed by the structure of the mesal process of the inferior appendages, which is very asymmetrically positioned, with its base strongly arched and armed with what are apparently short papillae. The apex of the process is acutely narrowed, unlike most of the new species of the leroda subgroup considered here. The asymmetric ventral process can be located on either the right or left side (which is typical of the asymmetry of the leroda subgroup). The figured specimen has the process on the left side; a specimen with the process on the right side would obviously appear different from the illustration. Adult — Length of forewing: male 2.6 - 3.0 mm; female 2.7 - 3.3 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection only suggestively developed, apicolateral lobes relatively short, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short, apically acute, dorsolateral projection (generally only unilaterally present), and elongate, very asymmetrically developed, ventromesal projection (variably appearing as emerging from either right or left side), ventromesal projection dorsally looped at base, with distinct papillae or sensilla, apex acute, tapering, not scabrous. Mesal pockets of inferior appendage with apical processes very short. Paramere appendage elongate, narrow, subequal in length to dorsal phallic spine, noticeably enlarged and indistinctly scabrous preapically, apex acute; base of appendage emerging from projecting membranous lobe. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin distinctly widened at about middle, rounded ventral projection articulating with base of phallicata, apex of spine acutely narrowed, only weakly dorsally curved; spine, in dorsal view, relatively narrow throughout length, apex tapering and very acutely narrowed. Phallicata with rounded sclerotized projections on dorsal margin. Endophallic membrane apparently simple in structure, without ventromesal spine; phallotremal spines small, spine-like, lightly sclerotized.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC4F83AFF01BC26451DFCCF.taxon	materials_examined	Holotype male (pinned) — ECUADOR: Pastaza: Puyo, 30. i. 1976, Spangler et al. (UMSP 000095068) (NMNH). Paratypes — ECUADOR: Pastaza: Puyo, 5. v. 1977, PJ Spangler and DR Givens – 9 males, 28 females (alcohol) (NMNH); same locality and collectors, 6. v. 1977 – 6 males, 15 females (alcohol) (NMNH); same locality and collectors, 8. v. 1977 – 4 males, 6 females (alcohol) (UMSP); same locality and collectors, 14. v. 1977 – 1 male, 1 female (alcohol) (NMNH); same data as Holotype – 7 males, 195 females (alcohol) (NMNH); Puyo, riverside, 29. v. 1975, Cohen and Langley – 3 males, 3 females (alcohol) (NMNH); Puyo (3 km N), 30. v. 1975, Cohen and Langley – 1 male, 1 female (pinned) (NMNH); Puyo (3 km W), 15. vii. 1976, J Cohen – 19 males, 127 females (alcohol) (NMNH). Etymology — This species is named M. draconis, from the Latin word draco, a fabulous lizard-like animal (dragon), and in this case pertaining to the very asymmetric and ornamented ventromesal process of the inferior appendages of this species, with some semblance of a dragon’s tail.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC5F839FF01BBA64531FDAF.taxon	description	Fig. 42 Among species in the leroda subgroup that are characterized by an asymmetric and apically modified ventromesal process on the inferior appendages, this species is unique in having the apex of this structure forked, with small spines or sensilla. It is also characterized by having paramere appendages that are relatively thick and downward curved, with the apices scabrous. The latter character is useful in separating it from M. parameralda, n. sp., with which it co-occurs. In M. parameralda the appendages are narrower and dorsally curved. Both species lack rounded basodorsal processes on the phallicata. Adult — Length of forewing: male 2.6 - 3.0 mm; female 2.8 - 3.0 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish brown (specimen faded and largely denuded). Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection only suggestively developed, apicolateral lobes short, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral projections, and elongate, asymmetric mesal projection, mesal projection forked preapically, apices with small spines or sensilla. Mesal pockets of inferior appendage with apical processes very short. Paramere appendages long, relatively thick, rod-like, distinctly ventromesally curved, subequal in length to dorsal phallic spine, apex scabrous and slightly enlarged. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin widened in basal ½, apex acute, slightly dorsally curved; spine in dorsal view, relatively narrow throughout, slightly widened preapically, apex abruptly and acutely narrowed. Phallicata without distinct basodorsal projections, ventral margin with short lateral sclerites. Endophallic membrane short, inflated, with short curved ventromesal spine; phallotremal spines consisting of 2 pairs of widely separated processes, dorsal ones lightly sclerotized, subtending apex of dorsal phallic spine, ventral ones submembranous, projecting apically.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC5F839FF01BBA64531FDAF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Los Rios: Quevedo (56 km N), 28 - 29. vii. 1976, J Cohen (UMSP 000124889) (NMNH). Paratypes — ECUADOR: Cotopaxi: Quevedo (36 km NE), 335 m, 21. vii. 1976, J Cohen – 2 males (alcohol) (NMNH); Los Rios: same data as Holotype – 2 males (alcohol) (NMNH); Pichincha: Santo Domingo (47 km S), Río Palenque Biol. Station, 29. vii. 1976, J Cohen – 27 males (alcohol) (NMNH), 5 males (alcohol) (UMSP); Tungurahua: Yanayacu, 300 m, 29 - 30. viii. 1977, LE Peña G – 4 males, 2 females (alcohol) (NMNH). Etymology — This species is named M. furcula, as a diminutive form of the Latin furca, meaning fork, and referring to the forked apex of the ventromesal process of the inferior appendages.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC6F838FF01BA0643FBFE2F.taxon	description	Fig. 43 This species is most similar to M. meralda (Mosely), agreeing in a number of details, especially in having an elongate, asymmetric mesal process on the inferior appendages, a phallicata lacking distinct upright lateral processes, and in the absence of a ventromesal endophallic spine. Also, in both species, the mesal process of the inferior appendages is uniformly narrow throughout, rather than distinctly enlarged apically. Mortoniella parameralda differs from M. meralda in the form of the paramere appendages; in both species they are relatively elongate, but they are more uniformly wide in M. meralda, flattened and ribbon-like, and lack scabrous apices, whereas in M. parameralda they are narrow, except preapically, scabrously developed, and distinctly upturned. The dorsal phallic spine in M. parameralda also appears to be more narrowed apically, very attenuate and needle-like. In the form of the paramere appendages, M. parameralda closely resembles M. ruedae, n. sp., but that species has distinct upright processes on the phallicata, and also a small ventromesal endophallic spine. Mortoniella parameralda was found to occur sympatrically with M. furcula, n. sp., which it also generally resembles. It can be distinguished from that species in lacking a forked ventromesal process of the inferior appendages and also in the orientation of the paramere appendages, broad and downward curved in M. furcula and upturned apically in M. parameralda. Adult — Length of forewing: male 2.3 - 2.6 mm; female 3.0 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish brown (specimens faded and largely denuded). Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection only suggestively developed, apicolateral lobes elongate, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral projections, and very elongate, asymmetric ventromesal projection, apex of ventromesal projection scabrous, projecting nearly straight, not or only indistinctly inflated. Mesal pockets of inferior appendage with apical processes very short. Paramere appendage elongate, narrow, subequal in length to dorsal phallic spine, distinctly enlarged and scabrous preapically, apex acute. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin widened in basal ½, apical 1 / 3 very acutely narrowed, needlelike, distinctly dorsally inflected; spine in dorsal view, widened through middle, apex very acutely narrowed. Phallicata relatively elongate and tube-like, without sclerotized dorsolateral projections. Endophallic membrane inflated, with pair of projecting lateral membranous lobes, ventromesal spine absent; phallotremal spines indistinct, rounded, weakly sclerotized.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC6F838FF01BA0643FBFE2F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Los Rios: Quevedo (56 km N), 28 - 29. vii. 1976, J Cohen (UMSP 000124888) (NMNH). Paratypes — ECUADOR: Cotopaxi: Latacunga (133 km W), 329 m, 2. vii. 1975, Langley and Cohen – 1 male, 1 female (alcohol) (NMNH); Quevedo (36 km NE), 335 m, 21. vii. 1976, J Cohen – 2 males (alcohol) (NMNH); Esmeraldas: La Union, 3. ii. 1979, JJ Anderson – 1 male (alcohol) (NMNH); Los Rios: same data as Holotype – 8 males (alcohol) (NMNH); Pichincha: Santo Domingo (47 km S), 29. vii. 1976, J Cohen – 15 males (alcohol) (NMNH), 3 males (alcohol) (UMSP). Etymology — This species is named M. parameralda from the Greek para, meaning beside or near, and referring to the close similarity of this species to M. meralda.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC7F847FF01B986433DFE6F.taxon	description	Fig. 44, 45	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC7F847FF01B986433DFE6F.taxon	description	Adult — Length of forewing: male 2.4 - 3.0 mm; female 2.5 - 3.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis relatively indistinct, marked with light brown setae. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection only suggestively developed, apicolateral lobes elongate, subacute, slightly mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral lobes, and very elongate, asymmetric ventromesal projection, apex of ventromesal projection scabrous, projecting nearly straight, not (or only indistinctly) inflated. Mesal pockets of inferior appendage with apical processes very short. Paramere appendage elongate, narrow, subequal in length to dorsal phallic spine, distinctly enlarged and scabrous preapically, apex acute. Dorsal phallic spine, as viewed laterally, with dorsal margin undulate in contour, ventral margin widened in basal ½, apex of spine acute, slightly dorsally inflected; spine in dorsal view, widened through middle, apex very acutely narrowed. Phallicata relatively elongate and tube-like, with rounded, sclerotized, dorsolateral projections, laterally with longitudinal, flattened, crease-like projection. Endophallic membrane relatively simple in structure, elongate, ventromesal spine very short; phallotremal spines present, short and spine-like.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFC7F847FF01B986433DFE6F.taxon	materials_examined	Holotype male (pinned) — BOLIVIA: La Paz: San Buenaventura-Ixiamas rd., km 23, Hacienda Chiquitos, Arroyo Chiquitos, 14.33470 ° S, 67.70340 ° W, el 284 m, 23. vii. 2003, Robertson and Blahnik (UMSP 000093907) (UASC). Paratypes — BOLIVIA: La Paz: same data as holotype – 1 male, 11 females (pinned) (UMSP); AMNI Madidi, Comun. San Miguel de Bala, Arroyo Bacuatra Grande, 14.51228 ° S, 67.52308 ° W, el 280 m, 17 - 19. vii. 2003, Robertson, Blahnik, Apaza – 3 males, 5 females (pinned) (UMSP), 1 male, 2 females (pinned) (NMNH). Etymology — This species is named M. ruedae for P. A. Rueda Martín, who recorded it from Bolivia under the name of M. simla and also illustrated it.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB8F846FF01BA464193FACF.taxon	description	Fig. 46, 47	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB8F846FF01BA464193FACF.taxon	description	Adult — Length of forewing: male 2.6 - 3.0 mm; female 2.7 - 3.2 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X elongate, lateral margins subparallel, apicomesal projection not, or only suggestively, developed, apicolateral lobes elongate, subacute, mesally curved; ventrolateral lobes rounded, weakly developed. Inferior appendages with short rounded dorsolateral projections, and very elongate, sinuous, asymmetric, ventromesal projection, apex of ventromesal projection scabrous, slightly inflated, distinctly bent or inflected. Mesal pockets of inferior appendage with apical processes very short. Paramere appendage moderately elongate, shorter than dorsal phallic spine, irregular in contour, scabrous apically. Dorsal phallic spine, as viewed laterally, with dorsal margin slightly undulate in contour, ventral margin widened in about middle, apex of spine acute, slightly dorsally inflected; spine, in dorsal view, widened through middle, apex very acutely narrowed, needle-like. Phallicata relatively elongate and tube-like, with rounded, sclerotized, dorsolateral projections, laterally with longitudinal, flattened, crease-like projection. Endophallic membrane relatively simple in structure, short, ventromesal spine prominent, curved; phallotremal spines present, narrow and needlelike. Material examined — TRINIDAD: Simla, 9 - 11. ii. 1966, Duckworths – 81 males, 116 females (Paratypes-alcohol) (NMNH); Tacarigua, Río Caura Rec. Area, 10 ° 43 ’ N, 61 ° 17 ’ W, 22. vi. 1993, Flint and Adams – 1 male (pinned) (NMNH); Verdant Vale, Arima R., 10.68333 ° N, 61.30000 ° W, 170 m, 19. vi. 1993, N Adams and W Mathis – 33 males, 252 females (alcohol) (NMNH); VENEZUELA: Sucre: Río Cocollar, 1.5 km SE Las Piedras de Cocollar, 10.16028 ° N, 63.79342 ° W, 810 m, 7 - 8. iv. 1995, Holzenthal and Flint – 30 males, 88 females (alcohol), 3 males, 5 females (pinned) (UMSP) (NMNH); Quebrada Zapateral, 1.5 km SE Las Piedras de Cocollar, 10.16255 ° N, 63.79312 ° W, 810 m, 9. iv. 1995, Holzenthal and Flint – 5 males, 4 females (alcohol) (UMSP) (NMNH). Distribution — Trinidad, Venezuela. — limona subgroup Included species: Mortoniella akrogeneios, n. sp.; M. auricularis, n. sp.; M. gracilis, n. sp.; M. guyanensis, n. sp.; M. limona (Flint); M. prolata, n. sp.; M. quadrispina, n. sp.; M. variabili s, n. sp. This subgroup is comprised of Mortoniella limona Flint and the seven new species recognized here. Species range from Guyana to Peru. In general, the dorsal phallic spine is compressed (laterally flattened) in species of this subgroup. The most diagnostic character defining this group is the presence of a pair of ventral (and usually rounded) apicomesal processes on tergum X, which more or less straddle the dorsal phallic spine (Fig. 52 A). Additionally, all of the species have 1 or 2 pairs of enlarged endophallic spines. When 2 pairs are present, it is obvious that the second pair is part of the same process. The spines possibly represent a modification of the small phallotremal spines bordering the phallotremal opening in a number of other species subgroups. Although these large paired spines are present, a ventromesal endophallic spine, which is a common feature in many subgroups of the leroda group, is absent. Species in the M. limona subgroup also lack dorsolateral sclerotized processes on the phallicata, but most of the species have the ventral margin of the phallicata modified into a kind of projecting lobe, only slightly developed in some species, but distinctly projecting in others. The ventral process of segment VI is generally large and subtriangular, decidedly ventrally projecting, and very wide basally. This compares to a more posteriorly directed process in members of the punensis subgroup. The anterior margin of this process is not retracted, as in many members of the florica / leroda subgroup, with the sole exception of M. guyanensis, n. sp., whose ventral process is more or less typical of species in that subgroup.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB9F845FF01BDA6455DFCEF.taxon	description	Fig. 48 Mortoniella akrogeneios, n. sp. is superficially most similar to M. limona (Flint), due to the shape of the apex of the dorsal phallic spine, which has a very pronounced rounded ventral projection in both species. However, the ventral projection of M. akrogeneios is narrower (and thus may appear more prominent). Additional diagnostic characters distinguishing it from M. limona include its apically projecting inferior appendages, short paramere appendages, and shallower emargination of tergum X. Also, M. akrogeneios has only small rounded, ventral projections on the phallicata, not nearly as projecting as those in M. limona and its two closely related species, M. gracilis, n. sp. and M. variabilis, n. sp. Adult — Length of forewing: male 3.0 - 3.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) light brown. Wing bar not evident Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apical margin concave, apicolateral lobes very weakly projecting; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, irregular, projecting. Inferior appendages without dorsolateral projections, apicoventral projections paired, projecting, upturned and acute apically. Mesal pockets of inferior appendage with apical processes short, posterodorsally curved. Paramere appendage short (much shorter than dorsal phallic spine) and relatively thick, apex acute. Dorsal phallic spine, as viewed laterally, with pronounced compressed apical expansion, upturned and acute dorsally, enlarged and rounded ventrally, dorsal and ventral projections connected almost linearly on posterior margin; spine, in dorsal view, very slightly widened in middle, apex acute. Phallicata with dorsal margin somewhat elevated and sclerotized, ventrally with pair of small rounded and sclerotized basal projections. Endophallic membrane short, with 2 pairs of enlarged spines (possibly phallotremal complex), 1 pair extending straight, the other strongly curved; ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB9F845FF01BDA6455DFCEF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pastaza: Puyo (27 km N), Est. Fluvia Metrica, 4. ii. 1976, Spangler, et al. (UMSP 000097078) (NMNH). Paratypes — ECUADOR: Pastaza: same data as holotype – 1 male (alcohol) (NMNH). Etymology — This species is named M. akrogeneios from a Greek word meaning “ with a prominent chin, ” and referring to the projecting ventral margin of the dorsal phallic spine in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBAF844FF01BBC6454DFE8F.taxon	description	Fig. 49 This species is very similar to M. quadrispina, n. sp. Both species have relatively elongate, narrow ventrolateral projections from the inferior appendages and a pair of apicolateral projections from the apical expansion of the dorsal phallic spine. Mortoniella auricularis differs in that the ventral margin of the expansion of the dorsal phallic spine is rounded, rather than angulate. Adult — Length of forewing: male 2.8 - 3.0 mm; female 3.1 - 3.7. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin nearly straight between apicolateral lobes, apicolateral lobes distinctly projecting, tapering, acute apically; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, irregular, projecting. Inferior appendages with very short rounded dorsolateral projections, apicoventral projections paired, elongate, narrow, nearly linear, acute apically. Mesal pockets of inferior appendage with apical processes moderately large, prominent, posterodorsally curved. Paramere appendage elongate (nearly as long as dorsal phallic spine), nearly uniformly narrow, apex acute. Dorsal phallic spine compressed apically, as viewed laterally, with distinct rounded apicoventral expansion, preapically (or apically) with pair of small lateral projections; spine, in dorsal view, nearly uniform in width, apicolateral projections evident. Phallicata with dorsal margin sclerotized, laterally with pair of protruding, apically rounded projections, ventrally with pair of small rounded sclerotized basal projections. Endophallic membrane short, with pair of enlarged spines (possibly phallotremal complex); ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBAF844FF01BBC6454DFE8F.taxon	materials_examined	Holotype male (pinned) — COLOMBIA: Meta: Quebrada Blanca, 3 km W Restrapo, 11. ii. 1983, OS Flint, Jr (UMSP 000146422) (NMNH). Paratypes — COLOMBIA: Meta: Quebrada Blanca, 3 km W Restrepo, 11. ii. 1983, OS Flint, Jr – 1 male, 4 females (pinned), 2 males, 27 females (alcohol) (NMNH). Etymology — This species is named M. auricularis, derived from the Latin word auricula, for ear, referring specifically to the apicolateral projections of the dorsal phallic spine in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBBF843FF01B9E64214FECF.taxon	description	Fig. 50 Mortoniella gracilis is very similar to M. variabilis, n. sp. and M. limona (Flint), as discussed in the diagnoses for those species. All of these species are characterized by elongate sclerotized projections from the ventral surface of the phallicata. The ventral projection is variable in length and shape, even within the respective species, and thus the exact shape is not reliably diagnostic. Mortoniella gracilis can generally be distinguished from the other two species by the much more slender, less projecting, ventral protrusion of the apex of the dorsal phallic spine. There is often a spine-like projection from the posterior margin of this apical expansion; this is more likely to be preapical and slightly downturned, rather than upturned from the ventral margin, which is more typical of M. variabilis. The armature of the endophallic membrane is most similar to M. limona in that it is composed of 2 pairs of distinctly sclerotized spines. In M. variabilis, 1 of the pairs of spines is typically either small and lightly sclerotized (and thus not readily visible), or possibly absent in some specimens. Adult — Length of forewing: male 2.9 - 3.4 mm; female 3.0 - 3.8. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length slightly greater than width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, apicomesal margin with V-shaped (usually) or U-shaped emargination between apicolateral lobes, emargination extending about 1 / 3 length of segment, apicolateral lobes tapered, acute apically, lobes very short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages with short rounded dorsolateral projections and very short acute paired apicoventral projections. Mesal pockets of inferior appendage with apical processes moderately large, posterodorsally curved. Paramere appendage elongate (subequal to dorsal phallic spine), narrow, nearly uniformly in width, apex acute. Dorsal phallic, as viewed laterally, compressed apically, with weakly rounded apicoventral expansion, posterior margin usually with evident acute, slightly down-turned, preapical projection; spine in dorsal view, nearly uniform in width, apex acute. Phallicata with dorsal margin sclerotized, extending into endophallic membrane, basoventrally with pair of very prominent curved, apically rounded sclerotized projections. Endophallic membrane short, with 2 pairs of enlarged sclerotized spines (possibly phallotremal complex); ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBBF843FF01B9E64214FECF.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Sucre: Peninsula de Paria, Puerto Viejo, “ Río el Pozo, ” 10.71788 ° N, 62.47615 ° W, el 20 m, 2 - 3. iv. 1995, Holzenthal, Flint, Cressa (UMSP 000001425) (UMSP). Paratypes — VENEZUELA: Barinas: 22 km NW Barinas, 24. ii. 1976, CM and OS Flint, Jr – 1 male, 8 females (pinned) (NMNH); Monagas: Guachero Cave N. P., 10.172033 ° N, 63.555250 ° W, el 1110 m, 20 - 21. vii. 2010, Holzenthal, Thomson, Cressa – 2 males, 1 female (pinned) (UMSP); Sucre: Quebrada Zapateral, 1.5 km SE Las Piedras de Cocollar, 10.16255 ° N, 63.79312 ° W, el 810 m, 9. iv. 1995, Holzenthal and Flint – 2 males, 4 females (pinned); 3 males, 2 females (alcohol) (NMNH); Peninsula de Paria, Santa Isabel, Río Santa Isabel, 10.67157 ° N, 62. 64923 ° W, el 20 m, 4. iv. 1995, Holzenthal, Flint, Cressa – 1 male, 12 females (pinned); 3 males, 45 females (alcohol) (UMSP); same data as Holotype – 9 males, 12 females (pinned), 23 males, 45 females (alcohol) (UMSP); Peninsula de Paria, Puerto Viejo, Río Puerto Viejo, 10.71895 ° N, 68.47905. ° W, el 15 m, 2. iv. 1995, Holzenthal, Flint, Cressa – 7 males, 11 females (alcohol) (MIZA); Río Cocollar, 1.5 km SE Las Piedras de Cocollar, 10.16028 ° N, 63.79342 ° W, el 810 m, 7 - 8. iv. 1995, Holzenthal and Flint – 9 males, 9 females (pinned), 66 males, 58 females (alcohol) (UMSP). Etymology — This species is named M. gracilis, from the Latin word for slender, and referring to the relatively narrow apex of the dorsal phallic spine, which character helps to distinguish it from its most closely related species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBCF843FF01B9A645AAF94F.taxon	description	Fig. 51 This species is very different from others in this group and was placed here largely because of the pair of apicomesal processes on tergum X and the enlarged pair of endophallic spines. It is the only species of Mortoniella currently recorded from Guyana. Additional collecting may prove that it would be better treated as the prototype of an additional species subgroup. It differs from other species of the limona subgroup in that the dorsal phallic spine is depressed (dorsoventrally flattened), rather than compressed. The ventral process of segment VI is more or less typical of species in the florica / leroda subgroup (and unlike species in the limona subgroup) in that it is truncate apically and slightly retracted anterobasally. Among species in the leroda group, it is also unusual in having a lightly sclerotized dorsomesal apodeme on the phallobase and a recurved dorsal lobe on the inferior appendages. The only other species of the leroda group with a strongly recurved dorsal lobe on the inferior appendages are members of the akantha and bolivica subgroups. The spined apex of the dorsal phallic spine, in combination with the characters discussed above, is especially distinctive and diagnostic for this species. Adult — Length of forewing: male 3.0 - 3.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish brown (specimen badly faded). Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, short, ventrally projecting, truncately rounded apically, length slightly greater than width at base, process slightly retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin V-shaped, apicolateral lobes short, truncate; apicomesal lobes short, narrow, with several apical setae, visible in dorsal and lateral views; ventrolateral lobes distinct, projecting, rounded. Inferior appendages with narrow, recurved dorsal lobes, ventral lobe prominent, setose, acutely projecting mesally. Mesal pockets of inferior appendage with apical processes short, posteriorly curved. Paramere appendage moderate in length (shorter than dorsal phallic spine), posteroventrally projecting, uniform in width, apex acute. Phallobase with lightly sclerotized, compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, curved basally, relatively narrow, only slightly widened ventrally, distinctly dorsally curved and nearly rectilinearly inflected in about apical ¼, apex depressed (flattened), with numerous small spines; spine, in dorsal view, nearly uniform in width throughout length, apex acute. Phallicata with dorsal margin sclerotized, ventrally with pair of very small rounded sclerotized basal projections. Endophallic membrane short, with pair of very prominent enlarged spines (possibly modified phallotremal spines); ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBCF843FF01B9A645AAF94F.taxon	materials_examined	Holotype male (alcohol) — GUYANA: Potaro River, Kaeteur Falls, 5.17500 ° N, 59.48167 ° W, 1350 ft., 21 - 23. viii. 1997, OS Flint, Jr (UMSP 000124864) (NMNH). Paratypes — GUYANA: same data as Holotype – 1 male (alcohol) (NMNH). Etymology. This species is named M. guyanensis for Guyana, the country in which it was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBCF842FF01BF264226F8CF.taxon	description	Fig. 52, 100	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBCF842FF01BF264226F8CF.taxon	description	The male holotype of Mexitrichia macuta Botosaneanu was examined. It differs in no substantial way from M. limona and we consider it to be a synonym. The setae on dorsal phallic spine, discussed by Botosaneanu as a distinguishing feature, appear to have been either an artifact, or minute striae that may have appeared to be setae. The dorsal phallic spine, in many species of the limona subgroup, is often minutely serrate or laciniate on the apicocaudal surface; this may alternatively account for what Botosaneanu described as setae. Adult — Length of forewing: male 2.9 - 3.3 mm; female 3.2 - 3.9. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, apicomesal margin concavely emarginate between apicolateral lobes, emargination extending about 1 / 3 length of segment, apicolateral lobes tapered, rounded apically, lobes short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages irregular in shape, with short tapering dorsolateral projections and short narrow paired apicoventral projections. Mesal pockets of inferior appendage with apical processes moderately large, posterodorsally curved. Paramere appendage elongate (subequal to dorsal phallic spine), narrow, nearly uniformly in width, apex acute. Dorsal phallic, as viewed laterally, compressed apically, with very prominent rounded apicoventral expansion, posterior margin often evidently serrulate or micro-laciniate; spine in dorsal view, nearly uniform in width, apex acute. Phallicata with dorsal margin sclerotized, extending into endophallic membrane, with apical depression to accommodate apicoventral projection of dorsal phallic spine, basoventrally with pair of very prominent sclerotized, curved, apically rounded projections. Endophallic membrane short, with 2 pairs of enlarged sclerotized spines (possibly phallotremal complex); ventromesal spine absent. Material examined — VENEZUELA: Macuto, Río del Teleferico, 26. ii. 1982, L Botosaneanu – male Holotype of Mexitrichia macuta (ZMA); Aragua: Est. Exp. Cataurito, 1. ii. 1983, OS Flint, Jr – 1 male (pinned) (NMNH): Falcón: P. N. Sierra de San Luis, Cataratas del Río Hueque, 11.17847 ° N, 69.56220 ° W, el 583 m, 6. vi. 2001, Holzenthal, Blahnik, Paprocki, Cressa – 14 males, 51 females (pinned), 22 males, 63 females (alcohol) (UMSP); Guarico: P. N. Guatopo, Quebrada Guatopo, 0.5 km N Est. La Colina, 10.014 ° N, 66.363 ° W, el 600 m, 22. i. 1994, Holzenthal, Cressa, Rincón – 1 male (pinned), 30 males, 21 females (alcohol) (UMSP); Lara: P. N. Terepaima, Quebrada San Antonio, 9.86257 ° N, 69.21830 ° W, el 631 m, 17. vi. 2001, Holzenthal, Blahnik, Paprocki, Cressa – 10 males, 4 females (pinned), 1 male, 1 female (alcohol) (UMSP). Distribution — Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBEF841FF01B8A6435DFB4F.taxon	description	Fig. 53 Among species in the limona subgroup, Mortoniella prolata most closely resembles M. auricularis, n. sp. and M. quadrispina, n. sp. in having elongate apicoventral projections from the inferior appendages; those of M. prolata differ in being extremely elongate. Additionally, it differs in lacking the distinctive apicolateral projections from the apex of the dorsal phallic spine that characterize the other two species. Adult — Length of forewing: male 3.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) yellowish brown. Wing bar not evident Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apical margin concave, apicolateral lobes very weakly projecting; apicomesal lobes visible in lateral view, broadly rounded apically; ventrolateral lobes distinct, irregular, projecting. Inferior appendages with short tapering dorsolateral projections, apicoventral projections paired, very elongate, narrow, acute apically. Mesal pockets of inferior appendage with apical processes short, posterodorsally curved. Paramere appendage short (much shorter than dorsal phallic spine), dorsally curved, apex acute. Dorsal phallic spine, as viewed laterally, with apex strongly upturned and acute, ventral margin extended as rounded expansion; spine, in dorsal view, very slightly widened in middle, apex acute. Phallicata with dorsal margin somewhat elevated and sclerotized (to accommodate ventral margin of dorsal phallic spine), ventrally with pair of small rounded, sclerotized basal projections; endophallic membrane with pair of elongate, apically acute spines (possibly phallotremal complex); ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBEF841FF01B8A6435DFB4F.taxon	materials_examined	Holotype male (alcohol) — PERU: Huanuco: Tingo Maria, premontane rain forest, 672 m, 1 - 6. ii. 1980, JB Heppner (UMSP 000097160) (MJP). Paratypes — PERU: Huanuco: same data as holotype – 1 male (alcohol) (NMNH). Etymology — This species is named M. prolata, from the Latin prolatus (to carry forward), used in the sense of being extended or elongate, and referring specifically to the very elongate apicoventral projections of the inferior appendages in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBEF840FF01BD2641B0FD0F.taxon	description	Fig. 54 This species is apparently very closely related to M. auricularis, n. sp., differing primarily in the shape of the apex of the dorsal phallic spine. Mortoniella quadrispina has two apicolateral projections, as in M. auricularis, but also has the ventral lobe sharply angulate, rather than rounded, thus producing 4 spine-like projections on the apex of the dorsal phallic spine (1 apicodorsal, 2 lateral, and 1 apicoventral). Adult — Length of forewing: male 3.5 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) medium brown. Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin with U-shaped emargination, extending about 1 / 3 length of tergum, apicolateral lobes tapering, acute apically, relatively short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages without dorsolateral projections, apicoventral projections paired, narrow, nearly linear, moderately elongate, acute apically. Mesal pockets of inferior appendage with apical processes large, prominent, posterodorsally curved. Paramere appendage moderately elongate (shorter than dorsal phallic spine), nearly uniformly narrow, apex acute. Dorsal phallic spine compressed apically; as viewed laterally, with acute apicoventral projection, preapically (or apically) with pair of small acute lateral projections, apicomesally with additional small acute projection; spine, in dorsal view, nearly uniform in width, apicolateral projections evident. Phallicata with pair of small rounded basoventral projections, and somewhat indistinct, rounded, apicoventral lobes. Endophallic membrane short, with pair of very enlarged, curved, sclerotized spines (possibly phallotremal complex); ventromesal spine absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBEF840FF01BD2641B0FD0F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pastaza: Puyo (27 km. N), Estación Fluvia Metrica, 4. ii. 1976, Spangler, et al. (UMSP 000097062) (NMNH). Etymology — This species is named M. quadrispina for the four acute projections on the apex of the dorsal phallic spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBFF84FFF01BB664297F9CF.taxon	description	Fig. 55	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFBFF84FFF01BB664297F9CF.taxon	materials_examined	Mortoniella variabilis is very similar to M. gracilis, n. sp. and M. limona (Flint). All of these species have elongate basoventral projections from the phallicata, a distinguishing feature of this complex. The shape of these projections is variable in each of these species, but in M. variabilis the projections tend to be narrower and longer than in M. gracilis. The most diagnostic feature distinguishing the species in this complex is the shape of the apex of the dorsal phallic spine. Unfortunately, this is somewhat variable, even within the species, and this is especially so for M variabilis. In general, the ventral projection at the apex of the dorsal phallic spine is less produced in M. variabilis than in M. limona, but more so than in M. gracilis. The ventral projection often (or perhaps usually) has a distinct, upturned, spine-like projection, not present in M. limona. The development of this spine is quite variable in the material examined, as is the length of the paramere appendages (see Fig. 55 D for an extreme variant from Colombia). We are interpreting this as variation within a species, but the issue may be worthy of reinvestigation when more material is available. There was no evident difference in the female genitalia of specimens from Colombia and Venezuela. Mortoniella variabilis is probably more likely to be confused with M. limona than M. gracilis, because the ventral projection of the dorsal phallic spine is distinctly produced in both species, but the character state in M. variabilis never reaches the extreme condition found in M. limona. Also, the apex of the dorsal phallic spine, above the ventral expansion, tends to be narrower and more attenuate in M. variabilis than in M. limona. In most material of M. variabilis, only 1 pair of endophallic spines is evident, but in specimens with the armature fully expanded, a second, very lightly sclerotized pair may also be evident; the second pair may be a consistent feature, not easily observed unless the armature is completely everted. In the other two species, 2 pairs of sclerotized spines are generally evident, even when the armature is not expanded. Adult — Length of forewing: male 2.7 - 3.4 mm; female 3.0 - 3.9. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments and basal segments of antennae pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, apicomesal margin with V-shaped or U-shaped emargination between apicolateral lobes, extending about 1 / 3 length of tergum; apicolateral lobes tapered, acute apically, short and narrow as viewed laterally; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, projecting. Inferior appendages somewhat variable shape, with short tapering dorsolateral lobes and apicoventral projections very short or absent. Mesal pockets of inferior appendage with apical processes moderately large, posterodorsally curved. Paramere appendage short or moderate in length (shorter than dorsal phallic spine), narrow, nearly uniformly in width, apex acute. Dorsal phallic, as viewed laterally, compressed apically, with prominent rounded apicoventral expansion and (usually) an acute, dorsally-oriented projection from ventral margin; apex of dorsal phallic spine narrowing and attenuate, often slightly posteriorly curved; spine, in dorsal view, nearly uniform in width, apex acute. Phallicata with dorsal margin sclerotized, extending into endophallic membrane, with apical depression to accommodate apicoventral projection of dorsal phallic spine, basoventrally with pair of very prominent curved, apically rounded, sclerotized, projections. Endophallic membrane short, with one pair of prominent sclerotized spines, and second pair of smaller, lightly sclerotized spines (possibly sometimes absent); ventromesal spine absent. Holotype male (pinned) — VENEZUELA: Zulia: Parque Nacional Perijá, Río Negro in Toromo, 10.051 ° N, 72.712 ° W, el 360 m, 15. i. 1994, Holzenthal, Cressa, Rincón (UMSP 000001386) (UMSP). Paratypes — COLOMBIA: Magdalena: Municipio Ciénaga, Río Cordoba, 25 km NS “ Estacion Exp. San Lorenzo, ” Sierra Nevada de Santa Marta, 11.03944 ° N, 74.03833 ° W, el 930 m, 12. xii. 1997, F Muñoz-Q et al. – 5 males, 14 females (pinned) (UMSP); Municipio de Santa Marta, Río Minca en Minca, 11.14306 ° N, 74.11611 ° W, el 570 m, 9. xii. 1997, F. Muñoz-Q et al. – 1 male, 1 female (pinned) (UMSP); Meta: Quebrada Blanca, 3 km W Restrepo, 11. ii. 1983, OS Flint, Jr – 1 male (alcohol) (NMNH); VENEZUELA: Barinas: Parque Nacional Sierra Nevada, Quebrada San Juan in Santa Rosa, 8.46450 ° N, 70.84867 ° W, el 1000 m, 21. iii. 1997, Holzenthal – 2 males, 5 females (alcohol) (UMSP); 22 km N Barinitas, 24. ii. 1976, CM & OS Flint, Jr – 1 male, 2 females (alcohol) (NMNH); Zulia: same data as Holotype – 2 males, 2 females (pinned), 3 males (alcohol) (UMSP), 2 males, 2 females (alcohol) (NMNH), 2 males, 2 females (alcohol) (MIZA). Etymology — This species is named M. variabilis because of the variability in the structure of the apex of the dorsal phallic spine and length of the paramere appendages. — pocita subgroup Included species: Mortoniella pocita (Flint). This subgroup was proposed by Blahnik and Holzenthal (2011) to contain only M. pocita (Flint). The species was originally described from Argentina and subsequently listed from Bolivia by Rueda Martín and Gibon (2008). It has a general similarity to several of the species listed below as leroda group species “ unplaced to subgroup, ” especially in having elongate inferior appendages and a modified apex of the dorsal phallic spine. However, none of these species have elongate dorsal processes on the phallicata, a character found in M. pocita and also used to define the punensis subgroup. Because of our uncertainty about whether this reflects a relationship between these 2 subgroups, and because of its unusual overall combination of characters, we are continuing to list M. pocita in its own subgroup.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB0F84EFF01BEA64121F9CF.taxon	description	Fig. 56	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB0F84EFF01BEA64121F9CF.taxon	description	This species was redescribed and reillustrated by Blahnik and Holzenthal (2011) and the reader is referred to that paper for the description. The illustration is included here for completeness of coverage. The species has the apex of the dorsal phallic spine enlarged, similar to some species of the limona subgroup. It differs from M. limona, and other species with this characteristic, in having a very elongate, curled dorsolateral processes on the phallicata. Distribution — Argentina, Bolivia. — punensis subgroup Included species: Mortoniella armata (Jacquemart)?; M. chalalan, n. sp.; M. dentiterga, n. sp.; M. emarginata, n. sp.; M. marini (Rueda Martín and Gibon); M punensis (Flint); M. sinuosa, n. sp. This subgroup was proposed by Blahnik and Holzenthal (2011) to contain specifically M. punensis (Flint) and M. marini (Rueda Martín and Gibon). A somewhat broader interpretation of the group is required to include the assemblage of species listed below and we are not altogether certain that all of the species placed here are closely related. The primary character used to indicate monophyly of the group is the structure of the dorsal processes of the phallicata, which have either elongate, digitate or spine-like processes, which seems to hold or direct the paramere appendages; in many of the species the paramere appendages are directed mesally, so that they cross over one another. Whether this is a native configuration, or an assumed state resulting from clearing the genitalia, is difficult to say. Upright processes from the ventral margin of the phallicata, previously discussed as a character of the group, seem to apply only to M. punensis and M. marini, which are undoubtedly closely related taxa. In general, the species in this group have the dorsal phallic spine somewhat depressed apically (dorsoventrally flattened); the apex may be somewhat asymmetrical in some species. The hind wing has either forks II and III present or fork II only (smaller species). The ventral process of segment VI is prominent and more distinctly posteriorly directed than in most other species subgroups of the leroda group (M. sinuosa an exception). There is considerable variation among the species included here in the armature of the endophallic membrane, as for instance, presence or absence of a ventromesal spine, and whether 1 or 2 spines are present. There is also variation in the structure of the inferior appendages, and in the length of the spine-like apical processes of the mesal pockets. Mortoniella sinuosa is perhaps the most divergent of the taxa placed here, as noted in its description; its placement is speculative and based primarily on the presence of spine-like processes on the phallicata. Although M. pocita (Flint) and M. rectiflexa, n. sp. also possesses elongate dorsal processes on the phallicata, they differ enough from species of the subgroup to make their placement here doubtful. The former was placed in its own species subgroup by Blahnik and Holzenthal (2011) and the latter is included in the species listed as “ unplaced to subgroup. ” Mortoniella armata (Jacquemart, 1963), which is very inadequately illustrated and known only from Argentina, is speculatively included in this subgroup, largely because it seems to possess the elongate hooked processes on the dorsal margin of the phallicata that characterizes this group. Only fragments of the genitalia were illustrated in the original description. Flint et al. (1999) noted that the genitalia are missing from the holotype slide mount. The structure of the spines of the phallicata in M. armata (and apparently also the upright lobes from the phallicata) are very similar to M. punensis itself, but the structure of the dorsal phallic spine seems to be different (based on the original illustration) in being narrower in lateral view and lobed or divided apically.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB1F84DFF01BEA64451FAEF.taxon	description	Fig. 57 This diminutive species has a relatively simple overall morphology, but is reasonably distinctive. In addition to the spine-like dorsal processes of the phallicata, which character was used to place it in the punensis subgroup, the species has two straight, narrow endophallic spines, 1 ventromesal spine and 1 apicodorsal spine. This combined configuration is not known for any other species. The dorsal phallic spine is symmetrical in shape and has its apex depressed (dorsoventrally flattened), as is typical of the subgroup. The shape of tergum X, which is short, with relatively broad apicolateral processes and a concave mesal invagination, is also distinctive. Adult — Length of forewing: male 2.6 - 2.7 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsal segments whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI relatively small, laterally compressed, posteriorly directed, length about 1 ½ times width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin somewhat concave between apicolateral lobes, apicolateral lobes relatively short and broad, subtruncate; ventrolateral lobes rounded, weakly projecting. Inferior appendages short, without dorsolateral projections, apicoventral projections short, acute apically, ventromesal margin with very short subtruncate projection. Mesal pockets of inferior appendage with spine-like apical processes short, posterodorsally curved. Paramere appendage moderately elongate (shorter than dorsal phallic spine), nearly uniformly narrow, apex acute, opposite appendages converging mesally. Dorsal phallic spine stout, nearly uniform in width, without distinct ventral projection, depressed apically, apex gradually upturned, acute in both lateral and dorsal views. Phallicata with dorsal margin sclerotized and produced anteriorly into anvil-like projection, laterally with pair of curved spine-like projections, projections hooked over paramere appendages; ventral margin of phallicata simple, without lobes or projections. Endophallic membrane elongate, with small rounded membranous preapical and apicomesal lobes; ventromesal spine nearly straight, very narrow and nail-like; endophallic membrane with additional apical dorsomesal spine, spine very narrow, attenuate, slightly longer than ventromesal spine; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB1F84DFF01BEA64451FAEF.taxon	materials_examined	Holotype male (pinned) — BOLIVIA: LaPaz: ANMI Madidi, Chalalan Ecolodge, Raya Mayo river at Anta Trail, 14.43557 ° S, 67.92935 ° W, el 264 m, 26. vii. 2003, Robertson and Blahnik (UMSP 000094778) (UASC). Paratypes — BOLIVIA: Santa Cruz: PN and ANMI Amboró, Guardia Parque Mataracú, Quebrada Verde Uno, 17.55389 ° S, 63.86917 ° W, el 374 m, 19 - 23. xi. 2004, Robertson, Garcia and Vidaurre – 2 males (alcohol) (UMSP); PN & ANMI Amboró, Guardia Parque Mataracú, Río Yapacani, 17.52072 ° S, 63.86795 ° W, 329 m, 26. xi. 2004, Roberson, Garcia, Vidaurre – 1 male (UMSP). Etymology — This species is named M. chalalan, used as a noun in apposition, for the name of the Ecolodge in Madidi National Park where the type specimen was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB2F84CFF01BDC643D5FB2F.taxon	description	Fig. 58 Mortoniella dentiterga is most readily diagnosed by the hooked, spine-like processes from the dorsal margin of the phallicata and by the characteristic structure of tergum X in dorsal view, which has the apicolateral processes subtruncate (tooth-like) and inset laterally. It lacks the upright processes from the ventral margin of the phallicata found in M. punensis (Flint) and M. marini (Rueda Martín and Gibon), and also has a narrower apex on the dorsal phallic spine. The asymmetry of the spine-like projections from the phallicata seems to be a consistent feature, although possibly variable as to whether the longer spine is on the right or left side (as is often the case for asymmetrical features). Adult — Length of forewing: male 2.3 - 2.9 mm; female 2.7 - 3.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) medium brown (specimens largely denuded and faded). Male genitalia — Ventral process of segment VI laterally compressed, short, posteriorly projecting, subtriangular, acute apically, length slightly greater than width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, mesal margin straight or slightly convex, apicolateral lobes short, tooth-like, inset from acute lateral angles of tergum; ventrolateral lobe short, rounded. Inferior appendages with very short rounded dorsal projections, apically with pair of narrow, attenuate ventral projections, ventromesal projection very short, rounded. Mesal pockets of inferior appendage with apical processes posteriorly projected, moderately elongate (shorter than ventral projections of inferior appendage). Paramere appendage elongate, narrow, nearly uniform in width, extending about same length as dorsal phallic spine, apex acute; appendages curved mesally and crossing over one another. Dorsal phallic spine, as viewed laterally, widened in about middle, gradually upturned and narrowed apically, apex slightly notched; in dorsal view, distinctly widened in middle, apex narrow, subacute. Phallicata with pair of curved, spine-like projections from dorsal margin, crossing over paramere appendages (one projection typically more curved than the other). Endophallic membrane relatively simple, with single curved ventromesal spine; phallotremal spines only suggestively developed, short, weakly sclerotized.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB2F84CFF01BDC643D5FB2F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Los Rios: Quevedo (56 km N), Río Palenque Biol. Station, 28 - 29. vii. 1976, J Cohen (UMSP 000124865) (NMNH). Paratypes — ECUADOR: Cotopaxi: Quevedo (36 km NE), Río Palenque Biol. Station, 335 m, 21. vii. 1976, J Cohen – 3 males, 3 females (alcohol) (NMNH); El Oro: Canton de Arenillas, Las Lajas, 600 m, 30. v. 1979, JJ Anderson – 2 males (alcohol) (NMNH); Esmeraldas: La Union, 3. ii. 1979, JJ Anderon – 2 males 1 female (alcohol) (NMNH); Loja: Río Puyango, 300 m, 17 - 18. viii. 1977, LE Peña G – 19 males (alcohol) (NMNH); Los Rios: same data as Holotype – 34 males (alcohol) (NMNH), 5 males (alcohol) (UMSP); Manabi: Santo Domingo (29 km SW), Rancho Ronald, 20. vii. 1978, JJ Anderson – 2 males (alcohol) (NMNH); Pichincha: Santo Domingo (47 km S), 29. vii. 1976, J Cohen – 8 males (alcohol) (NMNH). Etymology — This species is named M. dentiterga, from the Latin dens, for tooth, and tergum, for back (referring to the dorsal sclerites in Insecta), for the characteristic shape of tergum X of this species, which has somewhat tooth-like apicolateral projections.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB3F84BFF01BC86440CFC4F.taxon	description	Fig. 59 Mortoniella emarginata is probably most similar to M. dentiterga, n. sp., especially in the relatively spine-like dorsal projections from the phallicata. It is most readily distinguished by the structure of tergum X, which has a small V-shaped apicomesal emargination, and apicolateral projections of the tergum that are acute, rather than bluntly toothed. Neither the holotype specimen from Ecuador, nor paratypes from the same country, have the paramere appendages crossed over one another. The single specimen from Colombia has the spines crossed and held in place by the rounded posterior margin of the phallicata spines. This is an unusual configuration and whether it is normal is difficult to say without additional material. The apex of the dorsal phallic spine in the specimen from Colombia is also slightly less upturned. Despite these differences, a close comparison of specimens from the two countries reveals no significant structural differences; we infer that the differences only represent regional or individual variation. Adult — Length of forewing: male 3.2 - 3.6 mm; female 3.6 - 4.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color (in alcohol) medium brown. Tibial spurs darker than legs, contrasting in color. Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, short, posteriorly projecting, subtriangular, acute apically, length about 1 ½ times width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin somewhat produced dorsally, rounded laterally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel, apical margin with small, but distinct, V-shaped mesal emargination, apicolateral lobes short, subtriangular, acute apically; ventrolateral lobe short, rounded. Inferior appendages with dorsolateral margin rounded, apically with pair of relatively short, apically acute, ventral projections, ventromesal projection very short (about ½ length of ventral projections), acute apically. Mesal pockets of inferior appendage with apical processes short, posteriorly projected. Paramere appendage moderately elongate (shorter than dorsal phallic spine), narrow, nearly uniform in width, apex acute; appendages curved mesally, but not crossing over one another (except in specimen from Colombia). Dorsal phallic spine, as viewed laterally, relatively stout, undulate in contour, upturned in about apical ¼; in dorsal view, relatively broad, distinctly widened in apical 1 / 2, apex narrowed and notched mesally. Phallicata with pair of curved, spine-like projections from dorsal margin, basoventrally with pair of rounded, sclerotized lobes. Endophallic membrane with pair of short curved ventromesal spines (or ventromesal spine divided apically into pair of spines); phallotremal spines short, blunt, weakly sclerotized.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB3F84BFF01BC86440CFC4F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pichincha: Pachijal, 26. vi. 1976, J Cohen (UMSP 000097042) (NMNH). Paratypes — COLOMBIA: Antioquia: Km. 50, Río Aurra, San Jeronimo, 14. ii. 1983, OS Flint, Jr – 1 male, 1 female (alcohol) (NMNH); ECUADOR: Pastaza: Pachijal, 26. vi. 1976, J Cohen – 3 males, 2 females (alcohol) (NMNH); Pichincha: same data as Holotype – 2 males, 4 females (alcohol) (NMNH); Río Umachaca, Forest Station Maquipucuna, ca. 5 km E Nanegal, 0.12500 ° N, 78.61667 ° W, 1250 m, 4 - 5. ix. 1990, OS Flint, Jr – 2 males (alcohol) (NMNH); Nanegal, 335 m, 19 - 20. ix. 1977, LE Peña G – 1 male, 1 female (alcohol) (UMSP). Etymology — This species is named M. emarginata for tergum X of the male, which has a characteristic narrow, V-shaped apicomesal emargination or notch.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB4F84AFF01BC264244FAAF.taxon	description	Fig. 60	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB4F84AFF01BC264244FAAF.taxon	description	Adult — Length of forewing: male 3.5 - 4.4 mm; female 3.9 - 4.8 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments whitish or pale brown. Tibial spurs darker than legs, somewhat contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI large, laterally compressed, subtriangular (rather variable in shape), posteriorly or posteroventrally directed, length about 1 to 1 ½ times width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin distinctly produced and rounded in dorsal ½, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel, apicomesal margin straight or somewhat concave between apicolateral lobes, apicolateral lobes divided or forked into subacute dorsal and ventral lobes, dorsal lobes mesally curved from lateral margins, relatively narrowly separated mesally; ventrolateral lobes rounded, projecting. Inferior appendages short, without dorsolateral projections, apicoventral projections very short (shorter than ventromesal projection), ventromesal margin with short, apically rounded projection. Mesal pockets of inferior appendage with spine-like apical processes short, posteriorly curved. Paramere appendage moderately elongate (shorter than dorsal phallic spine), nearly uniformly narrow, apex acute; opposite appendages converging and crossing over one another mesally. Dorsal phallic spine stout, with distinct ventral projection in about middle, apex gradually upturned; as viewed dorsally, with apex of spine acutely bifid (depressed), lateral margins of bifid apex distinctly sclerotized. Phallicata with pair of curved, narrow, spinelike projections from dorsal margin, projections hooked over paramere appendages; basoventral margin of phallicata with large upright lobe, extending transversely to dorsal margin, apex of lobe subacute. Endophallic membrane relatively elongate, with rounded membranous lobes; ventromesal spine short, bluntly rounded apically; phallotremal spines lightly sclerotized, but distinct, projecting apicolaterally. Material examined — BOLIVIA: Cochabamba: rd. to Villa Tunari, Chapare, 1300 m, 8 - 10. xii. 1984, LE Peña G – 1 male, 11 females (pinned) (NMNH); Carrasco N. P., Río San Mato, at cable crossing to park, 17.06392 ° S, 65.47558 ° W, el 449 m, 12. xi. 2004, Robertson, Garcia, Vidaurre – 1 male (pinned) (UMSP); La Paz: Quebradas de Río Zongo, 1400 m, 24 - 30. x. 1984, LE Peña G – 2 males, 8 females (pinned) (NMNH); PN and ANMI Cotapata, Estación Biol. Tunquini, Río Huarinilla, 16.20272 ° S, 67.83748 ° W, el 1253 m, 8. xii. 2004, Robertson and Valdivia – 5 males, 44 females (pinned) (UMSP); PN and ANMI Cotopata, Estación Tunquini, Quebrada El Padrini, 16.20322 ° S, 67.84487 ° W, el 1343 m, 6 - 7. xii. 2004, Robertson and Valdivia, 3 males, 16 females (pinned) (UMSP); PN and ANMI Cotopata, Estación Biol. Tunquini, Río Santa Catarina, 16.19477 ° S, 67.86805 ° W, el 1525 m, 2 - 5. xii. 2004, Robertson and Valdivia – 27 males, 195 females (pinned) (UMSP); Santa Cruz: Caballero Prov., PN Amboró, 17.83556 ° S, 64.38972 ° W, el 2050 m, 15 - 21. x. 2001, S Spector and J Ledezma – 1 male (pinned) (UMSP); PERU: Huanuco, 16. ix. 1954, EI Schlinger and ES Ross – 7 males, 11 females (alcohol) (CAS); Cuzco: Paucartambo to Pilcopata rd., Río San Pedro at Puente San Pedro, 13.05500 ° S, 71.54633 ° W, 1445 m, 24. vi. 1993, Blahnik and Pescador – 1 male (pinned) (NMNH); Paucartambo to Pilcopata rd., Quebrada Quitacalzon at Puente Quitacalzon, 13.02617 ° S, 71.49950 ° W, 1050 m, 25 - 27. vi. 1993, Blahnik and Pescador – 1 male, 1 female (pinned) (NMNH); Buenos Aires, 53 km W Pilcopata, lower montane wet forest, 2280 m, 3 - 6. xii. 1979, JB Heppner – 1 male, 1 female (alcohol) (NMNH). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB5F84AFF01BD064299F8CF.taxon	description	Fig. 61	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB5F84AFF01BD064299F8CF.taxon	distribution	Distribution — Argentina, Bolivia.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB6F848FF01B8A640F1FD48.taxon	description	Fig. 62	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB6F848FF01B8A640F1FD48.taxon	description	Adult — Length of forewing: male 4.3 - 4.7 mm; female 4.5 - 5.1 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Overall color dark brown, apices of tarsal segments whitish or pale brown. Tibial spurs slightly darker than legs, not greatly contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI very large, laterally compressed, subtriangular, ventrally projecting, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin somewhat rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short to moderate in length, lateral margins subparallel, apicomesal margin with broad U-shaped invagination, extending nearly ½ length of segment; apicolateral lobes formed by mesal invagination, apices distinctly sclerotized, narrow, subtruncate; apex, as viewed laterally, truncate, with tiny acute projection from ventral margin; ventrolateral lobes nearly obsolete. Inferior appendages with short dorsolateral projections and very elongate, narrow, sinuous, ventral projections. Mesal pockets of inferior appendage with spine-like apical processes very short, posteriorly projected. Paramere appendage very short, spine-like, tapering from base, apex acute. Dorsal phallic spine relatively narrow throughout, without distinct ventral projection, apical ½ narrow, attenuate, gradually dorsally curved; as viewed dorsally, relatively narrow throughout, apex acute. Phallicata with pair of posterolaterally curved, horn-like projections from dorsal margin; basoventral margin of phallicata with short, apically rounded, posteriorly projecting, lobe. Endophallic membrane relatively short, rounded, with pair of prominent, dorsolaterally curved, spines from ventral margin, apparently subtending apical projections of inferior appendages; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB6F848FF01B8A640F1FD48.taxon	materials_examined	Holotype male (pinned) — BOLIVIA: La Paz: PN and ANMI Cotapata, Estación Biol. Turquini, Quebrada El Padríni, 16.20322 ° S, 67.84487 ° W, el 1343 m, 6 - 7. xii. 2004, Robertson and Valdivia (UMSP 000094220) (UASC). Paratypes — BOLIVIA: La Paz: same data as holotype – 4 females (pinned), 1 male, 1 female (alcohol) (UMSP); PERU: Cuzco: Paucartambo, Quita Calzon, ca. 30 km NW Pilcopata, km 164, 13.15000 ° S, 71.36667 ° W, 1030 m, 1 - 2. ix. 1989, N Adams, et al. – 8 females (alcohol) (NMNH); Paucartambo, Quita Calzon, ca. 30 km NW Pilcopata, km 164, streamlet 50 m E Quita Calzon, 13.15000 ° S, 71.36667 ° W, 1030 m, 2. ix. 1989, N Adams, et al. – 1 male, 3 females (alcohol) (NMNH); Paucartambo to Pilcopata rd., Quebrada Quitacalón at Puente Quitacalzón, 13.02617 ° S, 71.49950 ° W, el 1050 m, 25 - 27. vi. 1993, R Blahnik & M Pescador – 1 male, 6 females (pinned) (NMNH). Etymology — This species is named M. sinuosa for the elongate sinuous ventral projections from the inferior appendages, which is a distinctive characteristic for this species. — leroda group taxa, unplaced to subgroup The following species may have their closest relationships to the limona, pocita, or punensis subgroups, but the character evidence for allying them with those subgroups (or with one another) was too weak to be anything more than speculative. Despite this, the majority of the species grouped here do share some character similarities, which might be indicative of their relationship. All of the taxa, except M. pica, n. sp. have only fork II present in the hind wing. Within the region covered, this character state is otherwise found only in species of the atenuata subgroup and in two species placed in the punensis subgroup. Similar reduced hind wing venation also occurs in the albolineata and pumila subgroups from southeastern Brazil. An additional character similarity is that, females of all of the species, except M. applanata and M. pica (for which no female was associated), have well developed pheremonal sacks on both segments VI and VII. The character state was not observed in any other taxa in the leroda group, for which the most common state is for sacks to be present on only segment VI. It does, however, characterize at least some species in the bilineata group. Possibly it reflects a plesiomorphic state for the taxa grouped here; it could also represent a character reversal, since very minute and vestigial sacks are present on segment VII of various other subgroups of the leroda group. Of the species placed here, only M. rectiflexa has dorsolateral projections on the phallicata (as in the punensis subgroup). Three of the species, M. biramosa, M. membranacea, and M. rectiflexa have elongate projections from the mesal pockets of the inferior appendages, an unusual character for leroda group tax (inferred to be plesiomorphic, but possibly a character reversal for these taxa). In its unusual combination of characters, each of the taxa placed here is unique and interesting in its own way.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB7F857FF01BB2645AAFD2F.taxon	description	Fig. 63, 101	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB7F857FF01BB2645AAFD2F.taxon	description	Adult — Length of forewing: male 2.5 - 3.2 mm; female 3.0 - 3.4 mm. Forewing with forks I, II, and III present, hind wing fork II only. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments whitish or pale brown. Tibial spurs darker than legs, somewhat contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI large, laterally compressed, subtriangular, ventrally or posteroventrally directed, length slightly greater width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X short, lateral margins subparallel, apicomesal margin nearly straight, apicolateral lobes relatively broad and short, irregular in shape; ventrolateral lobes short, rounded. Inferior appendages distinctly setose, without dorsolateral projections, apicoventral projections relatively elongate, very distinctly depressed (dorsoventrally flattened), narrow in lateral view, broad in ventral view, with narrow, attenuate apices. Mesal pockets of inferior appendage with spine-like apical processes short, posteriorly curved. Paramere appendages apparently doubled, one pair very short, other pair moderately elongate, narrow (extending about as far as dorsal phallic spine), apices acute. Dorsal phallic spine relatively short and stout, ventrally deflected in about middle, apex broadly and asymmetrically bifurcate (more or less depressed), apices of bifurcation acute. Phallicata tubular, without dorsal projections, ventral margin distinctly sclerotized. Endophallic membrane short rounded, with paired membranous lobes; ventromesal spine short, apparently nearly continuous with ventral margin of phallicata; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFB7F857FF01BB2645AAFD2F.taxon	materials_examined	Holotype male (pinned) — PERU: Madre de Dios: Manu, Pakitza, 12.11667 ° S, 70.96667 ° W, malaise trap (day collection), 250 m, 14 - 23. ix. 1988, O Flint and N Adams (UMSP 000157355) (MJP). Paratypes — PERU: Madre de Dios: same data as Holotype – 3 males, 3 females (pinned) (NMNH); Manu, Pakitza, 11.93333 ° S, 71.30000 ° W, 250 m, 19 - 23. ix. 1989, N Adams et al. – 1 male (pinned) (NMNH); Manu, Pakitza, 12.11667 ° S, 70.96667 ° W, trail 2, 1 st stream, 250 m, 14 - 23. ix. 1988. O Flint and N Adams – 16 males, 3 females (alcohol) (NMNH), 3 males, 1 female (alcohol) (UMSP). Etymology — This species is named M. applanata from the Latin word applanatus, meaning flattened, and referring to the inferior appendages of this species, which are dorsoventrally flattened.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA8F857FF01BA864236F8CF.taxon	description	Fig. 64 Mortoniella barinasi is most readily diagnosed by the elongate, curved ventral projections of the inferior appendages and by the shape and enlarged apex of the dorsal phallic spine. In both of these characters it has a similarity to M. pocita, which differs in having elongate projections from the dorsal margin of the phallicata and in lacking a ventromesal endophallic spine. The similarity of the two species may be only superficial. Adult — Length of forewing: male 2.3 - 2.8 mm; female 3.0 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3 / 4: 4. Overall color light brown (in alcohol, specimens faded). Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, subtriangular, posteroventrally directed, length about 2 times width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively short, lateral margins subparallel, apicomesal margin nearly straight, apicolateral lobes relatively broad and short, subtruncate, apicolateral margin of each lobe with small, acute point; ventrolateral lobes short, rounded. Inferior appendages with short dorsolateral projections, apicoventral projections elongate, sinuous, apices acute and curved mesally. Mesal pockets of inferior appendage with spine-like apical processes narrow, moderate in length, posteriorly projecting. Paramere appendage elongate (extending about as far as dorsal phallic spine), somewhat irregular in contour, apex very narrow, acute, with sinuous dorsal inflection. Dorsal phallic spine stout, with ventral margin very sinuously indentate to accommodate rounded projection on endophallic membrane, apex enlarged and inflated, ventral margin of apex rounded, dorsal margin somewhat hood-like. Phallicata very short, with basodorsal projection, articulating with dorsal phallic spine, laterally with prominent sclerotized, posteriorly projecting, apically rounded, lobe. Endophallic membrane with narrow, curved ventromesal spine; phallotremal spines relatively elongate, narrow, narrowly divided, emerging from rounded basal projection that articulates with sinuous ventral indentation of dorsal phallic spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA8F857FF01BA864236F8CF.taxon	materials_examined	Holotype male (alcohol) — VENEZUELA: Barinas: Río Santo Domingo, 17. ii. 1976, CM and OS Flint, Jr (UMSP 000097090) (NMNH). Paratypes — VENEZUELA: Barinas: same data as holotype – 6 males, 71 females (alcohol) (NMNH), 2 males, 6 females (alcohol) (UMSP). Etymology — This species is named M. barinasi for the state of Barinas in Venezuela, where the type specimens were collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA9F856FF01B8A6441EFB6F.taxon	description	Fig. 65, 113 Mortoniella biramosa is best diagnosed by having 2 pairs of subequal paramere appendages and paired ventral endophallic spines. Paired endophallic spines are typical of members of the bolivica subgroup, which differ in having a reflexed dorsal lobe on the inferior appendages. The relatively short, but distinctly posteriorly directed ventral process of segment VI is similar to species in the punensis subgroup. The elongate, spine-like projections from the mesal pockets of the inferior appendages are also distinctive and represent a relatively unusual character within the leroda group, most of whose species have short curved projections. Adult — Length of forewing: male 2.6 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color light brown (in alcohol). Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, subtriangular, posteriorly directed, length slightly greater than width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel, apicomesal margin with shallow V-shaped emargination, apicolateral lobes narrow, moderately elongate, subacute; ventrolateral lobes short, rounded. Inferior appendages without dorsolateral lobes, apicoventral projections moderately elongate, upturned apically, apices acute. Mesal pockets of inferior appendage with spine-like apical processes elongate, narrow, posteriorly projected. Paramere appendages doubled, appendages on each side elongate, narrow, subequal in length, apices acute, extending about as far as dorsal phallic spine. Dorsal phallic spine undulate in contour, nearly uniform in width, strongly upturned in about apical 1 / 3, apex acute or subacute. Phallicata moderately elongate, without basodorsal projection, basoventral margin with short rounded lobes on either side. Endophallic membrane with membranous or lightly sclerotized basodorsal lobes and projecting, divided apex, ventrally with pair of prominent curved spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA9F856FF01B8A6441EFB6F.taxon	materials_examined	Holotype male (alcohol) — VENEZUELA: Barinas: Río Sinigüis in Caño Grande, 8.4000 ° N, 70.77417 ° W, el 520 m, 1997. iii. 22, Holzenthal (UMSP 000092464) (UMSP). Etymology — This species is named M. biramosa, from the Latin ramosus, or branch, and referring to the doubled paramere appendages that characterizes this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA9F855FF01BD4642E5FC4F.taxon	description	Fig. 66 This species has the distinctive feature of having the apex of the dorsal phallic spine modified into a membranous lobe, and can be diagnosed on that basis alone, since it is found in no other described species. Other characters, considered in combination, are also distinctive and do not suggest a close relationship of M. membranacea to any other described species. Characters of note, most probably plesiomorphic for the leroda group, are the simple tubular phallicata, without distinct processes, the elongate, spine-like projections from the mesal pockets of the inferior appendages, the absence of endophallic spines, the general shape of tergum X, which has a general similarity to species in the florica / leroda subgroups, and the short, posteriorly projected, ventral process of segment VI. It also lacks fork III in the hind wing, a presumably derived character state. Adult — Length of forewing: male 2.9 - 3.2 mm; female 3.2 - 3.6 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color light brown, apices of tarsal segments whitish or pale brown. Tibial spurs darker than legs, contrasting in color. Wing bar at anastamosis indistinct, marked with light brown setae, most evident at arculus. Male genitalia — Ventral process of segment VI laterally compressed, subtriangular, posteriorly directed, length slightly greater than width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel, apicomesal margin nearly straight, apicolateral lobes narrow, moderately elongate, acute or subacute apically, somewhat mesally curved from lateral margin; ventrolateral lobes short, rounded. Inferior appendages with short dorsolateral lobes, apicoventral projections moderately elongate, narrow, acute apically. Mesal pockets of inferior appendage with spine-like apical processes narrow, elongate, posteriorly projecting. Paramere appendage elongate (extending about as far as dorsal phallic spine), uniformly narrow, acute apically, dorsally inflected at base and ventrally curved apically. Dorsal phallic spine nearly uniform in width, strongly upturned in about apical ¼, upturned apex with projecting membranous posterior lobe. Phallicata simple in structure, elongate, tubular, without distinct projections. Endophallic membrane without spines, but with conspicuous membranous basodorsal lobes (apparently subtending apices of paramere appendages); phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA9F855FF01BD4642E5FC4F.taxon	materials_examined	Holotype male (pinned) — BOLIVIA: Santa Cruz: PN and ANMI Amboró, Guardia Parque Mataracú, Quebrada Verde Uno, 17.55389 ° S, 63.86917 ° W, el 374 m, 19 - 23. xi. 2004, Robertson, Garcia and Vidaurre (UMSP 000094105) (UASC). Paratypes — BOLIVIA: Santa Cruz: same data as Holotype – 6 males, 65 females (pinned); 7 males (alcohol) (UMSP), 1 male (pinned) (NMNH); PN and ANMI Amboró, Guardia Parque Mataracú, Río Yapacaní, 17.52072 ° S, 63.86795 ° W, el 329 m, 26. xi. 2004, Robertson, Garcia and Vidaurre – 21 males (alcohol) (UMSP); same locality and collectors, 28. xi. 2004 – 1 male, 12 females (pinned) (UMSP); PN and ANMI Amboró, Guardia Parque Mataracú, Las Cataratas, 17.56972 ° S, 63.85013 ° W, el 375 m, 24 - 25. xi. 2004, Robertson, Garcia and Vidaurre – 1 male, 9 females (pinned) (NMNH). Etymology — This species is names M. membranacea for the membranous apex of the dorsal phallic spine, a diagnostic character for this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAAF853FF01BC264446FE0F.taxon	description	Fig. 67 Mortoniella pica has the general appearance of species in the limona subgroup, with enlarged, paired spines (possibly modified phallotremal spines) on the endophallic membrane, and with the apex of the dorsal phallic spine compressed and modified. However, it lacks the rounded apicomesal processes of tergum X that characterize members of the limona subgroup, and it possesses a small ventromesal spine on the endophallic membrane, which is not present in any of described species of that subgroup. However, it is conceivable that it could be a relatively basal or unusually modified species of the limona subgroup. Regardless of its affinity, M. pica is readily diagnosed by the structure of the apex of the dorsal phallic spine, which has a very acute ventral projection, unlike any other species of Mortoniella. Adult — Length of forewing: male 3.6 mm. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0: 4: 4. Color (in alcohol) medium brown. Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, subtriangular, posteriorly directed, length slightly greater than width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin rounded in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, base with raised mesal process bearing long apical setae, lateral margins subparallel, apicomesal margin broadly U-shaped, apicolateral lobes narrow, moderately elongate, acute apically, somewhat mesally curved from lateral margin; ventrolateral lobes rounded, projecting. Inferior appendages with short, subtruncate dorsolateral lobes and short, subtriangular ventromesal projection. Mesal pockets of inferior appendage with spine-like apical processes short, posterodorsally curved. Paramere appendage moderate in length, shorter than dorsal phallic spine, relatively stout, uniform in width, acute apically, weakly arched and downcurved. Dorsal phallic spine with slight ventral protrusion in basal ½, upturned in about apical ¼, apex acute, ventral margin with acute projection at point of inflection. Phallicata short, weakly sclerotized, simple in structure, with short basodorsal projection. Endophallic membrane with short curved ventromesal spine and pair of large sclerotized endophallic spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAAF853FF01BC264446FE0F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pastaza: Puyo (27 km N), Estación Fluvia Metrica, 4. ii. 1976, Spangler et al. (UMSP 000097063) (NMNH). Etymology — The name M. pica was suggested by the apex of the dorsal phallic spine of this species, somewhat resembling a pike or a woodpecker’s bill (family Picidae). Mortoniella (Mortoniella) rectiflexa, new specie s Fig. 68 This is a very distinctive species, unlikely to be confused with any other described species. Especially diagnostic is the strongly upturned and widened apex of the dorsal phallic spine, which is acute apically and has minute spines on its posterior (ventral) surface. Equally diagnostic are the helical projections from the dorsal margin of the phallicata, which cause the paramere appendages to cross over one another. This character could indicate a relationship to the punensis subgroup, but other character evidence is equivocal. Adult — Length of forewing: male 2.9 - 3.4 mm; female 3.0 - 3.9 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 4: 4. Overall color (in alcohol) medium brown. Wing bar not evident. Male genitalia — Ventral process of segment VI large, laterally compressed, subtriangular, posteriorly directed, length about 1 ½ to 2 times width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin distinctly rounded in dorsal ½, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel, apicomesal margin with shallow indentation, apicolateral lobes moderately elongate, subacute apically, weakly mesally curved from lateral margins; ventrolateral lobes rounded, distinctly projecting. Inferior appendages short, without distinct dorsolateral or apicoventral projections. Mesal pockets of inferior appendage with spine-like apical processes relatively elongate, posteriorly projecting, subtending ventral margin of phallicata. Paramere appendage elongate (longer than apical inflection of dorsal phallic spine), nearly uniformly narrow, apex acute; opposite appendages converging and crossing one another mesally. Dorsal phallic spine stout, with only slight ventral bulge in about middle, apex very abruptly and nearly rectilinearly upturned in about apical 1 / 3, spine slightly widened at point of inflection, narrowing to acute apex, posterior margin of flexed apex with numerous small spines; as viewed dorsally, very distinctly widened in about middle, less so at apical inflection. Phallicata with pair of narrow, helically curved, spine-like projections from dorsal margin, projections hooked over paramere appendages, causing them to cross over one another mesally. Endophallic membrane without ventromesal spine; phallotremal spines very small. Holotype male (pinned) — ECUADOR: Pastaza: Puyo (27 km N) Estación Fluvia Metrica, 4. ii. 1976, Spangler, et al. (UMSP 000146419) (NMNH). Paratypes — ECUADOR: Esmeralda: La Union, 3. ii. 1979, JJ Anderson – 2 males, 1 female (alcohol) (NMNH); Napo: Tena (17 km SW), 28. v. 1977, PJ Spangler and DR Givens – 1 male, 8 females (alcohol) (NMNH); Puyo, 6. v. 1977, PJ Spangler and DR Givens – 1 male, 7 females (alcohol) (NMNH); Pastaza: Puyo, 5. v. 1977, PJ Spangler and DR Givens – 4 males, 20 females (alcohol) (NMNH); same locality and collectors, 6. v. 1977 – 2 males, 1 female (alcohol) (NMNH); same locality and collectors, 8. v. 1977 – 1 male (alcohol) (NMNH); same locality and collectors, 10. v. 1977 – 1 male, 1 female (alcohol) (NMNH); same locality and collectors, 13. v. 1977 – 1 male, 6 females (alcohol) (NMNH); same locality and collectors, 14. v. 1977 – 1 male, 2 females (alcohol) (NMNH); same locality and collectors, 16. v. 1977 – 1 male (alcohol) (NMNH); same locality and collectors, 21. v. 1977 – 5 males, 9 females (alcohol) (UMSP); Puyo, 30. i. 1976, Spangler et al. – 8 males, 11 females (alcohol) (NMNH); Puyo, malaise trap, 8 - 10. ii. 1976, Spangler et al. – 1 male (alcohol) (NMNH); Puyo (27 km N), Est. Fluvia Metrica, 4. ii. 1976, Spangler, et al. – 3 males, 1 female (pinned), 14 males, 6 females (alcohol) (NMNH); Puyo (3 km W) 15. vii. 1976, J Cohen – 6 males, 20 females (alcohol) (NMNH); Zamora- Chinchipe: Zamora, at lights, 1 - 5. vi. 1976, A Langley et al. – 1 male (pinned) (NMNH); Zamora, 4. xii. 1978, JJ Anderson – 6 males, 15 females (alcohol) (NMNH). Etymology — This species is named M. rectiflexa for the apex of the dorsal phallic spine, which is bent or flexed at nearly a right angle and is particularly diagnostic.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFACF852FF01BA66430EFEA8.taxon	description	— argentinica subgroup Included species: M. argentinica (Schmid); M. cornuta, n. sp.; M. croca, n. sp.; M. curvistylus, n. sp.; M. spinulata (Flint). The similarity of Mortoniella argentinica (Schmid) to M. spinulata (Flint) was discussed in a previous paper (Blahnik and Holzenthal 2011) and the reader is referred to that paper for a redescription and reillustration of M. argentinica. Mortoniella spinulata and 3 related new species are described below. Of the species assigned to this subgroup, only M. argentinica has an elongate, narrow ventral process on segment VI; the others have the process more or less trianguloid, relatively short and wide basally and more or less ventrally projecting, thus more like species in the leroda group. Females of M. argentinica and M. cornuta, n. sp., which are the only species of the subgroup for which females were available, have genitalia similar to those in the leroda group, without an invaginated dorsal margin of segment VIII. All of the species in the argentinica subgroup, except M. curvistylus, n. sp., are characterized by having 2 pairs of paramere appendages (considered an apomorphy for the subgroup); elongate, narrow, lance-like projections from the mesal pockets of the inferior appendages; and an elongate, recurved dorsal lobe on the inferior appendages. As discussed above, the latter characters may be plesiomorphic for the subgenus Mortoniella as a whole, though absent or lost in the majority of taxa, in both the leroda and bilineata groups. The general shape of segment IX of species in the argentinica subgroup resembles species of the leroda species group: posterior margin uniformly rounded and with the lobes narrowly separated dorsally, or relatively narrowly separated. Most of the species seem to have at least a small mesal apodeme on the dorsal margin of the phallobase, as in species of the bilineata group. Species lack distinct dorsal projections on the base of the phallicata and also lack a ventromesal endophallic spine, although sclerotized phallotremal spines or paired endophallic spines (possibly modified phallotremal spines) are present. On balance, the subgroup seems to be more closely related to the leroda group than to the bilineata group.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFADF852FF01B90644C7F9AF.taxon	description	Fig. 69, 99 Mortoniella cornuta, n. sp. is most similar to M. spinulata (Flint), especially in the shape of tergum X, which has a very narrow apicomesal cleft, apparently to accommodate a sharply upturned and much narrowed apex of the dorsal phallic spine. The most distinctive and diagnostic character of this species is a pair of conical horn-like projections emerging from the lateral margins of the dorsal phallic spine at about midlength. Mortoniella cornuta also differs from M. spinulata in that the reflexed dorsal branch of inferior appendages is longer and lacks spines; also, the paired paramere appendages are more or less subequal in length, rather than being distinctly different in length and shape. Adult — Length of forewing: male 3.6 - 4.0 mm; female 3.9 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Head distinctly small. Overall color dark brown, apices of mesotarsal segments whitish. Tibial spurs slightly darker than legs, weakly contrasting in color. Wing bar at anastamosis marked with white setae. Male genitalia — Ventral process of segment VI laterally compressed, large, subtriangular, ventrally directed, length subequal to width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively short, basally with rounded elevation, lateral margins converging apically, apicomesal margin with very narrow incision, apicolateral lobes very narrowly separated, acute apically; ventrolateral lobes prominent, rounded. Inferior appendages with very elongate, narrow posteriorly recurved, dorsolateral projections, apicoventral projections absent. Mesal pockets of inferior appendage with spine-like apical processes narrow, very elongate, posteriorly projecting. Paramere appendages doubled, both elongate (extending about as far as dorsal phallic spine), more lateral one very narrow basally, slightly widened in apical ½, mesal one narrow, more distinctly widened preapically, apices of both acute. Dorsal phallic spine widened in middle, with very distinct conical lateral projections, apical 1 / 3 distinctly dorsally inflected and very narrow, apex acute. Phallobase with small, but distinct, lightly sclerotized mesal apodeme. Phallicata simple in structure, elongate tubular, with short basodorsal projection. Endophallic membrane elongate, without ventromesal spine; phallotremal spines small, distinct, apical.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFADF852FF01B90644C7F9AF.taxon	materials_examined	Holotype male (pinned) — ECUADOR: Tungurahua: 13 km E Baños, el 1550 m, 15. ix. 1990, OS Flint, Jr (UMSP 000146416) (NMNH). Paratypes — ECUADOR: Tungurahua: same data as holotype – 3 males, 1 female (pinned) (NMNH). Etymology — This species is named M. cornuta from the Latin cornu, a horn, and referring to the conical horn-like processes on the lateral margin of the dorsal phallic spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFADF851FF01BE064364FB4F.taxon	description	Fig. 70	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFADF851FF01BE064364FB4F.taxon	description	Adult — Length of forewing: male 3.5 - 3.7 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Head distinctly small. Overall color dark brown, apices of mesotarsal segments whitish. Tibial spurs slightly darker than legs, weakly contrasting in color. Wing bar at anastamosis marked with white setae. Male genitalia — Ventral process of segment VI laterally compressed, large, subtriangular, posteroventrally directed, length subequal to width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, basally with rounded elevation, lateral margins subparallel, apicomesal margin with very wide, V-shaped emargination, apicolateral lobes subacute, slightly mesally curved; ventrolateral lobes relatively prominent, rounded. Inferior appendages with dorsal projection apparently vestigial, lightly sclerotized, very narrow, attenuate, posteriorly curved. Mesal pockets of inferior appendage with spine-like apical processes narrow, very elongate, posteriorly projecting. Paramere appendages doubled, one pair elongate (extending about as far as dorsal phallic spine), slightly widened in apical ½, apex acute, lateral pair about ½ length of other pair, very narrow, apex distinctly hooked. Dorsal phallic spine, in lateral view, narrow, attenuate, apical ½ retrorsely recurved; in dorsal view, relatively narrow throughout. Phallobase with small, but distinct, lightly sclerotized mesal apodeme. Phallicata simple in structure, with pair of short rounded basoventral projections. Endophallic membrane elongate, with prominent paired ventrolateral membranous lobes, phallotremal spines small, apical.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFADF851FF01BE064364FB4F.taxon	materials_examined	Holotype male (pinned) — PERU: Cuzco: Paucartambo, Puente San Pedro, ca. 50 km W Pilcopata, 1600 m, 2 - 3. ix. 1988, O Flint and N Adams (UMSP 000157345) (MJP). Paratypes — PERU: Cuzco: same data as Holotype – 1 male (pinned) (NMNH). Etymology — This species is named M. croca, from the Medieval Latin word croca, a shepherd’s crook (used as a noun in apposition), referring to the shorter of the two paramere appendages in this species, which is very distinctly hooked apically.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAEF850FF01BD2644C7FC8F.taxon	description	Fig. 71 Mortoniella curvistylus has a general similarity to other species in the argentinica subgroup, particularly in the possession of an elongate and strongly curved dorsal process on the inferior appendages and very elongate spine-like processes from the mesal pockets of the inferior appendages. Unlike the other species in this group, it seems to have only 1 pair of paramere appendages (possibly a secondary loss?). It has a relatively narrow V-shaped emargination of tergum X, but wider and more distinct than in either M. cornuta, n. sp. or M. spinulata (Flint). The most unusual and diagnostic feature of this species is that the dorsal phallic spine seems to be fused or semi-fused to the endophallic membrane, from which point a retrorse spine emerges that articulates with a depression on the basodorsal margin of the phallicata. Adult — Length of forewing: male 3.8 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Head distinctly small. Overall color dark brown. Tibial spurs slightly darker than legs, contrasting in color. Wing bar at anastamosis marked with white setae. anterolaterally, length greatest midlaterally, posterolateral margin slightly produced, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderate in length, basally with rounded elevation, lateral margins subparallel, apicomesal margin with narrow V-shaped incision, apicolateral lobes formed by apical incision, apices of lobes somewhat ventrally curved, subacute; ventrolateral lobes prominent, rounded. Inferior appendages with very elongate, narrow, posteriorly recurved, dorsolateral lobes, apicoventral projections absent. Mesal pockets of inferior appendage with spine-like apical processes narrow, very elongate, posteriorly projecting. Paramere appendage single on each side, narrow, elongate (extending about as far as dorsal phallic spine), distinctly widened preapically, widened preapical expansion with several spines, apex acute. Dorsal phallic spine, in lateral view, almost uniformly narrow, apical 1 / 3 weakly upturned, spine possibly fused or semifused to phallicata at inflection; as viewed dorsally, relatively broad in middle, gradually narrowing apically, apex acute. Phallobase with very small, lightly sclerotized, mesal apodeme. Phallicata relatively elongate, tubular, apicodorsal margin (or base of endophallic membrane) with retrorse, spine-like projection, emerging at inflection point of dorsal phallic spine, apex of spine articulating with basodorsal margin of phallicata. Endophallic membrane short, simple in structure, without spines; phallotremal spines apparently absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAEF850FF01BD2644C7FC8F.taxon	materials_examined	Holotype male (pinned) — ECUADOR: Zamora-Chinchipe: 30 km E Loja, el 2000 m, 23. ix. 1990, OS Flint, Jr (UMSP 000146418) (NMNH). Etymology — This species is named M. curvistylus from the Latin curvus, meaning bent, and stylus, a sharp pointed instrument, and referring to the unusual retrorse spine on the endophallic membrane that hooks backward to contact a depression in the dorsal margin of the phallicata.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAFF85FFF01BBE643C0F9AF.taxon	description	Fig. 72	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFAFF85FFF01BBE643C0F9AF.taxon	description	Adult — Length of forewing: male 4.1 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Head distinctly small. Overall color dark brown. Tibial spurs slightly darker than legs, contrasting in color. Wing bar at anastamosis indistinctly marked with white setae. Male genitalia — Ventral process of segment VI laterally compressed, large, subtriangular, ventrally directed, length subequal to width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short with very elongate setae, basally with rounded elevation, ventrolateral lobes prominent, projecting laterally; overall shape of tergum, in dorsal view, subquadrate, with mesal pair of very narrowly separated, spine-like, apicolateral lobes; apices of ventrolateral lobes angular, prominent. Inferior appendages with elongate, narrow, posteriorly recurved, dorsolateral lobes, with row of spines in apical ½, apex of dorsolateral lobe very narrowly attenuate; apicoventral projections absent. Mesal pockets of inferior appendage with spine-like apical processes elongate, posteriorly projecting. Paramere appendages doubled, one pair elongate, narrow (extending about as far as dorsal phallic spine), slightly widened in apical ½ and strongly mesally curved, widened apex covered with minute spines; second, lateral pair about ½ length of first pair, very narrow throughout, distinctly hooked apically, apex acute. Dorsal phallic spine distinctly widened in middle in both lateral and dorsal views, apical 1 / 3 dorsally inflected and very narrow, apex acute, ventral margin of spine at apical inflection with distinct small tubercle or protuberance, articulating with sclerotized depression in dorsal margin of endophallic membrane. Phallobase with small, lightly sclerotized, mesal apodeme. Phallicata tubular, with pair of short rounded ventrolateral projections and additional pair of rounded ventromesal projections near apex. Endophallic membrane elongate, with depressed dorsal sclerite, accommodating reflexed apex of dorsal phallic spine; phallotremal spines small, distinct, apical. Material examined — COLOMBIA: Antioquia: Quebrada Espadera, 7 km N Medellín, 24. ii. 1983, OS Flint, Jr – 2 male Paratypes (pinned) (NMNH). — esrossi subgroup Included species: Mortoniella esrossi, n. sp. The single species listed here is placed in its own subgroup because it is morphologically distinct from other species. It is characterized by having paramere appendages that lack typical rounded basal processes; instead, the appendages lie parallel to the dorsal phallic spine and are each accompanied by a projecting ventral lobe (which could be a modification of the usual basal process). Mortoniella esrossi is also unusual in that the dorsal phallic spine seems to have a ventral “ foot ” in its apical part. This may reflect a fusion of the dorsal phallic spine with dorsal margin of the endophallic membrane. A similar development seems to occur in M. curvistylus, n. sp. in the argentinica subgroup, but whether an actual fusion occurs in either species is not easily determinable. An additional unusual feature of M. esrossi is a large endophallic spine with two branches, which may be a modification of the small phallotremal spines found in some other taxa. Like a number of the “ unplaced species ” discussed here, the inferior appendages have narrow, reflexed dorsal lobes, and spine-like projections from the mesal pockets of the inferior appendages that are relatively elongate. The presence of these features in what are otherwise very divergent taxa suggests that they may be plesiomorphic for the subgenus Mortoniella in general, in which case they would have had to been lost in various lineages, including the majority of taxa in both the leroda and bilineata groups. Other aspects of the morphology of M. esrossi suggest a possible relationship to the tridens subgroup, but these are admittedly subject to interpretation. Among these are the paramere appendages, which seem to be in a transitional stage of fusing to the dorsal phallic spine; also, the projecting ventral lobe at the base of each of the appendages is similar to that from which a second pair of appendages emerges in the tridens subgroup. If the latter represents a homology, a second pair of appendages may also have been present in the ancestor of M. esrossi, but secondarily lost. An additional character suggesting a relationship is the more or less subquadrate tergum X found in both M. esrossi and members of the tridens subgroup.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA0F85EFF01BE064247F9EF.taxon	description	Fig. 73 This species is best diagnosed by the reflexed dorsal lobe of the inferior appendages, subquadrate tergum X, without distinct apicolateral processes, and by the structure of the dorsal phallic spine, which has a projecting ventral “ foot ” in its apical ½. Adult — Length of forewing: male 3.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V; both wings broadly rounded apically. Spur formula 0: 4: 4. Color (in alcohol) yellowish brown (specimen faded and partially denuded). Wing bar not evident. Male genitalia — Ventral process of segment VI laterally compressed, narrow, elongate, posteriorly directed, length about 2 times width at base, apex subacute. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X moderately elongate, subquadrate, lateral margins subparallel, apicomesal margin nearly straight, apicolateral lobes absent (or not evident in dorsal view); ventrolateral lobes elongate, more or less truncate apically. Inferior appendages with elongate, narrow, posteriorly recurved, dorsolateral projections, reflexed apices attenuate, lightly sclerotized, acute apically, apicoventral projections absent. Mesal pockets of inferior appendage with spine-like apical processes narrow, elongate, posteriorly projecting; as viewed ventrally, sinuously curved at base, subparallel apically. Paramere appendage elongate (extending about as far as dorsal phallic spine), narrow, nearly uniform in width, acute apically. Basal segment of parameres not evident as such, but with projecting, rounded, flattened lateral lobes. Dorsal phallic spine, as viewed laterally, nearly uniform in width, apex slightly upturned, acute, ventral margin at about apical 1 / 3, with foot-like projection, possibly fused to dorsal margin of endophallic membrane; spine, as viewed dorsally, relatively broad, apex very acutely narrowed. Phallicata simple in structure, without distinct projections. Endophallic membrane with prominent, forked sclerite (possibly modified phallotremal spines), apices of branches acute.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA0F85EFF01BE064247F9EF.taxon	materials_examined	Holotype male (alcohol) — COLOMBIA: 3 mi W Villavicencio, 11. iii. 1955. EI Schlinger and ES Ross (UMSP 000095053) (CAS). Etymology — This species is named M. esrossi in honor of Edward S. Ross, co-collector of the type specimen. — proakantha subgroup Included species: Mortoniella proakantha, n. sp. Except for its primitive hind wing venation (3 forks present, II, III, and V) and the very elongate, narrow ventral process of segment VI, this species could easily pass as a member of the akantha subgroup of the leroda species group. Characters suggestive of this include the overall structure of the inferior appendages, with a symmetrical ventromesal projection and e l o n g a t e, r e f l e x e d d o r s a l l o b e s. T h e l a t t e r c h a r a c t e r, a s d i s c u s s e d a b o v e, i s l i k e l y a plesiomorphic character for the subgenus Mortoniella (present in the immediate ancestor of the group). Other character similarities to the leroda group include a segment IX with uniformly rounded anterior margin and with lobes narrowly separated dorsally; short, curved, spine-like projections from the mesal pockets of the inferior appendages; and the general shape of tergum X, with a rounded mesal excavation and projecting lateral lobes. Female genitalia for the species are highly unusual, and unlike any other species of Mortoniella examined, in having a segment VIII with elongate, wiry setae and in having a segment X that is bulbous, rather than flattened and fused to segment IX (Fig. 114). It is conceivable that the female is incorrectly associated, but there is no other described species within Protoptilinae that it could be assigned to. Despite the name given to the included species, a basal or even close relationship to the akantha group cannot be inferred and we are uncertain about its actual relationship.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA1F85DFF01BEC6425BF8CF.taxon	description	Fig. 74, 114 Mortoniella proakantha bears an overall similarity to members of the akantha subgroup of the leroda group, particularly in the shape of the inferior appendages, which have a symmetrical mesal projection and narrow, recurved dorsolateral projections, and also have mesal pockets with short spine-like ventral projections. It differs diagnostically in having fork V of the hind wing present (Cu 1 forked apically) and in having a very elongate, narrow ventral process on segment VI (both probably plesiomorphic characters). Additionally, the dorsal phallic spine is unusual in having a long curved basal stalk (as in many members of the bilineata group) and an apex that is bifid and has many minute spines. Adult — Length of forewing: male 3.5 - 3.7 mm; female 4.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Color (in alcohol) brown. Tibial spurs darker than legs, contrasting in color. Wing bar not evident. Male genitalia — Ventral process of segment VI very narrow and elongate, posteriorly directed, length about 6 times width at base, apex acute. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin broadly rounded, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apicomesal margin with relatively shallow U-shaped or V-shaped emargination, apicolateral lobes short, inset from lateral margin, apices distinctly sclerotized and somewhat ventrally curved; ventrolateral lobes narrow, elongate, projecting, rounded apically, with elongate setae. Inferior appendages with very elongate, narrow, posteriorly recurved, dorsolateral projections, somewhat detached or inset from lateral margin, apices acute, with few minute spines, apicoventral projection of appendages prominent, elongate, apex subtruncate. Mesal pockets of inferior appendage with spine-like apical processes short, posterodorsally curved. Paramere appendage relatively short (much shorter than dorsal phallic spine), slightly widened in apical ½, apex acute, with numerous small spines-. Basal segment of paramere enlarged and elevated, subtending dorsal phallic spine at apex of long basal stalk. Dorsal phallic spine, as viewed laterally, with elongate, narrow basal stalk, apical ½ narrow, undulate in contour, apex with minute spines, slightly upturned; spine, as viewed dorsally, narrow throughout (basal ½ very narrow), apex acutely bifid, with minute spines. Phallicata narrow basally, expanded dorsally to form rounded elevation, apparently articulating with dorsal phallic spine. Endophallic membrane simple, without spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA1F85DFF01BEC6425BF8CF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Napo: OY 10 unnamed trib. to Oyacachi R., ca. 5.2 mi W of Oyacachi (UMSP 000097164) (NMNH). Paratypes — ECUADOR: Napo: same data as holotype – 1 male, 1 female (alcohol) (NMNH). Etymology — This species is named M. proakantha, from the Latin or Greek prefix pro -, meaning before, and referring to its retention of primitive characters and overall resemblance to species in the akantha subgroup. — santiaga subgroup Included species: Mortoniella acutiterga, n. sp.; M. santiaga Sykora. The members of this subgroup are very unusual and have no obvious close relationship to other subgroups. Mortoniella santiaga was placed by Sykora in the flinti subgroup of the bilineata group. However, it has very few character similarities to either members of the flinti subgroup or to the bilineata group, other than its hind wing venation (Cu 1 forked or fork V present) and the narrow, elongate ventral process of segment VI, both undoubtedly plesiomorphic characters. Characters suggestive of a relationship to the bilineata group include the short, paired, sclerotized, ventral lobes of the phallicata and the sharply angled ventrolateral lobe of tergum X. Characters suggestive of a relationship to the leroda group include the overall shape of segment X (broadly rounded laterally and with lobes narrowly separated dorsally), the very short spine-like projections from the mesal pockets of the inferior appendages, and the absence of a dorsal apodeme on the phallobase. None of these characters are decisive. The rounded, elevated base of the phallicata somewhat resembles that of M. proakantha, n. sp. and M. unilineata Sykora, but whether this reflects a relationship is also uncertain. All of these species also have a very elongate, narrow ventral process on segment VI.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA3F85CFF01B8A64136FAEF.taxon	description	Fig. 75 This species is apparently very closely related to M. santiaga Sykora. Unfortunately, the type of M. santiaga could not be located for a direct comparison. Both species are characterized by an unusual and expanded, almost laciniate, apicolateral margin of the dorsal phallic spine, and arched, spine-like apices of tergum X. Diagnostic differences, based on the description of M. santiaga, include the form of the paramere appendages, which lack projections in M. acutiterga, and the structure of tergum X, which has the ventrolateral projections more narrow and spine-like. Adult — Length of forewing: male 4.2 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 3: 4. Color (in alcohol) yellowish brown. Tibial spurs darker than legs, contrasting in color. Wing bar not evident. Male genitalia — Ventral process of segment VI very narrow elongate, posteriorly directed, length about 6 times width at base, apex somewhat depressed, acute as viewed laterally, rounded as viewed ventrally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X moderate in length, lateral margins somewhat converging apically, apicomesal margin with very short, acute projection, apicolateral lobes distinctly sclerotized, diverging as viewed dorsally, elongate tapering, acute apically, ventrally curved; ventrolateral lobes with one or two narrow projections with prominent apical setae. Inferior appendages short and very strongly fused to phallic ensemble, lateral margins slightly projecting, narrowing to acute ventromesal projection. Mesal pockets of inferior appendage with spine-like apical processes short, posteriorly projecting. Paramere appendage elongate (surpassing dorsal phallic spine), uniformly narrow, apex acute, dorsally curved apically. Dorsal phallic spine, as viewed laterally, relatively stout, curved basally, gradually dorsally curved apically, preapically with lateral expansion with numerous minute spines or scale-like projections; as viewed dorsally, relatively narrow throughout, except for preapical lateral expansion. Phallicata narrow basally, expanded dorsally to form rounded elevation, apparently articulating with dorsal phallic spine, ventrally with retrorse lateral projections. Endophallic membrane simple, without spines; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA3F85CFF01B8A64136FAEF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Napo: PAP 8 unnamed trib. Papallacta R., HwyE- 28, ca. 1.7 mi SW Papallacta, 0.385893 ° W, 78.143530 ° W, 25. i. 2012, B Gill (UMSP 000097163) (NMNH). Etymology — This species is named M. acutiterga for the very acutely angled apicolateral lobes of tergum X.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA3F85BFF01BDC643B5FC6F.taxon	distribution	Distribution — Ecuador. — tridens subgroup Included species: Mortoniella tridens, n. sp., M. triramosa, n. sp. The two species included here constitute a closely related species pair and are unusual in that the dorsal phallic spine is apparently 3 - partite, deeply divided to form a mesal and 2 lateral projections. A similar character state seems to be indicated in the illustration of M. armata (Jacquemart). The overall illustration of that species is so inadequate that it is difficult to make direct comparisons, but it does not seem to be closely related and we have speculatively placed it in the punensis subgroup (see for more extensive explanation). An unusual feature of the tridens subgroup is that the basal segment of the parameres is flattened and fused to the dorsal margin of the phallobase, appearing as a projecting, flattened lobe. This is connected to the lateral paramere appendage by an elongate membrane, such that the base of the paramere appendage emerges ventrally to the phallic ensemble, with the apical part strongly dorsally curved. A plausible explanation for the tripartite dorsal phallic spine is that the original paramere appendage structure was doubled, as in members of the argentinica subgroup, and the dorsolateral pair, along with the basal structure of the paramere, became fused with the dorsal phallic spine, producing the tripartite assemblage. On this basis, we speculate that the two groups may be related, with M. esrossi, n. sp. (in the esrossi subgroup) as a possible connecting species (which see for a more complete explanation). Like most members of the argentinica subgroup, the ventral process of the species in the tridens subgroup is large, subtriangular, and ventrally projected, thus resembling species in the leroda group. It seems likely that both of these subgroups are related to that lineage. Unlike species in the argentinica subgroup, the inferior appendages of species in the tridens subgroup are very strongly fused to the phallicata, forming a short setose ventromesal projection, and lack narrow, reflexed dorsolateral lobes. Additionally, the spine-like projections from the mesal pockets of the inferior appendages are not nearly so elongate.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA4F85AFF01BC4643C3FE2F.taxon	description	Fig. 76	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA4F85AFF01BC4643C3FE2F.taxon	description	Adult — Length of forewing: male 3.2 - 3.3 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color dark brown, Tibial spurs slightly darker than legs, weakly contrasting in color. Wing bar at anastomosis distinct, marked with white setae. Male genitalia — Ventral process of segment VI laterally compressed, very large, subtriangular, ventrally directed, length subequal to width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced in dorsal ½, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively short, subquadrate, with pair of small raised setose processes at about midlength, lateral margins subparallel, apicolateral lobes reduced to small trianguloid processes, inset from lateral margin, acute apically; ventrolateral lobes prominent, rounded. Inferior appendages short, strongly fused to phallicata, distinguishable as such mostly by setae, lateral lobes obsolete, apicomesal margin slightly projecting, bluntly truncate. Mesal pockets of inferior appendage with spine-like apical processes relatively short, narrow, sinuous, posteriorly projecting. Paramere appendage moderately elongate, narrow, with base displaced ventrally on membranous lobe, apex posterodorsally curved. Dorsal phallic spine cleft apically to form 3 narrow branches, mesal one dorsally inflected, lateral ones subparallel basally, divergently and laterally curved near apex. Phallicata with pair of small rounded ventral lobes near apex. Endophallic membrane elongate, without membranous lobes or ventral spine; phallotremal spines very small, indistinct, apical.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA4F85AFF01BC4643C3FE2F.taxon	materials_examined	Holotype male (pinned) — PERU: Cuzco: Paucartambo; Pilcopata to Atlaya, 4. ix. 1988, O Flint and N Adams (UMSP 000157321) (MJP). Paratype — PERU: Cuzco: same data a holotype – 1 male (pinned) (NMNH). Etymology — This species is named M. tridens, from the Latin word for a fork with three tines, and referring to the tripartite structure of the dorsal phallic spine.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA5F859FF01B9864114FDA8.taxon	description	Fig. 77 This species is closely related to M. triden s, n. sp., as discussed in the diagnosis for that species. It can be distinguished by its shorter paramere appendages and more elongate inferior appendages, as well as by having a decurved ventromesal projection near the apex of the phallicata, rather than a pair of apicolateral lobes. It also lacks the wart-like projections on tergum X found in M. tridens. Adult — Length of forewing: male 4.0 mm. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Overall color medium brown, apices of tarsal segments paler. Tibial spurs slightly darker than legs, weakly contrasting in color. Wing bar at anastomosis distinct, marked with white setae. Male genitalia — Ventral process of segment VI laterally compressed, very large, subtriangular, ventrally directed, length subequal to width at base, apex acute, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin slightly produced in dorsal ½, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X relatively short, subquadrate, lateral margins subparallel, apicolateral lobes reduced to small trianguloid processes, inset from lateral margin, acute apically, barely visible in dorsal view; ventrolateral lobes prominent, rounded. Inferior appendages strongly fused to phallicata, distinguishable as such mostly by setae, lateral lobes obsolete, apicomesal margin slightly projecting. Mesal pockets of inferior appendage with spine-like apical processes moderately elongate, sinuous, posteriorly projecting. Paramere appendage short, narrow, with base displaced ventrally on membranous lobe, apex posterodorsally curved. Dorsal phallic spine cleft apically to form 3 narrow branches, mesal one dorsally inflected, lateral ones subparallel basally, divergently and laterally curved near apex. Phallicata with small ventromesal projection at apex. Endophallic membrane elongate, without membranous lobes or ventral spine; phallotremal spines only suggestively developed, very lightly sclerotized.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA5F859FF01B9864114FDA8.taxon	materials_examined	Holotype male (pinned) — BOLIVIA: Yungas La Paz: Circuata to Cajuata, 2400 m, 3 - 5. xii. 1984, LE Peña G (UMSP 000118512) (NMNH). Etymology — This species is named M. triramosa, Latin for three-branched, and referring to tripartite dorsal phallic spine of this species. — unilineata subgroup The single unusual species placed here was placed in the argentinica subgroup by Sykora, albeit with some hesitation. Sykora did not discuss the characters he used to establish his subgroups. Blahnik and Holzenthal (2012) suggested that both species of the argentinica subgroup should probably be removed from the bilineata group and were possibly unrelated. The only obvious characters linking the two species are the broadly rounded anterior margin of segment IX and the reflexed dorsal lobe of the inferior appendages. It should be noted that Sykora described the reflexed dorsal processes of the inferior appendages as being lateral paired processes (paramere appendages). True paramere appendages seem to be absent in this species. Both of the character similarities listed are probably plesiomorphic characters, and the latter of these would not have been evident to Sykora, based on his interpretation of the structure. Mortoniella unilineata has the anterior margin of segment IX somewhat produced in its ventral ½, as in species of the bilineata group, and the lobes are also further apart dorsally than most species in the leroda group. On the other hand, the female genitalia lack the mesal invagination of segment VIII that characterizes the bilineata group (Fig. 115), implying that if it is related to this group, it must be in a more basal position than other described species. The very elongate ventral process of segment VI and elevated basal projection of the phallicata somewhat resembles the character states in M. proakantha, n. sp. and M. gilli, n. sp., both treated here as “ unplaced species. ” If these species are related, it cannot be a very close relationship, because they are all very apomorphically distinct. Possibly, they represent divergent members of a lineage basal to both the bilineata and leroda groups.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA6F859FF01BA064232F8CF.taxon	description	Fig. 78, 115	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA6F859FF01BA064232F8CF.taxon	description	Adult — Length of forewing: male 4.6 - 6.0 mm; female (undeterminable, from pharate pupa). Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0: 4: 4. Color (in alcohol) medium brown. Tibial spurs slightly darker than legs, contrasting in color. Wing bar not evident (one white band at midlength, in original description). Male genitalia — Ventral process of segment VI very narrow elongate, posteriorly directed, length about 6 times width at base, apex acute as viewed laterally and ventrally. Segment IX with anterolateral margin rounded, somewhat more produced in ventral ½, posterolateral margin slightly produced dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X short, apicomesal margin with weak emargination, apicolateral lobes not evident as such, broadly rounded; ventrolateral lobes prominent and projecting, continuous with apicolateral lobes. Inferior appendages with very elongate, narrow, reflexed dorsolateral projections, acute apically, ventromesal margin narrow, without projection. Mesal pockets of inferior appendage with spinelike apical processes moderately elongate, posteriorly projecting, apparently fused to and projecting below ventral margin of phallicata. Paramere appendages absent. Phallobase with short, lightly sclerotized, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, relatively uniform in width, distinctly upturned in about apical 2 / 5, apical inflection hollowed on ventral surface, apparently to accommodate large membranous lobe from dorsal margin of endophallic membrane; as viewed dorsally, with broadly rounded lateral expansions at apical inflection, narrowing apically, apex rounded. Phallicata with broadly rounded and distinctly sclerotized basodorsal expansion, ventrally with projecting mesal sclerite, arched above and fused to spine-like projections of mesal pockets of inferior appendages. Endophallic membrane elongate, with projecting lateral and dorsomesal membranous lobes, spines absent; phallotremal spines absent. Material examined — VENEZUELA: Merida: Laguna de Murubaji, Paramos zone, site 1, rocky stream below lake, el 3300 m, 10. vii. 1991, GS Vick – 2 males, 1 female (alcohol, pharate adults) (NMNH). Distribution — Venezuela.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FFA7F867FF01BC46412BF8CF.taxon	materials_examined	Type species: Mexitrichia pacuara Flint, 1974. This new subgenus can be distinguished from the subgenus Mortoniella by several characters considered collectively. One of these, the small size of the species, is suggested by the name assigned to the subgenus. The most distinctive character defining the subgenus is the reduction of the hind wing venation to include only fork II. A similar reduction occurs in some species and species subgroups of the leroda group of the subgenus Mortoniella, but generally the costal margin of these species has a distinct bend or inflection (Fig. 101 B). This is much less distinct in species of Nanotrichia, such that the overall wing shape is more spear-like (Fig. 102 B, 103 B, 104 B). Another character synapomorphy that seems to be consistent for the included species is that the middle legs have only 3 spurs (rarely 2). A similar reduction occurs in various species in the leroda and bilineata groups of the subgenus Mortoniella, but the usual spur formula in both of these groups is 0: 4: 4. Additionally, the anterior margin of segment IX of males in the subgenus Nanotrichia is distinctly produced in its ventral ½ and the posterior margin is nearly linear. The lobes of the segment are separated dorsally by ½ or more of the width of the segment (with several exceptions). Species in the bilineata group of the subgenus Mortoniella also have the anterior margin of segment IX produced in the ventral ½ and the lobes widely separated dorsally, but these species are larger and have retained fork V in the hind wing, and usually also fork III. In general, the very shortened phallobase and relatively simple structure of tergum X of species in Nanotrichia also support their separation from the subgenus Mortoniella. Two very distinctive species groups are assigned to Nanotrichia, the ormina and velasquezi groups. These are most convincingly defined by differences in the female genitalia. Of these, the ormina group is the more variable and the placement of M. rodmani Blahnik and Holzenthal and M. simplicis, n. sp. within this group is conjectural, since females are not known for these species. Some species in both the ormina and velasquezi groups have modified scale-like setae paralleling the major veins of the fore- and / or hind wings. A similar development occurs in a few species of the leroda group of the subgenus Mortoniella. The character is mostly hidden by the normal setae present and may not be readily evident. The scales seem to be readily lost in alcohol preserved specimens. In some species of the velasquezi group, there is a more general and readily apparent field of scale-like setae on the hind wing. These are of somewhat different morphology than the scale-like setae paralleling the veins (stalked basally and upright, rather than leaf-like and flattened). Whether the presence of scale-like setae represents an independent character development in the two groups, or an apomorphy for the group, lost in some taxa, is uncertain. Adult — Length of forewing: 1.7 - 4.0 mm; females slightly larger than males. Forewing with forks I, II, and III, forks I and II usually sessile, sometimes with fork II stalked (Fig. 104 A); hind wing with fork II only (Fig. 102 B, 103 B, 104 B), basal forks of Rs and M veins at about midlength and narrowly separated (ormina group), or with basal fork of Rs more proximal (velasquezi group). Crossveins of forewing (r, s, r-m, m, m-cu, cu) linear or nearly so and usually hyaline; hind wing with r-m only (ormina group, Fig. 103 B, 104 B), or with crossveins absent (velasquezi group, Fig. 102 B). Costal margin of hind wing without distinct inflection, wings more or less spear-shaped. Forewing rather broadly rounded apically in velasquezi group (Fig. 102 A), narrow and angulate in ormina group (Fig. 103 A, 104 A), apex of hind wing angulate. Fore- and / or hind wing frequently with adpressed scale-like setae paralleling veins; some species of velasquezi group also with patch of upright scale-like or hair-like setae on hind wing. Spur formula 0: 3: 4 (0: 2: 4, as variant in M. usseglioi). Overall color varying from light brown to dark brown; forewing usually marked with white or whitish setae at anastamosis, in some species of ormina group with small white spots apically. Male genitalia — Ventral process of segment VI relatively elongate, narrow, and posteriorly projecting (length usually 2 times width or more). Segment IX with anterolateral margin rounded and produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, lobes usually separated dorsally by ½ or more of width of segment. Tergum X well developed, setose, usually with deep V-shaped or U-shaped apicomesal invagination (M. aequalis an exception); apicolateral lobes usually simple in structure and not strongly sclerotized. Inferior appendages directly fused to one another and to ventral margin of phallic ensemble, generally short and reduced in ormina group, or in velasquezi group with more distinct dorsolateral process and apex sometimes elongate and fused to mesal spinelike process; appendages always with paired mesal pockets accommodating rod-like projections of phallobase, these very enlarged in velasquezi group, apices of pockets with relatively short to very elongate spine-like projections, typically narrow in ormina group, widened in velasquezi group. Basal segments of parameres variable, usually relatively simple in structure in velasquezi group (short, rounded and fused to one another), often more modified in ormina group, sometimes with elongate rod-like projections; apical segments (paramere appendages) usually present, short to elongate, typically more or less rod-like, often strongly curved. Phallobase short or very short, always with, paired rod-like projections from the apicoventral margin, these short in ormina group (sometimes with apex flared), long and with apices flared in velasquezi group; dorsal margin with moderately elongate mesal spine, relatively simple in structure in some, more often with lateral margins projecting; dorsal phallic spine without basal articulation with phallicata, apex strongly reflexed in most. Phallicata usually short and only indistinctly sclerotized, with distinctly sclerotized, rounded, lateral projection in velasquezi group. Endophallic membrane very variable, usually relatively short and simple in structure, sometimes with dorsal or lateral spines or spine-like structures or with membranous lateral lobes; in velasquezi group with pair of ventral sclerites or spine-like projections and dorsomesal projection, typically with small spines or short spine-like projection; phallotremal spines not usually evident (M. collegarum a notable exception). Etymology — This subgenus is named Nanotrichia from the Greek words nanos, meaning small, and referring to the small size of the species in the subgenus, and trich - for hair, a term generally applied to species in the order Trichoptera because of their hairy wings. The name is feminine in gender.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF99F865FF01BB4641FBFE2F.taxon	description	Fig. 79 Mexitrichia aequalis Flint: 1963: 472.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF99F865FF01BB4641FBFE2F.taxon	description	Adult — Length of forewing: male 2.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color (in alcohol) pale brown. Forewing with membrane pale at anastomosis. Presence of scale-like setae on fore- and hind wings of male not ascertained. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 ½ times width at base. Segment VIII relatively narrow. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins, as viewed dorsally, distinctly invaginated in middle; apex of tergum with very shallow apicomesal invagination, thus without distinct apicolateral lobes; tergum, in lateral view, with shallow rounded apicolateral incision, forming angularly projecting ventrolateral lobe. Inferior appendages small, with short, posteriorly directed lobes. Mesal pockets of inferior appendage large, with elongate and relatively wide, posteriorly-directed, spine-like, apicoventral projections, apices narrowed and slightly decurrent. Paramere with elongate, narrow projection, apparently from enlarged basal lobe of appendage, extending about as far as apical inflection of dorsal phallic spine; paramere appendage probably vestigial, composed of short, lightly sclerotized, projection, mesal to basal lobe. Phallobase with short, rounded, lightly sclerotized, dorsomesal apodeme; ventrally with short ventral rod-like projections, widely flared apically. Dorsal phallic spine, as viewed laterally, nearly uniform in width, strongly upturned in about apical ¼, slightly widened preapically, apex rounded; as viewed dorsally, nearly uniform in width, slightly enlarged preapically, apex subacute, entire. Structure of phallicata not completely evident, apparently tubular, unmodified. Endophallic membrane simple, without spines; phallotremal spines not evident. Material examined — PERU: Puerto Bermudez, Rio Pichis, 17. vii. 1920, male Holotype, type 3892 (CUIC). Distribution — Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9AF865FF01B9864218FA4F.taxon	description	Fig. 80 Mexitrichia aries Flint: 1963: 470.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9AF865FF01B9864218FA4F.taxon	description	Adult — Length of forewing: male ca. 2.0 mm. Forewing venation not evident (pharate adult). Spur formula 0: 3: 4. Overall color (in alcohol) yellowish-brown. Scale-like setae on wings of male not evident, probably absent. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 ½ times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X moderate in length, lateral margins subparallel; apex of tergum with deep Vshaped emargination, extending less than ½ length of tergum; apicolateral lobes not evident as such, formed by mesal invagination, apicolateral margin subtruncate; tergum, in lateral view, with weakly projecting ventrolateral lobes, more or less continuous with apicolateral margin. Inferior appendages hardly evident as such, strongly fused to phallic ensemble. Mesal pockets of inferior appendage with short, posterodorsally curved, spine-like, apicoventral projections. Paramere appendage short, narrow, nearly uniform in width, apex acute, extending about same length as apical inflection of dorsal phallic spine; Phallobase with short ventral rod-like projections, not flared apically. Dorsal phallic spine, as viewed laterally, short, strongly upturned in about apical 1 / 3, apex acute; as viewed dorsally, with distinct lateral wing-like appendages at about midlength. Phallicata very short and weakly sclerotized. Endophallic membrane with pair of prominent lateral, helically coiled, sclerites; phallotremal spines apparently absent. Material examined — ECUADOR: Napo-Pastaza: Río Chingual, 8 mi E of El Pun, 8000 ft., 4. v. 1958, RW Hodges – male Holotype (pharate adult, alcohol) (USNM type # 66021) (NMNH). Distribution — Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9AF864FF01BE2642E8FDCF.taxon	description	Fig. 81, 103, 116	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9AF864FF01BE2642E8FDCF.taxon	materials_examined	Material examined — BOLIVIA: Santa Cruz: PN and ANMI Amboró, Guardia Parque Mataracú, Río Yapacaní, 17.52072 ° S, 63.86795 ° W, el 329 m, 26. xi. 2004, Robertson, Garcia and Vidaurre – 9 males, 16 females (pinned), 1890 males, 408 females (alcohol) (UMSP); Provincia Florida, Bermejo, viejo caretera S. C. - CBBA, Río Bermejo @ Quebrada Chorro Viejo, 18.16642 ° S, 63.58023 ° W, el 749 m, 15. xi. 2004, Robertson and Vidaurre – 13 males, 32 females (pinned) (UMSP). Distribution — Argentina, Bolivia, Chile.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9BF863FF01BAA64293FD6F.taxon	description	Fig. 82, 118	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9BF863FF01BAA64293FD6F.taxon	description	Adult — Length of forewing: male 1.7 - 2.0 mm; female 1.9 - 2.3 mm. Forewing with forks I, II, and III present, fork II with very long stalk, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish-brown. Tibial spurs short, apparently same color as legs. Forewing with distinct wing bar at anastomosis. Males with scale-like setae paralleling veins of both fore- and hind wings. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 ½ times width at base. Segment VIII relatively narrow. Segment IX with anterolateral margin rounded and distinctly produced in ventral half, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X elongate, lateral margins subparallel, slightly invaginated in middle; apex of tergum with deep V-shaped emargination, extending less than ½ length of tergum; apicolateral lobes simple, subacute apically, formed by mesal invagination; tergum, in lateral view, with shallow, rounded, apicolateral incision, forming angularly projecting ventrolateral lobe. Inferior appendages with short upright dorsolateral lobes. Mesal pockets of inferior appendage small, with very elongate, narrow, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, extending about same length as apical inflection of dorsal phallic spine; basal segment of parameres, lateral to paramere appendage, forming rounded lateral sclerite with elongate spine-like projection from dorsal margin, slightly longer than paramere appendage. Phallobase with short rounded, lightly sclerotized, dorsomesal apodeme; ventrally with short ventral rod-like projections, not flared apically. Dorsal phallic spine, as viewed laterally, nearly uniform in width, strongly upturned in about apical 1 / 3, slightly widened preapically, apex acute; as viewed dorsally, slightly widened in basal ½, apex weakly divided mesally, forming 2 acute apical projections. Phallicata distinctly tube-like, without basal projection, apicolateral margins rounded. Endophallic membrane apparently short, without spines (but with indistinct apical sclerite); phallotremal spines absent. Material examined — ECUADOR: Napo: Pano, 580 m, 12. ix. 1990, OS Flint, Jr – 1 male (alcohol) (NMNH); 5.2 km SW Pano, 640 m, 13. ix. 1990, OS Flint, Jr – 1 male (alcohol) (NMNH); Santa Cecilia, 16. v. 1975, PJ Spangler – 2 males (alcohol) (NMNH); Lago Agrio (5 km N), 26. ix. 1975, A Langley – 1 male (pinned), 4 males, 2 females (alcohol) (NMNH); Pastaza: Puyo, 13. v. 1977, PJ Spangler and DR Givens – 3 males, 11 females (alcohol) (NMNH); same locality and collectors, 17. v. 1977 – 2 males, 1 female (alcohol) (NMNH); same locality and collectors, 21. v. 1977 – 13 males, 41 females (alcohol) (NMNH); Puyo (27 km N) Est. Fluvia Metrica, 4. ii. 1976, Spangler, et al. – 6 males, 15 females (alcohol) (NMNH); Zamora-Chinchipe: Río Chicaña, 9 km N Yanzatza, 880 m, 20. ix. 1990, OS Flint, Jr – 1 male, 1 female (alcohol) (UMSP). Distribution — Colombia, Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9CF863FF01BB464326FAEF.taxon	description	Fig. 83	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9CF863FF01BB464326FAEF.taxon	materials_examined	Additional material examined — ECUADOR: El Oro: 9 mi S Santa Rosa, 3.45000 ° S, 79.96667 ° W, 1.23.1955, EI Schlinger and ES Ross – 3 males, 1 female (alcohol) (CAS); Loja: Río Puyango, 300 m, 17 - 18. viii. 1977, LE Peña G – 3 males (alcohol) (NMNH). Distribution — Costa Rica, Colombia, Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9CF862FF01BDC64468FC8F.taxon	description	Fig. 84 This species is very similar to Mortoniella aries (Flint), especially in the shape of the dorsal phallic spine, which is strongly reflexed apically and has subquadrate lateral wings. It can be distinguished by lacking the spiral processes present on the endophallic membrane in M. aries; instead, it has a few scattered spines. Adult — Length of forewing: male 2.6 mm. Forewing with forks I, II, and III present, fork II with short stalk, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Tibial spurs short. Overall color (in alcohol) yellowish-brown (specimen faded and partially rubbed, wing bar not evident). Fore- and hind wings of male with scale-like setae paralleling major veins (except apically). Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 2 ½ times width at base. Segment IX with anterolateral margin rounded and produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X moderate in length, lateral margins subparallel; apex of tergum with deep U-shaped emargination, extending more than ½ length of tergum; apicolateral lobes not evident as such, formed by mesal invagination, apicolateral margin subtruncate; tergum, in lateral view, with ventrolateral lobes more or less continuous with apicolateral margin, not distinctly defined. Inferior appendages with short setose dorsolateral projections. Mesal pockets of inferior appendage with moderately elongate, posterodorsally curved, spine-like, apicoventral projections. Paramere appendage moderately elongate, narrow, nearly uniform in width, apex acute, extending slightly beyond apical inflection of dorsal phallic spine; basal segment of paramere rounded, with short spine-like projection from posterior margin. Phallobase with short ventral rod-like projections, not flared apically. Dorsal phallic spine, as viewed laterally, short, strongly upturned in about apical 1 / 3, apex subacute in lateral view, rounded in dorsal view; as viewed dorsally, with distinct lateral, wing-like, appendages in basal ½. Phallicata scarcely evident, very short and weakly sclerotized. Endophallic membrane with a couple of clusters of needle-like spines on either side (mostly in pairs); phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9CF862FF01BDC64468FC8F.taxon	materials_examined	Holotype male (alcohol) — PERU: Huánuco: 16. ix. 1954, EL Schlinger and ES Ross (UMSP 000130031) (CAS). Etymology — The species is named M. paucispina, Latin for few-spined, in reference to the scattered (and generally paired) spines on the endophallic membrane of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9DF861FF01BBE6412BFECF.taxon	description	Fig. 85 This species is probably most closely related to M. triangularis, n. sp. Both species have a deep mesal incision at the apex of the dorsal phallic spine. Mortoniella quadridactyla can be easily distinguished by having the dorsal phallic spine much narrower in lateral view, with the apex upturned, and with each half of the divided apex subdivided into a pair of acute spines. Tergum X is also simpler in construction, with the apicolateral lobes very broadly rounded. Adult — Length of forewing: male 1.7 - 1.9 mm; female 2.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish-brown (specimen faded and partially rubbed, wing bar not evident). Forewings of male with scale-like setae paralleling major veins (except apically). Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apicomesally with deep V-shaped emargination, extending more than ½ length of tergum; apicolateral lobes simple, rounded apically, formed by mesal invagination; tergum, in lateral view, with rounded apicolateral invagination, demarcating prominent and apically rounded ventrolateral lobe. Inferior appendages with small setose lateral projections, apparently fused ventrally to spine-like projections of mesal pockets. Mesal pockets of inferior appendage small, with very elongate, narrow, posteriorly-directed, spinelike, apicoventral projections. Paramere appendage elongate, strap-like, apex acute, extending about same length as dorsal phallic spine. Phallobase with very short ventral rod-like projections, each distinctly flared apically. Dorsal phallic spine, as viewed laterally, relatively narrow, strongly and sinuously upturned in about middle, apex distinctly forked into pair of acute projections; as viewed dorsally, distinctly widened in apical ½, apicomesally with deep U-shaped invagination, extending about ½ length of spine, each half of divided apex forked preapically to produce 4 apical spine-like projections. Phallicata relatively lightly sclerotized and difficult to discern. Endophallic membrane elongate, with large basodorsal, balloon-like lobe; phallotremal spines very small.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9DF861FF01BBE6412BFECF.taxon	materials_examined	Holotype male (alcohol) — VENEZUELA: Barinas: Barinas, Río Santo Domingo, 17. ii. 1976, CM and OS Flint, Jr (UMSP 000097089) (NMNH). Paratypes — VENEZUELA: Barinas: same data as holotype – 2 males, 2 females (alcohol) (NMNH). Etymology — This species is named Mortoniella quadridactyla, derived from Latin for four fingers or appendages, and referring to the four acute projections at the apex of the dorsal phallic spine of the species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9EF861FF01B9A643F7F8CF.taxon	description	Fig. 86 This is a very distinctive species with several character attributes similar to M. rodmani Blahnik and Holzenthal. The two species are probably related. Both are provisionally placed in the ormina group, because this is the group with which they share the greatest superficial similarity. Female genitalia, which are distinctive for the ormina group, would more conclusively demonstrate this, but are unknown for either species. Possibly, the two form the nucleus of an additional species group. Character similarities to M. rodmani include the possession of ventromesal lobes on tergum X, which probably serve as guides for the dorsal phallic spine; the general shape of the paramere appendages, which are curved ventrally from their base and dorsally curved apically; the basoventral projections from phallicata, which seem to form a ventral support for the paramere appendages (the projections in M. rodmani with an apical spine); and the prominent 3 - lobed apicoventral sclerite on the endotheca. Neither of the species seems to have scale-like setae paralleling the veins of the wings. Mortoniella simplicis is easily distinguished by the shape of the dorsal phallic spine, which is sinuously curved in the middle and rounded apically, and rests in a depression formed by the dorsal margin of the phallicata (or phallicata / endotheca), and by the shape of tergum X and its ventromesal lobes. Adult — Length of forewing: male 2.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color medium brown. Tibial spurs relatively short, darker than legs, contrasting in color. Forewing with distinct white wing bar at anastomosis. Males without scale-like setae on wings. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 2 times width at base, apex subacute. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X relatively short, lateral margins subparallel; apex of tergum with deep U-shaped emargination, extending about ½ length of tergum; apicolateral lobes formed by mesal invagination, apices of lobes rounded laterally, acute on margin bordering mesal invagination; tergum, in lateral view, with projecting, broadly rounded, non-setose, ventromesal lobes, apparently straddling dorsal phallic spine. Inferior appendages very small, setose, rounded dorsally. Mesal pockets of inferior appendage with moderately elongate, posterodorsally curved, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute; appendage uniformly curved, ventrally from base and dorsally projecting apically. Phallobase with short ventral rod-like projections, not flared apically. Dorsal phallic spine, as viewed laterally, nearly uniform in width, rounded apically, with pronounced sinuous deflection in about middle. Phallicata with raised dorsal projection, articulating with sinuous deflection of dorsal phallic spine, basoventrally with depressed, rounded, lateral projections (apparently as resting platform for ventral deflection of paramere appendage). Endophallic membrane with small internal sclerite (possibly phallotremal sclerite), and prominent elongate, narrow ventral sclerite, branched apically into 3 acute lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9EF861FF01B9A643F7F8CF.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Miranda: Parque Nacional Guatopo, Quebrada Macanilla at La Macanilla, 10.113 ° N, 66.516 ° W, el 550 m, 23. i. 1994, Holzenthal, Cressa, Rincón (UMSP 000041336) (UMSP). Etymology — This species is named M. simplicis, derived from the Latin word for simple, in reference to the relatively simple and unspecialized genitalia of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9FF860FF01B8A64435FACF.taxon	description	Fig. 87 This species is easily diagnosed by the shape of the dorsal phallic spine, which is trianguloid in lateral view, widening from the base to the apex, with apical spines on the dorsal and ventral margins. Like M. quadridactyla, n. sp., which is probably the most closely related species, the spine is deeply cleft mesally. In the case of M. triangularis, the cleft extends almost to the base of the spine. Adult — Length of forewing: male 2.0 mm; female 2.2 - 2.3 mm. Forewing with forks I, II, and III present, fork II with long stalk, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color medium brown, legs slightly paler. Tibial spurs short, slightly darker than legs, weakly contrasting in color. Forewing with white wing bar at anastomosis (evident in some specimens). Forewings of male with scale-like setae paralleling major veins (except apically). Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base, apex subacute. Segment IX with anterolateral margin rounded and slightly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by less than ½ width of segment. Tergum X moderately elongate, lateral margins slightly converging from base; apex of tergum with U-shaped emargination, extending less than ½ length of tergum; apicolateral lobes simple, with small angulate projection; tergum, in lateral view, with subtruncate apex and broadly rounded basal ventrolateral lobe. Inferior appendages with small setose dorsolateral projections, apices acute and posteriorly directed. Mesal pockets of inferior appendage with elongate, narrow, posteriorly-directed, spine-like, apicoventral projections, apparently fused to ventral margin of phallicata basally. Paramere appendage elongate, narrow, apex acute, strongly and nearly evenly curved, apex dorsally directed, extending about same length as dorsal phallic spine. Phallobase with short ventral rod-like projections. Dorsal phallic spine, as viewed laterally, subtriangular in shape, narrow basally, very wide apically, with apical spine-like projections from dorsal and ventral margins, and additional spine on dorsal margin at about midlength; as viewed dorsally, with deep mesal incision, extending almost entire length of spine, spine-like projections from ventral margin projecting laterally, dorsal spines posteriorly projecting. Phallicata relatively lightly sclerotized, simple in structur-e. Endophallic membrane somewhat inflated, with large, lightly sclerotized, dorsal lobes; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9FF860FF01B8A64435FACF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Zamora-Chinchipe: Zamora, 31. v. 1976, A Langley, et al. (NMNH) (UMSP 000095073). Paratypes — ECUADOR: Zamora-Chinchipe: Zamora, 4. xii. 1978, J Anderson – 2 males, 4 females (alcohol) (NMNH). Etymology — This species is named M. triangularis for the shape of the dorsal phallic spine of the male, which is distinctly trianguloid in appearance, as viewed laterally.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9FF86FFF01BDA64244FACF.taxon	description	Fig. 88, 104, 117	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF9FF86FFF01BDA64244FACF.taxon	description	Adult — Length of forewing: male 1.8 - 2.0 mm; female 1.8 - 2.2 mm. Forewing with forks I, II, and III present, fork II with very long stalk, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 2 / 3: 4. Overall color medium brown, legs slightly paler. Tibial spurs short, slightly darker than legs, weakly contrasting in color. Forewing with interrupted white wing bar at anastomosis, apices of wings with indistinct white spots. Males with scale-like setae paralleling veins of both fore- and hind wings. Male genitalia — Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 2 ½ times width at base. Segment VIII relatively narrow. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by slightly less than ½ width of segment. Tergum X elongate, lateral margins subparallel; apex of tergum with deep V-shaped mesal emargination, extending less than ½ length of tergum; apicolateral lobes simple, subtruncate apically, as viewed dorsally, formed by mesal invagination; tergum, in lateral view, with very shallow apicolateral incision, forming angularly projecting ventrolateral lobe. Inferior appendages scarcely evident, strongly fused to venter of phallic ensemble. Mesal pockets of inferior appendage small, with elongate, narrow, posteriorly-directed, spine-like, apicoventral projections. Paramere with 3 elongate, narrow, subequal appendages, 1 emerging from dorsal of margin of rounded basal segment, 1 evidently emerging mesal to basal segment (possibly from mesal surface), 1 (probably true paramere appendage) emerging lateral to basal segment and crossing over corresponding appendage from opposite side, ventral to phallic ensemble. Phallobase with short rounded, lightly sclerotized, dorsomesal apodeme; ventrally with short ventral rod-like projections, not flared apically. Dorsal phallic spine, as viewed laterally, nearly uniform in width, strongly upturned in about apical 1 / 3, slightly widened preapically, apex acute; as viewed dorsally, nearly uniform in width, apex weakly divided mesally, forming 2 acute apical projections. Phallicata distinctly tube-like, with pair of short spine-like basolateral projections, apicolateral margins flared. Endophallic membrane somewhat ballooned, with indistinct apical sclerite; phallotremal spines very small, but distinct. Material examined — BOLIVIA: La Paz: ANMI Madidi, Chalalan Ecolodge, Río Tuichi and tributary at entrance to lodge, 14.41695 ° S, 67.90630 ° N, 300 m, 27. vii. 2003, Robertson and Blahnik – 392 males, 972 females (alcohol) (UMSP); PERU: Avispas, - x. 1962, LE Peña G – 38 males, 15 females (alcohol) (NMNH); Junin: Mission Cutivireni at Río Namiri, 6 - 25. iii. 1985, HM Savage – 4 males, 1 female (alcohol) (NMNH); Madre de Dios: Tambopata Wildlife Res., 30 km SW Pto. Maldonado, 12.83333 ° S, 69.28333 ° W, 290 m, 1 - 14. i. 1983, JJ Anderson – 1 male (pinned) (NMNH); Manu, Erika (nr. Salvacion), 550 m, 4 - 6. ix. 1988, O Flint and N Adams – 1 male, 2 females (pinned), 35 males, 136 females (alcohol) (NMNH); Manu, Río Manu, Limonal (10 km N Boca Manu), 200 m, 7. ix. 1988, O Flint and N Adams – 8 females (alcohol) (NMNH); Manu, Pakitza, 12.11667 ° S, 70.96667 ° W, 250 m, 9. ix. 1988, O Flint and N Adams – 1 male (alcohol) (NMNH); Hostel Erica (nr. Salvacion), 12 ° 53 ’ S, 71 ° 14 ’ W, 550 m, 3 - 5. ix. 1989, RA Faitoute et al. – 2 males, 5 females (alcohol) (NMNH); same location and date, N Adams et al. – 1 male (alcohol) (NMNH); between Boca Manu and Romera, along Manu River, 250 m, 16. ix. 1989, RA Faitoute – 1 female (alcohol) (NMNH). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF90F86EFF01BDA64382FAAF.taxon	description	Fig. 89 The genital morphology of this species is rather difficult to interpret and we are not exactly sure whether the prominent lateral spines with enlarged bases should be considered paramere appendages, with the basal segment of the paramere modified into short spines, or if the short basal spines are paramere appendages and the lateral spine-like processes are endophallic sclerites, homologous to those in M. aries (Flint). The description follows the latter interpretation and, based on this interpretation, M. zamora is probably most similar to M. aries, differing in the structure of the lateral sclerites of the endothecal membrane and also in having the dorsal phallic spine more strongly reflexed, with its lateral wing-like processes posteriorly projected. Mortoniella zamora also has some (probably superficial) similarities to species in the velasquezi group, including rod-like appendages from the ventral margin of the phallobase that are inflated apically (although very short); relatively large mesal pockets on the inferior appendages; very elongate, curved, spine-like projections from the mesal pockets; and inferior appendages with short, upright dorsal lobes, fused apically to the lateral margin of the phallicata. The most diagnostic features of M. zamora are probably found in the form of the basal sclerites of the endophallic membrane and in the very elongate, curved, spine-like projections from the mesal pockets of the inferior appendages. The strongly recurved dorsal phallic spine and posteriorly projecting lateral wing-like projections from the same structure are also usefully diagnostic. Adult — Length of forewing: male 2.3 - 2.5 mm; female 2.6 mm. Forewing with forks I, II, and III present, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0: 3: 4. Overall color (in alcohol) yellowish-brown (specimen badly faded). Wing bar not evident. Males with at least some scale-like setae paralleling veins of forewing. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X moderate in length, lateral margins subparallel; apex of tergum with deep U-shaped emargination, extending nearly ½ length of tergum; apicolateral lobes not evident as such, formed by mesal invagination, apicolateral margin subtruncate; tergum, in lateral view, with broadly rounded ventrolateral lobes, more or less continuous with apicolateral margin. Inferior appendage with short setose, thumb-like, dorsal projection, evidently fused to phallicata posteriorly. Mesal pockets of inferior appendage large, with very elongate and strongly posterodorsally curved, spine-like, apicoventral projections. Paramere appendage very short, acute apically. Phallobase with short ventral rod-like projections, strongly flared apically. Dorsal phallic spine, as viewed laterally, short, strongly reflexed in about apical 1 / 3, apex acute; as viewed dorsally, with distinct lateral, posteriorly curved, wing-like appendages at about midlength. Phallicata short and membranous or weakly sclerotized, hardly evident as such, continuous with endophallic membrane. Endophallic membrane inflated, with pair of prominent lateral sclerites, enlarged basally and with apical spine-like projections from ventral margin; phallotremal spines absent.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF90F86EFF01BDA64382FAAF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Zamora-Chinchipe: Zamora, 4. xii. 1978, JJ Anderson (UMSP 000097035) (NMNH). Paratypes — COLOMBIA: Cauca: Municipio de Inzá, Quebrada San Andrés, ca 500 m W Restaurante “ La Portada, ” San Andrés de Pisimbalá, 2.58222 ° N, 76.04333 ° W, el 1750 m, 21. xii. 1997, F Muñoz-Q, et al. – 1 male (pinned) (UMSP); ECUADOR: Zamora-Chinchipe: same data as holotype – 2 males, 1 female (alcohol) (NMNH). Etymology — This species is names M. zamora, used as a noun in apposition, for the name of the collection locality where the type specimen was collected.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF92F86CFF01BA4644DCFE6F.taxon	description	Fig. 90, 120 Mortoniella cognata is very similar to M. velasquezi Flint. Both species have relatively short, bluntly rounded lobes on what appear to be inferior appendages, but probably represents elongate lateral lobes of the phallicata. The most readily diagnosed difference is that the hind wings of males of M. cognata do not have a distinct field of upright scale-like setae, although it does have a field of upright, narrow setae. There are several other differences. The dorsal phallic spine is narrower in lateral view (i. e. it lacks a distinct protruding “ belly ”), the apex is slightly less reflexed, and the angular projection on the posteroventral margin is less defined. Additionally, the paramere appendages are shorter and the dorsomesal projection on the endophallic membrane tends to be membranous or only weakly sclerotized, rather than distinctly sclerotized and spine-like. Adult — Length of forewing: male 2.5 - 3.0 mm; female 2.8 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsi slightly paler. Tibial spurs darker than legs, contrasting in color. Forewing with narrow white wing bar at anastomosis. Males with forewings densely covered with short, adpressed setae; hind wings with scale-like setae paralleling major veins and with field of elongate, upright setae, but without darkened, semi-erect scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow and slightly constricted basally, length about 3 ½ times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X, as viewed laterally, short and narrow, rod-like, basomesally with pair of setose sclerites; as viewed dorsally, short and very wide, apicomesally with deep and very wide Ushaped emargination, extending more than ½ length of tergum; ventrolateral lobes obsolete. Inferior appendage with short digitate, setose dorsolateral projections, fused apicoventrally to spine-like projections of mesal pockets. Mesal pockets of inferior appendage very large, with moderately elongate, thick, posterodorsally curved, spine-like, apicoventral projections. Paramere appendage very short, apex acute, emerging near dorsal margin of basal segment. Phallobase very short, with elongate ventral rod-like projections, strongly flared apically. Dorsal phallic spine, as viewed laterally, short and relatively narrow, dorsolateral margins expanded, spine weakly reflexed in about apical 2 / 5, basal part with weakly projecting “ belly, ” ventral margin with rounded indentation at apical inflection, reflexed apex with distinct, but rather weakly produced, angular projection on posterior margin, apex acute; as viewed dorsally, very wide, lateral margins subparallel, apex bluntly rounded. Phallicata short, continuous with endophallic membrane apically, laterally with relatively elongate, apically rounded, sclerotized projections, subequal in length to spine-like projections of mesal pockets of inferior appendage; projections appearing ventrally as broad, apically rounded lobes of inferior appendages. Endophallic membrane continuous with phallicata, without membranous lateral lobes, dorsomesally with membranous or blunt, weakly sclerotized projection, apparently articulating with indentation in dorsal phallic spine; ventrally with pair of short blunt sclerites (modified phallotremal spines?).	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF92F86CFF01BA4644DCFE6F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Barinas: Río Sinigüis in Caño Grande, 8.40000 ° N, 70.77417 ° W, el 520 m, 22. iii. 1997, Holzenthal (UMSP 000001557) (UMSP). Paratypes — ECUADOR: Napo: Río Jondachi, 30 km N Tena, 950 m, 10. ix. 1990, OS Flint, Jr – 2 males, 5 females (pinned), 2 males, 8 females (alcohol) (NMNH); Pastaza: Puyo, 30. i. 1976, Spangler, et al. – 1 male (alcohol) (NMNH); Puyo, 5. v. 1977, PJ Spangler and DR Givens – 2 males, 5 females (pinned); 1 male, 17 females (alcohol) (NMNH); VENEZUELA: Barinas: same data as holotype – 5 males (pinned), 47 males (alcohol) (UMSP), 15 males (alcohol) (MIZA); Portuguesa: Río Las Marias at Finca Los Cerajones, ca. 5 km NE Potrero, 9.20550 ° N, 69.70750 ° W, el 270 m, 25. iii. 1997, Holzenthal and Flecker – 1 male (alcohol) (UMSP). Etymology — This species is named M. cognata from the Latin word cognatus, meaning kindred or related, and referring to the close similarity between this species and M. velasquezi.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF93F86BFF01BA464113FD6F.taxon	description	Fig. 91, 125	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF93F86BFF01BA464113FD6F.taxon	description	Adult — Length of forewing: male 2.3 - 2.6 mm; female 2.6 - 2.9 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) medium brown, legs slightly paler. Tibial spurs darker than legs, contrasting in color. Forewing with distinct white wing bar at anastomosis, Males without scale-like setae on wings. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X, as viewed laterally, relatively short and narrow, rod-like, apex rounded; as viewed dorsally, short and wide, apicomesally with deep emargination, extending about ½ length of tergum; ventrolateral lobes obsolete. Inferior appendage with prominent digitate, setose dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets. Mesal pockets of inferior appendage very large, with elongate thick, spine-like, apicoventral projections. Paramere appendage elongate, narrow, extending nearly straight, nearly uniform in width, apex acute. Phallobase very short, with elongate ventral rodlike projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, short and wide, strongly reflexed in about apical 2 / 5; ventral margin with dimple-like indentation at point of inflection; reflexed apex strongly inflated, rounded on posterior margin, subacute apically; as viewed dorsally, with distinct rounded dorsolateral projections in basal ½, gradually narrowing apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with broadly rounded sclerotized projections, surrounding mesal pockets of inferior appendage, Endophallic membrane short and wide, continuous with phallicata, dorsomesally with strongly sclerotized, short, preapical, spine-like projection, articulating with dimple-like indentation in dorsal phallic spine; ventrally with pair of strongly sclerotized, curved, spine-like projections (modified phallotremal spines?).	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF93F86BFF01BA464113FD6F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pastaza: Puyo (3 km W), 15. vii. 1976, J Cohen (UMSP 000097141) (NMNH). Paratypes — ECUADOR: Napo: Puyo, 6. v. 1977, PJ Spangler and DR Givens – 1 male (alcohol) (NMNH); Archidona, Río Misahuallí, 650 m, 11. ix. 1990, PJ Spangler – 1 male, 2 females (alcohol) (NMNH); Pastaza: Puyo, riverside, 29. v. 1975, Cohen and Langley – 4 males (alcohol) (NMNH); Puyo, 5. v. 1977, PJ Spangler and DR Givens – 10 males 14 females (alcohol) (NMNH); same locality and collectors, 6. v. 1977 – 3 males, 6 females (alcohol) (NMNH); same locality and collectors, 7. v. 1977 – 3 males, 4 females (alcohol) (NMNH); same locality and collectors, 8. v. 1977 – 2 males, 5 females (alcohol) (NMNH); same locality and collectors, 10. v. 1977 – 4 male, 1 female (alcohol) (NMNH); same locality and collectors, 11. v. 1977 – 1 male (alcohol) (NMNH); same locality and collectors, 13. v. 1977 – 10 males, 15 females (alcohol) (NMNH); same locality and collectors, 14. v. 1977 – 14 males, 31 females (alcohol) (NMNH); same locality and collectors, 21. v. 1977 – 4 males (alcohol) (NMNH); Puyo, 30. i. 1976, Spangler, et al. – 170 males, 185 females (alcohol) (NMNH); same locality and collectors, 1. ii. 1976 – 1 male, 1 female (alcohol) (NMNH); same locality and collectors, 8 - 11. ii. 1976 – 1 male, 3 females (alcohol) (NMNH); Puyo (1.5 km S), 14. v. 1977, PJ Spangler and DR Givens – 14 males, 31 females (alcohol) (NMNH); same data as Holotype – 51 males, 52 females (alcohol) (NMNH) 5 males, 5 females (alcohol) (UMSP); PERU: Monson Valley, Tingo Maria, 18.1 x. 1954, EI Schlinger and ES Ross – 1 male (alcohol) (CAS); Cuzco: Pilcopata, premontane moist forest, 600 m, 11 - 14. xii. 1979, JB Heppner – 1 male (alcohol) (NMNH); Madre de Dios: Manu, Erika (near Salvación), 550 m, 4 - 6. ix. 1988, O Flint and N Adams – 2 males, 1 female (alcohol) (NMNH); Manu, Erika Hostel (near Salvación), 12.0500 ° S, 71.2333 ° W, 550 m, 3 - 5. ix. 1989, N Adams et al. – 1 male, 1 female (alcohol) (NMNH); same locality and date, RA Faitoute et al. – 2 males (alcohol) (NMNH). Etymology — This species is named M. coheni for Jeffrey Cohen, in recognition of the contribution he made in furthering the knowledge of the aquatic insects of Ecuador through his Peace Corps collecting efforts.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF94F86AFF01BB464244FC4F.taxon	description	Fig. 92, 102, 121	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF94F86AFF01BB464244FC4F.taxon	description	Adult — Length of forewing: male 2.6 - 3.0 mm; female 3.0 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color dark brown, apices of tarsi paler. Tibial spurs slightly darker than legs, not or only weakly contrasting in color. Forewing with distinct white wing bar at anastomosis. Males with scale-like setae paralleling veins in forewing, hind wing with extensive field of slightly darkened, semi-erect, scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, lobes separated dorsomesally by more than ½ width of segment. Tergum X, as viewed laterally, relatively short, narrowed and slightly ventrally curved apically; as viewed dorsally, short and wide, apicomesally with deep U-shaped emargination, extending more than ½ length of tergum; ventrolateral lobes slightly produced basally, obsolete apically. Inferior appendage with short digitate, setose, dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets; composite apical structure forming prominent elongate, curved, ventral lobes. Mesal pockets of inferior appendage very large, with elongate, thick, spine-like apicoventral projections. Paramere appendage moderately elongate, narrow, extending nearly straight, slightly swollen preapically, with short spine-like apical projection. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, relatively narrow, strongly reflexed in about apical 1 / 3, apex rounded; as viewed dorsally, with distinct subparallel dorsolateral projections, narrowed basally and apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with broadly rounded sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with slightly raised projection with numerous minute spines, laterally with projecting membranous lobes with minute spines, apically with narrow projecting lobe, ventrally with pair of sclerotized projections, each with v-shaped apical notch (modified phallotremal spines?). Material examined — BOLIVIA: Cochabamba: PN and ANMI Carrasco, Paracticito, Río San Rafaél, Puente Panchito, 800 m from Guarda Parque, 17.06077 ° S, 65.48272 ° W, 438 m, 9 - 10. xi. 2004, Robertson, Garcia, Valdiva – 2 males, 17 females (pinned), 1 male (alcohol) (UMSP); Río San Maleo, Carrasco N. P. at cable crossing to park, 17.06392 ° S, 65.47558 ° W, 449 m, 11. xi. 2004, Robertson, Garcia, Vidaurre – 27 males, 43 females (pinned), 15 males, 957 females (alcohol) (UMSP); Santa Cruz: PN and ANMI Amboró, Guardia Parque Mataracú, Río Yapacani, 17.52072 ° S, 63.86795 ° W, 329 m, 26. xi. 2004, Robertson, Garcia, Vidaurre – 2 males, 5 females (pinned), 352 males, 125 females (alcohol) (UMSP); PERU: Madre de Dios: Manu, Erika (nr. Salvacion), 550 m, 4 - 6. ix. 1988, O Flint and N Adams – 2 males, 9 females (pinned), 3 males, 27 females (alcohol) (NMNH); Manu, Hostel Erika (nr. Salvacion), 12.88333 ° S, 71.23333 ° W, 550 m, 3 - 5. ix. 1989, N Adams et al. – 1 male, 1 female (alcohol) (NMNH); same locality and date, RA Faitoute et al. – 2 males, 4 females (alcohol) (NMNH); Amazonia Lodge, Río Alto Madre de Dios, 12.87033 ° S, 71.37600 ° W, 500 m, 30. vi. 1993, R Blahnik and M Pescador – 2 males, 13 females (alcohol) (NMNH). Distribution — Bolivia, Peru.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF95F869FF01BC2645ABFC4F.taxon	description	Fig. 93, 122 Mortoniella licina is very closely related to M. eduardoi (Rueda Martín and Gibon) and M. venezuelensis, n. sp. All of these species have the apical spine-like projections from the mesal pockets of the inferior appendages very elongate and fused to the ventral part of the inferior appendages to form elongate, projecting lobes. All of the species also have an endophallic membrane that is ornamented with a dorsomesal projection with minute spines, membranous lateral lobes with minute spines, and paired ventral sclerites, each with a somewhat forked apex. Mortoniella licina differs from the other two species in lacking a field of upright scale-like setae on the hind wings of males. It most resembles M. venezuelensis in that the paramere appendages are uniform in width and do not have modified apices. Those in M. licina are usually distinctly curved apically, rather than being straight. A subtle, but distinctive character is that the ventral lobes of the inferior appendages and attached spine-like projections of the mesal pockets, are longer and more distinctly upturned in M. licina. Adult — Length of forewing: male 2.7 - 3.1 mm; female 2.8 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) medium brown. Forewing with narrow whitish wing bar at anastomosis. Males with scale-like setae paralleling veins in fore- and hind wings, hind wing without additional field of darkened, semi-erect, scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 ½ times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X, as viewed laterally, relatively short, narrowed and slightly ventrally curved apically, basomesally with pair of setose sclerites; as viewed dorsally, short and wide, apicomesally with deep U-shaped emargination, extending more than ½ length of tergum; ventrolateral lobes slightly produced basally, obsolete apically. Inferior appendage with setose, digitate dorsolateral projections, fused apicoventrally to spine-like projections of mesal pockets; composite apical structure forming prominent and very elongate, curved, ventral lobes. Mesal pockets of inferior appendage very large, with elongate, thick, strongly posterodorsally curved, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, apex acute, usually distinctly curved in apical part. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, somewhat widened basally, strongly reflexed in about apical 1 / 3, apex rounded; as viewed dorsally, with distinct dorsolateral projections, widest in basal ½, tapering apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with rounded sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with slightly raised projection with numerous minute spines, laterally with projecting membranous lobes with minute spines, apically with narrow projecting lobe, ventrally with pair of sclerotized projections, each with v-shaped apical notch (modified phallotremal spines?).	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF95F869FF01BC2645ABFC4F.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Zamora-Chinchipe: Yanzaza (5 km N), 29. xi. 1978, JJ Anderson (UMSP 000096933) (NMNH). Paratypes — ECUADOR: Napo: Puerto Nuevo, 8. vii. 1976, J Cohen – 1 male, 1 female (alcohol) (NMNH); Pastaza: Puyo, 21. v. 1977, PJ Spangler and DR Givens – 1 male (alcohol) (NMNH); Puyo, 30. i. 1976, Spangler et al. – 9 males (alcohol) (NMNH); Tewaeno, 500 m, 18. v. 1976, J Cohen – 1 male (alcohol) (NMNH); Tzapino, 32 km NE Tigueno, 1.31667 ° S, 77.46667 ° W, 400 m, 22. v. 1976, J Cohen – 1 male (alcohol) (NMNH); Zamora-Chinchipe: Zamora, 4. xii. 1978, JJ Anderson – 232 males, 280 females (alcohol) (NMNH); Río Chicaña, 9 km N Yanzatza, 880 m, 20. ix. 1990, OS Flint, Jr – 11 males, 32 females (alcohol) (UMSP); same data as Holotype – 13 males, 110 females (alcohol) (NMNH); 6 km E Zumbi, 980 m, 21. ix. 1990, OS Flint, Jr – 1 male (pinned), 1 male, 1 female (alcohol) (NMNH). Etymology — This species is named M. licina from the Latin word licinus, meaning bent or turned upward, in reference to the strongly upturned ventral lobes of the inferior appendages in this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF96F868FF01BC264409FBAF.taxon	description	Fig. 94, 124 This is a distinctive species, unlikely to be confused with any other species in the velasquezi group. Especially diagnostic is the structure of the dorsal phallic spine, which has a patch of minute spines on its posteroventral surface. Other useful diagnostic characters include the strongly curved paramere appendages, relatively short inferior appendages, and the distinctively arched tergum X (as viewed laterally). Additionally, males have an extensive field of darkened scale-like setae on the hind wings. The female is also distinctive in that it has an arched and projecting tergum VIII. Adult — Length of forewing: male 2.8 - 3.2 mm; female 3.0 - 3.5 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color (in alcohol) light brown. Tibial spurs slightly darker than legs, contrasting in color. Forewing with distinct white wing bar at anastomosis. Males with scale-like setae paralleling veins in fore- and hind wing, hind wing with extensive field of darkened, semi-erect, scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow and slightly constricted basally, length about 4 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by about ½ width of segment. Tergum X, as viewed laterally, relatively short and narrow, distinctly arched in middle; as viewed dorsally, short and moderate in width, lateral margins subparallel, apicomesally with deep emargination, extending more than ½ length of tergum, basomesally with pair of setose sclerites; ventrolateral lobes obsolete. Inferior appendage with digitate, setose, dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets. Mesal pockets of inferior appendage very large, with moderately elongate, thick, spine-like, apicoventral projections. Paramere appendage moderately elongate, narrow, distinctly curved, ventrally near base and dorsally at apex, apex acute. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, almost evenly dorsally curved from base, apex slightly inflated, ventral margin with distinct notch near middle, apparently to accommodate lightly sclerotized projection on endophallic membrane, upturned part with slight indentation bearing numerous small spines; as viewed dorsally, with distinct dorsolateral projections, widest in basal ½, narrowing apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with broadly rounded sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with lightly sclerotized projection, ventrally with pair of short sclerotized projections (modified phallotremal spines?), each bluntly rounded as viewed ventrally, apices with minute spines.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF96F868FF01BC264409FBAF.taxon	materials_examined	Holotype male (alcohol) — ECUADOR: Pastaza: Puyo, 5. v. 1977, PJ Spangler and DR Givens (UMSP 000097077) (NMNH). Paratypes — ECUADOR: Napo: Puyo, 6. v. 1977, PJ Spangler and DR Givens – 1 male, 1 female (alcohol) (NMNH); Lago Agrio (30 km E) Via a Tarapoa, 17. x. 1975, A Langley – 22 males, 25 females (alcohol) (NMNH); Pastaza: Puyo, 5. v. 1977, PJ Spangler and DR Givens – 5 males, 7 females (alcohol) (NMNH); same locality and collectors, 6. v. 1977 – 3 males, 2 females (alcohol) (UMSP); same locality and collectors, 7. v. 1977 – 3 females (alcohol) (NMNH); Puyo, 30. i. 1976, Spangler et al. – 1 male, 7 females (alcohol) (NMNH); same locality and collectors, 1 - 7. ii. 1976 – 1 male (alcohol) (NMNH); same locality and collectors, 8 - 11. ii. 1976 – 1 female (alcohol) (NMNH); Puyo (27 km N) Est. Fluvia Metrica, 4. ii. 1976, Spangler et al. – 3 females (alcohol) (NMNH); Puyo (16 km W), 3. ii. 1976, Spangler et al. – 18 males, 41 females (alcohol) (NMNH); Tungurahua: Baños, 1798 m, 28. v. 1975, Cohen and Langley – 3 males, 2 females (alcohol) (NMNH); Baños (34 km E), 25. i. 1976, 1280 m, Spangler et al. – 9 males, 15 females (alcohol) (NMNH); Zamora-Chinchipe: Zamora, at lights, 1 - 5. vi. 1976, A Langley, et al. – 1 male (pinned) (NMNH); Zamora, 4. xii. 1978, JJ Anderson – 3 males, 10 females (alcohol) (NMNH). Etymology — We take pleasure in naming this species for Dr. Paul Spangler, retired aquatic beetle specialist at the Smithsonian Institution, who also collected many specimens of Trichoptera during his career, including the majority of type specimens of this species.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF97F877FF01BC064293FACF.taxon	description	Fig. 95, 119	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF97F877FF01BC064293FACF.taxon	description	Adult — Length of forewing: male 2.5 - 2.9 mm; female 3.3 - 4.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsi slightly paler. Tibial spurs darker than legs, contrasting in color. Forewing with narrow white wing bar at anastomosis. Males with forewings densely covered with short, adpressed, and somewhat thickened setae; those paralleling major veins in hind wings scale-like; hind wing with additional field of slightly darkened, short, semi-erect, scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow and slightly constricted basally, length about 4 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X, as viewed laterally, relatively short and narrow, rod-like, declivous apically, basomesally with pair of setose sclerites; as viewed dorsally, short and very wide, apicomesally with deep and very wide U-shaped emargination, extending more than ½ length of tergum; ventrolateral lobes obsolete. Inferior appendage with short digitate, setose, dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets. Mesal pockets of inferior appendage very large, with moderately elongate, thick, posterodorsally curved, spine-like, apicoventral projections. Paramere appendage short, narrow, apex acute, emerging near dorsal margin of basal segment, weakly dorsally curved. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, short and relatively wide, dorsolateral margins expanded, spine weakly reflexed in about apical 2 / 5, basal part with distinct projecting “ belly, ” ventral margin with rounded indentation at apical inflection, reflexed apex with distinctly angular projection on posterior margin, apex acute; as viewed dorsally, very wide, lateral margins subparallel, apex bluntly rounded. Phallicata short, continuous with endophallic membrane apically, laterally with relatively elongate, apically rounded, sclerotized projections, projecting beyond spine-like projections of mesal pockets of inferior appendage; projections appearing ventrally as broad, apically rounded lobes of inferior appendages. Endophallic membrane continuous with phallicata, without membranous lateral lobes, dorsomesally with strongly sclerotized, short, spine-like projection, articulating with indentation in dorsal phallic spine; ventrally with pair of short blunt sclerites (modified phallotremal spines?). Material examined — COLOMBIA: Antioquia: Finca Velasquez, Sopetran, 14. ii. 1983, OS Flint, Jr – 1 male Paratype (pinned) (NMNH); Quebrada El Poso, 8 km W El Penol, 9. ii. 1983, OS Flint, Jr – 1 male Paratype (pinned) (NMNH); Cauca: Municipio de Inzá, Quebrada San Andrés, ca. 500 m W Restaurante “ La Portada, ” San Andrés de Pisimbalá, 2.58222 ° N, 76.04333 ° W, el 1750 m, 21. xii. 1997, F Muñoz-Q, et al. – 7 males, 11 females (pinned) (UMSP); Risaralda: 4 km E Santa Rosa de Cabal, 29. ii. 1984, CM and OS Flint, Jr – 1 male (alcohol) (NMNH); Valle: Municipio de Buga, Río Guadalajara, “ La Piscina, ” ca. 4 km SE La Habana, 3.87472 ° N, 76.16889 ° W, el 1620 m, 8. i. 1998, F Muñoz-Q, et al. – 1 male, 5 females (pinned) (UMSP); ECUADOR: Cotopaxi: Quevedo (36 km NE), 335 m, 21. vii. 1976, J Cohen – 15 males, 23 females (alcohol) (NMNH); Imbabura: Río Chota, 10. ix. 1977, LE Peña G – 3 males, 2 females (pinned) (NMNH); Pichincha: Río Umachaca, Forest Station Maquipucuna, 0.12500 ° N, 78.61667 ° W, 1250 m, 4 - 5. ix. 1990, OS Flint, Jr – 1 male (pinned), 12 males, 13 females (alcohol) (NMNH); Nanegal, 1100 m, 19 - 20. ix. 1977, LE Peña G – 7 males, 3 females (alcohol) (NMNH); N Perucho, 18 - 19. ix. 1977, LE Peña G – 1 male, 28 females (alcohol) (NMNH); Santo Domingo (47 km S) 29. vii. 1976, J Cohen – 106 males, 10 females (alcohol) (NMNH); Santo Domingo de los Colorados (14 km E), 5. vii. 1975, Langley and Cohen – 2 males (alcohol) (NMNH). Distribution — Colombia, Ecuador.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF88F876FF01BDA641A5FA2F.taxon	description	Fig. 96, 123 Mortoniella venezuelensis is very closely related to M. eduardoi Rueda Martín and Gibon and M. licina, n. sp. All of these species have the apical spine-like projections from the mesal pockets of the inferior appendages very elongate and fused to the ventral part of the inferior appendages to form elongate, projecting lobes. All of the species also have an endophallic membrane that is ornamented with a dorsomesal projection with minute spines, membranous lateral lobes with minute spines, and paired ventral sclerites, each with a somewhat forked apex. Mortoniella venezuelensis differs from M. eduardoi in having paramere appendages that are uniform in width and not modified apically. Additionally, the color is different, medium brown, rather than very dark brown. Mortoniella licina differs in having slightly longer paramere appendages, which are also usually curved apically, rather than projecting straight. It also differs in that the fused ventral lobes of the inferior appendages are more elongate and more strongly dorsally curved than in M. venezuelensis. Additionally, M. licina differs in lacking a field of upright scale-like setae on the hind wing, which characterizes both M. venezuelensis and M. eduardoi. Adult — Length of forewing: male 2.6 - 2.9 mm; female 3.0 - 3.3 mm. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0: 3: 4. Overall color medium brown, apices of tarsi paler. Tibial spurs slightly darker than legs, weakly contrasting in color. Forewing with narrow whitish wing bar at anastomosis. Males with scale-like setae paralleling veins in fore- and hind wings, hind wing with additional field of darkened, semi-erect, scale-like setae. Male genitalia — Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X, as viewed laterally, relatively short, narrow, slightly declivous; as viewed dorsally, short and wide, apicomesally with deep U-shaped emargination, extending more than ½ length of tergum; ventrolateral lobes obsolete. Inferior appendage with digitate, setose, dorsolateral projections, fused apicoventrally to spine-like projections of the mesal pockets; composite apical structure forming prominent, elongate ventral lobes. Mesal pockets of inferior appendage very large, with elongate, thick, spine-like, apicoventral projections. Paramere appendage elongate, narrow, nearly uniform in width, extending nearly straight, apex acute. Phallobase very short, with elongate ventral rod-like projections, each strongly flared apically. Dorsal phallic spine, as viewed laterally, somewhat widened basally, strongly reflexed in about apical 1 / 3, apex rounded; as viewed dorsally, with distinct dorsolateral projections, widest in basal ½, tapering apically, reflexed apex rounded. Phallicata short, continuous with endophallic membrane apically, laterally with broadly rounded, sclerotized projections, surrounding mesal pockets of inferior appendage. Endophallic membrane continuous with phallicata, dorsomesally with slightly raised projection with numerous minute spines, laterally with projecting membranous lobes with minute spines, ventrally with pair of sclerotized projections, each with v-shaped apical notch (modified phallotremal spines?).	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF88F876FF01BDA641A5FA2F.taxon	materials_examined	Holotype male (pinned) — VENEZUELA: Barinas: Río Sinigüis in Caño Grande, 8.40000 ° N, 70.77417 ° W, el 520 m, 22. iii. 1997, Holzenthal (UMSP 000001551) (UMSP). Paratypes — VENEZUELA: Barinas: same data as Holotype – 8 males, 7 females (pinned), 42 males (alcohol) (UMSP), 15 males (alcohol) (MIZA); Río Santo Domingo, Barinas, 17. ii. 1976, CM and OS Flint, Jr – 81 males, 198 females (alcohol) (NMNH); Zulia: Perija El Tucuco, Missión El Tucuco, small stream nr. church, 27. ix. 1979, HM Savage – 1 male, 1 female (alcohol) (NMNH); Perijo El Tucuco, Missión El Tucuco, Río El Tucuco, 1 / 2 km from church, 1 - 5. x. 1979, HM Savage – 47 males, 31 females (alcohol) (NMNH); El Tucuco, Sierra de Perija, montane forest, 28 - 29. i. 1978, JB Heppner – 4 males (alcohol) (NMNH); Zulia: Parque Nacional Perijá, Río Negro in Toromo, 10.0510 ° N, 72.7120 ° W, el 360 m, 15. i. 1994, Holzenthal, Cressa, Rincón – 3 males (alcohol) (UMSP). Etymology — This species is named M. venezuelensis for Venezuela, the country of origin of the holotype specimen.	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF78F887FF01B8A640D0FA6F.taxon	description	............................................................................... ormina group, subgenus (Nanotrichia)	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
03F687A7FF78F887FF01B8A640D0FA6F.taxon	description	Apicomesal invagination of tergum IX narrow, lateral lobes more quadrate (Fig. 119 C, 120) ..................................................................................................................................................... 7	en	Blahnik, Roger J., Holzenthal, Ralph W. (2017): Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *. Insecta Mundi 2017 (602): 1-251, DOI: 10.5281/zenodo.5170203
