taxonID	type	description	language	source
03F68791FFD1180F8FFE3876025EFFD1.taxon	discussion	Remarks. Although Horaeomorphus is highly diverse and includes species small and large, flattened and strongly convex, with the pronotum strongly elongate or nearly circular, with or without antebasal transverse groove, yet with some experience it is easy to identify the genus even without examining the ventral characters. Independently of the variable body shape, the general shape of the head is nearly identical in all species of Horaeomorphus: always flattened, with the vertex never projecting dorsocaudad and its posterior margin concave; the bristles on tempora, genae and postgenae are sparse and inconspicuous in dry-mounted specimens, visible only in transparent mounts under high magnifications of a compound microscope; the supraantenal tubercles always prominent and in nearly all species accompanied posteromesally by a pair of distinct pits (absent only in H. calcarifer (Franz) and barely discernible in some specimens of H. sarawakensis Franz (Jałoszyński 2012 a )). All Australian specimens seen by the author have these pits. Another unique character of Horaeomorphus is the inversely U-shaped groove connecting a pair of basal foveae on each elytron; the groove extends from each fovea anteriorly onto the elytral articulating lobe overlapped by the pronotal base, but is usually partly visible in intact specimens. Many species of Horaeomorphus show a distinct sexual dimorphism, frequently expressed in elongated metatrochanters of males, sometimes to such extent that the distal part of trochanter forms a long rod-like projection exceeding half length of femur (examples are shown in Figs. 2, 4 – 5, and details in Fig. 20). In two species males differ from females in metafemora with a small ventral tooth. Australian species, at least those known from males, do not have any modifications of the legs. A character previously unknown was discovered during the present study. Horaeomorphus sakishimanus has the mesocoxae with dense and relatively long setae on the dorsal surface (Figs. 12 – 13), and bunches of two-nine setae are inserted on small papillae (Fig. 14). Examination of 15 species of Oriental Horaeomorphus (cPJ) revealed that all of them have densely setose dorsal surface of mesocoxae. Among Australian Cyrtoscydmini bunches of setae on the dorsal surface of mesocoxae were found only in Scydmaenozila gen. n. (Fig. 66), but in this genus there are two groups of setae, each seta inserted separately, the surface of coxa is not papillate, and there are numerous pores visible on the coxal cuticle. Composition and distribution. Thirty-nine species (including those redescribed and described in the present paper) can be unambiguously placed in Horaeomorphus. Thirty six are Asian (Oriental, Himalayan and Far Eastern), one Melanesian (from Fiji) and two Australian. Numerous additional undescribed species were seen by the author, mostly represented by unidentifiable females, from SE Asia (including the Indonesian New Guinea), and two more undescribed species (known from females) occur in Australia. Australian findings (Fig. 193) are located near the eastern coastline and fill a distributional gap between SE Asia / New Guinea and eastern Melanesia.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFDA18188FFE3A8B000DFE24.taxon	materials_examined	Type material studied. Lectotype (here designated): AUSTRALIA: ♂: five labels (Fig. 24): " Sidney " [white, handwritten], [Stenichnus [sic!] / anthicoides / Lea / det. H. Franz " [white, handwritten and printed], " S. Aust. Museum / specimen " [orange, printed], " PHAGONOPHANA / anthicoides Lea, 1914 / P. Jałoszyński, 2014 / LECTOTYPUS " [white, printed], " HORAEOMORPHUS / anthicoides (Lea, 1914) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Paralectotypes (2 ♀♀): 1 ♀, thick cardboard on which the specimen was originally mounted with handwritten " TY " and five labels (Fig. 25): " 13811 / Phagonophana / anthicoides Lea / N. S. Wales / TYPE " [white, handwritten, " TYPE " in red], " SAMA Database / No. 25 - 037014 " [white, printed], " PHAGONOPHANA / anthicoides Lea, 1914 / P. Jałoszyński, 2014 / PARALECTOTYPUS " [white, printed], " HORAEOMORPHUS / anthicoides (Lea, 1914) / P. Jałoszyński, 2014 " [white, printed] (SAM); 1 ♀ (headless), same data as for the lectotype, except for white paralectotype label (SAM). Revised diagnosis. Male and female: BL <1.8 mm; pronotum with three antebasal pits not connected by transverse groove. Male: aedeagus in ventral view with apical part projecting distally in middle, pointed.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFDA18188FFE3A8B000DFE24.taxon	description	Redescription. Body of male (Fig. 22) flattened, elongate and slender, with moderately long appendages, BL 1.78 mm; cuticle glossy, body uniformly light brown with slightly lighter appendages, vestiture yellowish. Head (Fig. 22) broadest at eyes, HL 0.28 mm, HW 0.40 mm; tempora slightly longer than eyes, strongly curving posteromesally; vertex strongly transverse, more convex on sides than in middle, with distinctly concave posterior margin; frons subtrapezoidal, posteriorly confluent with vertex, anteriorly gradually lowering toward clypeus; supraantennal tubercles prominent, each distinctly demarcated from median part of frons but not demarcated laterally from vertex, accompanied by pair of distinct pits near their posteromesal margins. Eyes moderately large, bean-shaped, weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect, except for sparse and short erect bristles on tempora. Antennae (Fig. 22) slender, AnL 0.80 mm; antennomeres I – IV each strongly elongate, V – VI slightly elongate, VII about as long as broad, VIII – X strongly transverse, XI shorter than IX – X together, about 1.6 x as long as broad, with indistinctly pointed apex. Pronotum (Fig. 22) strongly elongate and weakly convex, broadest near anterior third, PL 0.50 mm, PW 0.44 mm; anterior and lateral margins rounded; sides strongly convergent posteriorly; hind pronotal corners distinct but obtuse and blunt; posterior margin weakly arcuate; base of pronotum with three small and shallow but distinct pits. Punctures on pronotal disc distinct and relatively large, in middle of disc separated by spaces about as wide as 1 – 1.5 x diameters of puncture; setae sparse, thin, short and suberect, sides of pronotum with bristles. Elytra (Fig. 22) oval, as convex as pronotum, broadest in anterior third, EL 1.00 mm, EW 0.63 mm, EI 1.60; basal impressions short; humeral calli developed as elongate protuberances distinctly demarcated mesally from basal part of elytra; elytra with indistinct circumsutural impression near anterior third; elytral apices rounded together. Punctures on elytra unevenly distributed, those on circumsutural impression larger and deeper than those on pronotum, sharply marked and separated by spaces comparable to diameters of punctures, punctures reducing in diameter, depth and sparser toward elytral margins; setae short, sparse and suberect, sides of elytra with bristles. Hind wings well developed, about twice as long as elytra. Legs (Fig. 22) moderately long and slender, without modifications. Aedeagus (Figs. 29 – 30, 33 – 34) moderately stout, AeL 0.40 mm, in ventral view apical part subtrapezoidal, apical margin projecting distally in middle and pointed; internal armature darkly sclerotized and complicated, symmetrical, with long subapical median tubular structure and bell-shaped central complex of sclerites; parameres slender, not exceeding apex of median lobe, with short apical setae. Female (Fig. 23). Externally indistinguishable from male; BL 1.78 mm; HL 0.28 mm, HW 0.38 mm, AnL 0.78 mm; PL 0.50 – 0.51 mm, PW 0.43 - 0.45 mm; EL 1.00 – 1.03 mm, EW 0.60 – 0.65 mm, EI 1.58 – 1.67.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFDA18188FFE3A8B000DFE24.taxon	distribution	Distribution (Fig. 193 a). South-eastern Australia (eastern New South Wales).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFDA18188FFE3A8B000DFE24.taxon	discussion	Remarks. Horaeomorphus anthicoides among Australo-Oriental congeners is most similar to H. verus sp. n. described below. Both species have slender and flat bodies with distinctly elongate pronotum, with unmodified hind trochanters and femora in males. These characters clearly separate the Australian species from those occurring in the Oriental region that have a broad, nearly circular pronotum, strongly convex body or male sexual dimorphic characters located on the hind legs. A similar body form to that of H. anthicoides and H. verus can be found in H. wailimae (Lhoste) and H. deformatus Jałoszyński, 2006, but these species have modified hind legs. Horaeomorphus anthicoides externally also resembles H. minor Jałoszyński, 2009 from the Philippines, but the latter species has a strongly asymmetrical internal armature of the aedeagus. Interestingly, H. fakfakensis Jałoszyński, 2009 from New Guinea is not so similar to the Australian congeners. Franz (1975) studied three specimens from SAM, and he treated one of them as a holotype (apparently the male, whose aedeagus was illustrated in Franz (1975 )). Lea (1914) did not specify the number of specimens used for the species description. Originally, all three specimens were presumably mounted on one card, and one of them was annotated as " TY " (Fig. 25). This is not a valid designation of the holotype, and all three specimens have a status of syntypes. The Franz's (1975) redescription does not contain any data that fulfill the requirements of the formal lectotype designation. The " holotype " bears labels written apparently by Franz, who also re-mounted the three specimens; the original set of labels is now pinned under one of the females. The male is here designated as a lectotype to provide a name-bearing type for Phagonophana anthicoides. Three females of Horaeomorphus representing two species (Figs. 27 – 28) were found in the ANIC collection. They are similar in the general body shape and measurements to H. anthicoides, but differ slightly in the shape of pronotum and elytra or punctation of various body parts. They were collected in the Blue Mountains and the Royal National Park. Lea (1915) gives further details concerning the collecting locality for H. anthicoides: Sydney, Glen Innes (the town of Glen Innes is located over 600 km north of Sydney).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: three labels: " 16.05 S 145.28 E QLD / CT 2 10 m, 2 km SSW / Cape Tribulation / 1 - 28 Feb 1996 / L. Umback / FI Trap JCU " [white, printed], " ANIC / Specimen " [green, printed], " HORAEOMORPHUS / verus m. /, det. P. Jałoszyński, 2013 / HOLOTYPUS " [red, printed] (ANIC).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	diagnosis	Diagnosis. Male: BL> 2 mm; pronotum with three antebasal pits connected by distinct transverse groove. Male: aedeagus in ventral view with apical part broadly rounded.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	description	Description. Body (Fig. 26) flattened, elongate and slender, with moderately long appendages, BL 2.05 mm; cuticle glossy, body uniformly dark brown with slightly lighter appendages, vestiture light brown. Head (Fig. 26) broadest at eyes, HL 0.33 mm, HW 0.41 mm; tempora slightly longer than eyes, strongly curving posteromesally; vertex strongly transverse, more convex on sides than in middle, with distinctly concave posterior margin; frons subtrapezoidal, posteriorly confluent with vertex, anteriorly gradually lowering toward clypeus; supraantennal tubercles prominent, each sharply demarcated from median part of frons and indistinctly demarcated laterally from vertex, each accompanied by pair of deep pits near posteromesal margin. Eyes moderately large, bean-shaped, weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect, except for sparse and short erect bristles on tempora. Antennae (Fig. 26) slender, AnL 0.88 mm; antennomeres I – VI each distinctly elongate, VII about as long as broad, VIII – X strongly transverse, XI shorter than IX – X together, about 1.5 x as long as broad, with indistinctly pointed apex. Pronotum (Fig. 26) distinctly elongate and weakly convex, broadest near anterior third, PL 0.58 mm, PW 0.53 mm; anterior and lateral margins rounded; sides moderately strongly convergent posteriorly; hind pronotal corners distinct but obtuse and blunt; posterior margin weakly arcuate; base of pronotum with three small but deep and distinct pits connected by sharply marked transverse groove. Punctures on pronotal disc small but distinct, in middle of disc separated by spaces about as wide as 1 - 1.5 x diameters of punctures; setae moderately dense, thin, short and suberect, sides of pronotum with bristles. Elytra (Fig. 26) oval, as convex as pronotum, broadest slightly behind anterior third, EL 1.15 mm, EW 0.75 mm, EI 1.53; basal impressions short; humeral calli developed as elongate protuberances distinctly demarcated mesally from basal part of elytra; elytra with indistinct circumsutural impression near anterior third, small area behind impression slightly raised; elytral apices rounded together. Punctures on elytra unevenly distributed, those on circumsutural impression larger and deeper than those on pronotum, sharply marked and separated by spaces as wide as 0.5 – 1 diameters of punctures, punctures reducing in diameter, depth and sparser toward elytral margins; setae moderately long and dense, suberect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 26) moderately long and slender, without modifications. Aedeagus (Figs. 31 – 32, 35 – 36) moderately stout, AeL 0.40 mm, in ventral view apical part subtrapezoidal, apical margin rounded in middle; internal armature darkly sclerotized and complicated, symmetrical, with long subapical median tubular structure and bell-shaped central complex of sclerites; parameres slender, not exceeding apex of median lobe, with long apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	etymology	Etymology. The Latin adjective verus (real, true, genuine) was chosen as a specific epithet, to underline that this is the first and only true Australian Horaeomorphus originally placed in a correct genus.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	distribution	Distribution (Fig. 193 b). North-eastern Australia (northern Queensland).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC718198FFE387001DFFDBB.taxon	discussion	Remarks. Horaeomorphus verus has the aedeagus similar to that of H. anthicoides, but clearly differs in the external structures; e. g., the body length, proportions of body parts, the sculpture of pronotal base and much longer and denser setae on the dorsum. For characters separating this species from Oriental congeners see the Remarks section at the redescription of H. anthicoides.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC6181D8FFE3BC002C9FDC8.taxon	materials_examined	Type species: Scydmaenus simplicicornis Lea, 1910: 185 (here designated).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC6181D8FFE3BC002C9FDC8.taxon	diagnosis	Diagnosis. Body (Figs. 46 – 48) elongate and strongly convex; head (Fig. 37) short and convex; vertex (Fig. 37; vt) not projecting dorsocaudad, its posterior margin convex or nearly straight; supraantennal tubercles barely marked and not accompanied posteriorly by pits; frontoclypeal suture or groove absent; eyes located in middle of head; tempora (Fig. 37; tm) long and strongly curved toward occipital constriction; tempora, genae and postgenae without bristles; ' neck region' distinctly demarcated from anterior part of head capsule by constriction and much narrower than vertex; hypostomal ridges (Fig. 38; hr) rudimentary, indistinctly marked only just behind cardines; posterior tentorial pits (Fig. 38; ptp) elongate and located distinctly in front of transverse groove demarcating ' neck' ventrally; submentum (Fig. 38; smn) without lateral sutures; each mandible with apical tooth sharply demarcated from basal part and densely setose prostheca (Fig. 40; pst), subapical teeth absent; maxillary palpomeres III and IV (Fig. 41; mxp 3 – 4) slender, strongly elongate; pronotum (Figs. 42, 46, 48) without antebasal pits or grooves, only with barely noticeable, shallow and diffused transverse impression; prosternum laterally demarcated from hypomera by pronotosternal sutures (Fig. 43; nss), without intercoxal process or carina; procoxal sockets (Fig. 43; pcs) narrowly open; prothoracic hypomera (Fig. 43; hy) with large internal (adcoxal) parts demarcated from sides of pronotum by incomplete hypomeral ridges (Fig. 43; hyr) developed only anteriorly; sides of prothorax (Fig. 43) with short and dense thick bristles located only in anteroventral portion and not visible or barely visible in dorsal view; mesoventrite with short anterior ridge (Fig. 45; ar) without posterior median projection; mesoventral process (Fig. 45; msvp) narrow and weakly expanding ventrally but distinct and clearly separating mesocoxae, interrupted near anterior margins of mesocoxal cavities; ventro- and dorsolateral foveae (Fig. 45; vlf, dlf) present; anterior metaventral process (Fig. 45; amvp) present; metaventral intercoxal process (Fig. 45; mtvp) broadly separating metacoxae, without spines, with shallowly concave posterior margin; each elytron with two asetose rudiments of basal foveae (Fig. 44; bef) not connected by groove and barely discernible in dry-mounted specimens; aedeagus (Figs. 49 – 56) with free and slender parameres.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC6181D8FFE3BC002C9FDC8.taxon	description	Description. Body (Figs. 46, 48) 2.20 – 2.58 in length, strongly convex, elongate but moderately slender, with long appendages, pigmentation from reddish-brown to nearly black, cuticle setose. Head (Figs. 37 – 38, 46, 48) with occipital constriction (Fig. 37; occ) much narrower than vertex and dividing the head capsule into exposed anterior part and narrow posterior ' neck region' retracted into prothorax; eyes small and slightly oval, located in median part of head; tempora (Fig. 37; tm) long and strongly curved posteromesally, without bristles; vertex (Fig. 37; vt) transverse, uniformly convex, not projecting dorso-caudad, with evenly rounded and convex posterior margin sharply demarcated from occiput; frons (Fig. 37; fr) confluent with vertex, transverse and subtrapezoidal, evenly convex, anteriorly steeply but not abruptly lowering toward labrum; frontoclypeal groove absent; antennal insertions broadly separated, located beneath feebly developed supraantennal tubercles. Labrum (Fig. 39) transverse with rounded anterior margin and narrow translucent marginal velum, with six thick anteroventral (anterior epipharyngeal) bristle-like sensilla projecting from under anterior margin, and with large number of moderately long dorsal setae. Mandibles (Fig. 40) symmetrical, subtriangular, each with broad base and curved, rapidly narrowing and pointed apical part; prostheca (Fig. 40; pst) present, with dense thin setae along mesal mandibular margin, slightly expanding onto dorsal surface of mandible and reaching mandibular base. Maxilla (Fig. 41) composed of small cardo (Fig. 41; cd) bearing several setae; subtriangular basistipes (Fig. 41; bst); elongate mediostipes (Fig. 41; mst); elongate galea (Fig. 41; gal) and lacinia (Fig. 41; lac); large, elongate palpifer (Fig. 41; pfp); and strongly elongate and moderately large maxillary palp (Fig. 38; mxp) composed of elongate palpomere I (Fig. 41; mxp 1), strongly elongate, clavate and slender palpomere II (Fig. 41; mxp 2), large and strongly elongate palpomere III (Fig. 41; mxp 3) broadest near distal third, and small, slender, subconical and pointed palpomere IV (Fig. 41; mxp 4). Labium (Fig. 38) with large submentum (Fig. 38; smn) not demarcated posteriorly from gular plate and not demarcated laterally from postcardinal parts of hypostomae; subrectangular and transverse mentum (Fig. 38; mn); and relatively long prementum (Fig. 38; pmn) bearing narrowly separated at bases long 3 - segmented labial palps and ligula with single median bristle. Hypostomal ridges (Fig. 38; hr) superficial and visible only just behind cardines. Gular plate (Fig. 38; gp) large and subtrapezoidal, with rapidly narrowed anterior portion; gular sutures (Fig. 38; gs) superficial; posterior tentorial pits (Fig. 38; ptp) narrow and arcuate, located clearly in front of transverse groove ventrally demarcating ' neck region'. Antennae (Figs. 46, 48) long and slender, gradually but only slightly thickening distally; antennomere XI conspicuously long. Prothorax (Figs. 42 – 43, 46, 48) in dorsal view subtrapezoidal, broadest anterior to middle, with rounded anterior margin and anterior parts of lateral margins, with weakly marked, blunt and obtuse anterior corners; posterior corners blunt and obtuse; posterior margin weakly arcuate. Pronotum without antebasal pits and grooves, and without sublateral carinae or lateral edges, with indistinct, shallow and diffused antebasal transverse impression (Fig. 42). Sides of pronotum mostly with thin setae, only anteroventral parts of hypomera with thick bristles (Fig. 43). Prosternum (Fig. 43) with moderately short basisternal part (Fig. 43; bst) indistinctly demarcated from procoxal cavities (Fig. 43; pcc); median part of sternum slightly raised but not carinate; procoxal sockets (Fig. 43; pcs) open; hypomera (Fig. 43; hy) elongate, each divided into broad lateral part confluent with pronotum and narrower but still broad internal (adcoxal) part; hypomeral ridges (Fig. 43; hyr) incomplete, visible only along anterior margins of adcoxal parts of hypomera; pronotosternal sutures (Fig. 43; nss) entire. Mesocutellum (Fig. 44; scl 2) subtriangular, in intact specimens hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture present. Mesoventrite (Fig. 45) with narrow and indistinctly demarcated anterior ridge (Fig. 45; ar); mesoventral intercoxal process (Fig. 45; msvp) carinate and moderately expanding ventrally, anteriorly separated from anterior ridge and interrupted near anterior margins of mesocoxal cavities; sides of mesoventrite with shallow concave procoxal rests (Fig. 45; pcr) filled with short setae (= setose impressions) but without asetose impressions; mesanepisternum with moderately long prepectus (Fig. 45; pre), sides of mesoventrite with two pairs of narrow, deep and setose ventrolateral and dorsolateral foveae (Fig. 45; vlf, dlf); mesocoxal projections (Fig. 45; mcp) prominent, with mesocoxal sockets located on their mesal surface and not visible in ventral view. Metaventrite (Fig. 45; v 3) subrectangular, anteriorly fused with mesoventrite, posteriorly moderately deeply bisinuate and with broad, shallowly emarginate metaventral intercoxal process (Fig. 45; mtvp); anteriorly with short and narrow anterior metaventral process (Fig. 45; amvp). Metanepisterna and metepimera narrow. Metafurca (Fig. 45) with short stalk and divergent lateral furcal arms (Fig. 45; lmfa). Elytra (Figs. 44, 46, 48) oval, each with two asetose rudiments of basal foveae (Fig. 44; bef) barely discernible in dry-mounted specimens; each humerus with small and blunt but distinct humeral denticle (Fig. 42; hd); subhumeral lines absent. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 46, 48) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; metatrochanters relatively long; all femora strongly clavate; tibiae and tarsi long and slender. Aedeagus (Figs. 49 – 56) elongate and darkly sclerotized, relatively thick-walled, with symmetrical median lobe but strongly asymmetrical and complicated subapical assemblage of internal sclerites; parameres free and slender, with apical setae.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC6181D8FFE3BC002C9FDC8.taxon	etymology	Etymology. Leascydmus is a name dedicated to Arthur Mills Lea, an Australian entomologist who described many Scydmaeninae and in the description of Scydmaenus simplicicornis wrote: " the species may eventually be regarded as belonging to a new genus ", which proved correct. Gender masculine. Composition and distribution. Two species of Leascydmus are known from south-eastern Australia (Victoria and New South Wales; Fig. 194).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC6181D8FFE3BC002C9FDC8.taxon	discussion	Remarks. Leascydmus is a remarkable genus because of its superficial similarity to Scydmaenus Latreille, 1802. The general shape of the pronotum and elytra, the broad metaventral intercoxal process and relatively long metatrochanters resemble some subgenera of Scydmaenus, from which Leascydmus differs in the diagnostic characters of Cyrtoscydmini (e. g., the subconical maxillary palpomere IV; discussed in Jałoszyński 2012 c). Among Australian Cyrtoscydmini open procoxal sockets can be found only in Palaeoscydmaenus Franz, but this genus has a clearly different habitus, with subtriangular head, the ' neck region' nearly as broad as vertex and much less distinctly demarcated from the anterior part of the head capsule, and the pronotum elongate and broadest in the anterior part. Besides, Palaeoscydmaenus belongs to a group of genera characterized by the submentum demarcated laterally by complete lateral sutures, which are absent in Leascydmus. Leascydmus differs from all Australian genera without lateral sutures of submentum in rudimentary hypostomal ridges (shared only by Microscydmus and Penicillidmus, but these genera include beetles not reaching 1 mm in length, with 3 - segmented antennal club, and antebasal pits on pronotum), tempora, genae and postgenae covered only with setae, without bristles, open procoxal sockets and broadly separated metacoxae (character shared with at least some subgenera of Euconnus, including Euconnus s. str., but the latter genus has pits and sometimes transverse grooves on the base of pronotum and long, continuous and strongly expanding ventrally, keel-shaped mesoventral intercoxal process). The short head with arcuate posterior margin of convex vertex, short pronotum without pits or grooves, humeral denticles and strongly elongate antennae with the antennomere XI as long as IX – X together are sufficient to identify Leascydmus.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC2181E8FFE38E00024FF0B.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: seven labels (Fig. 47): " simplicicornis / Lea, Type / Geelong " [white, handwritten and printed], " simplicicornis / Lea TYPE / Geelong " [white, handwritten], " 13295 / Scydmaenus / simplicicornis / Lea / Victoria / TYPE " [white, handwritten, " TYPE " on right margin in red], " Horaeomorphus / simplicicornis / (Lea) / det. H. Franz " [white, handwritten and printed], " SAMA Database / No. 25 - 037063 " [white, printed], " SAMA Digital Image / 4. 9. 2013 " [green, printed and handwritten], " LEASCYDMUS / simplicicornis (Lea, 1910) / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (SAM). Additional material: ♀, " Forrest V. ", leg. or coll. H. W. Davey (labeled as " Co-type "; see remarks below) (SAM). Revised diagnosis. Male and female: body nearly black with yellowish-brown tarsi. Male: aedeagus in lateral view with apex slightly curved dorsally.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC2181E8FFE38E00024FF0B.taxon	description	Redescription. Body of male strongly convex, elongate and slender, with long appendages, BL 2.58 mm; cuticle glossy, body nearly black, margins of elytra reddish, antennae and apices of tibiae brown, tarsi yellowishbrown, vestiture yellowish. Head broadest at eyes, HL 0.48 mm, HW 0.53 mm; tempora longer than eyes, strongly curved posteromesally; vertex and frons confluent and evenly convex. Eyes small and weakly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae short, sparse and suberect. Antennae slender, AnL 1.75 mm; all antennomeres elongate, antennomere XI as long as IX – X together. Pronotum in dorsal view approximately subquadrate with rounded sides, broadest near anterior third, PL 0.58 mm, PW 0.60 mm; anterior margin nearly straight; anterior corners obtuse and blunt; lateral margins rounded; posterior corners strongly obtuse and indistinctly marked; posterior margin nearly straight; pronotal base with shallow and indistinct transverse impression. Punctures on pronotal disc similar to those on frons and vertex; setae sparse, short and suberect, near posterior corners distinctly denser and thicker. Elytra oval, more convex than pronotum, broadest slightly anterior to middle, EL 1.53 mm, EW 0.98 mm, EI 1.56; basal impressions absent; humeral denticles tiny but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae short, sparse and suberect to erect. Hind wings well developed, about twice as long as elytra. Legs long and slender, without modifications; all tibiae broadest in proximal half. Aedeagus (Figs. 49 – 50, 53 – 54) elongate, darkly sclerotized, AeL 0.75 mm, in ventral view apex of ventral wall subtriangular and apex of dorsal wall evenly narrowing; internal armature composed of a complicated set of elongate sclerites; parameres slender, with moderately long apical and subapical setae. Female (Fig. 46). Similar to male; BL 2.58 mm; HL 0.45 mm, HW 0.53 mm, AnL 1.98 mm; PL 0.60 mm, PW 0.60 mm; EL 1.53 mm, EW 1.00 mm, EI 1.53.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC2181E8FFE38E00024FF0B.taxon	distribution	Distribution (Fig. 194 a). South-eastern Australia (southern Victoria).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC2181E8FFE38E00024FF0B.taxon	discussion	Remarks. Scydmaenus simplicicornis is represented in the SAM collection by two specimens (male and female), labeled as a type and co-type. The male bears the original handwritten label (Fig. 47) with the annotation " type ", and a female has printed labels and is marked as a co-type. Lea (1910) in his original descriptions mentioned only one, clearly male specimen and used the term " type " in reference to it. Therefore, in accordance with the ICZN, this male specimen was fixed as a holotype by original designation. The status of the female specimen as a syntype is clearly incorrect. Lea did not mention any specimens other than the male type, and although the collector's name (H. W. Davey) given in the original description appears on the female's label, the collecting locality is " Forrest V. ", which may stand for the Forrest River valley in Western Australia, while the holotype male comes from Geelong, Victoria. It seems unlikely that the same species was collected in SE and WN Australia, by the same collector, and Lea studied both specimens but mentioned only Geelong. The female must be of a later origin; also the collecting site does not seem correct. Lea (1910) mentioned that in the type " the abdomen is strongly notched at apex, the notch fringed with tiny pubescence ". The male is partly disarticulated and the terminal abdominal segments are missing, probably destroyed or lost during a previous preparation; the specimen was not removed from the mounting card during the present study to avoid further damage. The holotype is accompanied by three specimens of ants and Lea (1910) mentioned that it was collected in a nest of Iridomyrmex nitidus.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: two labels: " Megalong / N. S. Wales / J. Armstrong " [white, printed and handwritten], " LEASCYDMUS / armstrongi m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC). Paratypes: 2 ♂♂, 4 ♀♀, same data as for holotype, all with yellow " paratypus " labels (ANIC, cPJ).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	diagnosis	Diagnosis. Male and female: body uniformly brown, only tarsi lighter. Male: aedeagus in lateral view with apex slightly curved ventrally.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	description	Description. Body of male (Fig. 48) strongly convex, elongate and slender, with long appendages, BL 2.35 – 2.48 mm (mean 2.43 mm); cuticle glossy, body brown, tarsi lighter, yellowish-brown, vestiture yellowish. Head (Figs. 37 – 38, 48) broadest at eyes, HL 0.43 mm, HW 0.48 – 0.50 mm (mean 0.49 mm); tempora longer than eyes, strongly curved posteromesally; vertex and frons confluent and evenly convex. Eyes small and weakly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae short, sparse and suberect. Antennae (Fig. 48) slender, AnL 1.83 – 1.85 mm (mean 1.84 mm); all antennomeres elongate, antennomere XI as long as IX – X together. Pronotum (Figs. 42, 48) in dorsal view approximately subquadrate with rounded sides, broadest near anterior third, PL 0.58 – 0.60 mm (mean 0.58 mm), PW 0.55 mm; anterior margin nearly straight; anterior corners obtuse and blunt; lateral margins rounded; posterior corners strongly obtuse and indistinctly marked; posterior margin nearly straight; pronotal base with shallow and indistinct transverse impression. Punctures on pronotal disc similar to those on frons and vertex; setae sparse, short and suberect, near posterior corners distinctly denser and thicker. Elytra (Figs. 44, 48) oval, more convex than pronotum, broadest slightly anterior to middle, EL 1.35 – 1.45 mm (mean 1.42 mm), EW 0.83 – 0.98 mm (mean 0.91 mm), EI 1.49 – 1.64; basal impressions absent; humeral denticles tiny but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae short, sparse and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 48) long and slender, without modifications; all tibiae broadest in proximal half. Aedeagus (Figs. 51 – 52, 55 – 56) elongate, darkly sclerotized, AeL 0.73 mm, in ventral view apex of ventral wall subtrapezoidal and apex of dorsal wall rapidly narrowing; internal armature composed of a complicated set of elongate sclerites; parameres slender, with moderately long apical and subapical setae. Female. Similar to male; BL 2.20 – 2.40 mm (mean 2.30 mm); HL 0.40 – 0.43 mm (mean 0.42 mm), HW 0.48 – 0.50 mm (mean 0.49 mm), AnL 1.63 – 1.78 mm (mean 1.68 mm); PL 0.53 – 0.55 mm (mean 0.53 mm), PW 0.53 – 0.55 mm (mean 0.54 mm); EL 1.25 – 1.43 mm (mean 1.35 mm), EW 0.93 – 1.03 mm (mean 0.98 mm), EI 1.22 – 1.49.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	distribution	Distribution (Fig. 194 b). South-eastern Australia (Blue Mountains in New South Wales).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	etymology	Etymology. Leascydmus armstrongi is dedicated to J. W. T. Armstrong, who described many Australian beetle taxa and collected the type series of this species.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFC018108FFE3D20074FF944.taxon	discussion	Remarks. This species is highly similar to L. simplicicornis in the body shape, proportions of body parts, and even in the aedeagus. The striking difference in pigmentation, smaller body and the antennae distinctly longer in relation to body allow for unambiguous discrimination from L. simplicicornis.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCF18128FFE3FD007C7F9D9.taxon	materials_examined	Type species: Phagonophana macrosticta Lea, 1910: 189 (here designated).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCF18128FFE3FD007C7F9D9.taxon	diagnosis	Diagnosis. Body (Figs. 72 – 75) elongate and strongly convex; head short and convex; vertex not projecting dorsocaudad, its posterior margin nearly straight or indistinctly concave in middle; supraantennal tubercles distinct but weakly raised and not accompanied posteriorly by pits; frontoclypeal groove present; eyes located in middle of head; tempora long and strongly curved toward occipital constriction; tempora, genae and postgenae (Fig. 57) with dense bristles; ' neck region' distinctly demarcated from anterior part of head capsule by constriction and much narrower than vertex (Fig. 57); hypostomal ridges (Fig. 57; hr) sharply marked and long, reaching nearly to posterior tentorial pits and connected in front of them; posterior tentorial pits (Fig. 57; ptp) elongate and located distinctly in front of transverse groove demarcating ' neck' ventrally; submentum (Fig. 57; smn) without lateral sutures; each mandible with sharp and large subapical tooth (Fig. 60; sat) and densely setose dorsomesal prostheca (Fig. 60; pst); maxillary palpomeres III and IV (Fig. 61; mxp 3 – 4) slender, strongly elongate; pronotum with two distinct pairs of antebasal pits (Fig. 64; lp, ip) and additional, indistinct lateral pair of shallow impressions, without lateral and sublateral carinae or edges; prosternum laterally demarcated from hypomera by pronotosternal sutures (Fig. 63; nss), with indistinct intercoxal carina; procoxal sockets (Figs. 62 – 63; pcs) closed by posterolateral lobes of prosternum; prothoracic hypomera (Fig. 62 – 63; hy) with large internal (adcoxal) parts not demarcated from sides of pronotum; hypomeral ridges (Fig. 63; hyr) incomplete, visible only along anterior margins of adcoxal parts of hypomera; sides of pronotum with dense thick bristles (Fig. 62) well visible in dorsal view; mesoventrite with short anterior ridge (Fig. 67; ar) in middle connected with mesoventral process (Figs. 67 – 69; msvp) which is narrow and weakly expanding ventrally but distinct and clearly separating mesocoxae, interrupted in front of anterior margins of mesocoxal cavities; ventro- and dorsolateral foveae (Fig. 69; vlf, dlf) present; anterior metaventral process absent; metaventral intercoxal process (Fig. 71) broadly subtriangular, with two barely discernible tiny denticles, without median notch; metacoxae (Fig. 68; cx 3) narrowly separated in middle; each elytron with two deep and relatively large asetose basal foveae (Fig. 65; bef) not connected by groove and well visible in dry-mounted specimens; aedeagus with free and slender parameres (Fig. 70; pm).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCF18128FFE3FD007C7F9D9.taxon	description	Description. Body (Figs. 72 – 75) large (BL 2.58 – 3.18 mm), strongly convex, elongate but moderately slender, with long appendages, pigmentation dark brown or (in one species) bicolorous, light brown with dark brown elytra bearing variously contrasting light brown spots, cuticle setose. Head (Figs. 57, 72 – 75) with occipital constriction (Fig. 57; occ) much narrower than vertex and dividing the head capsule into exposed anterior part and narrow posterior ' neck region' retracted into prothorax; eyes large and nearly circular, located in anterior part of head; tempora long and strongly curved posteromesally, with dense bristles; vertex transverse, uniformly convex, not projecting dorsocaudad, with rounded, nearly straight or slightly concave posterior margin sharply demarcated from occiput, densely covered with bristles directed posteriorly; frons confluent with vertex, transverse and subtrapezoidal, anteriorly steeply and abruptly lowering toward labrum; frontoclypeal groove present; antennal insertions broadly separated, located beneath distinct but weakly raised supraantennal tubercles. Labrum (Fig. 59) transverse and subtrapezoidal with nearly straight anterior margin, with numerous short anteroventral (anterior epipharyngeal) trichia projecting from under anterior margin, and with small number of long dorsal setae. Mandibles symmetrical, subtriangular, each with broad base and large subapical tooth (Fig. 60; sat); prostheca (Fig. 60; pst) present, with dense and thin setae along mesal margin and extending onto a large portion of dorsal surface, reaching mandibular base. Each maxilla composed of small cardo (Fig. 61; cd) bearing two setae; subtriangular basistipes (Fig. 61; bst); elongate mediostipes (Fig. 61; mst); elongate galea (Fig. 61; gal) and lacinia (Fig. 61; lac); large, elongate palpifer (Fig. 61; pfp); and strongly elongate and moderately large maxillary palp (Fig. 57; mxp) composed of elongate palpomere I (Fig. 61; mxp 1), strongly elongate, slightly clavate and slender palpomere II (Fig. 61; mxp 2), large and strongly elongate palpomere III (Fig. 61; mxp 3) broadest near apex, and small, slender, subconical and pointed palpomere IV (Fig. 61; mxp 4). Labium (Figs. 57 – 58) with large submentum (Fig. 57; smn) not demarcated laterally from postcardinal parts of hypostomae; subtrapezoidal mentum (Fig. 57; mn); and short prementum (Fig. 58; pmn) bearing narrowly separated at bases long 3 - segmented labial palps (Fig. 57; lp) and a pair of median bristles inserted on small rounded ligula (Fig. 58; lg). Hypostomal ridges (Fig. 57; hr) sharply marked, long and extending nearly to posterior tentorial pits, connecting in front of pits. Gular plate (Fig. 57; gp) large and subtrapezoidal, with rapidly narrowed anterior portion; gular sutures (Fig. 57; gs) well-visible; posterior tentorial pits (Fig. 57; ptp) narrow and slightly arcuate, located clearly in front of transverse groove ventrally demarcating ' neck region'. Antennae (Figs. 72 – 75) long and slender, gradually but only slightly thickening distally; antennomere XI longest. Prothorax (Figs. 62 – 64, 72 – 75) in dorsal view strongly elongate and broadest near anterior third, with rounded anterior margin and anterior parts of lateral margins, with weakly marked, blunt and obtuse anterior corners; posterior corners distinct but blunt and obtuse; posterior margin deeply bisinuate. Base of pronotum with two pairs of distinct antebasal pits not connected by groove or impression (Fig. 64; lp, ip), additionally with indistinct lateral impressions, without lateral or sublateral carinae or edges. Sides of pronotum mostly with thick bristles (Fig. 62) well visible in dorsal view. Prosternum with moderately short basisternal part (Figs. 62 – 63; bst) indistinctly demarcated from procoxal cavities (Figs. 62 – 63; pcc); median part of sternum slightly raised but not carinate; procoxal sockets (Figs. 62 – 63; pcs) closed by posterolateral lobes of prosternum; hypomera (Figs. 62 – 63; hy) elongate, each divided into broad lateral part confluent with pronotum and narrower but still broad internal (adcoxal) part; hypomeral ridges (Fig. 63; hyr) incomplete, visible only along anterior margins of adcoxal parts of hypomera; pronotosternal sutures (Fig. 63; nss) entire, obscured by dense setation. Mesocutellum (Fig. 65; scl 2) subtriangular, in intact specimen hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture (Fig. 65; sss) present. Mesoventrite (Figs. 67 – 69) with narrow and distinctly demarcated anterior ridge (Figs. 67 – 69; ar); mesoventral process (Figs. 67 – 69; msvp) carinate and moderately expanding ventrally, anteriorly connected with anterior ridge and interrupted in front of anterior margins of mesocoxal cavities; sides of mesoventrite with shallow concave procoxal rests (Figs. 67 – 69; pcr) filled with short setae (= setose impressions) but without asetose impressions; mesanepisternum with long prepectus (Figs. 67 – 69; pre), sides of mesoventrite with moderately broad, deep and setose ventrolateral and dorsolateral fovea (Fig. 69; vlf, dlf); mesocoxal projections (Figs. 67 – 69; mcp) prominent, with mesocoxal sockets located on their mesoventral surface and visible in ventral view. Metaventrite (Figs. 68 – 69; v 3) subrectangular, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate and with broadly subtriangular metaventral intercoxal process (Fig. 71; mtvp) bearing two tiny denticles; anterior metaventral process absent. Metanepisterna and metepimera narrow. Metafurca (Fig. 69) with long stalk and divergent lateral furcal arms (Fig. 69; lmfa). Elytra (Figs. 65, 72 – 75) oval, each with two moderately large, deep and asetose basal foveae (Fig. 65; bef) not connected by groove; humeral denticles and sub-humeral lines absent. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 66, 72 – 75) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora strongly clavate; tibiae and tarsi long and slender. Mesocoxae (Fig. 66; cx 2) with largely asetose dorsal surface (adjacent to mesocoxal cavity) bearing numerous pores and bunches of long setae. Aedeagus (Figs. 70, 80 – 96) elongate and moderately darkly sclerotized, relatively thick-walled, with symmetrical median lobe and symmetrical, complicated assemblage of internal sclerites; parameres (Fig. 70; pm) free and slender, without apical setae.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCF18128FFE3FD007C7F9D9.taxon	etymology	Etymology. Scydmaenozila is a combination of Scydmaen - and Oz, with feminine suffix. Gender feminine. Composition and distribution. Four species of Scydmaenozila have a disjunct distribution: one is known from the southernmost part of Western Australia, and three others from the south-eastern part of the continent (New South Wales, Australian Capital Territory and South Australia) and Tasmania. One species is known both from the continent and Tasmania. Additionally, three undescribed species represented by female specimens were also seen in ANIC samples, collected in Walpole-Nornalup National Park (Western Australia), Esperance (Western Australia), and Blundells Creek (Australian Capital Territory).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCF18128FFE3FD007C7F9D9.taxon	discussion	Remarks. Scydmaenozila can be identified on the basis of dorsal characters: the large body, short head with distinct frontoclypeal groove, tempora and sides of pronotum with dense bristles; elongate pronotum broadest in anterior part and bearing two pairs of distinct antebasal pits, without grooves or carinae, and two distinct basal elytral foveae. The combination of medially connected and long hypostomal ridges; absent lateral sutures of submentum; incomplete hypomeral ridges; distinct mesoventral process separating mesocoxae; and metacoxae narrowly separated by subtriangular metaventral intercoxal process confirm identification.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCC18158FFE3A8B0760FB85.taxon	materials_examined	Type material studied. Lectotype (here designated): AUSTRALIA: ♂: five labels (Fig. 76): " Co-type " [white, printed], [Horaeomorphus / macrostictus / Lea / det. H. Franz " [white, handwritten and printed], " S. Aust. Museum / specimen " [orange, printed], " PHAGONOPHANA / macrosticta Lea, 1910 / P. Jałoszyński, 2014 / LECTOTYPUS " [white, printed], " SCYDMAENOZILA / macrosticta (Lea, 1910) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Paralectotype: ♀: seven labels (Fig. 77): " 14300 / Phagonophana / macrosticta / Tasmania / TYPE " [white, handwritten, " TYPE " in red ink], " macrosticta / Lea TYPE / Marrawah " [white, handwritten], " Horaeomorphus / macrostictus / Lea / det. H. Franz " [white, handwritten and printed], " SAMA Database / No. 25 - 037062 " [white, printed], " PHAGONOPHANA / macrosticta Lea, 1910 / P. Jałoszyński, 2014 / PARALECTOTYPUS " [white, printed], " SCYDMAENOZILA / macrosticta (Lea, 1910) / P. Jałoszyński, 2014 " [white, printed] (SAM). Additional material (38 exx.). 1 ♂, 1 ♀ (holotype and paratype of Horaeomorphus australiensis): S Australia, Bakewell (NHM); 1 ex., Australian Capital Territory, Blundells Creek, 3 km E of Piccadilly Circus, 850 m, Feb. 1984, " flight intercept window / trough trap ", leg. Lawrence & Johnson; 8 exx., same data except for Jan. 1985, leg. Weir, Lawrence, Johnson; 3 exx., same data, except for Mar. 1985; 5 exx., same data except for Apr. 1985; 1 ex., same data, except for May 1985; 2 exx., same data, except for Sep. 1985; 1 ex., same data, except for Oct. 1985; 1 ex., same data except for 6 km NE of Piccadilly Circus, 750 m, Feb. 1985; 2 exx., Blundells Creek, 3 km E of Piccadilly Circus, 850 m, 7 Sept. 1986, " Berlesate, ANIC 1066, leaf litter ", leg. J. F. Lawrence; Black Mountain near Canberra, " near light trap ", flight intercept trap, 9 - 30 Sept. 1996, leg. T. Weir & W. Dressler; 1 ex., Tasmania, Cradle Mountain, leg. J. Armstrong; 1 ex., Tasmania, 7 km SWW Derwent Bridge, pantrap, 6 Jan. - 2 Feb. 1983, leg. I. Naumann & J. Cardale; 1 ex., Tasmania, Hobart, Mt. Nelson, dry sclerophyll, under fallen branch, 2 - 5 Feb. 1992, leg. C. Reid; 2 exx., Tasmania, Anson's Bay Road, pitfall trap, " Leptospermum, in wet heath ", 20 - 25. ii. 1993, leg. J. Trueman et al.; 2 exx., same data except for Peters Link Road, dry sclerophyll; 1 ex., same data, except for 22 - 27. v. 1993; 2 exx., same data, except for Eddystone Point, 20 - 25. ii. 1993, coastal heath; 1 ex. same data, except for 22 - 27. v. 1993 (specimens in ANIC and cPJ). Revised diagnosis. Male and female: elytra with variously distinct pattern: base of elytra and subtriangular subapical spots dark brown, remaining surface light brown. Male: aedeagus slender, in ventral view with broadly rounded apex; apical part of median lobe abruptly demarcated.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCC18158FFE3A8B0760FB85.taxon	description	Redescription. Body of male (Fig. 72) strongly convex, elongate and slender, with long appendages, BL 2.58 – 2.95 mm (mean 2.80 mm); cuticle glossy, head, pronotum and appendages light to dark brown, each elytron with dark suture, base and subtriangular subapical spot, remaining surface lighter (in some cases indistinctly but the pattern still discernible), vestiture dark brown, on elytra slightly darker than on remaining body parts. Head (Figs. 57, 72) broadest at eyes, HL 0.50 – 0.58 mm (mean 0.51 mm), HW 0.50 – 0.60 mm (mean 0.56 mm); tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (Fig. 72) slender, AnL 1.48 – 1.73 mm (mean 1.51 mm); antennomeres I – VII distinctly elongate, VIII slightly longer than wide, IX – X each about as long as broad, XI as long or nearly as long as IX – X together, 2.5 x as long as broad, pointed. Pronotum (Figs. 64, 72) in dorsal view strongly elongate, broadest near anterior third, PL 0.70 – 0.85 mm, (mean 0.79 mm), PW 0.55 – 0.63 mm (mean 0.59 mm); anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae. Elytra (Figs. 65, 72) regularly oval, broadest near middle, EL 1.38 – 1.60 mm (mean 1.49 mm), EW 0.90 – 1.13 mm (mean 1.04 mm), EI 1.33 – 1.53 (mean 1.44); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae long, curved sparse and erect, in lightly pigmented specimens setae on light elytral spots contrastingly dark. Hind wings well developed, about twice as long as elytra. Legs (Figs. 66, 72) long and slender, without modifications. Aedeagus (Figs. 70, 80 – 82, 89 – 90) elongate and relatively slender; AeL 0.58 mm; in ventral view short apical part abruptly demarcated, with broadly and weakly rounded apical margin; in lateral view apical part subtriangular and sharply demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with indistinctly broadened apices. Aedeagus erected (Fig. 82) with all components of internal sac extruded and still easily recognizable submedian darkly sclerotized subtriangular central complex. Female. Similar to male, on average slightly larger, but body lengths and proportions of body parts overlap with those measured in males; distinguished by antennomere VIII about as long as broad and antennomeres IX – X slightly transverse. BL 2.73 – 2.98 mm (mean 2.86 mm); HL 0.50 – 0.58 mm (mean 0.52 mm), HW 0.55 – 0.63 mm (mean 0.58 mm), AnL 1.40 – 1.75 mm (mean 1.50 mm); PL 0.75 – 0.83 mm (mean 0.80 mm), PW 0.55 – 0.63 mm (mean 0.60 mm); EL 1.43 – 1.65 mm (mean 1.54 mm), EW 1.00 – 1.15 mm (mean 1.07 mm), EI 1.33 – 1.53 (mean 1.44).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCC18158FFE3A8B0760FB85.taxon	distribution	Distribution (Fig. 195 a). South-eastern Australia: South Australia, Australian Capital Territory and Tasmania.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFCC18158FFE3A8B0760FB85.taxon	discussion	Remarks. Franz (1975) studied two specimens from SAM, and he treated one of them as a holotype (without specifying the sex), the other one as a male co-type. Lea (1910) clearly stated that two specimens were available to him: " One specimen was taken from an ants' nest under a stone, a second under a near by stone but not associated with ants ". These two specimens have a status of syntypes, and Franz's (1975) redescription does not contain any data that fulfill the requirements of the formal lectotype designation. The two specimens presumably were originally mounted on one card (a frequent case in the Lea Collection) and separated in a later time. The " holotype " mentioned by Franz as a specimen bearing the original set of labels is a female, and the " co-type " without original labels is a male, distinctly smaller than the female, and with the aedeagus mounted by Franz in euparal (remounted in Canada balsam during the present study) and illustrated (Franz 1975; Fig. 125). The aedeagus is erected and distorted (Fig. 82), but internal structures are highly similar to those found in all additional males from the Australian Capital Territory and Tasmania, and also in the holotype of Horaeomorphus australiensis. The 25 specimens from Blundells Creek and 11 from various localities in Tasmania show a considerable variation in the body length and pigmentation, ranging from small (BL 2.63 mm) to large (BL 2.98 mm) individuals and from sharply contrasting light and dark elytral pattern to only indistinct (but still discernible) spots with diffused margins. The aedeagus of the holotype of H. australiensis does not differ from those found in all males from Blundells Creek and Tasmania, and the specimen's external morphology falls within the range of interindividual variability of H. macrostictus. Horaeomorphus australiensis is therefore placed as a junior synonym of Scydmaenozila macrosticta (= Horaeomorphus macrostictus). The male syntype of Phagonophana macrosticta that does not bear the original labels (Fig. 76) is here designated as a lectotype to provide a name-bearing type for this name. As mentioned above, this species is relatively variable in the body length, proportions of body parts and the elytral pattern. Specimens from Tasmania and Australia are similarly variable and do not differ in the range of variability. However, since these populations are isolated, an analysis of larger samples may reveal some subtle differences. The elytral pattern, although variably distinct, is observable even in the most darkened specimens, in which the contrast between lighter and darker spots is low. In all remaining species of Scydmaenozila no traces of elytral pattern can be seen and the elytra are uniformly brown. Two specimens from NHM (the type series of H. australiensis) bear labels with " S Australia, Bakewell ". There is Bakewell Bore in South Australia (over 630 km NW of Adelaide), but Bakewell on the labels is most likely a person name (Adam Ślipiński, in an email to the author dated 06.03.2014). The exact collecting place of Scydmaenozila macrosticta in South Australia remains unknown, but judging from the distribution in the Australian Capital Territory and in Tasmania (where it is known from scattered localities from nearly entire island), this species may occur in SE part of South Australia and possibly also in Victoria.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF7182B8FFE3A8B0035FD59.taxon	materials_examined	Type material studied. Lectotype (here designated): AUSTRALIA: ♂: seven labels (Figs. 78 – 79): " latipennis / Lea TYPE / Bridgetown " [thick mounting card on which originally four syntypes were mounted; white, handwritten]; " latipennis / Lea, Type / Bridgetown " [white, handwritten and printed]; " 13794 / Phagonophana / latipennis Lea / W. Australia / TYPE " [white, handwritten, " TYPE " in red ink], " SAMA Database / No. 25 - 037011 " [white, printed], " SAMA Digital Image / 14. 8. 2013 " [green, printed and handwritten], " PHAGONOPHANA / latipennis Lea, 1910 / det. P. Jałoszyński, 2014 / LECTOTYPUS " [white, printed], " SCYDMAENOZILA / latipennis (Lea, 1910) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Paralectotypes (7 exx.): 2 ♀♀ originally mounted on one card with the lectotype, remounted during present study and labeled with white, printed copies of the lectotype labels (SAM); 1 ♂, originally mounted on one card with the lectotype, remounted by Franz, three labels: " Bridgetown " [white, handwritten], " Horaeomorphus / latipennis / Lea / det. H. Franz " [white, handwritten and printed], " S. Aust. Museum / specimen " [orange, printed] (SAM); 1 ♀, four labels: " Australia / 1910 – 172. " [white, printed, first line underlined in red], " Co-type " [circular, white with yellow margin, printed], " Phagonophana / latipennis Lea / W. Aust. / Co-type Lea " [white, handwritten and printed], " ♀ " [white, printed] (NHM); 1 ♂, 1 ♀ (mounted on separate cards but on one pin): six labels: " Bridgetown / W. A. Lea " [white, printed], " Phagonophana / latipennis Lea / W. Australia / Co-type " [white, handwritten, " Co-type " in red ink], " On permanent loan from / MACLEAY MUSEUM / University of Sydney " [white, printed], " PARATYPE " [blue, printed], " ANIC Database No. / 25 062435 " [white, printed], " Aust. Nat. / Ins. Coll. " [green, printed] (ANIC); 1 ♀, seven labels: " WESTERN AUSTRALIA " [white, handwritten], " E. W. Ferguson / Collection " [white, printed], " Phagonophana / latipennis Lea / W. Australia / Co-type " [white, handwritten, " Co-type " in red ink], " PARATYPE " [blue, printed], " PARATYPE " [grey, printed], " ANIC Database No. / 25 062434 " [white, printed], " ANIC / specimen " [green, printed] (ANIC). All paralectotypes additionally with two white, printed labels: " PHAGONOPHANA / latipennis Lea, 1910 / det. P. Jałoszyński, 2014 / PARALECTOTYPUS ", and " SCYDMAENOZILA / latipennis (Lea, 1910) / det. P. Jałoszyński, 2014 ". Additional material (4 exx.). 2 ♂♂, Western Australia, Millbrook, W Millstream Road - Millbrook Nature Reserve, 26. iv. 2001, " JVAP # 31 pitfall " (ANIC, cPJ); 1 ♂, 1 ♀, Western Australia, side track off Pemberton Northcliffe Road, sifted litter, 11. x. 2010, 112 m, leg. K. Will (ANIC, cPJ). Revised diagnosis. Male and female: body uniformly brown. Male: aedeagus stout, in ventral view with distinctly emarginate apex.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF7182B8FFE3A8B0035FD59.taxon	description	Redescription. Body of male (Fig. 73) strongly convex, elongate and slender, with long appendages, BL 2.80 – 2.95 mm (mean 2.87 mm); cuticle glossy, uniformly dark brown with slightly reddish hue, vestiture reddishbrown. Head (Fig. 73) broadest at eyes, HL 0.48 – 0.50 mm (mean 0.50 mm), HW 0.55 – 0.60 mm (mean 0.58 mm); tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (Fig. 73) slender, AnL 1.75 – 1.90 mm (mean 1.83 mm); all antennomeres distinctly elongate, XI slightly shorter than IX – X together, 2.5 x as long as broad, pointed. Pronotum (Fig. 73) in dorsal view strongly elongate, broadest near anterior third, PL 0.78 – 0.85 mm, (mean 0.80 mm), PW 0.58 – 0.63 mm (mean 0.60 mm); anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae. Elytra (Fig. 73) slightly rhomboidal, broadest in or slightly behind middle, EL 1.50 – 1.63 mm (mean 1.57 mm), EW 1.13 – 1.23 mm (mean 1.17 mm), EI 1.30 – 1.40 (mean 1.35); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 73) long and slender, without modifications. Aedeagus (Figs. 83 – 84, 91 – 92) elongate but relatively stout; AeL 0.53 mm; in ventral view apex not demarcated, with distinctly emarginate apical margin; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, apices barely noticeably broadened. Female. Similar to male, on average slightly larger, but body lengths and proportions of body parts overlap with those measured in males, distinguished by antennomere VIII about as long as broad and antennomeres IX – X slightly transverse. BL 2.85 – 3.05 mm (mean 2.95 mm); HL 0.48 – 0.50 mm (mean 0.50 mm), HW 0.58 – 0.60 mm (mean 0.58 mm), AnL 1.58 – 1.75 mm (mean 1.69 mm); PL 0.78 – 0.85 mm (mean 0.81 mm), PW 0.58 – 0.63 mm (mean 0.60 mm); EL 1.55 – 1.70 mm (mean 1.64 mm), EW 1.13 – 1.23 mm (mean 1.18 mm), EI 1.38 – 1.42 (mean 1.39).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF7182B8FFE3A8B0035FD59.taxon	distribution	Distribution (Fig. 195 b). South-western Australia: Western Australia.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF7182B8FFE3A8B0035FD59.taxon	discussion	Remarks. Franz (1975) studied four specimens originally mounted on a single card, on which one male was annotated as " TY " (Fig. 78). Franz treated this male as a holotype and remaining specimens as paratypes, but studied the aedeagus only in a single remounted " paratype " male (Franz 1975). In fact, Lea (1910) clearly stated that he collected " numerous specimens obtained under a log on sandy ground in the company of ants ", and in his description none of them is treated as a type. Therefore, four specimens preserved at SAM, one at NHM and three at ANIC, all identifiable as coming from the original type series, are syntypes. The male originally mounted together with three other specimens on one card and annotated as " TY " is here designated as lectotype, to provide a name-bearing type for Phagonophana latipennis. Three more specimens labeled as Phagonophana latipennis with the same hand but not belonging to the type series (collected in different localities) (ANIC) belong to other species: two are designated here as the holotype and paratype of Scydmaenozila inexpectata sp. n. (from the Ferguson Collection), and one is a Syndicus (from the Ropes Collection). Scydmaenozila latipennis is similar to Scydmaenozila mallacootana sp. n. and Scydmaenozila inexpectata sp. n.; it clearly differs from these species in the broader elytra in relation to the pronotum (PW / EW 0.48 – 0.56 vs. 0.60 vs. 0.58 – 0.61, respectively), and in the structures of the aedeagus.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA / Victoria / Mallacoota N. P. / 26 May 1978 / S. & J. Peck " [white, printed], " mushroom / litter' [white, printed], " Aust. Nat. / Ins. Coll. " [green, printed], " SCYDMAENOZILA / mallacootana m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	diagnosis	Diagnosis. Male and female: body uniformly brown. Male: aedeagus slender, in ventral view with broadly rounded apex; apical part of median lobe abruptly demarcated.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	description	Description. Body of male (Fig. 74) strongly convex, elongate and slender, with long appendages, BL 2.75 mm; cuticle glossy, dark brown with reddish hue, elytra slightly lighter than head and pronotum, vestiture dark brown. Head (Fig. 74) broadest at eyes, HL 0.50 mm, HW 0.60 mm; tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (Fig. 74) slender, AnL 1.65; antennomeres I – VII distinctly elongate, VIII slightly longer than wide, IX – X each about as long as broad, XI as long as IX – X together, 2.5 x as long as broad, pointed. Pronotum (Fig. 74) in dorsal view strongly elongate, broadest near anterior third, PL 0.80 mm, PW 0.63 mm; anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae. Elytra (Fig. 74) slightly rhomboidal, broadest near middle, EL 1.45 mm, EW 1.05 mm, EI 1.38; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 74) long and slender, without modifications. Aedeagus (Figs. 85 – 86, 93 – 94) elongate and relatively slender; AeL 0.63 mm; in ventral view short apical part abruptly demarcated, with broadly and weakly rounded apical margin; in lateral view apical part subtriangular and sharply demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with distinctly broadened apices. Female. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	distribution	Distribution (Fig. 195 c). South-eastern Australia: Victoria.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	etymology	Etymology. Locotypical, after the Mallacoota National Park, where this species occurs.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182C8FFE3A8B0664FA73.taxon	discussion	Remarks. Scydmaenozila mallacootana has the aedeagus very similar to that of Scydmaenozila macrosticta; differences can be seen only in the arrangement of internal structures. However, externally these species are distinctly different: Scydmaenozila macrosticta has the bicolorous pattern on elytra, while Scydmaenozila mallacootana is uniformly brown; also the shape of the pronotum and elytra is different (Fig. 72 vs. 74).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: five labels: " Blue Mts. N. S. Wales " [white, printed], " F. W. Ferguson / Collection " [white, printed], " Phagonophana / latipennis Lea / N. S. Wales " [white, handwritten], " Aust. Nat. / Ins. Coll. " [green, printed], " SCYDMAENOZILA / inexpectata m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC). Paratype: 1 ♀, originally mounted on one card together with the holotype, labeled with white, printed copies of original labels, and with standard yellow " PARATYPUS " label (ANIC).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	diagnosis	Diagnosis. Male and female: body uniformly brown. Male: aedeagus slender, in ventral view with rounded and subtriangular apex; apical part of median lobe not demarcated.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	description	Description. Body of male (Fig. 75) strongly convex, elongate and slender, with long appendages, BL 2.98 mm; cuticle glossy, uniformly reddish-brown, vestiture slightly lighter than cuticle. Head (Fig. 75) broadest at eyes, HL 0.50 mm, HW 0.60 mm; tempora longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes large and strongly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine, separated by spaces 1.5 – 2 x as wide as diameters of punctures; setae on frons and anterior portion of vertex short, sparse and suberect, tempora and posterior portion of vertex densely covered with long bristles directed posteriorly and posterolaterally. Antennae (Fig. 75) slender, AnL 1.78; antennomeres I – VIII distinctly elongate, IX – X each about as long as broad, XI slightly shorter than IX – X together, slightly less than twice as long as broad, pointed. Pronotum (Fig. 75) in dorsal view strongly elongate, broadest near anterior third, PL 0.88 mm, PW 68 mm; anterior and lateral margins in anterior half rounded; sides slightly concave in posterior third; posterior margin distinctly bisinuate; base of pronotum with two pairs of large and deep pits, additionally with a pair of shallow but distinct impressions located dorsolaterally. Punctures on pronotal disc fine and inconspicuous; entire dorsal and lateral surface of pronotum covered with moderately dense bristles, dorsally with admixture of thin setae. Elytra (Fig. 75) regularly oval, broadest near middle, EL 1.60 mm, EW 1.10 mm, EI 1.45; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct than those on pronotum, separated by spaces 2 – 3 x as wide as diameters of punctures; setae long, curved, sparse and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 75) long and slender, without modifications. Aedeagus (Figs. 87 – 88, 95 – 96) elongate and slender; AeL 0.60 mm; in ventral view apex not demarcated and subtriangular, with narrowly rounded tip; in lateral view apical part subtriangular and not demarcated from ventral wall; internal armature complicated, with submedian complex of elongate and darkly sclerotized structures and subapical finely denticulate lateral lobes of internal sac; parameres long and slender, with strongly broadened apices. Female. Slightly larger than male and with shorter antennae, but does not differ in proportions of antennomeres. BL 3.18; HL 0.53 mm, HW 0.63 mm, AnL 1.65 mm; PL 0.95 mm, PW 0.73 mm; EL 1.70 mm, EW 1.25 mm, EI 1.36.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	distribution	Distribution (Fig. 195 d). South-eastern Australia: New South Wales.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	etymology	Etymology. The epithet inexpectata refers to the surprising finding of this new species among specimens identified previously by Lea as Phagonophana latipennis.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF3182D8FFE3C890677F931.taxon	discussion	Remarks. Scydmaenozila inexpectata, although externally similar to the remaining uniformly brown species, differs clearly in the aedeagus, which has the apex subtriangular, and not emarginated or broadly rounded.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF218228FFE3F4301B4FE63.taxon	materials_examined	Type species: Horaeomorphus puncticeps Franz, 1975: 142 (here designated).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF218228FFE3F4301B4FE63.taxon	diagnosis	Diagnosis. Body (Figs. 112, 114 – 116) elongate and flattened; head short and convex; vertex not projecting dorsocaudad, its posterior margin slightly to distinctly concave; supraantennal tubercles distinct but weakly raised and not accompanied posteriorly by pits; frontoclypeal groove present; eyes located in anterior part of head; tempora long and strongly curved toward occipital constriction; tempora, genae and postgenae with dense bristles; ' neck region' distinctly demarcated from anterior part of head capsule by constriction and as narrow as nearly half width of vertex; hypostomal ridges (Fig. 97; hr) sharply marked and long, reaching nearly to posterior tentorial pits but not connected in middle; posterior tentorial pits (Fig. 97; ptp) elongate and located distinctly in front of transverse groove demarcating ' neck' ventrally; submentum (Fig. 97; smn) without lateral sutures; each mandible with robust subapical tooth (Fig. 100; sat) and densely setose dorsomesal prostheca (Fig. 100; pst); maxillary palpomeres III and IV (Fig. 101; mxt 3 – 4) slender, strongly elongate; pronotum with one pair of small internal antebasal pits (Fig. 103; ip) which are separated by narrow longitudinal wrinkle or carina (Fig. 103; mc), external pair of diffused impressions (Fig. 103; li) and distinct sublateral carinae (Fig. 103; slc) extending from posterior pronotal corners to 1 / 5 – 1 / 4 PL; prosternum laterally demarcated from hypomera by pronotosternal sutures (Fig. 102; nss), with indistinct intercoxal carina; procoxal sockets (Fig. 102; pcs) closed by posterolateral lobes of prosternum; prothoracic hypomera (Fig. 102; hy) with large internal (adcoxal) parts demarcated from sides of pronotum by complete hypomeral ridges (Fig. 102; hyr); sides of pronotum with dense thick bristles (Fig. 102) well visible in dorsal view; mesoventrite with short anterior ridge (Fig. 105; ar) medially connected with mesoventral process (Fig. 105; msvp) which is narrow and weakly expanding ventrally and reaching only to middle of mesocoxal cavities, so that mesocoxae are indistinctly separated; ventrolateral foveae (Fig. 110; vlf) present, dorsolateral foveae absent; anterior metaventral process (Fig. 106; amvp) present; metaventral intercoxal process (Fig. 111; mtvp) broadly subtriangular, with small median notch; metacoxae (Fig. 109; cx 3) narrowly separated in middle; each elytron with two deep and relatively large asetose basal foveae (Fig. 104; bef) not connected by groove and well visible in dry-mounted specimens; metafemora in males with ventral projection (Figs. 117 – 120); aedeagus (Figs. 121 – 128) with free and slender parameres.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF218228FFE3F4301B4FE63.taxon	description	Description. Body (Figs. 112, 114 – 116) moderately large (BL 1.78 – 2.45 mm), flattened and slender, with long appendages, pigmentation brown, cuticle setose. Head (Figs. 97, 112, 114 – 116) with occipital constriction (Fig. 97; occ) much narrower than vertex and dividing the head capsule into exposed anterior part and narrow posterior ' neck region' retracted into prothorax; eyes small and slightly oval, located in anterior part of head; tempora long and in posterior half strongly curved posteromesally, with sparse bristles; vertex transverse, uniformly convex, not projecting dorso-caudad, with posterior margin slightly concave and distinctly demarcated from occiput; frons confluent with vertex, transverse and subtrapezoidal, convex, strongly but not abruptly lowering toward labrum; frontoclypeal groove present; antennal insertions broadly separated, located beneath distinct but weakly raised supraantennal tubercles. Labrum (Fig. 99) transverse and semioval with slightly emarginate anterior margin, with small number of long dorsal setae. Mandibles symmetrical, subtriangular, each with broad base and large subapical tooth (Fig. 100; sat), also distal mesal margin of basal part forms broad and blunt tooth in front of densely setose dorsomesal prostheca reaching mandibular base. Each maxilla composed of small cardo (Fig. 101; cd) bearing three setae; subtriangular basistipes (Fig. 101; bst); elongate mediostipes (Fig. 101; mst); elongate galea (Fig. 101; gal) and lacinia (Fig. 101; lac); large, elongate palpifer (Fig. 101; pfp); and strongly elongate and moderately large maxillary palpus (Fig. 97; mxp) composed of elongate palpomere I (Fig. 101; mxp 1), strongly elongate, slightly clavate and slender palpomere II (Fig. 101; mxp 2), large and strongly elongate, slender palpomere III (Fig. 101; mxp 3) broadest near distal third, and small, slender, subconical and pointed palpomere IV (Fig. 101; mxp 4). Labium (Figs. 97 – 98) with large submentum (Fig. 97; smn) not demarcated laterally from postcardinal parts of hypostomae; subtrapezoidal mentum (Fig. 97; mn); and short prementum bearing narrowly separated at bases long 3 - segmented labial palpi and a pair of median bristles inserted on small subtrapezoidal ligula (Fig. 98; lg). Hypostomal ridges (Fig. 97; hr) sharply marked, long and extending nearly to posterior tentorial pits but not connecting in middle. Gular plate (Fig. 97; gp) large and subtrapezoidal, with rapidly narrowed anterior part; gular sutures (Fig. 97; gs) superficial; posterior tentorial pits (Fig. 97; ptp) narrow and straight, located clearly in front of transverse groove ventrally demarcating ' neck region'. Antennae (Figs. 112, 114 – 116) long and slender, gradually but only slightly thickening distally; antennomere XI unremarkable. Prothorax (Figs. 102, 112, 114 – 116) in dorsal view weakly to strongly elongate and broadest near anterior third, with rounded anterior margin and anterior parts of lateral margins, with barely marked, blunt and strongly obtuse anterior corners; posterior corners distinct, obtuse; posterior margin shallowly bisinuate. Base of pronotum with distinct sublateral carinae (Fig. 103; slc) extending from posterior pronotal corners and as long as 1 / 5 – 1 / 4 of PL; base with one pair of small internal pits (Fig. 103; ip) in middle separated by narrow longitudinal wrinkle or carina (Fig. 103; mc), and with variously distinct lateral impressions (Fig. 103; li) adjacent to sublateral carinae. Sides of pronotum with thick and sparse bristles (Fig. 103). Prosternum with long basisternal part (Fig. 102; bst) distinctly demarcated from procoxal cavities (Fig. 102; pcc); median part of sternum slightly raised and forming narrow and indistinct prosternal carina; procoxal sockets (Fig. 102; pcs) closed by posterolateral lobes of prosternum; hypomera (Fig. 102; hy) elongate, each divided into broad lateral part confluent with pronotum and narrower internal (adcoxal) part; hypomeral ridges (Fig. 102; hyr) complete; pronotosternal sutures (Fig. 102; nss) entire. Mesocutellum (Fig. 104; scl 2) subtriangular, in intact specimen hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture (Fig. 104; sss) present. Mesoventrite (Fig. 105) with narrow and distinctly demarcated anterior ridge (Fig. 105; ar); mesoventral process (Fig. 105; msvp) carinate and moderately expanding ventrally, anteriorly connected with anterior ridge and terminated near middle of mesocoxal cavities, indistinctly separating mesocoxae; sides of mesoventrite with shallow procoxal rests (Fig. 105; pcr) filled with bristles (= setose impressions) but without asetose impressions; mesanepisternum with moderately long prepectus (Fig. 105; pre), sides of mesoventrite with moderately broad and deep setose ventrolateral foveae (Figs. 108, 110; vlf); dorsolateral foveae absent; mesocoxal projections (Figs. 105, 107, 109 – 110; mcp) prominent, with mesocoxal sockets (Figs. 105, 107, 109 – 110) located on their mesoventral surface and visible in ventral view. Metaventrite (Figs. 109 – 110; v 3) subrectangular, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate and with broadly subtriangular metaventral intercoxal process (Fig. 111; mtvp) bearing small median notch; anterior metaventral process present (Fig. 106; amvp). Metanepisterna and metepimera narrow. Metafurca with long stalk and divergent lateral furcal arms (Fig. 110; lmfa). Elytra (Figs. 104, 112, 114 – 116) oval, each with two moderately large, deep and asetose basal foveae (Fig. 104; bef) not connected by groove; humeral denticles and sub-humeral lines absent. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 112, 114 – 116) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora moderately strongly clavate; tibiae and tarsi long and slender. Males of all species have variously shaped projections on ventral margin of metafemora (Figs. 117 – 120). Aedeagus (Figs. 121 – 128) elongate and moderately darkly sclerotized, relatively thin-walled, with symmetrical median lobe and symmetrical, complicated assemblage of internal sclerites; parameres free and slender, with apical setae.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF218228FFE3F4301B4FE63.taxon	etymology	Etymology. Spinosciacharis is a combination of the stem Sciacharis (a genus name) with the prefix spino - referring to the femoral spines found in males of all species. Gender feminine. Composition and distribution (Fig. 196). Three species of Spinosciacharis are distributed in the southeastern part of the Australian continent (New South Wales and Victoria) and one in Tasmania.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF218228FFE3F4301B4FE63.taxon	discussion	Remarks. Among all Australian genera of Cyrtoscydmini, Spinosciacharis is most similar to Horaeomorphus in the general body form, but to Sciacharis s. str. in ventral structures. The flattened and slender body with weakly rounded sides of elytra, the shape of pronotum, and the femoral projections in males may be mistaken with similar features of some Horaeomorphus species. Spinosciacharis clearly differs from Horaeomorphus in numerous characters: the absence of pits on posteromesal margins of weakly marked supraantennal tubercles (in Horaeomorphus pits present and tubercles prominent); posterior tentorial pits located far in front of the anterior margin of ' neck region' (in Horaeomorphus pits are adjacent to the anterior margin of ' neck region'); much narrower occipital constriction; absence of median antebasal pronotal pit (present in Horaeomorphus); presence of sublateral pronotal carinae (absent in Horaeomorphus); narrow, ridge-like prosternal intercoxal carina (carina narrow but subrectangular with distinct lateral margins in Horaeomorphus); basal elytral foveae not connected by inversely U-shaped groove (in Horaeomorphus groove present); narrow anterior ridge of mesoventrite (broad in Horaeomorphus); presence of ventrolateral mesothoracic foveae (absent in Horaeomorphus); mesocoxae indistinctly separated by narrow mesoventral process only in their anterior half (distinctly separated in Horaeomorphus); presence of anterior metaventral process (absent in Horaeomorphus); and metaventral intercoxal process without spines (with a pair of long spines in Horaeomorphus). Spinosciacharis is similar to Sciacharis s. str. in the general body shape (especially the shape of the head), all ventral characters of the head; the presence of pronotal internal antebasal pits and sublateral carinae; feebly developed prosternal intercoxal carina; presence of two distinct basal elytral foveae; presence of only ventrolateral mesothoracic fovea; and the shape of metaventral intercoxal process. Spinosciacharis differs from Sciacharis s. str. in robust subtriangular mandibles with two subapical teeth (one small subapical tooth beneath narrow apical tooth and above strongly broadened mandibular base in Sciacharis s. str.); sublateral pronotal carinae extending much farther anteriorly; prosternum completely demarcated from adcoxal parts of hypomera (in Sciacharis s. str. prosternum partly fused with hypomera); the mesoventral intercoxal process not interrupted (in Sciacharis s. str. interrupted near anterior margins of mesocoxal cavities); and the presence of anterior metaventral process (absent in Sciacharis s. str.). The same characters, except for the prosternum completely separated vs. partly fused with hypomera differentiate Spinosciacharis from Sciacharis (Maorinus). The latter taxon has an elongated head, distinctly thickened maxillary palpi, hypomeral ridges not reaching posterior margins of adcoxal parts of hypomera; and long mesoventral intercoxal process extending from just behind anterior ridge of mesoventrite up to posterior margins of mesocoxal cavities (all these characters different in Spinosciacharis). The similarities between Spinosciacharis and Sciacharis may in future justify placing the former as a subgenus of Sciacharis.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFC18238FFE3BE807C9FBF6.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: six labels (Fig. 113): " Dorrigo / N. S. Wales " [white, printed], " Horaeomorphus / puncticeps m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " SAMA Database / No. 25 - 037012 " [white, handwritten], " SAMA Digital Image / 14. 8. 2013 " [green, printed and handwritten], " Spinosciacharis / puncticeps (Franz, 1975) / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (SAM). Revised diagnosis. Male: BL nearly 2.5 mm; elytra rhomboidal; EI <1.4; metafemur with straight and pointed ventral projection, without distal ventral lobe. Female and its diagnostic characters unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFC18238FFE3BE807C9FBF6.taxon	description	Redescription. Body of male (Fig. 112) flattened, elongate and slender, with long appendages, BL 2.45 mm; cuticle glossy, body dark brown with reddish hue, appendages slightly lighter, vestiture yellowish. Head (Fig. 112) rounded, broadest just behind eyes, HL 0.48 mm, HW 0.53 mm; tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum distinct, small but sharply marked, separated by spaces 1 – 1.5 x as wide as diameters of punctures; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae (Fig. 112) slender, AnL 1.10 mm; antennomeres I – VI elongate, VII – X about as long as broad; XI much shorter than IX – X together, 1.7 x as long as broad, pointed. Pronotum (Fig. 112) in dorsal view elongate, broadest near anterior third, PL 0.65 mm, PW 0.55 mm; anterior and lateral margins rounded; posterior margin distinctly bisinuate; base of pronotum with distinct sublateral carinae accompanied mesally by shallow impressions, median part of pronotal base with two small but distinct pits separated by narrow carina as long as 3 x diameter of pit and extending anteriorly and posteriorly beyond pits, posteriorly reaching posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous, less distinct than those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles. Elytra (Fig. 112) slightly rhomboidal, broadest slightly anterior to middle, EL 1.33 mm, EW 0.95 mm, EI 1.39; basal impressions short but deep; circumsutural area in anterior fourth slightly more convex than surrounding cuticle and bordered posteriorly by indistinct short impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum, less distinct than those on frons and vertex; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra. Legs (Figs. 112, 117) long and slender, metafemur (Fig. 117) with long, pointed and straight ventral projection, without ventral distal lobe. Aedeagus (Figs. 121 – 122) elongate, AeL 0.40 mm, in ventral view sides of median lobe concave in middle, apical portion subtrapezoidal; internal armature lightly sclerotized, with two pairs of subapical microdenticulate structures; parameres strongly broadening distally and their apices far exceeding apex of median lobe. Female. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFC18238FFE3BE807C9FBF6.taxon	distribution	Distribution (Fig. 196 a). South-eastern Australia: New South Wales.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFC18238FFE3BE807C9FBF6.taxon	discussion	Remarks. Spinosciacharis puncticeps is most conspicuous species of this genus; its large and dark body with broad elytra and unique metafemoral modifications allow for unambiguous identification.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: three labels: " 42.08 S 147.54 E TAS / Big Sassy Creek, 400 m / 17 May 1989 Site 1 / rotten log, Tube 252 / J. Diggle " [white, printed], " ANIC / Specimen " [green, printed], " Spinosciacharis / sassy m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC). Paratypes: 2 ♀♀, same data as for the holotype, with standard yellow " PARATYPUS " label (ANIC, cPJ).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	diagnosis	Diagnosis. Male and female: BL 2.00 – 2.10 mm; elytra oval; EI> 1.5. Male: metafemur with slightly curved and pointed ventral projection, with distinct distal ventral lobe.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	description	Description. Body of male (Fig. 114) flattened, elongate and slender, with long appendages, BL 2.10 mm; cuticle glossy, body light brown, with slightly lighter appendages, vestiture yellowish. Head (Fig. 114) rounded, broadest just behind eyes, HL 0.38 mm, HW 0.50 mm; tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum inconspicuous, fine and sparse, not larger than those on pronotum; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae (Fig. 114) slender, AnL 1.00 mm; antennomeres I – VI distinctly elongate, VII only slightly longer than broad, VIII – X slightly transverse; XI much shorter than IX – X together, 1.8 x as long as broad, pointed. Pronotum (Fig. 114) in dorsal view elongate, broadest near anterior third, PL 0.55 mm, PW 0.50 mm; anterior and lateral margins rounded; posterior margin distinctly bisinuate; base of pronotum with distinct but short sublateral carinae accompanied mesally by relatively deep impressions, median portion of pronotal base with two small but deep pits separated by narrow carina as long as two diameters of pit and only posteriorly extending beyond pits to reach posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous, similar to those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles. Elytra (Fig. 114) slightly rhomboidal, broadest distinctly anterior to middle, EL 1.18 mm, EW 0.78 mm, EI 1.52; basal impressions short but deep; circumsutural area in anterior fourth distinctly more convex than surrounding surface and accentuated posteriorly by distinct transverse impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra. Legs (Figs. 114, 118) long and slender, metafemur (Fig. 118) with long, pointed and slightly curved ventral projection femur, with distinct ventral distal lobe. Aedeagus (Figs. 123 – 124) elongate, AeL 0.35 mm, in ventral view sides of median lobe rounded, apical margin broadly rounded; internal armature in the only known male partly everted, lightly sclerotized, with one pair of lateral apical microdenticulate structures and two sublateral lobes, each with distinct external projection, median part of everted armature occupied by long subtriangular projection protruding distally; parameres not broadening distally and their apices only slightly exceeding apex of median lobe. Female. Similar to male, differs in unmodified metafemora and more slender elytra; BL 2.00 – 2.08 mm; HL 0.40 mm, HW 0.48 mm, AnL 0.95 mm; PL 0.50 mm, PW 0.45 – 0.49 mm; EL 1.10 - 1.18 mm, EW 0.68 – 0.73 mm, EI 1.62 – 1.63.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	etymology	Etymology. Locotypical, after the Big Sassy Creek.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	distribution	Distribution (Fig. 196 b). South-eastern Australia: Tasmania.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFFA18278FFE394B01B6FE5C.taxon	discussion	Remarks. Spinosciacharis sassy has the metafemoral projections similar to those of Spinosciacharis puncticeps, but it is smaller and lightly pigmented. Moreover, among all species, Spinosciacharis sassy has the shortest and broadest pronotum in relation to the head.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA: NSW / Clyde Mt. 18 km / SE Braidwood / 500 m 28 May 78 / S. & J. Peck " [white, printed], " wet sclerophyll / ravines, log / litter and fungi " [white, printed], " Aust. Nat. / Ins. Coll. " [green, printed], " Spinosciacharis / clydensis m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	diagnosis	Diagnosis. Male: BL 1.90 mm; elytra oval; EI <1.5; metafemur with distinctly curved and broadly rounded ventral projection and with distinct distal ventral lobe. Female and its diagnostic characters unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	description	Description. Body (Fig. 115) flattened, elongate and slender, with long appendages, BL 1.90 mm; cuticle glossy, body light brown, with slightly lighter appendages, vestiture yellowish. Head (Fig. 115) rounded, broadest just behind eyes, HL 0.38 mm, HW 0.40 mm; tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum inconspicuous, fine and sparse, not larger than those on pronotum; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae (Fig. 115) slender, AnL 0.85 mm; antennomeres I – VI distinctly elongate, VII – VIII about as long as broad, IX strongly and X slightly transverse, XI much shorter than IX – X together, 1.6 x as long as broad, pointed. Pronotum (Fig. 115) in dorsal view elongate, broadest near anterior third, PL 0.50 mm, PW 0.45 mm; anterior and lateral margins rounded; posterior margin indistinctly bisinuate; base of pronotum with distinct but short sublateral carinae accompanied mesally by shallow impressions, median part of pronotal base with two small but deep pits separated by narrow wrinkle as long as pits and not extending beyond pits, but posterior pronotal margin with indistinct median protuberance, so that the median wrinkle between pits can be described as interrupted behind pits. Punctures on pronotal disc fine and inconspicuous, similar to those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles and in anterior half with several extremely long, erect setae directed laterally. Elytra (Fig. 115) slightly rhomboidal, broadest distinctly anterior to middle, EL 1.03 mm, EW 0.70 mm, EI 1.46; basal impressions short but deep; circumsutural area in anterior fourth barely noticeably more convex than surrounding surface and posteriorly accompanied by distinct and large but shallow impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra. Legs (Figs. 115, 119) long and slender, metafemur (Fig. 119) with long and distinctly curved ventral projection slightly broadening toward broadly rounded apex, with distinct ventral distal lobe. Aedeagus (Figs. 125 – 126) in the only known male with completely everted internal armature, AeL (without everted parts) 0.33 mm, in ventral view sides of median lobe slightly concave in subapical region, apical margin broadly rounded; internal armature lightly sclerotized, with two strongly elongate and subtriangular lateral projections protruding distally, in lateral view with a pair of dorsally-directed hook-like projections; parameres distinctly broadening distally and their apices only slightly exceeding apex of median lobe. Female. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	etymology	Etymology. Locotypical, after the Clyde Mountain.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	distribution	Distribution (Fig. 196 c). South-eastern Australia: New South Wales.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFF818388FFE38BA0716FE24.taxon	discussion	Remarks. Spinosciacharis clydensis resembles Spinosciacharis acheron sp. n. described below; both species are small and have a similar body silhouette and proportions of body parts, also both have several extremely long lateral setae on sides of pronotum. The shape of metafemoral projection is clearly different (Fig. 119 vs. 120): curved and slightly broadening distally in Spinosciacharis clydensis vs. recurved and narrowing distally in Spinosciacharis acheron. Moreover, the antennae in Spinosciacharis clydensis are stouter in their distal half, the antennomere VII is nearly as long as broad, while in Spinosciacharis acheron the antennomere VII is distinctly elongate; also the antebasal median longitudinal carina or wrinkle separating internal pair of pits in Spinosciacharis clydensis is interrupted behind pits, while in Spinosciacharis acheron it is continuous.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA / Victoria / Acheron Gap / 750 m nr. / Warburton / 28 - 30 Apr. 78 / S. & J. Peck " [white, printed], " Berlesate frass / under Nothophagus / logs " [white, printed], " Aust. Nat. / Ins. Coll. " [green, printed], " Spinosciacharis / acheron m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] (ANIC). Paratypes (2 ♂♂, 3 ♀♀): same data as for the holotype, all with standard yellow " PARATYPUS " label (ANIC, cPJ).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	diagnosis	Diagnosis. Male and female: BL 1.78 – 1.98 mm; elytra oval; EI 1.46 – 1.56. Male: metafemur with distinctly recurved and broadly rounded ventral projection, with distinct distal ventral lobe.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	description	Description. Body of male (Fig. 116) flattened, elongate and slender, with long appendages, BL 1.90 – 1.98 mm (mean 1.93 mm); cuticle glossy, body light brown, with slightly lighter appendages, vestiture yellowish. Head (Fig. 116) rounded, broadest just behind eyes, HL 0.38 – 0.40 mm (mean 0.38 mm), HW 0.40 – 0.43 mm (mean 0.41 mm); tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum inconspicuous, fine and sparse, not larger than those on pronotum; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae (Fig. 116) slender, AnL 0.90 – 0.93 mm (mean 0.92 mm); antennomeres I – VII distinctly elongate, VIII about as long as broad, IX-X distinctly transverse, XI much shorter than IX – X together, 1.6 x as long as broad, pointed. Pronotum (Fig. 116) in dorsal view elongate, broadest near anterior third, PL 0.50 mm, PW 0.40 – 0.45 mm (mean 0.42 mm); anterior and lateral margins rounded; posterior margin indistinctly bisinuate; base of pronotum with distinct but short sublateral carinae accompanied mesally by shallow impressions, median part of pronotal base with two small but deep pits separated by narrow carina twice as long as pits and extending beyond pits only posteriorly, up to posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous, similar to those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles and in anterior half with several extremely long, erect setae directed laterally. Elytra (Fig. 116) slightly rhomboidal, broadest distinctly anterior to middle, EL 1.03 – 1.08 mm, EW 0.68 – 0.73 mm, EI 1.48 – 1.56; basal impressions short but deep; circumsutural area in anterior fourth slightly more convex than surrounding surface and posteriorly accompanied by distinct, shallow and short impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra. Legs (Figs. 116, 120) long and slender, metafemur (Fig. 120) with long and distinctly recurved ventral projection slightly narrowing toward broadly rounded apex, with distinct ventral distal lobe. Aedeagus (Figs. 127 – 128) relatively stout, AeL 0.48 mm, in ventral view sides of median lobe rounded, apical margin subtriangular and broadly rounded; internal armature lightly sclerotized, composed of elongate paired sclerites; parameres not broadening distally and their apices only slightly exceeding apex of median lobe. Female. Similar to male, differs in unmodified metafemora; BL 1.78 – 1.95 mm (mean 1.87 mm); HL 0.38 mm, HW 0.38 – 0.43 mm (mean 0.40 mm), AnL 0.83 – 0.90 mm (mean 0.86 mm); PL 0.43 – 0.50 mm (mean 0.47 mm), PW 0.40 – 0.43 mm (mean 0.42 mm); EL 0.98 – 1.08 mm (mean 1.03 mm), EW 0.63 – 0.70 mm (mean 0.68 mm), EI 1.46 – 1.56 (mean 1.52).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	etymology	Etymology. Locotypical, after the Acheron Gap.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	distribution	Distribution (Fig. 196 d). South-eastern Australia: Victoria.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE718398FFE387003A3FD59.taxon	discussion	Remarks. This species is most similar to Spinosciacharis clydensis; see Remarks section under the latter species.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE6183E8FFE3BAB0063FA9A.taxon	materials_examined	Type species: Horaeomorphus thompsoni Franz, 1975: 145 (here designated).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE6183E8FFE3BAB0063FA9A.taxon	diagnosis	Diagnosis. Body (Figs. 160 – 163) stout to moderately elongate, moderately to strongly convex; head short and convex; vertex not projecting dorsocaudad, its posterior margin slightly convex, nearly straight or slightly concave; supraantennal tubercles from barely discernible to feebly raised, not accompanied posteromesally by pits; frontoclypeal groove absent; eyes located in anterior or median part of head; tempora long and strongly curved toward occipital constriction; tempora, genae and postgenae with moderately dense bristles; ' neck region' distinctly demarcated from anterior part of head capsule by constriction and much narrower than vertex; hypostomal ridges (Fig. 129; hr) sharply marked just behind cardines and variously long, extending at least to middle between anterior margin of submentum and posterior tentorial pits or farther up to posterior tentorial pits, not connected in middle; posterior tentorial pits (Fig. 129; ptp) elongate and located distinctly in front of transverse groove demarcating ' neck' ventrally; submentum (Fig. 129; smn) without lateral sutures; each mandible subtriangular, with variously developed subapical tooth, from rudimentary (Fig. 132, 151) to distinct (Fig. 142; sat) but always small, and with densely setose mesal prostheca (Fig. 132; pst); maxillary palpomeres III and IV (Fig. 133; mxp 3 – 4) slender, strongly elongate; pronotum with various antebasal sculpture (Figs. 135, 145, 154) composed of pits or / and transverse groove, without sublateral carinae and lateral edges; prosternum laterally demarcated from hypomera by pronotosternal sutures (Fig. 134; nss), in middle between procoxal cavities (Fig. 134; pcc) variously distinctly raised and forming low and elongate intercoxal carina; procoxal sockets (Fig. 134; pcs) closed by posterolateral lobes of prosternum; prothoracic hypomera (Fig. 134; hy) with large internal (adcoxal) parts demarcated from sides of pronotum by hypomeral ridges that can be nearly complete (Fig. 134; hyr) or indistinct in median (Fig. 153; hyr) or median and posterior portion (Fig. 146; hyr); sides of pronotum with variously dense thick bristles (Figs. 134, 146, 153) well visible in dorsal view; mesoventrite with short anterior ridge (Fig. 137; ar) medially connected with mesoventral process (Fig. 137; msvp) which is narrow and weakly expanding ventrally and reaching only to anterior margin of mesocoxal cavities, so that mesocoxae are not separated; ventrolateral mesothoracic foveae (Fig. 139; vlf) present, dorsolateral foveae absent; anterior metaventral process absent; metaventral intercoxal process (Fig. 140; mtvp) broadly subtriangular, without median notch; metacoxae (Fig. 138; cx 3) contiguous or nearly contiguous; each elytron (Fig. 136; bef) with two asetose rudiments of basal foveae not connected by groove and barely discernible or not discernible in dry-mounted specimens; aedeagus (Figs. 168 – 173) with free and slender parameres.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE6183E8FFE3BAB0063FA9A.taxon	description	Description. Body (Figs. 160 – 163) moderately large (BL 1.45 – 1.85 mm), relatively stout, with long appendages, pigmentation brown, cuticle setose. Head (Figs. 129, 141, 148) with occipital constriction (Figs. 129, 141, 148; occ) much narrower than vertex and dividing head capsule into exposed anterior part and narrow posterior ' neck region' retracted into prothorax; eyes located in anterior part of head or nearly in middle, oval or kidney-shaped, weakly protruding from the head silhouette; tempora long and strongly curved posteromesally, with sparse bristles; vertex transverse, uniformly convex, not projecting dorso-caudad, with posterior margin nearly straight or slightly concave; frons confluent with vertex, transverse and subtrapezoidal; frontoclypeal groove absent; antennal insertions broadly separated, located beneath distinct but weakly raised supraantennal tubercles. Labrum (Figs. 131, 142, 150) transverse, subtrapezoidal (Fig. 131), subrectangular (Fig. 142) or semioval (Fig. 150), with straight or slightly emarginate anterior margin, with short and dense lateral groups of anteroventral setae (Fig. 131), or long and dense anterior setae (Fig. 142), or six pairs of thick and pointed anteroventral sensilla (Fig. 150) projecting from beneath anterior margin, and with small number of long dorsal setae. Mandibles (Figs. 132, 143, 151) symmetrical, subtriangular, each with broad base and one variously distinct subapical tooth, from rudimentary (Figs. 132, 151) to small but distinct (Fig. 143); setose prostheca (Figs. 132, 143, 151; pst) present, mesal, not expanding onto dorsal surface of mandible and reaching mandibular base. Each maxilla composed of small cardo (Figs. 129, 141, 148; cd) bearing two setae; subtriangular basistipes (Figs. 133, 144, 152; bst); elongate mediostipes (Figs. 133, 144, 152; mst); elongate galea (Figs. 133, 144, 152; gal) and lacinia (Figs. 133, 144, 152; lac); large, elongate palpifer (Figs. 133, 144, 152; pfp); and strongly elongate and moderately large maxillary palp (Figs. 129, 141, 148; mxp) composed of elongate palpomere I (Figs. 129, 141, 148; mxp 1), strongly elongate, slightly clavate and slender palpomere II (Figs. 129, 141, 148; mxp 2), large and strongly elongate, slender palpomere III (Figs. 129, 141, 148; mxp 3) broadest near middle or in distal third, and small, slender, subconical and pointed palpomere IV (Figs. 129, 141, 148; mxp 4). Labium (Figs. 129 – 130, 141, 148 – 149) with large submentum (Figs. 129, 141, 148; smn) not demarcated laterally from postcardinal parts of hypostomae; subtrapezoidal mentum (Figs. 129, 141, 148 – 149; mn); and short prementum (Fig. 141; pmn) bearing narrowly separated at bases long 3 - segmented labial palps (Figs. 129, 141, 148; lp) and a pair of long lateral bristles inserted on small subrectangular or inversely subtriangular ligula (Figs. 130, 141, 149; lg); additionally ligula with 2 – 4 short setae in middle. Hypostomal ridges (Figs. 129, 141, 148; hr) sharply marked, extending to middle of distance between anterior margin of submentum and posterior tentorial pits (Figs. 129, 149) or farther (Fig. 141) nearly up to posterior tentorial pits, but not connecting in middle. Gular plate (Figs. 129, 141, 148; gp) large and subtrapezoidal, with rapidly narrowed anterior part (between transverse impression in front of ' neck region' and posterior tentorial pits; lateral margins of this anterior part of gular plat are not marked); gular sutures (Figs. 129, 141, 148; gs) superficial; posterior tentorial pits (Figs. 129, 141, 148; ptp) narrow and straight or slightly arcuate, located clearly in front of transverse impression ventrally demarcating ' neck region'. Antennae (Figs. 160 – 163) moderately long and slender, gradually and only slightly thickening distally or with indistinct club composed of five terminal antennomeres; antennomere XI unremarkable. Prothorax (Figs. 134 – 135, 145 – 146, 153 – 154, 160 – 163) in dorsal view slightly to strongly elongate and broadest near anterior third, with rounded anterior margin and anterior parts of lateral margins, with barely marked, blunt and strongly obtuse anterior corners; posterior corners distinct, obtuse; posterior margin nearly evenly arcuate or shallowly bisinuate. Pronotum without sublateral carinae, with transverse groove (Figs. 135, 145, 154; tg), in some species accompanied by internal and lateral pair of pits (Fig. 135; ip, lp) or interrupted in middle (Fig. 135). Sides of pronotum with thick and sparse bristles (Figs. 134, 146, 153). Prosternum with short basisternal part (Figs. 134, 146, 153; bst) distinctly demarcated from procoxal cavities (Figs. 134, 146, 153; pcc); median part of sternum with narrow and variously distinct prosternal intercoxal carina; procoxal sockets (Figs. 134, 146, 153; pcs) closed by posterolateral lobes of prosternum; hypomera (Figs. 134, 146, 153; hy) elongate, each divided into broad lateral part confluent with pronotum and narrower internal (adcoxal) part; hypomeral ridges (Figs. 134, 146, 153; hyr) weakly marked in median and / or posterior portion; pronotosternal sutures (Figs. 134, 146, 153; nss) entire. Mesoscutellum (Fig. 155; scl 2) subtriangular, in intact specimens hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture (Fig. 155; sss) present. Mesoventrite with narrow and distinctly demarcated anterior ridge (Figs. 137, 147, 156; ar); mesoventral process (Figs. 137, 147, 156; msvp) carinate and moderately expanding ventrally, anteriorly connected with anterior ridge and not reaching anterior margins of mesocoxal cavities, so that mesocoxae are not separated; sides of mesoventrite with shallow procoxal rests (Figs. 137, 147, 156; pcr) filled with bristles (= setose impressions) but without asetose impressions; mesanepisternum with short prepectus (Figs. 137, 147, 156; pre), sides of mesoventrite with narrow but deep setose ventrolateral foveae (Figs. 139, 147, 158; vlf, dlf); dorsolateral foveae absent; mesocoxal projections (Figs. 137, 147, 156; mcp) prominent, with mesocoxal sockets located on their mesoventral surface and visible in ventral view. In one species mesanepisterna each with thick and long bunch of dense setae (Fig. 147; seb), under stereoscopic microscope appearing as solid projection (Fig. 162). Metaventrite (Figs. 138 – 139, 147, 157 – 158; v 3) subrectangular, anteriorly fused with mesoventrite, posteriorly nearly straight with broadly subtriangular and rounded metaventral intercoxal process (Figs. 140, 147, 159; mtvp) without median notch; anterior metaventral process absent. Metanepisterna and metepimera narrow. Metafurca with short stalk and divergent lateral furcal arms (Figs. 139, 147, 158; lmfa). Elytra (Figs. 136, 155, 160 – 163) oval, each with two rudiments of asetose basal foveae (Figs. 136, 155; bef) not connected by groove; humeral denticles and subhumeral lines absent. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 160 – 163) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora moderately strongly clavate; tibiae and tarsi long and slender. Aedeagus (Figs. 168 – 173) elongate and lightly sclerotized, thin-walled, with symmetrical median lobe and symmetrical assemblage of internal sclerites containing H-shaped subapical median structures and often with bunches of long hair-like structures on surface of internal sac; parameres free and slender, with apical setae.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE6183E8FFE3BAB0063FA9A.taxon	etymology	Etymology. The name Sciacharoides is derived from Sciacharis, to underline similarities between these taxa. Gender masculine. Composition and distribution. Four species of Sciacharoides are known from the easternmost part of Australia (Queensland and New South Wales) (Fig. 197). The genus may be broader distributed, but some species might have been misplaced in other genera, e. g. Sciacharis or Euconnus.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE6183E8FFE3BAB0063FA9A.taxon	discussion	Remarks. Sciacharoides differs from Sciacharis in the following characters: the sublateral pronotal carinae absent (present in Sciacharis); the mesoventral process developed only in front of mesocoxal cavities (extending between mesocoxal cavities in Sciacharis) and in consequence the mesocoxae not separated (clearly separated by mesoventral intercoxal process in Sciacharis); and the basal elytral foveae rudimentary (well-developed in Sciacharis). Species of Sciacharoides are also stouter and more convex than Sciacharis. It is possible that other species from Australia, New Zealand and New Caledonia placed by Franz in Sciacharis may in fact belong to Sciacharoides; all members of this complex require a comprehensive revision. Among Australian Cyrtoscydmini only Palaeoscydmaenus has the mesocoxae contiguous and not separated by mesoventral process (Jałoszyński 2012 c), but this genus has lateral sutures of submentum, open procoxal sockets, asetose impressions of mesoventrite, lacks thick bristles on the head and prothorax, and lacks basal elytral foveae. Species included hereby in Sciacharoides show minor but intriguing differences in the antebasal sculpture of the pronotum, length of hypostomal ridges, and in the general body shape. These differences may in future be used to subdivide Sciacharoides into subgenera; currently too few species are known to draw conclusions about their possible relationships. Some species described from Australia and New Zealand under Sciacharis (or its junior synonyms) may in fact belong to Sciacharoides. For instance, aedeagi illustrated by Franz (1975) for Euconnus (Allomaoria) parvicollis (Lea, 1914), E. quinarius Franz, 1975, E. relatus (Broun, 1893 b) (all currently in Sciacharis s. str.) seem to contain structures of the internal sac similar to those of Sciacharoides, including the H-shaped sclerite. For this reason only four species previously misplaced in Horaeomorphus are revised here, and although several other, presumably undescribed species, are known to the author, they will be treated separately, when all Australian species of Sciacharis have been verified (Jałoszyński, in preparation).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE018308FFE3C73072DFB4C.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: six labels (Fig. 164): " Umg. [i. e., env.] Maipoton [sic!] / Queensld., Austr. / lg. H. Franz 1970 " [white, printed], " Horaeomorphus / thompsoni / m / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " SAMA Database / No. 25 - 037021 " [white, printed], " SAMA Digital Image / 20. 8. 2013 " [green, printed and handwritten], " SCIACHAROIDES / thompsoni (Franz, 1975) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Additional material (7 exx.). 2 ♀♀, New South Wales, Wild Cattle Creek State Forest, " in moss on fallen tree ", 30. v. 1993, leg. T. Gush; 1 ♂, New South Wales, Pines Creek State Forest, " in upright dead tree stump ", 31. v. 1993, leg. T. Gush; 1 ♂, New South Wales, Chichester State Forest, " in debris on log ", 6. vi. 1992, leg. T. Gush; 1 ♀, New South Wales, Wang Wauk State Forest, " in moss on fallen tree and stump ", 26. iv. 1993, leg. T. Gush; 1 ♂, Queensland, Mt. Glorious, leaf and log litter with fungi (Berlesate), 21. xii. 1990, leg. J. Lawrence; 1 ♂, Queensland, Mt. Glorious, 800 m, bark and log litter, 28. vi. 1978, leg. S. & J. Peck (specimens in ANIC and cPJ). Revised diagnosis. Male: BL 1.68 – 1.80 mm; pronotum with antebasal transverse groove interrupted in middle and with distinct internal pair of pits; mesanepisterna without setal brushes; subtriangular apical region of aedeagus in ventral view distinctly demarcated from basal capsule by lateral constriction. Females are unremarkable and may not be possible to identify if not collected together with males.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE018308FFE3C73072DFB4C.taxon	description	Redescription. Body of male (Fig. 160) strongly convex, elongate but relatively stout, with long appendages, BL 1.68 – 1.80 mm (mean 1.74 mm); cuticle glossy, body uniformly light brown with slightly lighter appendages, vestiture yellowish. Head (Fig. 129, 160) in dorsal view rounded, broadest at eyes, HL 0.30 – 0.33 mm (mean 0.31 mm), HW 0.33 – 0.38 mm (mean 0.34 mm); tempora slightly longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes moderately large and moderately projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, on frons and anterior part of vertex sparse and suberect, tempora and posterior margin of vertex with dense bristles directed posteriorly and slightly posterolaterally. Antennae slender (Fig. 160), with five terminal antennomeres forming indistinct club, AnL 0.83 – 0.90 mm (mean 0.86 mm); antennomeres I – VI elongate (but only II strongly), VII – VIII about as long as broad or barely discernibly longer than broad; IX – X slightly transverse; XI only slightly longer than X, 1.4 x as long as broad, with blunt apex. Pronotum (Fig. 160) indistinctly elongate, broadest between middle and anterior third, PL 0.40 – 0.45 mm (mean 0.44 mm), PW 0.38 – 0.40 mm (mean 0.39 mm); anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with shallow and narrow transverse groove interrupted in middle and with two pairs of pits: larger internal pair and smaller external pair. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect, sides of pronotum with moderately dense bristles. Elytra (Fig. 160) oval, broadest distinctly anterior to middle, EL 0.98 – 1.03 mm (mean 0.99 mm), EW 0.70 – 0.80 mm (mean 0.75 mm), EI 1.28 – 1.39 (mean 1.33); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc much more distinct than those on pronotum, in anterior median area separated by spaces comparable to diameters of punctures, on sides and posteriorly punctures reducing in diameter and depth; setae much longer than those on head and pronotum, moderately dense, curved and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 160) long and slender, without modifications. Aedeagus (Figs. 168 – 170) relatively stout, AeL 0.33 mm, in ventral view drop-shaped with demarcated and strongly narrowing subtriangular apical region; internal armature composed of H-shaped subapical structure proximally connected to two lateral bunches of thin hair-like sclerites and with a similar bunch of shorter sclerites in middle; H-shaped structure in erected aedeagus (Figs. 169 – 170) protruding distally and hair-like sclerites projecting outwards from the surface of endophallus; parameres slender, each with three apical setae. Female. Similar to male, externally indistinguishable; BL 1.55 – 1.73 mm (mean 1.67 mm); HL 0.28 – 0.30 mm (mean 0.28 mm), HW 0.30 – 0.33 mm (mean 0.31 mm), AnL 0.78 – 0.80 mm (mean 0.79 mm); PL 0.38 – 0.43 mm (mean 0.41 mm), PW 0.35 – 0.38 mm (mean 0.36 mm); EL 0.90 – 1.03 mm (mean 0.98 mm), EW 0.70 – 0.75 mm (mean 0.73 mm), EI 1.29 – 1.37 (mean 1.33).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE018308FFE3C73072DFB4C.taxon	distribution	Distribution (Fig. 197 a). South-eastern Australia: Queensland and New South Wales.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE018308FFE3C73072DFB4C.taxon	discussion	Remarks. The name " Maipoton " on the locality label is misspelled Mapleton. This species highly resembles Sciacharoides eucalypti comb. n., but is clearly larger. Small differences in the shape of the aedeagus and its internal structures and in proportions of antennomeres may fall within an interindividual variability of a single species. However, a small available sample is insufficient to decide whether Sciacharoides thompsoni and Sciacharoides eucalypti are synonymous names; this problem is pending further study.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFED18368FFE3ED40275FF04.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: six labels (Fig. 165): " Umg. [i. e., env.] Brisbane / Queensld., Austr. / lg. H. Franz 1970 " [white, printed], " Horaeomorphus / eucalypti m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " male symbol " [white, printed], " SAMA Database / No. 25 - 037015 " [white, printed], " SCIACHAROIDES / eucalypti (Franz, 1975) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Revised diagnosis. Male: BL 1.45 mm; pronotum with antebasal transverse groove interrupted in middle and with distinct internal pair of pits; mesanepisterna without setal brushes; subtriangular apical region of aedeagus in ventral view laterally confluent with basal capsule. Female and its diagnostic characters unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFED18368FFE3ED40275FF04.taxon	description	Redescription. Body of male (Fig. 161) strongly convex, elongate but relatively stout, with long appendages, BL 1.45 mm; cuticle glossy, body uniformly light brown with slightly lighter appendages, vestiture yellowish. Head (Fig. 161) in dorsal view rounded, broadest at eyes, HL 0.25 mm, HW 0.30 mm; tempora slightly longer than eyes, strongly curved posteromesally; vertex weakly convex; frons subtrapezoidal. Eyes moderately large and moderately projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, on frons and anterior part of vertex sparse and suberect, tempora and posterior margin of vertex with dense bristles directed posteriorly and slightly posterolaterally. Antennae (Fig. 161) slender, with five terminal antennomeres forming indistinct club, AnL 0.68 mm; antennomeres I – II distinctly elongate, III – VI each only slightly longer than broad, VII – X distinctly transverse, XI only slightly longer than X, 1.2 x as long as broad, with blunt apex. Pronotum (Fig. 161) indistinctly elongate, broadest between middle and anterior third, PL 0.38 mm, PW 0.33 mm; anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with shallow and narrow transverse groove interrupted in middle, and with two pairs of pits: larger internal pair and smaller external pair. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect, sides of pronotum with moderately dense bristles. Elytra (Fig. 161) oval, broadest distinctly anterior to middle, EL 0.83 mm, EW 0.60 mm, EI 1.38; basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc much more distinct than those on pronotum, in anterior median area separated by spaces comparable to diameters of punctures, on sides and posteriorly punctures reducing in diameter and depth; setae much longer than those on head and pronotum, moderately dense, curved and erect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 161) long and slender, without modifications. Aedeagus (Fig. 171) relatively stout, AeL 0.25 mm, in ventral view drop-shaped with strongly narrowing subtriangular apical region laterally confluent with sides of median lobe; internal armature composed of H-shaped subapical structure proximally connected to two lateral bunches of thin hair-like sclerites; parameres slender, each with three apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFED18368FFE3ED40275FF04.taxon	distribution	Distribution (Fig. 197 b). South-eastern Australia: Queensland.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFED18368FFE3ED40275FF04.taxon	discussion	Remarks. This species looks like a miniature Sciacharoides thompsoni; s ee Remarks section at the latter species above.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE918378FFE399000A9FA6C.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: six labels (Fig. 166): " Umg. [i. e., env.] Maipoton [sic!] / Queensld., Austr. / lg. H. Franz 1970 " [white, printed], " Horaeomorphus / maipotonensis / m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " ♂ " [white, printed], " SAMA Database / No. 25 - 037018 " [white, printed], " SCIACHAROIDES / maipotonensis (Franz, 1975) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Additional material (5 exx.). 4 ♂♂, Queensland, Mt. Spec, 875 - 880 m, flight intercept trap, 6. xii. 1995, leg. M. Cermak; 1 ♂, same data except for 9. iii. 1995, Malaise trap (ANIC, cPJ). Revised diagnosis. Male and female: BL 1.70 – 1.85 mm; pronotum with complete antebasal transverse groove, without internal pair of pits; mesanepisterna with dense rod-like setal brushes projecting anteriorly.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE918378FFE399000A9FA6C.taxon	description	Redescription. Body of male (Fig. 162) strongly convex, elongate but relatively stout, with long appendages, BL 1.70 – 1.85 mm (mean 1.77 mm); cuticle glossy, body uniformly light brown, vestiture yellowish. Head (Figs. 141, 162) in dorsal view rounded, broadest at eyes, HL 0.30 mm, HW 0.33 – 0.35 mm (mean 0.35 mm); tempora slightly longer than eyes, strongly curved posteromesally; vertex weakly convex with rounded and convex posterior margin; frons subtrapezoidal. Eyes moderately large and weakly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine but dense, separated by spaces comparable to diameters of punctures; setae short, on frons and anterior part of vertex sparse and suberect, tempora and posterior margin of vertex with moderately dense bristles directed posteriorly and slightly posterolaterally. Antennae (Fig. 162) slender, with five terminal antennomeres forming indistinct club, AnL 0.83 – 0.93 mm (mean 0.90 mm), proportions of antennomeres slightly variable in the studied sample (see remarks below); in holotype antennomeres I – II distinctly elongate, III barely noticeable longer than broad, IV – X about as long as broad, XI only slightly longer than X, about 1.6 x as long as broad, with blunt apex; in males with longest antennae antennomeres I – II and V – VI are distinctly elongate, III – IV and VII – VIII slightly or barely noticeably longer than broad, IX – X about as long as broad, and XI about 1.7 x as long as broad. Pronotum (Fig. 162) strongly elongate and strikingly narrow in relation to elytra, broadest near anterior fourth, PL 0.43 – 0.45 mm (mean 0.43 mm), PW 0.33 – 0.38 mm (mean 0.34 mm); anterior and lateral margins rounded; sides in posterior 3 / 4 nearly straight and strongly convergent posteriorly; posterior margin shallowly bisinuate; base of pronotum with narrow and sharply marked transverse groove slightly deepened at each end, without pits. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect, sides of pronotum with moderately dense bristles. Elytra (Fig. 162) oval, broadest near middle, EL 0.98 – 1.13 mm (mean 1.04 mm), EW 0.73 – 0.80 mm (mean 0.76 mm), EI 1.33 – 1.41 (mean 1.36); basal impressions short but distinct; elytral apices separately rounded. Punctures on elytral disc much more distinct than those on pronotum but not particularly conspicuous, unevenly distributed and separated by spaces 0.5 – 2 x diameters of punctures; setae longer than those on head and pronotum, moderately dense, curved and suberect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 162) long and slender, without modifications. Aedeagus (Figs. 172 – 173) in all available specimens variously distorted (see remarks below), in holotype partly erected (Fig. 172), AeL 0.38 mm (including everted internal sac), in ventral view drop-shaped with subtriangular apical region; erected internal armature composed of large H-shaped and microdenticulate structure protruding distally, median lobe subapically with subtrapezoidal component of internal sac with its surface densely covered with denticles; the same structures can be recognized in not erected, but slightly distorted aedeagi of nontype males (Fig. 173); they are shorter than the erected copulatory organ (AeL 0.30 mm), the H-shaped structure is located subapically and components of the subtrapezoidal structure are in fact two separate parts, one located more basally and the other one more distally. Parameres slender, each with three-four apical setae. Female. Externally indistinguishable from male; BL 1.73 mm; HL 0.28 mm, HW 0.33 mm, AnL 0.90 mm; PL 0.43 mm, PW 0.35 mm; EL 1.03 mm, EW 0.73 mm, EI 1.41.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE918378FFE399000A9FA6C.taxon	distribution	Distribution (Fig. 197 c). South-eastern Australia: Queensland.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE918378FFE399000A9FA6C.taxon	discussion	Remarks. The name " Maipoton " on the locality label (and used to derive the specific epithet from) is misspelled Mapleton. The original description of this interesting species (Franz 1975) is an excellent example how hopeless are attempts to identify Australian species basing only on the available literature. The most striking character, the rod-like bunches of setae protruding from beneath the humeri and well-visible in dorsal view (Figs. 147, 162) were not even mentioned by Franz (1975). Moreover, the pronotum is described as devoid of antebasal transverse groove (clearly present in the holotype; Fig. 145), but in the middle in front of the basal margin with a blunt keel and on the sides near corners with a pit ― neither of these actually present. The illustration of the aedeagus (Franz 1975, Fig. 121) shows a partly erected internal sac. Sciacharoides maipotonensis is one of the most remarkable and easily identifiable Australian cyrtoscydmines, but when new specimens are confronted with the Franz's monograph it is not possible to even suspect that this may be the same species. The variability found in the proportions of antennomeres among studied specimens requires farther attention and a morphometric analysis of a larger material. It seems that smaller specimens have shorter antennomeres while the largest ones have more elongate antennae. The variability was found among specimens collected in the same locality (Mt. Spec) and the possibility of a co-existence of cryptic species seems rather low, but the small available sample does not allow drawing any firm conclusions. All available non-type specimens come from flight intercept traps and the dry-mounted males all had the aedeagi protruding entirely or partly from the genital opening, with distorted walls of median lobes (Fig. 173). This is probably an effect of a preservative used in the traps. The general shape of the median lobe is therefore not possible to describe in this species. However, the components of the relatively complicated internal armature of the aedeagus between the partly everted copulatory organ of the holotype and those of the non-type males seem identical, which supports the determination of all specimens as conspecific.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE818488FFE3C850740FAA4.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♀: six labels (Fig. 167): " Tamborine Mt. / Queensld., Austr. / lg. H. Franz 1970 " [white, printed], " Horaeomorphus / montis - tam - / borini [sic!] m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " SAMA Database / No. 25 - 037020 " [white, printed], " SAMA Digital Image / 20. 8. 2013 " [green, printed and handwritten], " SCIACHAROIDES / montistamborinensis (Franz, 1975) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Additional material. 1 ♀, Queensland, Joalah National Park, Tamborine Mt., litter in rainforest (Berlesate), 18 - 21. x. 1978, leg. J. Lawrence & T. Weir (ANIC). Revised diagnosis. Female: BL 1.48 – 1.53 mm; pronotum with complete antebasal transverse groove and moderately distinct external pair of pits or impressions; mesanepisterna without setal brushes.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE818488FFE3C850740FAA4.taxon	description	Redescription. Body of female (Fig. 163) strongly convex, elongate and relatively slender, with long appendages, BL 1.48 – 1.53 mm; cuticle glossy, body uniformly light brown with slightly reddish hue, appendages slightly lighter, vestiture yellowish. Head (Figs. 148, 163) in dorsal view rounded, broadest at eyes, HL 0.25 – 0.28 mm, HW 0.28 – 0.30 mm; tempora slightly longer than eyes, strongly curved posteromesally; vertex weakly convex, its posterior margin slightly concave in middle; frons subtrapezoidal. Eyes moderately large and weakly projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, on frons and anterior part of vertex sparse and suberect, tempora and posterior margin of vertex with dense bristles directed posteriorly and slightly posterolaterally. Antennae (Fig. 163) slender, gradually thickened distally, AnL 0.70 – 0.73 mm; antennomeres I – II distinctly elongate, III – V each slightly longer than broad, VI about as long as broad, VII – X each slightly transverse, XI much shorter than IX – X together, about twice as long as broad, with indistinctly pointed apex. Pronotum (Figs. 154, 163) indistinctly elongate, broadest between middle and anterior third, PL 0.38 mm, PW 0.35 – 0.38 mm; anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with narrow transverse groove connected at each end with distinct external pit or impression. Punctures on pronotal disc fine and inconspicuous; setae moderately dense, long and suberect to erect, sides of pronotum with moderately dense long bristles. Elytra (Fig. 163) oval, broadest distinctly anterior to middle, EL 0.83 – 0.90 mm, EW 0.63 mm, EI 1.32 – 1.44; basal impressions barely marked; elytral apices separately rounded. Punctures on elytral disc slightly more distinct than those on pronotum but still inconspicuous; setae moderately dense, long and suberect. Hind wings well developed, about twice as long as elytra. Legs (Fig. 163) long and slender, without modifications. Male. Unknown.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE818488FFE3C850740FAA4.taxon	distribution	Distribution (Fig. 197 d). South-eastern Australia: Queensland.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FFE818488FFE3C850740FAA4.taxon	discussion	Remarks. This species, known from females only, in dorsal view highly resembles members of various subgenera of Euconnus. Only examination of the ventral characters reveals that it in fact does not differ in any fundamental structures from other species placed here in Sciacharoides. The shape of head is clearly different from that of any Euconnus, but this character may be difficult to examine as the head is often declined and only partly visible in dry-mounted specimens. Similar species were described and placed by Franz (1975) in Euconnus, without paying attention to any ventral structures; it seems possible that more Sciacharoides (or Sciacharis) species can be found among other genera of Australian-Pacific Cyrtoscydmini. The specimen studied and illustrated in the present paper (Figs. 148 – 159) is a female highly similar to the holotype of Horaeomorphus montistamborinensis. If collected in the Tamborine Mountain, it might have been identified as belonging to the same species. However, this female comes from near the Cape Tribulation, over 1400 km NW, and it is here identified as Sciacharoides sp., near montistamborinensis. Another similar species with the complete antebasal pronotal groove and without pits was seen from the Lamington National Park in Queensland; this is a male with interestingly modified elytral apices. As some misplaced species of Sciacharoides may hide among previously described Australian Euconnus or Sciacharis, I refrain from describing new species of Sciacharoides until these two genera have been at least preliminarily surveyed.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF97184E8FFE3CF0016EFDC9.taxon	description	Redescription. Body (Figs. 185 – 186) moderately large (in studied specimens about 2.2 – 2.5 mm of BL), strongly convex, moderately slender, with long appendages, pigmentation brown, cuticle setose. Head (Figs. 174, 185 – 186) elongate, with occipital constriction (Fig. 174; occ) much narrower than vertex and dividing the head capsule into exposed anterior part and narrow posterior ' neck region' retracted into prothorax; eyes moderately large and nearly circular but strongly protruding from head silhouette, located in anterior part of head; tempora long and strongly curved posteromesally, with dense bristles; vertex transverse, convex, slightly projecting dorsocaudad, with posterior margin convex and densely covered with bristles; frons confluent with vertex, transverse and subtrapezoidal, its anterior portion steeply but not abruptly lowering toward labrum; frontoclypeal groove absent; antennal insertions broadly separated, located beneath distinctly raised supraantennal tubercles. Labrum (Fig. 175) transverse with rounded lateral margins and shallow median emargination in anterior margin bordered at each side by angulate projection, with six long and pointed anteroventral (anterior epipharyngeal) sensilla and short fine trichia projecting from beneath anterior margin; dorsal setae sparse and long. Mandibles symmetrical, subtriangular, each with broader than long basal portion and one small subapical tooth (Fig. 176; sat); prostheca (Fig. 176; pst) present, dorsomesal, expanding far onto dorsal surface of mandible and reaching mandibular base. Each maxilla composed of small cardo (Fig. 177; cd) bearing two setae; subtriangular basistipes (Fig. 177; bst); elongate mediostipes (Fig. 177; mst); elongate galea (Fig. 177; gal) and lacinia (Fig. 177; lac); large, elongate palpifer (Fig. 177; pfp); and strongly elongate and moderately large maxillary palp (Fig. 174; mxp) composed of elongate palpomere I (Fig. 177; mxp 1), strongly elongate, slightly clavate and slender palpomere II (Fig. 177; mxp 2), large and moderately elongate, distinctly thickened palpomere III (Fig. 177; mxp 3) broadest nearly in middle, and small, slender, subconical and pointed palpomere IV (Fig. 177; mxp 4). Labium with large submentum (Fig. 174; smn) not demarcated laterally from postcardinal parts of hypostomae; subtrapezoidal mentum (Fig. 174; mn); and moderately long prementum bearing narrowly separated at bases long 3 - segmented labial palps (Fig. 174; lp) and a pair of long bristles on small inversely subtriangular ligula (Fig. 174; lg). Hypostomal ridges (Fig. 174; hr) sharply marked and extending to near middle between anterior margin of submentum and posterior tentorial pits but not connecting in middle. Gular plate (Fig. 174; gp) large and subtrapezoidal, with rapidly narrowed anterior part distinctly demarcated laterally by longitudinal grooves; gular sutures (Fig. 174; gs) superficial; posterior tentorial pits (Fig. 174; ptp) narrow and slightly arcuate, located clearly in front of transverse impression ventrally demarcating ' neck region'. Antennae (Figs. 185 – 186) moderately long and slender, gradually and distinctly thickening distally; antennomere XI unremarkable. Prothorax (Figs. 178 – 179, 185 – 186) in dorsal view strongly elongate, broadest near anterior third, with slightly concave anterior margin; anterior corners relatively distinct, strongly obtuse and blunt; anterior parts of lateral margins rounded; posterior corners blunt but distinct, nearly right; posterior margin deeply bisinuate. Pronotum with distinct sublateral carinae (Fig. 179; slc), without antebasal transverse groove, with two pairs of large internal and lateral pits (Fig. 179; ip, lp) and indistinct median longitudinal wrinkle or carina (Fig. 179; mc). Sides of pronotum with thick and dense bristles (Fig. 178) well visible in dorsal view. Prosternum with short basisternal part (Fig. 178; bst) distinctly demarcated from procoxal cavities (Fig. 178; pcc); median part of sternum with narrow intercoxal carina; procoxal sockets (Fig. 178; pcs) closed by posterolateral lobes of prosternum; hypomera (Fig. 178; hy) elongate, each divided into broad lateral part confluent with pronotum and narrower internal (adcoxal) part; hypomeral ridges (Fig. 178; hyr) incomplete, marked only in posterior portion of internal part of each hypomeron; pronotosternal sutures (Fig. 178; nss) entire. Mesocutellum (Fig. 180; scl 2) subtriangular, in intact specimens hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture (Fig. 180; sss) present. Mesoventrite with narrow and distinctly demarcated anterior ridge (Fig. 181; ar); mesoventral process (Fig. 181; msvp) carinate and moderately expanding ventrally, anteriorly connected with anterior ridge and reaching posterior margins of mesocoxal cavities, so that mesocoxae are clearly separated; sides of mesoventrite with shallow procoxal rests (Fig. 181; pcr) filled with bristles (= setose impressions) but without asetose impressions; mesanepisternum with moderately long prepectus (Fig. 181; pre), sides of mesoventrite with narrow and deep setose ventrolateral and dorsolateral foveae (Fig. 183; vlf, dlf); mesocoxal projections (Fig. 181; mcp) prominent, with mesocoxal sockets located on their mesal surface and in ventral view only partly visible. Metaventrite (Figs. 182 – 183; v 3) subrectangular, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate with broadly subtriangular and metaventral intercoxal process (Fig. 184; mtvp) with short median notch; metacoxae (Fig. 183; cx 3) nearly contiguous; anterior metaventral process absent. Metanepisterna and metepimera narrow. Metafurca with long stalk and divergent lateral furcal arms (Fig. 183; lmfa). Elytra (Figs. 180, 185 – 186) oval, each with two distinct and deep asetose basal foveae (Fig. 180; bef) not connected by groove; humeral denticles and subhumeral lines absent. Hind wings well-developed, about twice as long as elytra. Legs (Figs. 185 – 186) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora moderately strongly clavate; tibiae and tarsi long and slender. Aedeagus (Figs. 189 – 192) elongate and lightly sclerotized, thin-walled, with symmetrical median lobe and symmetrical internal armature; parameres free and slender, with apical setae. Spermatheca (Fig. 183; sp) large and globular, located deeply in metathorax. Composition and distribution. Sciacharis (Maorinus) currently includes 46 species (Jałoszyński 2014 b), only five of them occur in Australia (four only in Tasmania), while the rest inhabits New Zealand. However, these numbers are valid combinations under Sciacharis (Maorinus), while most of these species were assigned to Maorinus (originally to Euconnus (Maoria )) apparently without examination of important ventral characters and their generic status requires verification. Moreover, Kuschel (1990) synonimized two pairs of names of New Zealand Maorinus (as Euconnus) without giving any reasons for doing so (Jałoszyński 2014 b) and these decisions require verification.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF97184E8FFE3CF0016EFDC9.taxon	discussion	Remarks. One deviation from the body plan described previously (Jałoszyński 2014 b) was found in the species studied during the present work. The mesoventrite of Sciacharis (Maorinus) alacer (Broun, 1915), the type species of Maorinus, and that of Maorinus sp. illustrated in Jałoszyński (2014 b; Fig. 21; species highly similar to M. alacer and misidentified as such by Franz) has only the ventrolateral foveae, while in Sciacharis (Maorinus) tasmaniensis comb. n. there are two distinct pairs of foveae (Fig. 183). All other characters do not differ. The presence vs. absence of the dorsolateral foveae does not seem a sufficient criterion to place this species in a separate genus or subgenus. The character variability within Sciacharis and its subgenera is too poorly known to propose conclusive solutions other than used here, i. e. placing Horaeomorphus tasmaniensis in Sciacharis (Maorinus). Other Australian species of Maorinus (not misplaced in Horaeomorphus) will be revised in a separate paper (Jałoszyński, in preparation).	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF91184F8FFE3821004CFF29.taxon	materials_examined	Type material studied. Holotype: AUSTRALIA: ♂: seven labels (Fig. 188): " Griffith Collection Id. by A. M. Lea " [white, printed], " moss " [white with red margins, typed], " Horaeomorphus / tasmaniensis m. / det. H. Franz " [white, handwritten and printed], " Typus " [red, handwritten], " SAMA Database / No. 25 - 037013 " [white, printed], " SAMA Digital Image / 14. 8. 2013 " [green, printed and handwritten], " SCIACHARIS (MAORINUS) / tasmaniensis (Franz, 1975) / det. P. Jałoszyński, 2014 " [white, printed] (SAM). Additional material (7 exx.). 3 ♂♂, Tasmania, Bathurst Harbour, closed forest, flight intercept trap, 7. xii. 1990 - 15. i. 1991, leg. E. Nielsen & T. Edwards; 1 ♂, same data except for 15. i. - 20. ii. 1991, leg. A. Calder & W. Dressler; 1 ♂, 1 ♀, Tasmania, 4 km SE Weldeborough, pantrap, 13. i. - 7. ii. 1983, leg. I. Naumann & J. Cardale; 1 ♂, Tasmania, Mount Victoria Forest reserve, leaf litter, 3. ii. 1992, leg. T. Gush; (ANIC and cPJ). Revised diagnosis. Male and female: elytra with strongly waving pattern of setae. Male: each elytron with variously distinct small flattening or impression in subapical region near suture, covered with recumbent setae directed posteromesally; aedeagus in ventral view moderately stout, with rapidly narrowing subtriangular apical region and complicated internal armature containing two submedian lateral groups of hair-like sclerites broadly separated in middle and subapical groups of similar sclerites connecting in middle.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF91184F8FFE3821004CFF29.taxon	description	Redescription. Body of male (Figs. 185 – 186) strongly convex, elongate and slender, with long appendages, BL 2.23 – 2.45 mm (mean 2.33 mm); cuticle glossy, head pronotum and elytral suture reddish-brown (in some specimens also median parts of sides of elytra indistinctly darkened), remaining body parts light brown; vestiture yellowish. Head (Figs. 174, 185 – 186) elongate, broadest at eyes, HL 0.48 – 0.50 mm (mean 0.50 mm), HW 0.38 – 0.40 mm (mean 0.40 mm); tempora nearly three times as long as eyes, strongly curving posteromesally; vertex convex and slightly projecting dorsocaudad; frons subtrapezoidal. Eyes moderately large and nearly hemispherical, strongly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum distinct and dense but obscured by dense and moderately long suberect setae posteriorly gradually replaced by dense thick bristles covering tempora and nearly entire vertex. Antennae (Figs. 185 – 186) slender, AnL 1.03 – 1.25 mm (mean 1.14 mm); antennomeres I – II strongly elongate, III – VI about as long as broad, VII – X slightly but distinctly transverse, XI nearly as long as IX – X together, twice as long as broad, indistinctly pointed at apex. Pronotum (Figs. 179, 185 – 186) elongate, broadest near anterior third, PL 0.55 – 0.65 mm (mean 0.61 mm), PW 0.48 – 0.53 mm (mean 0.50 mm); anterior margin slightly concave; lateral margins in anterior half rounded; posterior margin distinctly bisinuate; base of pronotum with distinct sublateral carinae, two pairs of distinct pits (external located in large shallow impressions and internal) and short distinct median longitudinal carina. Punctures on pronotal disc fine and inconspicuous; entire pronotum covered with moderately dense suberect to erect bristles, dorsally with admixture of thin setae. Elytra (Figs. 185 – 187) oval, broadest distinctly anterior to middle, EL 1.15 – 1.33 mm (mean 1.23 mm), EW 0.80 – 0.90 mm (mean 0.86 mm), EI 1.34 – 1.52 (mean 1.43); basal impressions short but distinct; narrow adsutural area slightly impressed so that suture is raised; each elytron in subapical region near suture with small nearly round flattening or indistinct impression (setal patch; Fig. 187) covered with fine coarse microgranulation; elytral apices separately rounded. Punctures on elytral disc distinct and dense but shallow, with slightly raised margins so that surface of elytra appears slightly coarse; setae moderately dense, long and suberect, forming a waving pattern; setae on subapical impressions recumbent and directed posteromesally. Hind wings well developed, about twice as long as elytra. Legs (Figs. 185 – 186) moderately long and slender, without modifications. Aedeagus (Figs. 189 – 192) in ventral view moderately stout, AeL 0.43 mm, with rapidly narrowing subtriangular apical region forming thick plate; internal armature complicated and partly darkly sclerotized, with two paired groups of hair-like sclerites and lightly sclerotized, large subapical lobes; parameres slender, each with numerous apical and subapical setae. In erected condition (Figs. 191 – 192) subapical lobes fully extended laterodistally and forming large ' rabbit ear' - like projections and hair-like sclerites forming two longitudinal and one arcuate transverse rows on the surface of erected endophallus. Female. Similar to male, differs in elytra without subapical impressions; BL 2.38 mm, HL 0.50 mm, HW 0.40 mm, AnL 1.05 mm; PL 0.65 mm, PW 0.53 mm; EL 1.23 mm, EW 0.90 mm, EI 1.36.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF91184F8FFE3821004CFF29.taxon	distribution	Distribution (Fig. 198). South-eastern Australia: Tasmania.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
03F68791FF91184F8FFE3821004CFF29.taxon	discussion	Remarks. In Tasmania at least one highly similar species occurs, also with a waving setal pattern on the elytra, but differing in proportions of the antennomeres (especially the antennomere XI is distinctly shorter) and stouter aedeagus. Australian Maorinus will be revised separately; in the present paper only Sciacharis tasmaniensis, previously misplaced in Horaeomorphus, is dealt with.	en	Jałoszyński, Paweł (2014): ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 3828 (1): 1-76, DOI: 10.11646/zootaxa.3828.1.1
