identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E98784FFF6FFD1FF58433AFD80F9ED.text	03E98784FFF6FFD1FF58433AFD80F9ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glanochthon Schoch and Witzmann 2009	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Glanochthon Schoch and Witzmann, 2009</p>
            <p>Diagnosis</p>
            <p>(1) Preorbital region 1.8–2.2 times as long as postorbital skull table, (2) tabular horn prominent, (3) postglenoid region posteriorly longer than the articular facet, (4) jugal narrower than greatest orbit width, and (5) interclavicle slender and at least twice as long as wide (modified from Schoch and Witzmann, 2009b).</p>
            <p>Type species</p>
            <p> Glanochthon latirostre (Jordan, 1849). </p>
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	https://treatment.plazi.org/id/03E98784FFF6FFD1FF58433AFD80F9ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Schoch Correspondence, Rainer R.	Schoch Correspondence, Rainer R. (2021): Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships. Fossil Record 24 (1): 49-64, DOI: 10.5194/fr-24-49-2021, URL: http://dx.doi.org/10.5194/fr-24-49-2021
03E98784FFF6FFD1FF5844CEFD31FC3A.text	03E98784FFF6FFD1FF5844CEFD31FC3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Temnospondyli Zittel 1888	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Order  Temnospondyli Zittel, 1888</p>
            <p>Rhachitomi Watson, 1919 sensu Schoch, 2013</p>
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	https://treatment.plazi.org/id/03E98784FFF6FFD1FF5844CEFD31FC3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Schoch Correspondence, Rainer R.	Schoch Correspondence, Rainer R. (2021): Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships. Fossil Record 24 (1): 49-64, DOI: 10.5194/fr-24-49-2021, URL: http://dx.doi.org/10.5194/fr-24-49-2021
03E98784FFF6FFD5FF584176FAA6FC90.text	03E98784FFF6FFD5FF584176FAA6FC90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cheliderpeton lellbachae Kratschmer 2006	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Glanochthon lellbachae (Krätschmer, 2006) comb. nov. (Figs. 2–5) </p>
            <p> Cheliderpeton lellbachae Krätschmer, 2006</p>
            <p>Holotype</p>
            <p> NHMM 2006 / 14, 102 mm SL, complete skeleton with skin impression (Fig. 2).</p>
            <p>Type of locality and age</p>
            <p>The Klauswald southwest of Odernheim am Glan, Rhineland-Palatinate, Germany (Fig. 1). M9 sequence, Klauswald facies, Odernheim Subformation, Meisenheim Formation, lower Rotliegend, Autunian, lowermost Permian.</p>
            <p>Referred material</p>
            <p> Altogether, five additional specimens are referred to as  Glanochthon lellbachae (see material section). There remains a substantial number of specimens in private collections. </p>
            <p>Diagnosis</p>
            <p> Autapomorphies: (1) preorbital region in adults 1.8– 2.0 times as long as postorbital skull table, (2) dermal ornament with continuous and relatively tall radial ridges in the snout, frontals and cheek (contrasting the more polygonal arrangement in  S. nobilis ), (3) prefrontal anterolaterally expanded to form a more equant pentagon, (4) squamosal posteriorly only half as wide as quadratojugal, (5) phalanges of manus and pes slightly longer and more gracile than in  S. nobilis and (6) tail substantially longer than skull and trunk combined (shorter than that measurement in  S. nobilis ). </p>
            <p>Taxonomic assignment</p>
            <p> This taxon was originally erected as  Cheliderpeton lellbachae by Krätschmer (2006), who made reference to its resemblance to  Cheliderpeton latirostre as described and referred to by Boy (1993). Schoch and Witzmann (2009b) suggested the new genus name  Glanochthon for  Cheliderpeton latirostre after the type species  Cheliderpeton vranyi had been redescribed by Werneburg and Steyer (2002) and was found not to be closely related to  G. latirostre (Schoch and Witzmann, 2009b) . Based on its below-demonstrated close relationship to  G. latirostre ,  Cheliderpeton lellbachae is also referred to as  Glanochthon as a new combination. </p>
            <p> G. lellbachae co-occurs with  Sclerocephalus nobilis in the same locality and horizon (Krätschmer and Resch, 2005; Schoch and Witzmann, 2009a). The combined list of autapomorphic characters 1–5 for the genus distinguish this taxon from  S. nobilis and all other species of  Sclerocephalus . Hypothetical larvae and small juveniles of  G. lellbachae and  S. nobilis may not be distinguished on the basis of the mentioned features, but  S. nobilis generally has a wider jugal and more rounded orbit even at small stages. The presence of a third eryopiform temnospondyl at the Klauswald locality (suggested by Krätschmer, 2006) cannot be confirmed. With a maximum skull length of 17 cm (specimens in private collections),  G. lellbachae appears to have been smaller than the more heavily ossified  S. nobilis (24 cm), but admittedly the number of available specimens is very limited. </p>
            <p>Phylogeny and taxonomy</p>
            <p> In the analysis reported below,  S. nobilis and the  Glanochthon clade are found to be sister taxa. This recognized topological pattern leads to the phylogenetically problematic situation that  Sclerocephalus forms a grade towards  Glanochthon , which under strict application of cladistic principles would prompt erection of new genera at each node. An alternative option would be to define  Sclerocephalus more broadly, encompassing all of  Sclerocephalidae including  Glanochthon . At the present stage, I consider any such step premature as long as morphological effects of microevolution cannot be distinguished from other effects, especially plasticity (ecophenotypes), which is beyond the scope of the present study. </p>
            <p>Comment</p>
            <p> Based on the current phylogenetic findings,  Glanochthon is referred to as the family  Sclerocephalidae , which contains a monophyletic group including a long  Sclerocephalus grade and a terminal  Glanochthon clade. The former referral of  Glanochthon (as  Cheliderpeton latirostre ) within the  Intasuchidae (Schoch and Milner, 2000) was based on a suite of characters which appear to be convergent in the present light of evidence. </p>
            <p>Occurrence</p>
            <p> Although  S. nobilis and  G. lellbachae co-occur in the same locality and several successive horizons, they differ in sample size per horizon. Most notably,  G. lellbachae is more common in the upper fish beds (Obere Fischschiefer, “kalkige Papierschiefer”, K5), whereas  S. nobilis peaks in the lower fish beds (Untere Fischschiefer, K2) as defined by Krätschmer (2004). In the other beds (K3, 4), the two taxa co-occur, mostly represented by large juveniles. </p>
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	https://treatment.plazi.org/id/03E98784FFF6FFD5FF584176FAA6FC90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Schoch Correspondence, Rainer R.	Schoch Correspondence, Rainer R. (2021): Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships. Fossil Record 24 (1): 49-64, DOI: 10.5194/fr-24-49-2021, URL: http://dx.doi.org/10.5194/fr-24-49-2021
