identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EB87F8F171FFB6FF31F955798E4DDB.text	03EB87F8F171FFB6FF31F955798E4DDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium fusculum (Erichson 1839)	<div><p>Key to fusculum group</p><p>1 Punctation of each elytron not forming longitudinal rows of punctures........................................... 2</p><p>- Punctation of each elytron forming several vague and tangled longitudinal rows of punctures in middle................. 3</p><p>2 Pronotum about twice as wide as long, with anterior angles not or indistinctly protruded anteriad. Elytra moderately narrow, about as long as wide. Body reddish brown to brown. Larger: 2.26–3.25 mm. Aedeagus narrow (Fig. 60). Habitus as in Fig. 5. India, Nepal ................................................................... A. cavicrus (Champion, 1925)</p><p>- Pronotum slightly more than twice as wide as long, with anterior angles distinctly protruded anteriad. Elytra wide, longer than wide. Body yellow-brown. Smaller: 1.81–2.18 mm. Aedeagus wide (Fig. 79). Habitus as in Fig. 74. China: Sichuan ............................................................................. A. liliputense Shavrin &amp; Smetana, sp.n.</p><p>3 Mediobasal portion of pronotum with distinct transverse moderately coarse microsculpture between punctures............ 4</p><p>- Pronotum without microsculpture between punctures......................................................... 5</p><p>4 Pronotum less than twice as wide as long. Body darker, brown to black. Aedeagus wide, with short parameres (Figs. 38–43, Shavrin 2013: Figs. 1–6). Body length: 2.20–3.40 mm. Habitus as in Fig. 1. Europe, Middle East, Central Asia......................................................................................... A. fusculum (Erichson, 1839)</p><p>- Pronotum about one and half times as wide as long. Body paler, yellow-brown, with reddish-brown head. Aedeagus narrow, with significantly elongate parameres (Fig. 66). Body length: 2.75–3.48 mm. Habitus as in Fig. 46. India ...................................................................................... A. cuccodoroi Shavrin &amp; Smetana, sp.n.</p><p>5 Elytra about twice as long as pronotum.................................................................... 6</p><p>- Elytra more than twice as long as pronotum................................................................. 7</p><p>6 Pronotum markedly less than twice as wide as long. Aedeagus shorter and wider; internal sac with lateral rows of strongly sclerotized large teeth (Fig. 55). Body length: 2.30–3.30 mm. Habitus as in Fig. 47. Nepal ................................................................................................... A. denticulatum Shavrin &amp; Smetana, sp.n.</p><p>- Pronotum about twice as wide as long. Aedeagus slightly longer and narrower; internal sac without strongly sclerotized teeth (Fig. 52). Body length: 1.95–2.74 mm. Habitus as in Fig. 3. Nepal ................. A. besucheti Shavrin &amp; Smetana, sp.n.</p><p>7 Pronotum massive in general appearance, with widely rounded apical and basal margins. Head reddish brown; pronotum, elytra and abdomen yellow-brown. Aedeagus with median lobe wide, with wide, truncate apex (Fig. 71). Body length: 2.56–2.95 mm. Habitus as in Fig. 49. China: Yunnan .......................... A. latissimum Shavrin &amp; Smetana, sp.n.</p><p>- Pronotum narrower. Aedeagus different.................................................................... 8</p><p>8 Elytra longer than wide. Head reddish brown; pronotum, elytra and abdomen yellow-brown. Aedeagus as in Fig. 57. Body length: 2.85–2.95 mm. Habitus as in Fig. 4. China: Yunnan ....................... A. capitale Shavrin &amp; Smetana, sp.n.</p><p>- Elytra as long as or broader than long...................................................................... 9</p><p>9 Anterior angles of pronotum slightly protruded anteriad. Internal sac of aedeagus with rows of strongly sclerotized teeth (Fig. 50). Body length: 3.25–3.60 mm. Habitus as in Fig. 2. India, Nepal ...................... A. altivagans (Cameron, 1941)</p><p>- Anterior angles of pronotum strongly protruded anteriad. Internal sac of aedeagus without modifications............... 10</p><p>10 Middle portion of each elytron with irregular, transverse and diagonal elevations between punctures. Body reddish brown, with lateral portions of pronotum and elytra yellow-brown. Aedeagus as in Fig. 84. Body length: 2.65–2.75 mm. Habitus as in Fig. 76. India, Nepal ............................................................... A. rugosum (Champion, 1925)</p><p>- Middle portion of each elytron with indistinct to distinct longitudinal elevations between punctures................... 11</p><p>11 Elytra moderately narrow, slightly widened apicad. Body yellow-brown to reddish brown. Aedeagus as in Fig. 86; parameres widened apically. Body length: 2.35–3.20 mm. Habitus as in Fig. 77. India, Nepal ............. A. sociale (Cameron, 1941)</p><p>- Elytra distinctly widened apicad. Parameres narrow, not widened apically (Figs. 62, 68, 81, 89)....................... 12</p><p>12 Lateral portion of pronotum evenly narrowed posteriad and anteriad............................................ 13</p><p>- Lateral portion of pronotum more narrowed posteriad than anteriad............................................. 14</p><p>13 Elytra about as long as wide. Body reddish brown, with head and abdomen dark-brown. Aedeagus moderately wide, parameres vaguely exceeding apex of median lobe (Fig. 81). Body length: 3.10–4.50 mm. Habitus as in Fig. 75. Pakistan, India, Nepal .............................................................................. A. monticola (Cameron, 1924)</p><p>- Elytra distinctly broader than long. Body yellow-brown, with head reddish brown to brown. Aedeagus narrow, parameres distinctly exceeding apex of median lobe (Fig. 68). Body length: 2.35–3.65 mm. Habitus as in Fig. 48. China: Yunnan ................................................................................. A. laozii Shavrin &amp; Smetana, sp.n.</p><p>14 Grooves in front of ocelli long, reaching level of middle or anterior third of eyes. Parameres slightly exceeding apex of median lobe of aedeagus (Fig. 62). Body reddish brown to brown. Body length: 3.00– 3.45 mm. Habitus as in Fig. 45. China: Sichuan, Hubei .................................................................. A. confucii Shavrin &amp; Smetana, sp.n.</p><p>- Grooves in front of ocelli short, as long as or slightly longer than diameter of ocellus. Parameres very long, significantly exceeding apex of median lobe of aedeagus (Fig. 89). Body length: 2.49–3.34 mm. Body yellow-brown to reddish brown. Habitus as in Fig. 78. China: Yunnan .................................... A. splendidulum Shavrin &amp; Smetana, sp.n.</p></div>	https://treatment.plazi.org/id/03EB87F8F171FFB6FF31F955798E4DDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F170FFB9FF31F9867F29494E.text	03EB87F8F170FFB9FF31F9867F29494E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium fusculum (Erichson 1839)	<div><p>Anthobium fusculum (Erichson, 1839)</p><p>(Figs. 1, 6, 8, 10, 12, 14, 16, 18, 20, 22, 24, 26, 28–32, 38–43, 44)</p><p>Lathrimaeum fusculum Erichson, 1839: 626</p><p>Lathrimaeum fusculum: Erichson, 1840: 870; Redtenbacher, 1849: 829; Kelch 1852: 8; Fairmaire and Laboulbène, 1856: 637; Kraatz, 1857: 947; Lentz 1857: 57, 1879: 20; Gutfleisch &amp; Bose 1859: 179; Clasen 1861: 188; Hochhuth 1862: 103; Preller 1862: 48; Stierlin &amp; Gautard 1867: 106; Wilken 1867: 46; Fauvel, 1871: 120 [= 1872: 94]; Letzner 1871: 99, 1886: 142; Bertolini 1872 -1878: 242; Brüggemann 1873: 475; Redtenbacher, 1874: 268; Kellner 1876: 58; Seidlitz, 1875: 239; Dalla Torre 1879: 73; Mulsant and Rey, 1880: 131; Westhoff 1881: 98; Cornelius 1884: 25; Kittel 1884: 170; Cherkunov 1889: 164; Favre 1890: 129; Fauconnet 1892: 121; Ganglbauer, 1895: 722; Klima 1902: 71; Koltze 1901: 60; Stierlin 1900: 407; Fauconnet &amp; Viturat 1903 -1904: 251; Weber 1903: 38; Luze 1905: 54, 58, 67; Stierlin 1906: 179; Linke 1907: 11; Jakobson 1908: 459; Reitter, 1909: 188; Bernhauer and Schubert, 1910: 64; Gerhardt 1910: 53; Prossen 1910: 174;</p><p>Röttgen 1911: 71; Łomnicki 1913: 65; Tenenbaum 1913: 17; Johansen 1914: 596; Ammann &amp; Knabl 1922: 61; Porta</p><p>1926: 26; Fleischer 1927 -1930: 104; Zirk 1928: 6; Portevin, 1929: 442; Nowotny &amp; Polentz 1933: 14; Rapp 1933 -1935:</p><p>223; Scheerpeltz, 1933: 1055, 1961: 79, 1968: 13; Sainte-Claire Deville 1935: 83; Walles 1936: 176; Borchert 1937: 150,</p><p>1938: 22; Lysholm 1937: 154; Hölzel 1946: 71; Lohse 1948: 199; 1964: 52; Palm 1948: 100, 102; Wörndle 1950: 127;</p><p>Borchert 1951: 70; Hansen 1952: 79, 1964: 98; Horion 1951: 109, 1961: 243, 1963: 107; Strand 1953: 65; Porta 1959: 49;</p><p>Friden 1960: 30; Kirschenblat 1965: 163; Koch 1968: 91, 1989: 229; Franz 1970: 286; Heiss 1971: 37; Medvedev &amp;</p><p>Sosnina 1973: 825; Peez &amp; Kahlen 1977: 129; Coiffait, 1978: 274; Focarile 1978: 27, 1981: 35; Bercio &amp; Folwaczny</p><p>1979: 98; Burakowski et al. 1979: 55; Kofler 1980: 122; Uhlig &amp; Zerche 1981: 150; Berstorf et al 1982: 24; Korge 1989:</p><p>30, 1991: 284, 2005: 4; Toth 1982: 89; Terlutter 1984: 27; Zanetti, 1987: 297; Schatz et al. 1990: 205; Schülke et al. 1992:</p><p>167; Meybohm 1994: 48; Gürlich et al. 1995: 26; Kahlen 1995: 21, 2011: 180; Zanetti, 1995: 8; Böhme 1996: 46; Schatz</p><p>1996: 257; Scholze 1999: 380, 2009: 229; Geiser 2001: 188 &lt;?!&gt; Lathrimaeum ? fusculum: Baudi di Selve 1870: 401 Lathrymaeum [sic!] fusculum: Fröhlich 1897: 44 Lathrimaeum (Lathrimaeum) fusculum: Scheerpeltz, 1961: 79; Tichomirova 1973: 139 Olophrum fusculum: Redtenbacher, 1857: 249 Anthobium fusculum: Silfverberg 1979: 18; Rutanen 1983: 213; Lundberg 1988: 83; Zerche 1991: 314; Stoltze &amp; Pihl 1998:</p><p>109; Kubisz et al 1998: 234; Derunkov 1999: 8; Direktoratet for Naturforvaltning 1999: 85; Herman 2001: 232; Andersen</p><p>&amp; Olberg 2003: 61; Assing 2004: 676, 2005: 58, 2006: 226, 2007: 6, 2013: 106; Scholze et al. 2004: 275; Andersen &amp;</p><p>Hanssen 2005: 3505; Barndt 2005: 81; Bohač 2005: 436, 660, 2016: 4; Gutowski et al. 2006: 129; Schatz 2007: 90;</p><p>Schülke 2007: 139; Strobl 2007: 24; Bäse 2008: 130; Gontarenko 2008: 22; Szujecki 2008: 44, 116; Esser 2009: 27, 2017:</p><p>12; Semenov 2009: 24; Piryugin 2010: 281; Derunkov &amp; Bacal 2011: 106; Samin et al. 2011: 1236; Solodovnikov 2012:</p><p>67; Zanetti 2012: 56; Vogel 2013: 37; Petrenko et al. 2014: 6; Odegaard et al. 2015: 21; Hlavač et al., 2016: 2 &lt;?!&gt; Anthobium fusculum: Bacal et al. 2013: 423 Anthobium (Anthobium) fusculum: Gamarra &amp; Outerello 2008: 212; Smetana 2004: 238; Schülke &amp; Smetana 2015: 306 Lathrimaeum melanochromum Iablokoff-Khnzorian, 1961: 140 Lathrimaeum (Lathrimaeum) melanochromum: Tichomirova, 1973: 139 Lathrimaeum melanochromum: Herman 2001: 235 Anthobium (Anthobium) melanochromum: Smetana 2004: 239; Lathrimaeum mesasiaticum Kirschenblat, 1961: 363 syn. nov. Lathrimaeum (Lathrimaeum) mesasiaticum: Tichomirova 1973: 139 Anthobium mesasiaticum: Herman, 2001: 235; Hlavač et al., 2016: 2 Anthobium (Anthobium) mesasiaticum: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 306 Olophrum kondarense Iablokoff-Khnzorian, 1964: 157 Olophrum kondarense: Herman 2001: 349; Smetana 2004: 249 Lathrimaeum kashmiricum Coiffait, 1982: 150 Anthobium kashmiricum: Herman, 2001: 233 Anthobium (Prionothorax) kashmiricum: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 307 Deliphrosoma ivanovi Shavrin, 2012: 17 Deliphrosoma kirgizicum Shavrin, 2012: 19 Deliphrosoma sp.: Shavrin, 2012: 19</p><p>Type material examined: Holotype of Lathrimaeum mesasiaticum Kirschenblat 1961 ♀ [specimen without right antennomeres 3–11 and left antennomeres 10–11]: TAJIKISTAN: ‘Cтaлинaбaд [Stalinabad (= Dushanbe)] | Taджик. [Tajik[istan].] | Киpичeнкo [Kirichenko] 28 X [handwritten] 943’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | mesasiaticum sp. n. | Type | J Kirschenblatt det.’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘ HOLOTYPE | Lathrimaeum | mesasiaticum | Kirschenblatt J.D. det. 1961’ &lt;red rectangular label, printed&gt;, ‘ Anthobium | fusculum (Erichson, 1839) | Shavrin A.V. det. 2016’ &lt;rectangular label, printed&gt; (ZIN).</p><p>Paratypes of L. mesasiaticum Kirschenblat 1961: 1 ♀ [apical part of the abdomen is glued under the specimen]: TAJIKISTAN: ‘Пeнджикcит [Пeнджикeнт] | 10.XII. [1]943. | Киpичeнкo. [Pendzhiksit [Panjakent] 10.XII.1943 Kirichenko.]’ &lt;rectangular label, handwritten&gt;, ‘ Lathrimaeum | mesasiaticum sp.n. | JKirschenblatt det.’ &lt;rectangular label, handwritten in black Indian ink&gt; (ZIN); 1 ♀ [specimen without right antennomeres 3–11, left elytron and left middle leg]: UZBEKISTAN: ‘cт Гoлoднaя Cтeпь | Cp[eднe-] Aз[иaтcкaя].ж[eлeзнaя].д[opoгa]., Xoдж[eнтcкий].y[eзд] | Г Якoбcoн 5.[handwritten]IV.[19]03 [Golodnaya Step`st. Middle-Asian Railway, Khodzhentskiy uezd [Samarkand Area of Turkestan Territory of the Russian Empire] G. Yakobson 5.IV.03]’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | mesasiaticum sp. n. | J Kirschenblatt det.’ &lt;rectangular label, handwritten in black Indian ink&gt; (ZIN). Paratypes with additional labels: ‘ PARATYPE | Lathrimaeum | mesasiaticum | Kirschenblatt J.D. det. 1961’ &lt;red rectangular label, printed by the senior author&gt;, ‘ Anthobium | fusculum (Erichson, 1839) | Shavrin A.V. det. 2016’ &lt;rectangular label, printed&gt;.</p><p>Additional material: UKRAINE: 2 ♂: Odessa obl., Berekhovka Distr., vic. Raukhovka. 0 1.11.2008. A.V. Gontarenko (CS); RUSSIA: 1 ♂, 2 ♀: Volgograd Area, Elton Lake . 01- 10.08.2006. K. Makarov &amp; A. Matalin (MSPU) ; 1 ♂: Rostov Area, Alexandrovka vill., Leninskiy Forest . 17.03.1977. E. Khachikov (CK) ; TURKEY: Niğde, Ulukişla, 5 km SW Kilan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.431946&amp;materialsCitation.latitude=37.445" title="Search Plazi for locations around (long 34.431946/lat 37.445)">Güney Mountains</a>, 37°26'42''N, 34°25'55''E. 13.04.2018. S. Anlaş (CSA) ; IRAN: 1 ♀: Kopet-Dag, Khugdi env. 21.04.2005. lag3 [without collector`s name] (CS) ; KAZAKHSTAN: 2 ♂♂, 1 ♀: West Tian Shan, Chimkent Reg., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.383335&amp;materialsCitation.latitude=70.583336" title="Search Plazi for locations around (long 42.383335/lat 70.583336)">Aksu-Dzhabagly State Nature Reserve</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.383335&amp;materialsCitation.latitude=70.583336" title="Search Plazi for locations around (long 42.383335/lat 70.583336)">Unken-Kaindi</a>, 70°35'N 42°23'E. 11.06.1999. 2200 m a.s.l., in bear dung. J. Cooter leg. (CR) ; 6 ♂♂, 6 ♀♀: same Natural Reserve, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.416668&amp;materialsCitation.latitude=70.7" title="Search Plazi for locations around (long 42.416668/lat 70.7)">Kalbaidjailau</a>, 70°42'N 42°25'E. 16.06.1999. 2500 m a.s.l., grubbing—snow melt. J. Cooter leg. (CR) ; 9 ♂, 6 ♀: same Natural Reserve, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.375&amp;materialsCitation.latitude=70.65" title="Search Plazi for locations around (long 42.375/lat 70.65)">Ulken Kaindi Pass</a>, 70°39'N 42°22'30''E. 14.06.1999. Ca 3000 m a.s.l., at snow melt. J. Cooter leg. (CR) ; 1 ♂: Karatau Mts. (plateau), 36 km W Karatau Mt., N 43°10'09'' E 69°56'32''. 1130 m a.s.l., 0 6.05.2015. A.S. Prosvirov (CS) ; KYRGYZSTAN: 2 ♂♂, 4 ♀♀: Kyrgyzskiy Alatau, Ala Artcha. 1650 m. 29.10.2000 (CSC) ; TAJIKISTAN: 1 ♀: Varzob District, Khodja obi Garm. 2000 m a.s.l. 21- 24.03.2015. A. Barševskis (CS) ; 1 ♀: Kondar Pass, Kvak. 2200 m a.s.l., near snow field. 10.05.1988. V.G. Shilenkov (CS) ; PAKISTAN: 1 ♂, 1 ♀: Swat, Malam Jabba. 2500-2600 m a.s.l. 18.05.1983. Besuchet &amp; Löbl (MHNG) ; WITHOUT EXACT LOCALITY: 1 specimen [without apical segments of abdomen]: ‘34’ (ZMM; collection of A.L. Tichomirova) .</p><p>Redescription. Measurements (n=50): HW: 0.55–0.64, HL: 0.30–0.41; AL: 0.77; OL: 0.12–0.17; PL: 0.47–0.57; PW: 0.80–1.02; ESL: 1.14–1.25; EW: 1.05–1.25; AW: 1.05–1.17; MTbL: 0.57, MTrL: 0.32 (MTrL 1–4: 0.20; MTrL 5: 0.12); AedL: 0.31–0.52; TL: 2.10–3.40 (holotype of Lathrimaeum mesasiaticum: 2.70; holotypes of Lathrimaeum cachemiricum and Deliphrosoma ivanovi: 2.80; holotype of Deliphrosoma kirgizicum: 3.40).</p><p>FIGURES 14–27. Body parts of Anthobium fusculum group (14, 16, 18, 20, 22, 24, 26— A. fusculum, 15, 17, 19, 21, 23, 25, 27— A. monticola): 14–15—ventral aspect of head, 16–17—ventral aspect of prothorax, 18–19—scutellum (dorsal view), 20–21—ventral aspect of elytron, 22–23—ventral aspect of mesothorax, 24–25—ventral aspect of metathorax (without metanepisternum), 26–27—metendosternite (ventral view). Scale bar: 0.1 mm.</p><p>Body elongate, subconvex. Body and antennae brown to dark brown or black, usually with lateral portions of pronotum and elytra yellow-brown to reddish-brown; mouthparts and legs yellow to reddish-yellow; basal portion of antennomere 1 and tarsi (in some specimens) yellow. Body shiny, without microsculpture on elytra; head between supra-antennal elevation and anterior margin of eye with distinct diagonal cellular-shaped meshes, median part of head without or with transverse microsculpture beween punctures, more coarse in posterior portion between ocelli, infraorbital ridges of head with dense, small and coarse microsculpture (some specimens with very fine microsculpture on middle portion of head except of posterior part of infraorbital ridges with distinct transverse meshes between punctures); middle part of neck and abdominal tergites with distinct isodiametric microsculpture; pronotum with indistinct to distinct transverse microsculpture stronger in middle of anterior third (some specimens without microsculpture here); scutellum with indistinct to distinct transverse microsculpture or without it. Head with small or moderately large and deep punctation, markedly denser on posterior portion of infraorbital ridges; middle part of neck with dense smooth punctation; pronotum with irregular punctation as that on head, denser in middle, with interspaces between punctures as long as about diameter of two-four nearest punctures, sometimes sparser in mediobasal and lateral portions; scutellum with several small smooth to deep punctures; punctation of elytra as that in middle part of pronotum, but distinctly denser and deeper and/or smaller on parascutellar portion and along suture in some specimens, with interspaces between punctures in middle as long as diameter of one or two nearest punctures, on each elytron not forming longitudinal rows of punctures and more or less irregularly scattered, or with very indistinct three to four vague and tangled longitudinal rows along suture; abdominal tergites without or with sparse and very indistinct small punctures. Body glabrous; clypeus with several long tactile setae; anterior part of frons with several irregular small setae; middle part of pronotum and each elytron with irregular, very sparse, small and erect setae; abdominal tergites with regular sparse and small setae. Habitus as in Fig. 1.</p><p>Head 1.5–1.8 times as wide as long; middle portion of head slightly flattened or irregularly elevated, with indistinct to distinct semicircular impression between eyes at posterior third, usually with distinct and deep elongate grooves in front of ocelli, reaching middle or anterior third of eye; postocular ridges smooth and indistinct in some specimens; anterior portion between antennal insertion and anterior margin of eye with moderately deep semicircular notch. Ocelli large, situated at level or slightly behind of level of postocular ridges; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Labrum with widely rounded membranous latero-apical portions, moderately deep apical emargination and long thin basolateral projections (Fig. 6). Mandibles short, with narrow basal and elongate apical portions, with indistinct small tooth in middle on right mandible (Fig. 8). Labium and labial palpomeres moderately narrow, submentum small, elongate (Fig. 12). Apical segment of maxillary palp slightly more than twice as long as preceding segment (Fig. 11). Gular sutures with widely rounded apical parts, narrowly diverging posteriad; shortest distance between sutures located at level posterior third of eyes (Fig. 14). Antenna moderately short, slightly exceeding shoulders of elytra when reclined; length × width of antennomeres (holotype of L. mesasiaticum): 1: 0.11 × 0.07; 2: 0.08 × 0.06; 3: 0.08 × 0.04; 4: 0.07 × 0.04; 5–7: 0.06 × 0.04; 8–9: 0.05 × 0.05; 10: 0.05 × 0.06; 11: 0.10 × 0.06.</p><p>Pronotum 1.7 times as wide as long, 1.4–1.5 times as wide as head, widest at or about middle, evenly or slightly more narrowed posteriad than anteriad; apical margin distinctly shorter than posterior margin, with straight or slightly rounded apical margin; anterior angles widely rounded, not or slightly protruded anteriad; posterior angles widely rounded; lateral edges of pronotum with irregular and small, flattened crenulation; pronotum with very wide elevated middle portion, with wide and deep longitudinal impression (indistinct in some), with pair of indistinct to moderately deep semioval impression on mediobasal third; lateral portions slightly explanate, each with deep oval pit in front of middle. Prothorax (Fig. 16) with elongate intercoxal process and moderately wide slightly protruded mesosternal processes. Mesothorax (Fig. 22) transverse, narrow in middle, with thin and acute intercoxal process reaching posterior margin of mesocoxae. Scutellum (Fig. 18) with widely rounded apex. Metathorax (Fig. 24) narrow, with wide and deep intercoxal cavities and narrow intercoxal process. Metendosternite (Fig. 26) with strongly reduced paired medioapical protrusions.</p><p>Elytra about as long as wide, long, markedly more than twice as long as pronotum, more or less subparallelsided and indistincly widened apicad from middle, reaching apical margin of tergite IV or V, with straight or slightly rounded apical margins, truncated at suture; shoulders of elytra widely rounded, not protruded apicad (Fig. 20); lateral portions of elytra narrow, slightly explanate, latero-apical margins with small, irregular and smooth crenulation; each elytron with indistinct transverse elevations between punctures in parascutellar portions or with distinct transverse and diagonal elevations, significantly coarser in mediobasal portions in some.</p><p>Legs moderately long and slender, tibiae covered by elongate setae, distinctly stronger on lateral margin, with a few strong setae on apical margins near apex; first four protarsomeres of both sexes dilated; metatarsus less than half as long as metatibia.</p><p>Abdomen about as wide as elytra or slightly narrower, with pair of rounded tomentose wing-folding spots in middle of tergite V.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly emarginate (Fig. 2 8–29). Aedeagus wide and suboval (Fig. 38–43; Shavrin 2013: Fig. 1–6), with widely rounded apex; parameres wide, slightly exceeding apex of median lobe, with four short apical setae; internal sac variable, consisting of numerous acute thorns, spiralled in basal part in some specimens. Aedeagus laterally see in Hlaváč et al. (2016: 2).</p><p>Female. Apical margin of abdominal tergite VIII vaguely sinuate (Fig. 30). Apical margin of abdominal sternite VIII rounded (Fig. 31). Genital segment as in Fig. 32.</p><p>Comparative notes. Anthobium fusculum differs from the remaining species of the fusculum group by the dark brown to black coloration of the body, by the shape of the suboval aedeagus, wide parameres, and by the structure of the internal sac.</p><p>Distribution. Anthobium fusculum is widely distributed in the western Palaearctic Region (Fig. 44), from the middle (Austria, Czech Rebublic, France, Germany, Italy, Switzerland), eastern (Belarus, Poland, Russia, Ukraine) and northern Europe (Denmark, Finland, Norway, Sweden) to Caucasus (Armenia), Middle East (Turkey, Iran) and Central Asia (Kasakhstan, Kyrgyzstan, Tajikistan, Uzbekistan, Pakistan and India (Kashmir)).</p><p>The record of the species from Spain (Smetana 2004; Schülke &amp; Smetana 2015) is most likely based on the result of old misidentification (see also Gamarra &amp; Outerelo 2008).</p><p>Bionomics. According to published data, A. fusculum can be regarded as hygrophilous species; it was noted as psammophilous and phytodetriticolous (Uhlig &amp; Zerche 1981; Koch 1989; Vogel 2013), stenotopic ripicolous alpine (Schatz 2007) and riparian species (Andersen &amp; Hanssen 2005). The species is recorded from intermediate elevations beginning from about 100 m a.s.l. to high elevations about 3000 m a.s.l. It inhabits riparian and forest habitats, where it can be found under stones, mosses, litter and debris. Some specimens were observed under bark of trees, occasionally taken from bear dung, mushrooms, etc. Many populations in the Alps and some specimens from Aksu-Dzhabagly State Nature Reserve (Kazakhstan) and Kondar Pass (Tajikistan) were collected near the snow. The beetles have been collected from March to midwinter in the European part of its distribution where it was noted as winter active species (Schatz 2007).</p><p>Remarks. Lathrimaeum fusculum was originally described from unspecified number of types from “Mark Brandenburg ”, northern Germany. More than 150 years from the description, the species has been many times recorded from Europe, included in the regional European keys (see above) and it was redescribed several times (Fairmaire &amp; Laboulbène 1849, Luze 1905, Zanetti 1987, etc.). Zerche (1991) noted that he studied specimens of A. fusculum from Uzbekistan (Chimgan) that in habitus were somewhat different from European specimens and remotely looked like some species of the genus Deliphrosoma Reitter, 1909 . The external variability of the species was a long-standing base for descriptions of new taxa, which were later synonymised with A. fusculum: Assing (2005) studied holotypes of L. melanochromum Iablokoff-Khnzorian, 1961 (described from Armenia) and Olophrum kondarense Iablokoff-Khnzorian, 1964 (described from Tajikistan), and synonymised them with A. fusculum; Shavrin (2013) studied holotype of L. kashmiricum Coiffait, 1982 (described from Kashmir, India), and, together with Deliphrosoma ivanovi Shavrin, 2012 and D. kirgizicum Shavrin, 2012, which were described from the Central Asia, synonymised them with A. fusculum . Another new synonymy is presented here. Lathrimaeum mesasiaticum Kirschenblat, 1961 was described from the holotype and two “cotypes” (all females) from “Stalinabad” (= Dushanbe; Tajikistan), “Pendzhiksit” (=Panjakent, Tajikistan) and “Golodnaya Step`” (Uzbekistan). The first author compared this species with L. fusculum and noted that L. mesasiaticum “...easily differs [from L. fusculum] by more long elytra, more gentle and denser punctation of pronotum and many other characters [translated from Russian]”. The first author studied these types in ZIN and found no morhological differences between these and other specimens of A. fusculum . The photograph of habitus (Fig. 7) of A. fusculum in Boháč (2016) shows Olophrum fuscum (Gravenhorst, 1806) .</p><p>The species is reported from Pakistan for the first time.</p></div>	https://treatment.plazi.org/id/03EB87F8F170FFB9FF31F9867F29494E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F17FFFBAFF31F88778264C96.text	03EB87F8F17FFFBAFF31F88778264C96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium altivagans (Cameron 1941)	<div><p>Anthobium altivagans (Cameron, 1941)</p><p>(Figs. 2, 50–51, 54)</p><p>Lathrimaeum (Prionothorax) altivagans Cameron, 1941: 59</p><p>Anthobium altivagans: Herman, 2001: 227</p><p>Anthobium (Prionothorax) altivagans: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 307</p><p>Type material examined: Holotype by monotypy (“Type in my collection”) ♂: ‘Type’ &lt;round printed label with red margin&gt;, ‘Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ &lt;rectangular printed label&gt;, ‘L. | altivagans | TYPE [in red] Cam.’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘M. Cameron | Bequest. | B.M. 1955-147.’ &lt;rectangular printed label&gt;, ‘ Anthobium | altivagans (Cameron, 1941) | Shavrin A.V. det. 2016’ &lt;rectangular printed label&gt; (BMNH).</p><p>Additional material: INDIA: KASHMIR: 1 ♂, 5 ♀♀: ‘Para- | type’ &lt;round printed label with red margin&gt;, ‘ Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ &lt;rectangular printed label&gt;, ‘ M. Cameron | Bequest. | B.M. 1955- 147.’ &lt;rectangular printed label&gt; (BMNH) ; 1 ♀: ‘ Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’, ‘ W. Steel coll. | B.M. 1969-532’ (BMNH) ; NEPAL: 1 ♀: SE Annapurna mts. Telbrung Danda. 3200 m a.s.l. 0 9.06.1997. O. Jäger leg. (SNSD) ; 1 ♀: SE Annapurna mts., Rambrong Danda. S-slope, 3700 m a.s.l. NN, Rhododendron forest. 0 7.06.1997. O. Jäger leg. (SNSD) ; 1 ♂: SW Ganesh Himal, Abuthurn Lekh upp. Keronja. 3600-4000 m a.s.l. 19- 20.05.1996. D. Ahrens, J. Kulbe, B. Rulik leg. (NME) .</p><p>Redescription. Measurements (n=10): HW: 0.72–0.75, HL: 0.42–0.50; AL(holotype): 1.33; OL: 0.19–0.22; PL: 0.52–0.62; PW: 1.12–1.30; ESL: 1.42–1.62; EW: 1.48–1.72; AW: 1.27–1.45; MTbL(holotype): 0.82, MTrL(holotype): 0.40 (MTrL 1–4: 0.25; MTrL 5: 0.15); AedL: 0.57; TL: 3.25–3.60(holotype).</p><p>In external appearance similar to A. monticola . Head, antennomeres 4–11, median part of pronotum and abdomen reddish brown to brown; elytra yellow-brown to brown; mouthparts, antennomeres 1–3, apical and basal margins and lateral portions of pronotum, and legs yellow. Head with irregular large punctation, markedly denser and deeper on median portion between eyes and on posterior half of infraorbital ridges; middle part of neck with irregular, deep punctures variable in size; pronotum with irregular punctation similar to that on middle part of head, sometimes smaller and denser on mediobasal portion, larger and sparser on lateral portions, with variable, more or less wide impunctated area after the middle of disc; scutellum with several very small punctures; punctation of elytra denser and significantly coarser than that on pronotum, smaller on parascutellar area, each elytron with six vague and tangled longitudinal rows of punctures, remaining elytral punctures irregularly scattered laterally (some specimens with markedly coarse surface between punctures without longitudinal rows of punctures); abdominal tergites without distinct punctation. Habitus as in Fig. 2.</p><p>Head 1.5–1.7 times as wide as long; vertex and posterior parts of infraorbital ridges slightly elevated; median elevation without or with indistinct to distinct transverse impression between eyes at posterior third, with elongate, diagonally stretching grooves in front of ocelli, reaching anterior third of eye; postocular ridges small and obtuse. Ocelli situated at level of postocular ridges; distance between ocelli slightly less than twice of distance between ocellus and posterior margin of eye. Apical segment of maxillary palp about two and half times as long as preceding segment, widest in basal third, gradually narrowing apicad (Fig. x). Length × width of antennomeres (holotype): 1: 0.17 × 0.08; 2: 0.11 × 0.05; 3: 0.15 × 0.05; 4–7: 0.11 × 0.05; 8–9: 0.10 × 0.06; 10: 0.10 × 0.07; 11: 0.16 × 0.08.</p><p>Pronotum widest at middle; anterior angles widely rounded, slightly protruded anteriad; middle elevation of pronotum with idistinct, wide longitudinal portion on medioapical third and indistinct to distinct transverse impression on mediobasal third.</p><p>Elytra slightly broader than long, gradually widened apicad, reaching apical margin of abdominal tergite IV; basal and middle portions of elytra with variable indistinct to distinct elevations between punctures or without them, some specimens with very indistinct three longitudinal elevations in middle.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 50) narrow, with subtruncate apex; parameres distinctly exceeding apex of median lobe, with two short apical and preapical setae; internal sac long and wide, spiralled in basal portion, with characteristic lateral long rows of large, strongly scletotized teeth. Aedeagus laterally as in Fig. 51.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the general shape of pronotum and elytra with the presence of longitudinal elevations between punctures, A. altivagans is most similar to A. monticola, from which it differs by the shape of anterior angles of the pronotum slightly protruded anteriad, and by the shape and internal structure of the internal sac of the aedeagus, armed by rows of sclerotized teeth.</p><p>Distribution. The species is known from Kashmir (India) and central Nepal (Fig. 54).</p><p>Bionomics. Specimens were collected at elevations from 2400 to 4000 m a.s.l. One specimen in Nepal (Rambrong Danda) was taken by sifting forest floor litter in Rhododendron forest. Remarks. Anthobium altivagans is here recorded for the first time from Nepal.</p></div>	https://treatment.plazi.org/id/03EB87F8F17FFFBAFF31F88778264C96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F17CFFA5FF31FB0D7D824E96.text	03EB87F8F17CFFA5FF31FB0D7D824E96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium besucheti Shavrin & Smetana 2018	<div><p>Anthobium besucheti Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 3, 52–53, 54)</p><p>Type material examined: Holotype ♂: ‘ NEPAL Parbat | Distr. Ridge E | Ghoropani Pass | 3100m 7. X. 1983 | [A.] Smetana &amp; [I.] Löbl’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | besucheti sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (MHNG) .</p><p>Paratypes (23 specimens): 6 ♂♂ [two specimens dissected], 6 ♀♀: same data as the holotype (5 ♂, 5 ♀: MHNG; 1 ♂, 1 ♀: CS); 1 ♂: ‘ NEPAL: Lamjung, S- | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.34917&amp;materialsCitation.latitude=28.3825" title="Search Plazi for locations around (long 84.34917/lat 28.3825)">Sundar Danda</a>, 3140m, | 28°22'57''N, 84°20'57''E, | 21.IX.2013 | leg. [J.]Hagge &amp; [J.]Schmidt’ (NME) ; 1 ♀: ‘E. NEPAL: KOSI [Khandbari District] | Goru Dzure Dara | 3600 m, 9.IV.84 | [I.]Löbl-[A.]Smetana’ (MHNG) ; 4 ♂♂ [one specimen dissected], 5 ♀♀: ‘NEPAL—Goropani For. | Crête Est Goropani | 3100m 7.X.1983 | [I.]Löbl—[A.]Smetana’ (MHNG) . All paratypes with additional red rectangular printed label: ‘ PARATYPE | Anthobium | besucheti sp.n. | Shavrin A. &amp; Smetana A. des. 2018’.</p><p>Description. Measurements (n=24): HW: 0.55–0.57, HL: 0.32–0.36; AL(holotype): 0.93; OL: 0.12–0.14; PL: 0.41–0.48; PW: 0.81–0.95; ESL: 0.87–1.02; EW: 1.07–1.15; AW: 0.80–0.96; MTbL(holotype): 0.52; MTrL(holotype): 0.22 (MTrL 1–4: 0.14; MTrL 5: 0.08); AedL: 0.42; TL: 1.95–2.74 (holotype: 2.40).</p><p>Body elongate, convex. Body yellow to yellow-brown, some specimens with reddish-brown head, antennomeres 7–11 and abdomen; mouthparts, antennomeres 1–6 and legs yellow. Posterior portions of infraorbital ridges and area between ocelli with slightly rugose transverse sculpture between punctures, neck and abdominal tergites with distinct isodiametric microsculpture. Head with irregular, small and deep punctation, denser on posterior portion; middle part of neck with irregular and sparse punctation; pronotum with irregular punctation, markedly denser, larger and coarser than that on posterior part of head, sparser on lateral portions and posterior third of middle elevation, some specimens with small impunctate area between mediobasal impression and/or on basolateral portions; visible part of scutellum impunctated or with several small punctures; punctation of elytra dense, distinctly larger and deeper than that on pronotum, smaller and denser on parascutellar area, each elytron with five to eight very vague and tangled longitudinal rows of punctures, replaced by confluent punctures in some paratypes; abdominal tergites without visible punctures or with small and dense punctation on abdominal tergites VII–VIII. Habitus as in Fig. 3.</p><p>Head 1.5–1.7 times as wide as long; middle portion of head between eyes slightly elevated, usually with middle transverse impression between eyes at posterior third, with indistinct to distinct small and narrow grooves in front of ocelli, sometimes reaching middle length of eyes; postocular ridges usually smooth; anterior portion between antennal insertion and anterior margin of eye with wide and shallow semicircular notch. Ocelli small, situated at level of postocular ridges; distance between ocelli one and half times as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp about three times as long as preceding segment, widest in basal part and conically narrowed toward acute apex. Antenna moderately short, reaching shoulders of elytra when reclined; apical antennomeres widened, as wide as basal antennomere; length × width of antennomeres (holotype): 1: 0.13 × 0.05; 2: 0.07 × 0.04; 3–4: 0.07 × 0.03; 5–6: 0.08 × 0.03; 7: 0.07 × 0.03; 8: 0.07 × 0.04; 9: 0.07 × 0.05; 10: 0.07 × 0.05; 11: 0.15 × 0.05.</p><p>Pronotum about twice as wide as long, 1.4–1.6 times as wide as head, widest in front of middle, indistinctly and evenly narrowed both posteriad and anteriad; apical margin slightly shorter than posterior margin, widely rounded on anterior margin; anterior angles widely rounded, slightly protruded anteriad, posterior angles obtuse; lateral edges of pronotum with irregular, small and smooth crenulation; pronotum with very wide, markedly elevated middle portion, with very indistinct to distinct and deep longitudinal impression on anterior half of middle elevation, with pair of distinct, slightly or moderately deep transverse impressions on mediobasal third.</p><p>Elytra slightly broader than long, about twice as long as pronotum, slightly widened apicad, reaching apical margin of abdominal tergite IV or V; each elytron in middle with irregular, distinct elevations between punctures, forming indistinct two or three longitudinal elevations in some specimens.</p><p>Abdomen with indistinctly elevated paratergites.</p><p>Male. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (Fig. 52) with median lobe moderately wide basally, gradually narrowed toward subtruncate apex; parameres wide, significantly exceeding apex of median lobe, with two small apical and preapical setae; internal sac very narrow and long. Aedeagus laterally as in Fig. 53.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the body size, coloration, shape of the body with moderately short elytra, A. besucheti Shavrin &amp; Smetana, sp.n. is most similar to A. denticulatum Shavrin &amp; Smetana, sp.n., from which it differs by the slightly wider pronotum, by the narrower and smaller aedeagus with internal sac missing sclerotized teeth.</p><p>Distribution. The species is known from several localities in central and eastern Nepal (Fig. 54).</p><p>Bionomics. Specimens were collected at elevations from 3100 to 3600 m a.s.l. Specimens from Parbat District were taken by sifting of Carex mixed with other vegetation along edges of the pond in an Abies and Rhododendron forest; a male from Sundar Danda (Nepal) was taken by sifting leaf litter in Rhododendron - Abies forest, together with A. monticola (see below). Specimens from Goru Dzure Dara were collected together with A. cavicrus (see below) and were taken by sifting of various debris, moss and Rhododendron leaf litter in a small gorge with still remnants of snow.</p><p>Etymology. Patronymic, the species is named to honour our colleague, coleopterologist Claude Besuchet (Genève, Switzerland).</p></div>	https://treatment.plazi.org/id/03EB87F8F17CFFA5FF31FB0D7D824E96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F163FFA7FF31F9007E6C4F43.text	03EB87F8F163FFA7FF31F9007E6C4F43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium capitale Shavrin & Smetana 2018	<div><p>Anthobium capitale Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 4, 57–59)</p><p>Type material examined: Holotype ♂ [specimen dissected]: ‘ CHINA: N-Yunnan Diqing Tibet | Aut.Pr.Deqin Co. Meli Xue | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.7725&amp;materialsCitation.latitude=28.457834" title="Search Plazi for locations around (long 98.7725/lat 28.457834)">Shan</a> E-side 14km W Deqin | 28°27.47'N 98°46.35'E 2580m | 11.VI.2005 A.Smetana [C158]’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | capitale sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (NSMT) .</p><p>Paratype ♀: same data as the holotype, ‘ PARATYPE | Anthobium | capitale sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;rectangular label, printed&gt; (CS) .</p><p>Description. Measurements (n=2): HW: 0.62–0.65, HL: 0.42; AL(holotype): 1.05; OL: 0.17; PL: 0.50; PW: 0.97; ESL: 1.37; EW: 1.27–1.30; AW: 1.06–1.17; MTbL(holotype): 0.67, MTrL(holotype): 0.35 (MTrL 1–4: 0.23; MTrL 5: 0.12); AedL: 0.68; TL: 2.85(holotype)–2.95.</p><p>Body moderately elongate, subconvex. Head reddish brown; pronotum, elytra and abdomen yellow-brown; antennomeres 5–11 brown; mouthparts, antennomeres 1–4 and legs yellow. Head between supra-antennal elevation and anterior margin of eye with diagonal wavy meshes and coarse sculpture on posterior portion between eyes and ocelli; middle part of neck and abdominal tergites with distinct isodiametric microsculpture. Head with irregular, small and moderately deep punctation, markedly coarser on posterior portions of infraorbital ridges; middle part of neck with small, more or less regular punctation, markedly coarser near occipital furrow; pronotum with irregular, large and deep punctation, denser on apical half of middle elevation, sparser laterally, smaller along mediobasal margin, with small impunctate area between mediobasal impression; punctation of elytra dense and deep, smaller on parascutellar area and along suture, each elytron with six vague and tangled longitudinal rows of punctures, remaining elytral punctures forming very indistinct diagonal rows and becoming irregularly scattered laterally; abdominal tergites with indistinct, sparse and small punctation. Habitus as in Fig. 4.</p><p>Head 1.4–1.5 times as wide as long; middle portion of head slightly and irregularly elevated, with middle transverse triangular impression between eyes at posterior third, with narrow grooves in front of ocelli, reaching about middle length of eyes or slightly shorter (holotype); postocular ridges smooth; anterior portion of head between antennal insertion and anterior margin of eye with uneven, semicircular notch slightly deeper apically. Ocelli very large, situated at level of postocular ridges; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp two and half times as long as preceding segment, widest in basal part, gradually narrowed apicad. Antenna reaching anterior third of elytra when reclined, with antennomeres gradually widened apicad; length × width of antennomeres (holotype): 1: 0.12 × 0.06; 2: 0.10 × 0.05; 3: 0.11 × 0.04; 4–5: 0.10 × 0.05; 6: 0.09 × 0.05; 7–8: 0.08 × 0.05; 9: 0.08 × 0.06; 10: 0.07 × 0.06; 11: 0.12 × 0.06.</p><p>Pronotum slightly less than twice as wide as long, 1.5 times as wide as head, widest at middle, slightly more narrowed posteriad than anteriad; apical margin as wide as posterior margin; anterior angles moderately rounded, not protruded anteriad; posterior angles obtuse; lateral edges of pronotum with irregular, smooth crenulation; pronotum with very wide, markedly elevated middle portion, with indistinct (holotype) to distinct longitudinal impression on apical part of middle elevation, with deep and wide semioval impression on mediobasal third.</p><p>Elytra longer than wide, distinctly less than three times as long as pronotum, slightly widened apicad, reaching basal margin of abdominal tergite V; each elytron in middle with irregular, longitudinal elevations between third to sixth longitudinal row of punctures, small parascutellar area of each elytron slightly convex.</p><p>Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (Fig. 57) elongate, with widely trucated apex; parameres significantly exceeding apex of median lobe, with two very short apical and preapical setae; internal sac narrow and long, with two characteristic lateral rows of large sclerotized teeth. Aedeagus laterally as in Fig. 58.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Anthobium capitale Shavrin &amp; Smetana, sp.n. differs from remaining species of the monticola group by the elytra slightly longer than wide, as well as by the shape and internal structure of the aedeagus.</p><p>Distribution. The new species is at present known only from the type locality (Fig. 59) in the Meili-Xue Shan range in Yunnan, China.</p><p>Bionomics. Specimens were taken by sifting wet leaf litter, debris and moss on and around small seepages and on edges of a small creek in a mixed broadleaved forest (locality: C158).</p><p>Etymology. The specific epithet is the Latin adjective capitalis, - e (concerning the head). It brings attention to the fact that the head is of different colour.</p></div>	https://treatment.plazi.org/id/03EB87F8F163FFA7FF31F9007E6C4F43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F160FFA3FF31FBB278CE4AA8.text	03EB87F8F160FFA3FF31FBB278CE4AA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium cavicrus (Champion 1925)	<div><p>Anthobium cavicrus (Champion, 1925)</p><p>(Figs. 5, 60–61, 64)</p><p>Lathrimaeum cavicrus Champion, 1925: 102</p><p>Lathrimaeum cavicrus: Cameron, 1930: 153; Scheerpeltz, 1933: 1055</p><p>Lathrimaeum (Lathrimaeum) cavicrus: Scheerpeltz, 1961: 79</p><p>Anthobium cavicrus: Herman, 2001: 230</p><p>Anthobium (Anthobium) cavicrus: Smetana, 2004: 238; Schülke &amp; Smetana, 2015: 306</p><p>Type material examined: Lectotype of Lathrimaeum cavicrus Champion, 1925 ♂ [beetle is glued on rectangular plate]: ‘SYN- | TYPE’ &lt;round label with blue margin, printed&gt;, ‘Type | H. T.’ &lt;round label with red margin, printed&gt;, ‘ ♂ ’ &lt;rectangular label, printed&gt;, ‘2465’ &lt;rectangular label, printed&gt;, ‘W. Almora Divn | Kumaon U. P. | Oct. 1918. HGC.’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | cavicrus Ch. | type ♂ ’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘ Lathrimaeum | championi Bern.’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘ Lathrimaeum | cavicrus, | Champ.’ &lt;rectangular label, printed&gt;, ‘ LECTOTYPE | Lathrimaeum | cavicrus Champion, 1925 | Shavrin A.V. des. 2016’ &lt;red rectangular label, printed&gt;, ‘ Anthobium | cavicrus (Champion, 1925) | Shavrin A.V. det. 2016’ &lt;rectangular label, printed&gt; (BMNH)</p><p>Paralectotype of Lathrimaeum cavicrus Champion, 1925 ♀ [beetle is glued on rectangular plate]: ‘SYN- | TYPE’ &lt;round label with blue margin, printed&gt;, ‘ ♂ &lt;rectangular label, printed&gt;, ‘Kumaon U. P. | Oct. [handwritten] 1918. HGC. | W. Almora Divn &lt;rectangular label, printed&gt;, ‘2465’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | cavicrus Ch. ’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘G.C.Champion | Brit. Mus. | 1925-42.’ &lt;rectangular label, printed&gt;, ‘ PARALECTOTYPE | Lathrimaeum | cavicrus Champion, 1925 | Shavrin A.V. des. 2016’ &lt;red rectangular label, printed&gt;, ‘ Anthobium | cavicrus (Champion, 1925) | Shavrin A.V. det. 2016’ &lt;rectangular label, printed&gt; (BMNH); 1 ♀: ‘SYN- | TYPE’ &lt;round label with blue frame, printed&gt;, ‘ ♀ ’ &lt;rectangular label, printed&gt;, ‘W. Almora | Kumaon, | India. H.G.C.’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | cavicrus Ch. ’ &lt;rectangular label, handwritten&gt;, ‘ Lathrimaeum | championi Ch. | n.sp.’ &lt;rectangular label, handwritten&gt;, ‘ Lathrimaeum | cavicrus, | Champ.’ &lt;rectangular label, printed&gt;, ‘E.M.M. 1925. | det. G.C.C.’ &lt;rectangular label, printed&gt;, ‘G.C. Champion. | Brit. Mus. | 1925-42.’ &lt;rectangular label, printed&gt; (BMNH).</p><p>Additional material. INDIA: PUNJAB PROVINCE: 1 ♀: ‘Jalori Pass, Seraj Punjab. 10,800ft. H.G.C.’, ‘ Lathrimaeum ? n. sp.’, ‘H.G. Champion Coll. B.M. 1953-156’, ‘ Lathrimaeum ? cavicrus Champ. P.M. Hammond det. 1969’ (BMNH); UTTAR PRADESH PROVINCE: 1 ♀: Garhwal, 4 km au Sud de Bhatwari. 1400 m a.s.l. 23.10.1979. I. Löbl leg. (MHNG); 1 ♀: Garhwal, au-dessus Pauri. 1900 m a.s.l. 28.10.1979. I. Löbl leg. (MHNG); WEST BENGAL PROVINCE: 1 ♀: Darjeeling dist. Tigerhill. 18,10.1978. 2500-2600 m a.s.l. Besuchet &amp; Löbl leg. (MHNG); NEPAL: 1 ♂, 1 ♀: Kosi, Goru Dzure Dara, 0 9.04.1984. 3600 m a.s.l. I. Löbl &amp; A. Smetana leg. (MHNG); 1 ♀: Mere Dara. 3100-3300 m a.s.l. 0 7.04.1981. I. Löbl &amp; A. Smetana leg. [northern slope of a pass with solitary Abies and Rhododendron growth, still remnants of snow. Under rocks] (NSMT); 1 ♀: same data. 3200 m a.s.l. 0 8.04.1981. I. Löbl &amp; A. Smetana leg. 14A [mixed Abies and Rhododendron forest. Sifting of of wet Rhododendron leaf litter, moss and various debris on a seepage] (NSMT); 1 ♀: near Mere Dara. 3000 m a.s.l. 0 7.04.1981. I. Löbl &amp; A. Smetana leg. [Old Rhododendron forest, shady and cool, sifting of leaf litter and various debris] (MHNG); 1 ♂, 1 ♀: below Thare Pati. 3500 m a.s.l. 12.04.1981. I. Löbl &amp; A. Smetana leg. [ Acer enclave in a Rhododendron forest on a northern slope, still with snow around, sifting of maple leaf litter mixed with moss] (CS); 2 ♂♂, 1 ♀: Ridge between Mere Dara and Thare Pati. 3500 m a.s.l. 0 9.04.1981. I. Löbl &amp; A. Smetana leg. [mountain pasture, under rocks] (MHNG).</p><p>Redescription. Measurements (n=17): HW: 0.60–0.67, HL: 0.37–0.42; AL(lectotype): 1.03; OL: 0.15–0.19; PL: 0.41–0.57; PW: 0.87–1.02; ESL: 1.15–1.35; EW: 1.15–1.30; AW: 1.05–1.30; MTbL(lectotype): 0.65; MTrL(lectotype): 0.35 (MTrL 1–4: 0.23; MTrL 5: 0.12); AedL: 0.45–0.46; TL: 2.26–3.25(lectotype).</p><p>Body moderately elongate, subconvex. Body reddish brown to brown; lateral portions and basal margin of pronotum and some specimens with shoulders of elytra yellow-brown; antennomeres 9–11 brown; mouthparts, antennomeres 1–8 and legs yellow. Middle of vertex of head with distinct transverse microsculpture (some specimens), posterior portions of infraorbital ridges with slightly rugose sculpture; abdomen with fine isodiametric microsculpture. Head with irregular, large and deep punctation, markedly denser on middle elevation and coarser on posterior parts of infraorbital ridges; middle part of neck with dense punctation, as that on middle of head; pronotum with irregular dense punctation, as that on posterior part of head, sparser on lateral portion, smaller on medioapical and mediobasal margins; scutellum with several small punctures or without them; punctation of elytra larger and markedly denser than that on pronotum, smaller on parascutellar area, not forming longitudinal rows of punctures, interspaces between punctures in middle as long as about diameter of two nearest punctures; abdominal tergites with very indistinct smooth punctation. Habitus as in Fig. 5.</p><p>Head 1.5–1.6 times as wide as long; middle portion of head slightly elevated, middle with indistinct to distinct transverse impression between eyes at posterior third, with indistinct or distinct and moderately deep (specimens with more elevated middle portion of head) small grooves in front of ocelli, reaching level of posterior third of eye; postocular ridges smooth; anterior portions between antennal insertion and anterior margin of eye with wide superficial notch. Ocelli large, situated at level of postocular ridges; distance between ocelli one and a half times as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp two and half times as long as preceding segment, widest in basal third, gradually narrowed toward acute apex. Antenna slightly exceeding shoulders of elytra when reclined; length × width of antennomeres (lectotype): 1: 0.13 × 0.06; 2: 0.10 × 0.05; 3: 0.12 × 0.03; 4: 0.07 × 0.04; 5–9: 0.08 × 0.05; 10: 0.08 × 0.06; 11: 0.13 × 0.07.</p><p>Pronotum about twice as wide as long, 1.4–1.5 times as wide as head, widest about middle, more or less evenly narrowed posteriad and anteriad; apical margin distinctly narrower than posterior margin; anterior angles widely rounded, not or indistinctly protruded anteriad; posterior angles widely rounded or obtuse; lateral edges of pronotum with regular and small, acute or smooth crenulation; pronotum with wide middle portion markedly elevated, with distinct deep longitudinal impression located on apical part of elevation, with pair of deep semioval impressions on mediobasal third.</p><p>Elytra about as long as wide, about two and half times as long as pronotum, slightly widened apicad, reaching apical margin of abdominal tergite IV or V, with widely rounded apical margins, some specimens with small parascutellar area of each elytron slightly convex.</p><p>Abdomen as wide as elytra or slighty narrower, with pair of small or moderately large, rounded tomentose wing-folding spots in middle of tergite V.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 60) long and moderately narrow, median lobe markedly narrowed at about apical fourth, with wide, vaguely angulate apex; parameres wide, significantly exceeding apex of median lobe, with two small apical and preapical setae; internal sac narrow and long. Aedeagus laterally as in Fig. 61.</p><p>Female. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the irregularly arranged punctation of elytra, not forming longitudinal rows, A. cavicrus is most similar to the Chinese A. liliputense sp.n., from which it differs by darker coloration, larger body, narrower pronotum, elytra and aedeagus with significantly shorter internal sac.</p><p>Distribution. The species is known from Punjab, Uttarakhand and West Bengal provices of northern India and from central and eastern Nepal (Fig. 64).</p><p>Bionomics. Specimens were collected at elevations from 1400 to 3600 m a.s.l. Specimens from Mere Dara were collected under rocks or taken by the sifting of leaf litter, moss and various debris in mixed Abies and/or Rhododendron forest. Two specimens from Goru Dzure Dara (Nepal) were collected together with A. besucheti Shavrin &amp; Smetana, sp.n. (see above).</p><p>Remarks. The species was compared by Champion with A. rugosum, A. monticola and Eudeliphrum gracilipalpe Champion, 1920 . Anthobium cavicrus is here recorded for the first time from Nepal.</p></div>	https://treatment.plazi.org/id/03EB87F8F160FFA3FF31FBB278CE4AA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F165FFA2FF31F99979904FB7.text	03EB87F8F165FFA2FF31F99979904FB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium confucii Shavrin & Smetana 2018	<div><p>Anthobium confucii Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 45, 62–63, 65)</p><p>Type material examined: Holotype ♂: ‘ CHINA, Sichuan, Gongga | Shan, Lake abv. Camp 2 | 2750m, 25.VII.1994 | A.Smetana [C23]’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | confucii sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (NSMT) .</p><p>Paratypes (7 specimens): CHINA: SICHUAN: 1 ♂ [specimen dissected], 1 ♀: same data as the holotype (1 ♂: CS; 1 ♀: NSMT); 1 ♂ [specimen dissected]: ‘ CHINA: W-Sichuan 20. VI. 1999 | Ya `an Prefecture, Tianguan Co. | E Erlang Shan Pass, 2900 m | 9 km SE Luding, 29°52N, | 102°18E, Bachufer, Moss + | Schotter, leg. M. Schülke’ &lt;rectangeular label, printed&gt;, ‘ Sammlung | M. Schülke | Berlin’ &lt;green rectangular label, printed (CSC) ; HUBEI: 1 ♂ [specimen dissected]: ‘ CHINA: W-Hubei (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.36667&amp;materialsCitation.latitude=31.5" title="Search Plazi for locations around (long 110.36667/lat 31.5)">Daba Shan</a>) | creek valley 11 km NW Muyuping, | 31°30'N, 110°22'E, 1960 m, | 18. VII. 2001, | leg. M. Schülke [C01-17]’ &lt;rectangular label, printed&gt;, ‘creek valley, mixed deciduous | forest (sifted) [C01-17]’ &lt;rectangular label, printed&gt;, ‘ Sammlung | M. Schülke | Berlin’ &lt;rectangular label, printed&gt; (CSC) ; 1 ♀ [without right elytron]: ‘ CHINA: W-Hubei (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.36667&amp;materialsCitation.latitude=31.483334" title="Search Plazi for locations around (long 110.36667/lat 31.483334)">Daba Shan</a>) | creek valley 8 km NW Muyuping, | 31°29' N, 110°22' E, 1550- | 1650 m, 18. VII. 2001, | leg. M. Schülke [C01-16A]’ &lt;rectangular label, printed&gt;, ‘creek valley, deciduous | forest, moss | (sifted) [C01-16A]’ &lt;rectangular label, printed&gt; (CSC) ; 2 ♀♀: ‘ CHINA: W-Hubei (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.35&amp;materialsCitation.latitude=31.5" title="Search Plazi for locations around (long 110.35/lat 31.5)">Daba Shan</a>) | pass E of Mt. Da Shennongjia, | 12 km NW Muyuping, 31°30' N, | 110°21' E, 22. VII. 2001, | leg. M. Schülke [C01-13E]’ &lt;rectangular label, printed&gt;, ‘dry creek valley, mixed deciduous | forest, dead wood, mushrooms, | moss, 1950–2050 m (sifted) | [C01-13E]’ (1 ♀: CS; 1 ♀: CSC). All paratypes with additional red rectangular printed label : ‘ PARATYPE | Anthobium | confucii sp.n. | Shavrin A. &amp; Smetana A. 2018 ’.</p><p>Description. Measurements (n=7): HW: 0.72–0.75, HL: 0.42–0.45; AL(holotype): 1.08; OL: 0.20–0.22; PL: 0.52–0.55; PW: 1.17–1.24; ESL: 1.42–1.50; EW: 1.55–1.59; AW: 1.15–1.17; MTbL(holotype): 0.60, MTrL(holotype): 0.32 (MTrL 1–4: 0.21; MTrL 5: 0.11); AedL: 0.45–0.50; TL: 3.00–3.45 (holotype: 3.15).</p><p>In external appearance similar to A. monticola . Head and antennomeres 4–11 brown; middle portion of pronotum, elytra and abdomen reddish brown to brown; lateral portions of pronotum and sometimes of elytra, apical segments of abdomen yellow to yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Basolateral portions of head on posterior parts of infraorbital ridges and between ocelli with rugose transverse microsculpture in some specimens; middle part of neck with traces of indistinct isodiametric microsculpture. Middle portion of head with irregular small punctation, punctation between ocelli and on infraorbital ridges distinctly larger and deeper, rugose on posterior half of infraorbital ridges; pronotum with irregular, dense and deep punctation, sparser and smaller on mediobasal third, some specimens with small impunctate area between mediobasal impression; punctation of elytra slightly denser, markedly larger and deeper than that on pronotum, smaller on parascutellar portions and in specimen from Erlang Shan along suture. Habitus as in Fig. 45.</p><p>Head 1.6–1.7 times as wide as long; middle portion of head slightly elevated, with deep transverse impression between eyes at posterior third; anterior portion between antennal insertion and anterior margin of eye with moderately deep semicircular notch. Ocelli situated slightly behind level of acute postocular ridges. Length × width of antennomeres (holotype): 1: 0.12 × 0.07; 2–3: 0.10 × 0.04; 4: 0.10 × 0.05; 5: 0.09 × 0.05; 6–8: 0.09 × 0.06; 9: 0.08 × 0.06; 10: 0.07 × 0.07; 11: 0.15 × 0.07.</p><p>Pronotum more than twice as wide as long, 1.6 times as wide as head, widest in middle, more narrowed posteriad than anteriad; anterior angles distinctly protruded anteriad; posterior angles acute; pronotum with markedly convex middle portion, with indistinct longitudinal impression on medioapical part and deep semioval impression on mediobasal third.</p><p>Elytra slightly broader than long, markedly widened apicad, with wide and distinctly explanate lateral portions; middle part of each elytron with indistinct longitudinal elevations between punctures.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 62) with median lobe wide basally and gradually narrowed toward subtruncate apex; parameres distinctly exceeding apex of median lobe; internal sac wide and very long, spiralled in basal portion. Aedeagus laterally as in Fig. 63.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the shapes of pronotum and elytra, A. confucii Shavrin &amp; Smetana, sp.n. is most similar to Himalayan A. altivagans and A. monticola, and Chinese A. laozii Shavrin &amp; Smetana, sp.n. From A. altivagans it differs by the shape of anterior angles of pronotum strongly protruded anteriad; from A. monticola by acute posterior angles of pronotum and from A. laozii sp.n. by darker coloration of the forebody. In addition, it differs from A. monticola and A. laozii sp.n. by slightly wider elytra. From all these species, A. confucii sp.n. differs by indistinct microsculpture of basolateral portions of head, by larger punctation of elytra, by the lateral portion of pronotum more narrowed posteriad than anteriad, and by details of both external and internal structures of the aedeagus.</p><p>Distribution. The new species is known from several locations in Gongga Shan and Erlang Shan ranges in Sichuan and Daba Shan range in Hubei, China (Fig. 65).</p><p>Bionomics. Specimens from Hubei were collected at elevations from 1550 to 2900 m a.s.l. by sifting wet moss and grassy vegetation on large fallen trees close to the water near shallow lake in a primary deciduous forest (location: C23) and by sifting moss, dead wood and mushrooms in mixed deciduous forests.</p><p>Etymology. Patronymic, the species is named in honour of the great ancient Chinese philosopher Confucius.</p></div>	https://treatment.plazi.org/id/03EB87F8F165FFA2FF31F99979904FB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F16BFFAFFF31FBD37D354BAB.text	03EB87F8F16BFFAFFF31FBD37D354BAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium cuccodoroi Shavrin & Smetana 2018	<div><p>Anthobium cuccodoroi Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 46, 64, 66–67)</p><p>Type material examined: Holotype ♂ [specimen dissected; right antennomeres 6–11 are glued on the side of beetle]: ‘ INDE Garhwal (UP[Uttar Pradesh]) | 4 km au Sud de [handwritten] | Bhatwari 1400 m [handwritten] | I. Löbl 23.[handwritten]X.[19]79’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | cuccodoroi sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (MHNG) .</p><p>Paratypes: 3 ♀♀: same data as the holotype, ‘ PARATYPE | Anthobium | cuccodoroi sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;rectangular label, printed&gt; (2 ♀♀: MHNG; 1 ♀: CS) .</p><p>Description. Measurements (n=4): HW: 0.63–0.67, HL: 0.43–0.49; AL(holotype): 0.95; OL: 0.17–0.20; PL: 0.55–0.61; PW: 0.92–1.02; ESL: 1.27–1.36; EW: 1.22–1.36; AW: 1.07–1.12; MTbL(holotype): 0.65, MTrL(holotype): 0.35 (MTrL 1–4: 0.22; MTrL 5: 0.13); AedL(holotype): 0.62; TL: 2.75(holotype)–3.48.</p><p>Body moderately elongate, subconvex. Body and antennomeres 5–11 yellow-brown; head reddish brown; mouthparts, antennomeres 1–4, basal, lateral and apical margins of pronotum, anteriolateral portions of elytra, legs, paratergites and apical segments of abdomen yellow. Middle part of vertex with smoothed transverse meshes, middle portion of head between middle impression with irregular transverse meshes, lateral portions of head with coarsely rugose sculpture, significantly stronger on median and posterior portions of infraorbital ridges, with moderately strong transverse wrinkles on posterior part; dorsal part of neck with large cellular microsculpture; medioapical part of pronotum with transverse moderately coarse microsculpture larger than that on middle part of head; basal portion of scutellum with fine wavy microreticulation; lateral portion of pronotum and elytra without microsculpture; abdominal tergites with fine isodiametric microsculpture. Frons and middle portion of head with sparse, moderately small and deep, irregular punctation, denser and larger on rugose surface of infraorbital ridges; middle part of neck with more or less regular, large and deep punctation; pronotum with dense irregular punctation, distinctly larger and deeper on lateral portions; visible part of scutellum with several large and deep punctures; punctation of elytra as that on pronotum, markedly denser on parascutellar area and larger on medioapical portion, each elytron near suture with four vague and tangled longitudinal rows of punctures, adjacent median punctures, separated by impunctated longitudinal surface, forming indistinct and tangled rows, becoming irregularly scattered laterally; abdominal tergites with indistinct small punctures. Habitus as in Fig. 46.</p><p>Head 1.3–1.4 times as wide as long; vertex irregularly or evenly elevated, some specimens with middle transverse triangular impression between posterior margins of eyes (one paratype with well-developed triangular elevations behind impression); deep and moderately short grooves in front of ocelli reaching posterior margin of eye; postocular ridges moderately acute; anterior portion between antennal insertion and anterior margin of eye with small, superficial rounded notch. Ocelli very large and convex, situated at level of postocular ridge; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp two and half times as long as preceding segment, from markedly widened middle gradually narrowed to apex. Antenna slightly exceeding shoulders of elytra when reclined; length × width of antennomeres (holotype): 1: 0.13 × 0.06; 2: 0.10 × 0.04; 3: 0.11 × 0.04; 4–7: 0.08 × 0.04; 8: 0.07 × 0.05; 9: 0.06 × 0.06; 10: 0.05 × 0.06; 11: 0.11 × 0.06.</p><p>Pronotum 1.6 times as wide as long, 1.4–1.5 times as wide as head, widest about middle, markedly more narrowed posteriad than anteriad; apical margin distinctly narrower than posterior margin; anterior angles widely rounded, slightly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with serrate, regular and small crenulation; pronotum with wide middle elevation, with irregular and indistinct longitudinal impression on anterior part, and with a pair of moderately deep and large oval impressions on mediobasal third and moderately deep irregular impressions of very deep lateral pits.</p><p>Elytra narrow, slightly widened apicad, reaching apical margin of abdominal tergite IV or V, as long as wide or slightly longer, more than twice as long as pronotum; each elytron with slightly elevated middle portions in basal part.</p><p>Male. First four protarsomeres slightly dilated. Apical margin of abdominal tergite VIII slightly, that of sternite VIII distinctly emarginated. Aedeagus (Fig. 66) robust, with median lobe widened apicad toward wide, irregularly truncate apex; parameres robust, each widened around middle, significantly exceeding apex of median lobe, with two short apical and preapical setae; internal sac with elongate field of thorns, rolled in basal portion. Aedeagus laterally as in Fig. 67.</p><p>Female. First four protarsomeres not dilated. Apical margin of abdominal tergite VIII and sternite VIII straigth to rounded.</p><p>Comprarative notes. Based on the general shape of pronotum and elytra, and on the presence of distinct transverse microsculpture on medioapical part of the pronotum, A. cuccodoroi Shavrin &amp; Smetana, sp.n. is most similar to the western Palaearctic A. fusculum, from which it differs by the paler coloration of the slightly wider body with somewhat more transverse pronotum, by more elongate aedeagus with significantly longer, around middle widened parameres, and by the structure of the internal sac.</p><p>Distribution. The new species is at present known only from the type locality (Fig. 64) in Uttarakhand, northern India.</p><p>Bionomics. All specimens were taken by sifting forest floor litter at elevation 1400 m a.s.l.</p><p>Etymology. Patronymic, the species is named to honour our colleague, Giulio Cuccodoro (Genève, Switzerland).</p></div>	https://treatment.plazi.org/id/03EB87F8F16BFFAFFF31FBD37D354BAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F169FFAEFF31FC017EE44B8E.text	03EB87F8F169FFAEFF31FC017EE44B8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium denticulatum Shavrin & Smetana 2018	<div><p>Anthobium denticulatum Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 47, 54–56)</p><p>Type material examined: Holotype ♂: ‘ NEPAL, D: Kaski, Wl. | Madi Khola valley | above Sikles, 3200m’ &lt;rectangular label, printed&gt;, ‘ 28°22'48''N, 84°03' | 54''E, 13.IX.2013, leg. | [J.] Hagge &amp; [J.] Schmidt’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | denticulatum sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (NME) .</p><p>Paratypes (19 specimens): 6 ♂♂ [three specimens dissected], 13 ♀♀: same data as the holotype, ‘ PARATYPE | Anthobium | denticulatum sp.n. | Shavrin A. &amp; Smetana A. 2017 ’ &lt;red rectangular label, printed&gt; (5 ♂, 11 ♀: NME; 1 ♂, 2 ♀: CS) .</p><p>Description. Measurements (n=20): HW: 0.55–0.58; HL: 0.33–0.40; AL(holotype): 0.85; OL: 0.12–0.14; PL: 0.42–0.49; PW: 0.78–0.87; ESL: 0.85–0.93; EW: 0.96–1.12; AW: 0.86–0.97; MTbL: 0.45, MTrL: 0.20 (MTrL 1–4: 0.13; MTrL 5: 0.07); AedL: 1.10; TL: 2.30–3.30 (2.70– holotype).</p><p>In external appearance similar to A. besucheti . Body and antennomeres 9–11 yellow-brown to reddish brown, with darker head and abdomen in some specimens; mouthparts, antennomeres and legs yellow to yellow-brown. Head with distinct microsculpture, transverse in middle and between ocelli, isodiametric on anterior portions of infraorbital ridges, posterior portions of infraorbital ridges with slightly rugose vertical waves of sculpture; anterior part of middle elevation of pronotum with indistinct transverse meshes in some specimens; scutellum with fine isodiametric microsculpture. Head with irregular small punctation, denser and coarser on posteriolateral portions; pronotum with irregular, large and deep punctation, sparser on lateral portions, with small impunctate area between mediobasal impression in some specimens; apex of scutellum with several small punctures; punctation of elytra as that on pronotum, some specimens with distinctly smaller punctures on parascutellar area and along suture, each elytron with five to eight vague and tangled longitudinal, and with diagonal rows of punctures in some specimens. Habitus as in Fig. 47.</p><p>Head 1.4–1.6 times as wide as long; middle portion of head slightly and irregularly elevated, with indistinct to distinct transverse impression between eyes at posterior third, with narrow grooves in front of ocelli reaching posterior third or middle length of eyes; postocular ridges indistinct, very smooth. Length × width of antennomeres (holotype): 1: 0.12 × 0.06; 2: 0.07 × 0.04; 3: 0.08 × 0.03; 4–7: 0.07 × 0.04; 8: 0.06 × 0.04; 9–10: 0.06 × 0.05; 11: 0.12 × 0.05.</p><p>Pronotum 1.7–1.8 times as wide as long, 1.4–1.5 times as wide as head; lateral edges of pronotum with regular, small and smooth crenulation; pronotum usually with deep longitudinal impression in apical part, and with very deep transverse impression in mediobasal third in some specimens.</p><p>Elytra 1.1–1.2 times as wide as long, twice as long as pronotum; middle part of each elytron usually with irregular longitudinal elevations between punctures.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 55) with moderately wide median lobe with subtruncate apex; parameres significantly exceeding apex of median lobe, with two apical and two preapical setae; internal sac narrow and long, with rows of sclerotized widened teeth in both middle and basal part. Aedeagus laterally as in Fig. 56.</p><p>Female. Apical margin of abdominal tergite VIII slightly sinuate. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the body size, coloration, shape of the body with moderately short elytra, A. denticulatum Shavrin &amp; Smetana, sp.n. is most similar to A. besucheti Shavrin &amp; Smetana, sp.n., from which it differs by the slightly narrower pronotum, by wider and larger aedeagus, and by the presence of rows of sclerotized teeth in internal sac.</p><p>Distribution. The new species is at present known only from the type locality (Fig. 54) in the Madi Khola valley, central Nepal.</p><p>Bionomics. Specimens were taken by sifting leaf litter in Rhododendron forest at elevation 3200 m a.s.l. (J. Schmidt, pers. comm.)</p><p>Etymology. The specific epithet is the Latin adjective denticulatus, - a, - um (toothed). It refers to lateral rows of sclerotized teeth on the internal sac of the aedeagus.</p></div>	https://treatment.plazi.org/id/03EB87F8F169FFAEFF31FC017EE44B8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F168FFA8FF31FC387D754D3E.text	03EB87F8F168FFA8FF31FC387D754D3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium laozii Shavrin & Smetana 2018	<div><p>Anthobium laozii Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 48, 59, 68–70)</p><p>Type material examined: Holotype ♂: ‘P.R. CHINA, Yunnan, E | slope Cangshan at Dali, | N 25°40'48.5'' E 100°07' | 40.8'', 12.v.2010, 2724m, | sifting07, V. Grebennikov’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | laozii sp.n. | Shavrin A. &amp; Smetana A. 2017 ’ &lt;red rectangular label, printed&gt; (NSMT) .</p><p>Paratypes (83 specimens): 23 ♂ [three specimens dissected], 33 ♀: same data as the holotype (17 ♂, 27 ♀: NSMT; 3 ♂, 3 ♀: CS; 3 ♂, 3 ♀: CNC); 1 ♀: ‘P.R. CHINA, Yunnan, E | slope Cangshan at Dali, | N 25°40'15.1'' E 100°07'. | 39.9'', 10.v.2010, | sifting04, V.Grebennikov’ &lt;rectangular label, printed&gt; (NSMT) ; 2 ♀: ‘ CHINA, Yunnan, | E slope Cangshan at Dali | N 25°40'13.2'' E 100°07' | 54.8'', 2728m, 13.v.2010. | sift 09-11, V. Grebennikov’ &lt;rectangular label, printed&gt; (NSMT) ; 9 ♂ [two specimens dissected], 8 ♀: ‘ CHINA: Yunnan [CH07-01], | Dali Bai Auton. Pref., Diancang Shan W | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.10889&amp;materialsCitation.latitude=25.685833" title="Search Plazi for locations around (long 100.10889/lat 25.685833)">Dali</a>, 25°41'09''N, 100°06'32''E, 3000- | 3200 m, cleft in mixed forest, litter, | debris sifted, 27.V.2007, M. Schülke’ &lt;rectangular label, printed&gt; (2 ♂, 2 ♀: CS; 7 ♂, 6 ♀: CSC) ; 3 ♂ [one specimen dissected], 2 ♀: ‘ CHINA (Yunnan) | Dali Bai Auton. Pref., | Diancang Shan W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.10889&amp;materialsCitation.latitude=25.685833" title="Search Plazi for locations around (long 100.10889/lat 25.685833)">Dali</a> | 25°41'09''N / 100°06'32''E | 3000-3200 m (creek cleft in | mixed forest, slope with | moss) | 27.V.2007. D.W. Wrase [01B]’ &lt;rectangular label, printed&gt; (CSC) ; 1 ♀: ‘ CHINA (N-Yunnan) Dali Bai Nat. | Aut. Pref., Diancang Shan, | 4 km W Dali old town, | 2900-3000 m’ &lt;rectangular label, printed&gt;, ‘25°41.4'/ 100°06.7'E, E slope, | former stone pit (in overgrown | gravel/soil/plant roots/und[er]. stones) | 18.VI.2005 D.W. Wrase [12]’ &lt;rectangular label, printed&gt; (CSC) ; 1 ♀: ‘CHINA-YUNNAN | Yanmen | 13.6- 23.6.2005 | lgt. E. Kučera’ &lt;rectangular label, printed&gt; (CSC) . All paratypes with additional red rectangular printed label : ‘ PARATYPE | Anthobium | laozii sp.n. | Shavrin A. &amp; Smetana A. 2017 ’.</p><p>Description. Measurements (n=84): HW: 0.65–0.72, HL: 0.36–0.45; AL(holotype): 1.23; OL: 0.17–0.20; PL: 0.43–0.55; PW: 0.97–1.20; ESL: 1.25–1.50; EW: 1.38–1.62; AW: 1.11–1.39; MTbL(holotype): 0.55, MTrL(holotype): 0.35 (MTrL 1–4: 0.23; MTrL 5: 0.12); AedL: 0.40–0.47; TL: 2.35–3.65 (holotype: 3.20).</p><p>In external appearance similar to A. monticola and A. confucii sp.n. Head and abdomen, except for paratergites, reddish-brown to brown; antennomeres 3–11 or 4–11 brown; middle portion of pronotum and elytra yellow-brown; mouthparts, antennomeres 1 and 2 or 1–3, lateral portions of pronotum, elytra, and sometimes paratergites of abdomen and legs yellow. Basolateral portions of head without sculpture or with traces of transverse waves of microsculpture between ocellus and posterior margin of eye. Middle portions of head with irregular, sparse and small punctation, some specimens with deeper and more rugose punctation on posterior portions of infraorbital ridges, usually with impunctate small areas on middle elevation between anterior margins of eyes; neck with sparse, deep and small punctures; punctation of pronotum as that on head, denser on medioapical and mediobasal parts, distinctly sparser on latero-apical portions and smaller on mediobasal margin; punctation of elytra slightly larger and deeper as that on pronotum, smaller on parascutellar parts and along suture, each elytron with indistinct to distinct six to eight vague and tangled longitudinal rows of punctures somewhat more clear but smaller that in A. monticola . Habitus as in Fig. 48.</p><p>Head transverse, 1.6–1.8 times as wide as long; middle portion of head slightly elevated, with indistinct to distinct and deep transverse impression between eyes at posterior third, with long grooves in front of ocelli usually reaching anterior third of eye; postocular ridges distinct, acute or slightly smooth; anterior portion between antennal insertion and anterior margin of eye with moderately deep semicircular notch. Ocelli situated at level or slightly behind level of postocular ridges. Length × width of antennomeres (holotype): 1: 0.15 × 0.12; 2: 0.11 × 0.07; 3: 0.12 × 0.07; 4–8: 0.10 × 0.08; 9: 0.10 × 0.09; 10: 0.10 × 0.10; 11: 0.15 × 0.10.</p><p>Pronotum more than twice as wide as long, 1.4–1.6 times as wide as head, widest in middle, more or less evenly narrowed both posteriad and anteriad; posterior angles obtuse; anterior angles strongly protruded anteriad; pronotum with markedly convex middle portion, with small, indistinct longitudinal impression on medioapical third, with moderately deep semioval impression on mediobasal third.</p><p>Elytra distinctly broader than long, markedly widened posteriad, with wide explanate lateral portions; each elytron with indistinct to distinct longitudinal elevations between punctures.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 68) with narrow median lobe with subtruncate, rather narrow apex; parameres slightly (Fig. 70) or distinctly (Fig. 68) exceeding apex of median lobe, slightly attenuate around middle, with two apical and two preapical setae; internal sac simple, narrow anteriorly. Aedeagus laterally as in Fig. 69.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the shape of pronotum, A. laozii Shavrin &amp; Smetana, sp.n. is most similar to the Himalayan A. monticola, from which it differs by the paler coloration, by the elytra distinctly broader than long and by very narrow aedeagus.</p><p>Distribution. The species is known from several localities in Diancang Shan, and one in the environs of Yanmen, all of them in Yunnan Province, China (Fig. 59).</p><p>Bionomics. Specimens were collected at elevations from 2724 to 3200 m a.s.l. by sifting of forest litter, mosses and debris.</p><p>Etymology. Patronymic, the species is named to honour the ancient Chinese philosopher and writer Laozi, the founder of Taoism.</p></div>	https://treatment.plazi.org/id/03EB87F8F168FFA8FF31FC387D754D3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F16EFFAAFF31FA927D134D5A.text	03EB87F8F16EFFAAFF31FA927D134D5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium latissimum Shavrin & Smetana 2018	<div><p>Anthobium latissimum Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 49, 59, 71–73)</p><p>Type material examined: Holotype ♂: ‘ CHINA: Yunnan [CH 07-24], Nujiang | Lisu Aut. Pref., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.50361&amp;materialsCitation.latitude=27.798332" title="Search Plazi for locations around (long 98.50361/lat 27.798332)">Gaoligong Shan</a>, valley 18 | km W Gongshan, 3020 m, 27°47'54''N, | 98°30'13''E, mixed forest, litter, moss, | wood sifted, 7.VI.2007, M. Schülke’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | latissimum sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ &lt;red rectangular label, printed&gt; (CSC) .</p><p>Paratype ♂ [specimen dissected; left side of pronotum with artificial dent]: same data as the holotype, with additional red rectangular printed label: ‘ PARATYPE | Anthobium | latissimum sp.n. | Shavrin A. &amp; Smetana A. des. 2018’ (CS) .</p><p>Description. Measurements (n=2): HW: 0.60; HL: 0.35; AL(holotype): 0.92; OL: 0.13–0.15; PL: 0.50–0.58; PW: 1.00–1.02; ESL: 1.15–1.17; EW: 1.17–1.23; AW: 1.07–1.08; MTbL(holotype): 0.60, MTrL(holotype): 0.27 (MTrL 1–4: 0.16; MTrL 5: 0.11); AedL(paratype): 0.62; TL: 2.56–2.95(holotype).</p><p>Body elongate, subconvex. Pronotum, elytra and antennomeres 6–11 yellow-brown; head reddish-brown; mouthparts, antennomeres 1–5, lateral and basal margins of pronotum and legs yellow. Lateral portions of head on infraorbital ridge with coarsely rugose sculpture, sometimes with distinct small wrinkles on posterior portions (holotype); dorsal part of neck with transverse microsculpture; abdominal tergites with fine isodiametric microsculpture. Frons and middle portion of head with irregular, sparse, small and deep punctation, denser on lateral parts of infraorbital ridges and between ocelli; middle part of neck with regular, small and deep punctation; pronotum with irregular and deep punctation, significantly denser than that on middle part of head; scutellum with several small punctures; punctation of elytra as that on pronotum, slightly denser on parascutellar area, not forming longitudinal rows, with indistinct elevations between punctures on middle part of each elytron; abdominal terites without distinct punctation. Habitus as in Fig. 49.</p><p>Head transverse, 1.7 times as wide as long, slightly elevated in middle portion, with moderately flattened infraorbital ridges; middle part of head between posterior part of eyes with irregular shallow impression, more distinct in holotype; deep, narrow and moderately short grooves in front of ocelli reaching posterior third of length of eyes; posterior portions with distinct, moderately acute postocular ridges; anterior portion between antennal insertion and anterior margin of eye with superficial notch. Ocelli situated at level of postocular ridge; distance between ocelli distinctly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp elongate, about three times as long as preceding segment, markedly narrowed apicad from widest basal portion. Antenna slightly exceeding prescutellar area of elytra when reclined; length × width of antennomeres (holotype): 1: 0.13 × 0.06; 2: 0.10 × 0.05; 3: 0.08 × 0.04; 4: 0.08 × 0.05; 5: 0.08 × 0.07; 6–8: 0.07 × 0.05; 9: 0.07 × 0.06; 10: 0.06 × 0.07; 11: 0.11 × 0.07.</p><p>Pronotum wide and massive in general appearance, with widely rounded apical and basal margins, twice as wide as long, slightly narrower than elytra,1.6–1.7 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin about as long as posterior margin; anterior angles widely rounded, distinctly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with irregular small crenulation; pronotum with very wide middle elevation, with very small and indistinct longitudinal impression and irregular transverse impressions on mediobasal third; disc of pronotum with moderately strong small elevations between punctures in middle part.</p><p>Elytra narrow and elongate, indistinctly widened apicad from about middle, reaching basal margin or middle part (paratype) of abdominal tergite V, about as long as wide, more than twice as long as pronotum; each elytron with indistinctly elevated, small parascutellar and latero-apical portions.</p><p>Male. First four protarsomeres distinctly dilated. Apical margins of abdominal tergite VIII and sternite VIII slightly emarginated. Aedeagus (Fig. 71) with median lobe wide, with wide, truncate apex; parameres each with widened apical portion, slightly assymetrical, exceeding apex of median lobe, with two short apical and preapical setae; internal sac elongate, spiralled in basal portion. Aedeagus laterally as in Fig. 72.</p><p>Female unknown.</p><p>Comparative notes. Anthobium latissimum Shavrin &amp; Smetana, sp.n. may be distinguished from all remaining species of the fusculum group by the shape of the massive pronotum, with markedly rounded apical and basal margins, and by the shape and internal structure of the aedeagus.</p><p>Distribution. The new species is at present known only from the type locality in the Gaoligong Shan range in Yunnan, China (Fig. 59).</p><p>Bionomics. The specimen was taken by sifting litter, moss and wood in forest at elevation of 3020 m a.s.l. (locality DSCN7372: Fig. 73).</p><p>Etymology. The specific epithet is the Latin latissimus, -a, -um (the broadest). It refers to the very wide pronotum.</p></div>	https://treatment.plazi.org/id/03EB87F8F16EFFAAFF31FA927D134D5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F16CFF97FF31FA717CF74D03.text	03EB87F8F16CFF97FF31FA717CF74D03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium liliputense Shavrin & Smetana 2018	<div><p>Anthobium liliputense Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 65, 74, 79–80)</p><p>Type material examined: Holotype ♂ [specimen dissected]: ‘P.R. CHINA, Sichuan, | Emei Shan, N 29°33.6' | E103°20.6', 27.vi.-5.vii. | 2009, 1800-2400m, sifti | ng 11-17, V.Grebennikov’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | liliputense sp.n. | Shavrin A. &amp; Smetana des. A. 2018’ &lt;red rectangular label, printed&gt; (NSMT) .</p><p>Paratypes (6 specimens): 4 ♀♀: same data as the holotype (1 ♀: MHNG; 2 ♀♀: NSMT; 1 ♀: CNC; 1 ♀: CS); 1 ♀: ‘P.R. CHINA, Sichuan | EmeiShan, N29°33'36.3'' | E103°20'38.0'', | 15.vi.2010, 1947m, | sifting33, V.Grebennikov’ &lt;rectangular label, printed&gt; (NSMT) ; 1 ♀: ‘P.R. CHINA, Sichuan, | EmeiShan, N29°33'36.3'' | E103°20'38.0'', | 22.vi.2010, 1947m, | sifting39, V.Grebennikov’ (NSMT) . All paratypes with additional red rectangular printed label : ‘ PARATYPE | Anthobium | liliputense sp.n. | Shavrin A. &amp; Smetana A. des. 2018’.</p><p>Description. Measurements (n=7): HW: 0.52–0.55, HL: 0.29–0.35; AL: 0.73; OL: 0.12; PL: 0.40–0.42; PW: 0.85–0.92; ESL: 1.10–1.12; EW: 1.05–1.22; AW: 0.74–0.80; MTbL(holotype): 0.40, MTrL(holotype): 0.23 (MTrL 1–4: 0.15; MTrL 5: 0.08); AedL(holotype): 0.47; TL: 1.81(holotype)–2.18.</p><p>Body suboval, subconvex. Body and antennomeres 7–11 yellow-brown, head usually slightly darker; mouthparts, antennomeres 1–6 and legs yellow. Posterior portions of infraorbital ridges with rugose diagonal sculpture and small irregular wrinkles between punctures; middle part of neck and abdominal tergites with distinct wavy microsculpture. Frons and middle portion of head with irregular, sparse and small punctation, distinctly coarser and deeper on posterior parts of infraorbital ridges; neck with several very sparse, small punctures; pronotum with very dense rugose punctation, larger and deeper than that on middle part of head, sparser laterally, some specimens with small impunctate area on basal portion; punctation of elytra as that on pronotum, slightly denser and deeper on parascutellar area and along suture in some specimens, smoothed on apical third, punctures not forming longitudinal rows of punctures; abdominal tergites without distinct punctation. Habitus as in Fig. 74.</p><p>Head 1.5–1.7 times as wide as long, head indistinctly elevated in middle; occipital furrow between ocelli not distinct; grooves in front of ocelli very short and indistinct, very small and deep or slightly elongate, about as long as diameter of ocellus; postocular ridges indistinct, acute or obtuse; anterior portion between antennal insertion and anterior margin of eye with small superficial notch. Ocelli very large and convex, situated at level of posterior margin of eye or postocular ridge; distance between ocelli as long as or slightly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp about two and half times as long as preceding segment. Antenna moderately short, reaching shoulders of elytra when reclined; length × width of antennomeres (holotype): 1: 0.11 × 0.05; 2: 0.07 × 0.03; 3: 0.07 × 0.02; 4: 0.05 × 0.02; 5–6: 0.06 × 0.03; 7–8: 0.06 × 0.04; 9–10: 0.05 × 0.05; 11: 0.10 × 0.05.</p><p>Pronotum transverse, slightly more than twice as wide as long, 1.6 times as wide as head, widest in middle, about evenly narrowed both posteriad and anteriad; apical margin distinctly narrower than posterior margin; anterior angles rounded, distinctly protruded anteriad; posterior angles more or less rounded; lateral edges of pronotum with indistinct, smoothed small crenulation; pronotum with wide middle elevation slightly widened posteriad, with irregular and usually indistinct transverse impression on mediobasal third.</p><p>Elytra wide, slightly widened apicad, reaching apical parts of abdominal tergite V to VII, distinctly longer than wide, significantly more than twice as long as pronotum, with widely rounded apical angles; each elytron with indistinct slightly elevated middle portions close to parascutellar area in some.</p><p>Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (Fig. 79) with median lobe wide, gradually narrowed from about middle toward subtruncate apex; parameres narrow, each from about middle markedly turned mediad, exceeding apex of median lobe, with two short apical and preapical setae; internal sac with numerous elongate thorns, spiralled in basal portion. Aedeagus laterally as in Fig. 80.</p><p>Female. Apical margins of abdominal tergite VIII and sternite VIII straight to slightly rounded.</p><p>Comparative notes. Based on the irregularly arranged punctation of elytra, A. liliputense Shavrin &amp; Smetana, sp.n. is most similar to the Himalayan A. cavicrus, from which it differs by paler coloration, smaller body, wider pronotum, elytra and aedeagus, and spiraled basal portion of the internal sac.</p><p>Distribution. Anthobium liliputense sp.n. is known from two locations (Fig. 65) in the Emei Shan range in Sichuan, China.</p><p>Bionomics. All specimens were taken by sifting forest litter at elevations from 1800 to 2400 m.</p><p>Etymology. The specific epithet is derived from Lilliput, the fictional island with small people that appear in the novel “Gulliver's Travels” by the famous English writer Jonathan Swift. It refers to the very small size of the species.</p></div>	https://treatment.plazi.org/id/03EB87F8F16CFF97FF31FA717CF74D03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F151FF90FF31FAB678D748F6.text	03EB87F8F151FF90FF31FAB678D748F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium monticola (Cameron 1924)	<div><p>Anthobium monticola (Cameron, 1924)</p><p>(Figs. 7, 9, 11, 13, 15, 17, 19, 21, 23, 25, 27, 33–37, 75, 81–83)</p><p>Lathrimaeum monticola Cameron, 1924: 169</p><p>Lathrimaeum monticola: Cameron, 1930: 154</p><p>Lathrimaeum monticola: Scheerpeltz, 1933: 1056</p><p>Lathrimaeum (Lathrimaeum) monticola: Scheerpeltz, 1961: 78</p><p>Anthobium monticola: Herman, 2001: 236</p><p>Anthobium (Anthobium) monticola: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 307</p><p>Type material examined: Lectotype (here designated) of Lathrimaeum monticola Cameron, 1924 ♂: ‘SYN- | TYPE’ &lt;round label with blue margin, printed&gt;, ‘Type | H. T.’ &lt;round label with red margin, printed&gt;, ‘Chakrata Dist., | Korawa Khud 9100`’ &lt;rectangular label, printed&gt;, ‘Dr. Cameron. | 4. V. 22.’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | monticola | Cam.’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘E. I. 40 ’ &lt;rectangular label, handwritten in black&gt;, ‘M. Cameron | Bequest. | B.M. 1955-147.’ &lt;rectangular printed label&gt;, ‘ LECTOTYPE | Lathrimaeum | monticola | Cameron, 1924 | Shavrin A.V. des. 2016’ &lt;red rectangular label, printed&gt;, ‘ Anthobium | monticola (Cameron, 1924) | Shavrin A.V. det. 2016’ &lt;rectangular label, printed&gt; (BMNH).</p><p>Paralectotypes of Lathrimaeum monticola Cameron, 1924: 1 ♂: ‘SYN- | TYPE’ &lt;round label with blue frame, printed&gt;, ‘United Prov., | N. India. [underlined by yellow] | B.M. 1924-335.’ &lt;rectangular label, printed&gt;, ‘Chakrata Dist. | Korawa Khud 9100`| Dr. Cameron. | 4. V. 22.’ &lt;rectangular label, printed&gt;, ‘ Lathrimaeum | monticola | Cam. | Co-type’ &lt;rectangular label, handwritten in black Indian ink&gt; (BMNH); 3 ♂♂, 1 ♀: ‘SYN- | TYPE’ &lt;round label with blue frame, printed&gt;, ‘Gahan 7000`| Simla Hills’ &lt;rectangular label, printed&gt;, ‘Dr. Cameron. IX.1921.’ &lt;rectangular label, printed&gt;, ‘M. Cameron. | Bequest. | B.M. 1955-147.’ &lt;rectangular label, printed&gt; (BMNH); 1 ♀: ‘SYN- | TYPE’ &lt;round label with blue frame, printed&gt;, ‘Bindal Gadh. | Konain.’ &lt;rectangular label, printed&gt;, ‘Dr. Cameron. | 5. V. 21., M. Cameron. | Bequest. | B.M. 1955-147.’ &lt;rectangular label, printed&gt; (BMNH); 1 ♂ [specimem without left elytron]: ‘Gahan 7000`| Simla Hills’ &lt;rectangular label, printed&gt;, ‘Dr. Cameron. IX.1921.’ &lt;rectangular label, printed&gt;, ‘COTYPE’ &lt;red rectangular label, handwritten&gt;, ‘ Lathrimaeum | monticola | Cam’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘ Lathrimaeum | monticola | Cameron’ &lt;rectnagular label, printed&gt;, ‘Coll. W. Champman | in Coll. P. Griveau | MHNG - 2007’ &lt;rectangular label, printed&gt;, ‘monticola | Cam’ &lt;rectnagular label in yellow frame, handwritten in black Indian ink&gt; (MHNG); 2 ♂♂, 3 ♀♀: ‘SYN- | TYPE’ &lt;round label with blue frame, printed&gt;, ‘Theog 7600 | Simla Hills.’ &lt;rectangular label, printed&gt;, ‘Dr. Cameron | IX. 1921.’ &lt;rectangular label, printed&gt;, ‘M. Cameron. | Bequest. | B.M. 1955-147.’ &lt;rectangular label, printed&gt; (BMNH).</p><p>One paralectotype is deposited in the collection of FMNH; the photograph of habitus and labels are available in Arthropod Collections Database of FMNH: http://emuweb.fieldmuseum.org/arthropod/coleop.php (last Access: 18.06.2018).</p><p>Additional material: PAKISTAN: 1 ♀: AZAD JAMMU AND KASHMIR: Pakistan (AJK) Leepa-Top, Muzaffarābad. 3200-3300 m a.s.l. 14.06.1997. [W.] Heinz leg. (CAP) ; INDIA: HIMACHAL PRADESH: 4 ♂♂: ‘ Parbatti V. Kulu. Punjab. 6-8,000ft. H. G.C. ’, ‘ Lathrimaeum monticola Cam. ’ (BMNH) ; UTTARANCHAL PRADESH: 1 ♂: ‘ Konain, 7800. Chakrata, U.P. ’, ‘ M. Cameron. Bequest. B.M. 1955-147.’ (BMNH) ; NEPAL: 1 ♂: SW Dhaulagiri Himal env. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=83.25&amp;materialsCitation.latitude=28.5" title="Search Plazi for locations around (long 83.25/lat 28.5)">Jaljala La</a>, 28°30'N 83°15'E. 3300-3500 m a.s.l. 12-13, 21.05.2012. J. Schmidt leg. (NME) ; 1 ♀: Mahakali / Darchula, 14 km NNE Ghusa, river before <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.94334&amp;materialsCitation.latitude=29.935" title="Search Plazi for locations around (long 80.94334/lat 29.935)">Api</a>, 29°56'06''N 80°56'36''E. 3450 m a.s.l. 10.06.2005. A. Weigel leg. (CS) ; 1 ♂: Karnali Province, Jumla District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=82.30167&amp;materialsCitation.latitude=29.201666" title="Search Plazi for locations around (long 82.30167/lat 29.201666)">Gothichaur</a>, 29°12.1'N 82°18.1'E. 2850 m a.s.l., forest. 13.06.1997. A. Weigel leg. (NME) ; 1 ♀: same Province and District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=82.391&amp;materialsCitation.latitude=29.355" title="Search Plazi for locations around (long 82.391/lat 29.355)">Maharigaon</a>, 29°21.3'N 82°23.46'E. 3750 m a.s.l. NN. 0 6.07.1999. M. Hartmann leg. (NME) ; 1 ♀: P: Seti /D: Bajhang, 42 km NE Chainpur, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=81.48444&amp;materialsCitation.latitude=29.806389" title="Search Plazi for locations around (long 81.48444/lat 29.806389)">Kalapani Khola</a>, 29°48'23''N 81°29'04''E. 3750 m NN (#27). 23.06.2009. D. Mattern leg. (NME) ; 1 ♂: Lamjung, S-Sundar <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.34917&amp;materialsCitation.latitude=28.3825" title="Search Plazi for locations around (long 84.34917/lat 28.3825)">Danda</a>, 28°22'57''N 84°20'57''E. 3140 m a.s.l. 21.09.2013. [J.] Hagge &amp; [J.] Schmidt leg. (NME) ; 1 ♂: Manang District, For. W Bagarchhap. 22.09. 1983. 2250 m a.s.l. A. Smetana &amp; I. Löbl (MHNG) .</p><p>Redescription. Measurements (n=20): HW: 0.67–0.80, HL: 0.40–0.55; AL: 1.62; OL: 0.17–0.25; PL: 0.50–0.60; PW: 1.02–1.37; ESL: 1.37–1.75; EW: 1.40–1.85; AW: 1.11–1.41; MTbL(lectotype): 0.90, MTrL(lectotype): 0.40 (MTrL 1–4: 0.25; MTrL 5: 0.15); AedL: 0.60–0.67; TL: 3.10–4.50(lectotype).</p><p>Body more or less elongate to moderately wide, subconvex. Body reddish brown, with dark-brown head and abdomen, some specimens with elytra paler, yellow-brown to brown; mouthparts, antennomeres 1–3, lateral and basal margins of pronotum (some specimens) and legs yellow to yellow-brown. Body shiny, without microsculpture on pronotum and elytra; head between supra-antennal elevation and anterior margin of eye with very fine, indistinct or well-developed diagonal cellular-shaped meshes, basolateral portions of head behind eyes and posterior portions of infraorbital ridges with indistinct to distinct coarse microsculpture, some specimens with small elevated wrinkles between punctures; middle part of neck and abdominal tergites with distinct isodiametric microsculpture. Head with irregular and large punctation, denser and deeper on middle portion between eyes and on posterior half of infraorbital ridges; middle part of neck with more or less regular, small and deep punctures; pronotum with irregular punctation as that on middle part of head, distinctly sparser and smaller on mediobasal third between impressions and latero-apical portions; scutellum with several small punctures; punctation of elytra dense, markedly larger, deeper and sometimes coarser than that on pronotum, smaller on parascutellar area and along basal margins, each elytron with five to seven vague and tangled longitudinal rows of punctures, remaining elytral punctures forming indistinct and tangled diagonal rows and becoming irregularly scattered laterally; abdominal tergites with sparse, indistinct small punctures. Habitus as in Fig. 75.</p><p>Head 1.4–1.6 times as wide as long; vertex irregularly elevated, with middle transverse triangular impression between eyes at posterior third, with distinct occipital furrow between ocelli, with elongate grooves in front of ocelli, reaching middle or anterior third of eye; postocular ridges slightly protruded, acute, distance between postocular ridge and posterior margin of eye as long as two nearest facets; anterior portion between antennal insertion and anterior margin of eye with wide semicircular notch. Eyes large and convex. Ocelli very large, equal to diameter of two-three nearest punctures, convex, situated at level of posterior margin of eyes; distance between ocelli usually slightly longer than distance between ocellus and posterior margin of eye. Labrum moderately wide, with widely rounded membranous latero-apical portions, with deep apical emargination and with very long basolateral projections (Fig. 7). Mandibles short, with moderately narrow basal portion, with small distinct preapical tooth on right mandible (Fig. 9). Labium wide, with elongate submentum (Fig. 13). Apical segment of maxillary palp about twice as long as preceding segment (Fig. 11). Gular sutures with widely rounded apical part, more or less narrowly diverging posteriad; shortest distance between sutures located at level of posterior third of eyes (Fig. 15). Antenna moderately long, reaching anterior third of length of elytra when reclined; length × width of antennomeres (lectotype): 1: 0.25 × 0.07; 2: 0.16 × 0.05; 3: 0.17 × 0.05; 4–8: 0.13 × 0.05; 9–10: 0.11 × 0.07; 11: 0.17 × 0.07.</p><p>Pronotum transverse, twice as wide as long, 1.5–1.7 times as wide as head, widest in front of middle, evenly narrowed posteriad and anteriad; apical margin distinctly narrower than posterior margin, sinuately emarginate and widely rounded; anterior angles more or less widely rounded, distinctly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with regular small, smooth crenulation; pronotum with wide, slightly elevated middle portion, with wide, indistinct to distinct, short or long longitudinal impression reaching basal third of pronotum, with a pair of moderately deep semioval impressions on mediobasal third; lateral portions distinctly explanate, each with deep oval pit at middle. Prothorax (Fig. 17) with elongate intercoxal process and wide, strongly protruded mesosternal processes. Mesothorax (Fig. 23) transverse, moderately narrow in middle, with acute intercoxal process almost reaching posterior margin of mesocoxae. Scutellum (Fig. 19) with triangular apex. Metathorax (Fig. 25) wide, with wide and deep intercoxal cavities and wide intercoxal process. Metendosternite with long paired medioapical protrusions (Fig. 27).</p><p>Elytra about as long as wide, slightly less than three times as long as pronotum, slightly or distinctly widened apicad, reaching apical margin of abdominal tergite IV or VI, with widely rounded apical margins truncated at suture; shoulders of elytra indistinctly protruded anteriad (Fig. 21); lateral portions of elytra narrow to moderately wide, slightly explanate, latero-apical margins with small acute crenulation, flattened and reduced in middle; middle part of each elytron with irregular, indistinct to distinct elevations between punctures, sometimes forming three very indistinct longitudinal elevations. Wings fully developed.</p><p>Abdomen distinctly narrower than elytra, with pair of small rounded tomentose wing-folding spots in middle of tergite V; intersegmental membranes between tergites IV–VII narrow to wide, covered by brickwall-like sculpture.</p><p>Male. Apical margin of abdominal tergite VIII (Fig. 33) and sternite VIII (Fig. 34) vaguely sinuate. Aedeagus (Fig. 81) with median lobe very wide basally, gradually, markedly narrowed toward rounded apex; parameres narrow, slightly exceeding apex of median lobe, with two small apical and preapical setae; internal sac very narrow and moderately long, with curved basal portion. Aedeagus laterally as in Fig. 82.</p><p>Female. Apical margin of abdominal tergite VIII (Fig. 35) straight. Apical margin of abdominal sternite VIII (Fig. 36) narrowly rounded. Genital segment (Fig. 37) with narrow elongate sternite IX, gonocoxites moderately narrow, slightly widened basally, styli elongate and narrow.</p><p>Comparative notes. Based on the general shape of the distinctly transverse pronotum and the wide elytra with indistinct to distinct longitudinal elevations between punctures, A. monticola is most similar to the Himalayan A. altivagans and Chinese A. confucii Shavrin &amp; Smetana, sp.n. and A. laozii Shavrin &amp; Smetana, sp.n. From A. altivagans it differs by anterior angles of pronotum strongly protruded anteriad and by the lack of sclerotized teeth in internal sac of aedeagus; from A. confucii sp.n. by smaller punctation of elytra and by obtuse posterior angles of pronotum; from A. laozii sp.n. by darker elytra, coarser microsculpture of basolateral portions of head and larger punctation of distinctly longer elytra. Additionally, from A. altivagans and A. confucii sp.n. it differs by slightly longer elytra; from A. confucii sp.n. and A. laozii sp.n. by the lateral portions of narrower pronotum evenly narrowed posteriad and anteriad. From all these species, A. monticola differs by wider aedeagus and the very narrow internal sac.</p><p>Distribution. The species is known from Pakistan, India (Himachal Pradesh, Uttarakhand), and from western and central Nepal (Fig. 83).</p><p>Bionomics. Specimens were collected at elevations from 2100 to 3750 m a.s.l. Specimens from Nepal were taken by sifting leaf litter in Rhododendron forest and Rhododendron - Abies forest, by sifting wet moss, various</p><p>debris and fallen leaves along a small brook (J. Schmidt, pers. comm.). Specimen from Jaljala La was sifted together with A. sociale, specimen from Sundar Danda with A. besucheti sp.n. (see below). Remarks. Anthobium monticola is here recorded for the first time from Pakistan and Nepal.</p></div>	https://treatment.plazi.org/id/03EB87F8F151FF90FF31FAB678D748F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F156FF93FF31FBD178414E06.text	03EB87F8F156FF93FF31FBD178414E06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium rugosum (Champion 1925)	<div><p>Anthobium rugosum (Champion, 1925)</p><p>(Figs. 54, 76, 84–85)</p><p>Lathrimaeum rugosum Champion, 1925: 103</p><p>Lathrimaeum rugosum: Cameron, 1930: 154; Scheerpeltz, 1933: 1056; Coiffait, 1977: 244</p><p>Lathrimaeum (Lathrimaeum) rugosum: Scheerpeltz, 1961: 85</p><p>Anthobium rugosum: Herman, 2001: 238</p><p>Anthobium (Anthobium) rugosum: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 307</p><p>Type material examined: Holotype of Lathrimaeum rugosum Champion, 1925 [Immature specimen, without abdomen from tergites/sternites V] unsexed: ‘U. Gumti Val. | W. Almora Dn. | Apr.`19. H.G.C.’ &lt;rectangular printed label&gt;, ‘2836’ &lt;rectangular printed label&gt;, ‘Type | H. T.’ &lt;round printed label with red margin&gt;, ‘ Lathrimaeum | rugosum, | Champ.’ &lt;rectangular printed label&gt;, ‘E. M. M. 1925. | det. G.C.C.’ &lt;rectangular printed label&gt;, ‘ Lathrimaeum | rugicolle [sic!] | 1925 Ch. | Type’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘G.C. Champion | Brit. Mus. | 1925–42.’ &lt;rectangular printed label&gt;, ‘ HOLOTYPE | Lathrimaeum | rugosum Champion 1925 | det. R.G. Booth 2015 [printed]’ &lt;rectangular label, handwritten in black&gt;, ‘ Anthobium | rugosum (Champion, 1925) | Shavrin A.V. det. 2016’ (BMNH).</p><p>Additional material: INDIA: Punjab Province: 1 ♀: ‘Parbatti V. | Kulu. Punjab. | 6-8,000ft. H.G.C’, ‘H.G. Champion Coll. | B.M. 1953-156’, ‘ Lathrimaeum ? rugosum Champ. P.M.Hammond det. 1969’ (BMNH); NEPAL: 1 ♂, 1 ♀: W Dhaulagiri, river below Thankur, 28°37'25''N 83°00'32''E. 2800 m a.s.l. J. Schmidt leg. (NME, CS).</p><p>Redescription. Measurements (n=4): HW: 0.60; HL: 0.33–0.40; AL(holotype): 0.98; OL: 0.15–0.17; PL: 0.42–0.50; PW: 0.90–0.95; ESL: 1.21–1.25; EW: 1.16–1.22; AW: 0.95–1.05; MTbL(holotype): 0.55, MTrL(holotype): 0.30 (MTrL 1–4: 0.20; MTrL 5: 0.10); AedL: 0.43; TL: 2.65–2.75 (holotype without abdomen: 2.25).</p><p>Body moderately wide, subconvex. Head, antennomeres 4–11, middle portion of pronotum, visible part of scutellum and abdomen reddish brown; apical and basal margins and lateral portions of pronotum and elytra yellow-brown; mouthparts, antennomeres 1–3, legs and apical part of abdomen yellow. Infraorbital ridges with moderately strong sculpture and small elevated wrinkles between punctures, more rugose on posterior portions, with small diagonal impunctate areas behind middle impression; middle part of neck with distinct, dense microsculpture; scutellum with indistinct wavy meshes; abdomen with fine isodiametric microsculpture. Head with irregular, moderately large and deep microsculpture, denser on infraorbital ridges of head; middle part of neck with dense small punctures; pronotum with irregular punctation, markedly larger and deeper than that on head, sparser on lateral portions and basal margin and markedly sparser and smaller on mediobasal third, with impunctate small area between impression in some specimens; scutellum with several small punctures; punctation of elytra as that on pronotum, distinctly denser and smaller on parascutellar area, each elytron with six to eight vague and tangled longitudinal rows of punctures; abdominal tergites without punctation. Habitus as in Fig. 76.</p><p>Head 1.5–1.8 times as wide as long; middle portion slightly elevated, with deep transverse impression between eyes at posterior third, with long and deep grooves in front of ocelli, reaching apical third of eye; postocular ridges smooth; anterior portion between antennal insertion and anterior margin of eye with wide and moderately deep notch. Ocelli very large, situated at level of postocular ridges; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp about two and half as long as preceding segment. Antenna slightly exceeding shoulders of elytra when reclined; length × width of antennomeres (holotype): 1: 0.17 × 0.06; 2: 0.07 × 0.05; 3: 0.10 × 0.03; 4–6: 0.08 × 0.04; 7–9: 0.07 × 0.05; 10: 0.07 × 0.07; 11: 0.12 × 0.07.</p><p>Pronotum transverse, about twice as wide as long, 1.5 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin slightly narrower than posterior margin; anterior angles strongly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with irregular smooth crenulation; pronotum with wide, slightly elevated middle portion, with wide, indistinct longitudinal impression on medioapical third and a pair of moderately deep semioval impressions on mediobasal third.</p><p>Elytra about as long as wide, about two and half times as long as pronotum or slightly longer, slightly widened apicad, reaching abdominal tergite V; surface between punctures on medioapical and middle portions of each elytron with irregular, transverse and diagonal elevations.</p><p>Male. Apical margin of abdominal tergite VIII widely rounded. Apical margin of abdominal sterbite VIII slightly sinuate. Aedeagus (Fig. 84) short, median lobe with widely rounded apex; parameres short, moderately wide, slightly exceeding apex of median lobe, with two small apical and preapical setae; internal sac narrow, moderately long. Aedeagus laterally as in Fig. 85.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Anthobium rugosum may be distinguished from all remaining species of the fusculum group by the characteristic rugose surface on middle portion of elytra, with irregular transverse and diagonal elevations between punctures and by the internal and external structures of the aedeagus.</p><p>Distribution. The species is known from Punjab and Uttarakhand of India and from central Nepal (Fig. 54).</p><p>Bionomics. Specimens were collected at elevations from 1800 to 2800 m a.s.l. Specimens from Nepal were taken by sifting leaf litter in mixed forest near river bank (J. Schmidt, pers. comm.). Champion (1925) noted that the type specimen was collected in dead leaves.</p><p>Remarks. Smetana (2004) and Schülke &amp; Smetana (2015) cited the species only from Nepal.</p><p>Anthobium rugosum is here recorded for the first time from Punjab State of India.</p></div>	https://treatment.plazi.org/id/03EB87F8F156FF93FF31FBD178414E06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F155FF9CFF31F9B07D9749A3.text	03EB87F8F155FF9CFF31F9B07D9749A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium sociale (Cameron 1941)	<div><p>Anthobium sociale (Cameron, 1941)</p><p>(Figs. 64, 77, 86–87)</p><p>Lathrimaeum (Prionothorax) sociale Cameron, 1941: 59</p><p>Anthobium sociale: Herman, 2001: 239</p><p>Anthobium (Prionothorax) sociale: Smetana, 2004: 239; Schülke &amp; Smetana, 2015: 307</p><p>Type material examined: Holotype (by monotypy) ♀: ‘SYN- | TYPE’ &lt;round printed label with blue margin&gt;, ‘Type’ &lt;round printed label with red margin&gt;, ‘Kashmir | Gulmarg | vi-vii-31 | Dr.Cameron’ &lt;rectangular printed label&gt;, ‘L. | sociale | TYPE [in red] cam.’ &lt;rectangular label, handwritten in black Indian ink&gt;, ‘M. Cameron | Bequest. | B.M. 1955-147.’ &lt;rectangular printed label&gt;, ‘ Anthobium | sociale (Cameron, 1941) | Shavrin A.V. det. 2016’ (BMNH).</p><p>Additional material: INDIA: KASHMIR: 3 ♂, 3 ♀: ‘SYN- | TYPE’ &lt;round printed label with blue margin&gt;, ‘ Kashmir | Gulmarg | vi-vii-31 | Dr.Cameron’ &lt;rectangular printed label&gt;, ‘ M. Cameron | Bequest. | B.M. 1955-147.’ &lt;rectangular printed label&gt; (BMNH); UTTARAKHAND: 2 ♂, 1 ♀: 5 km NW Ghangaria, Valley of Flowers, 30°43.366'N 79°85.309'E. 3417 m a.s.l. A.V. Shavrin (CS) ; WEST BENGAL: Darjeeling District, Tonglu. 3100 m a.s.l. 16.10.1978. Besuchet &amp; Löbl (MHNG) ; NEPAL: 1 ♂, 1 ♀: Parbat District, Ghoropani Pass, SN slope. 2700 m a.s.l. 0 6.10.1983. A. Smetana &amp; I. Löbl (MHNG); 2 ♀: Manaslu Mts., SW Meme Pokhari, 28°21'41N 84°30'42E. 3200-3300 m a.s.l., Quellgebiet, mittl. Nebenfluss des Ngadi Khola. 11.05.2005. J. Schmidt leg. (NME) ; 1 ♀: SW Dhaulagiri Himal env., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=83.25&amp;materialsCitation.latitude=28.5" title="Search Plazi for locations around (long 83.25/lat 28.5)">Jaljala La</a>, 28°30'N, 83°15'E. 3300-3500 m a.s.l. 12-13,21.05.2012. J. Schmidt leg. (NME) .</p><p>Redescription. Measurements (n=14): HW: 0.55–0.62; HL: 0.32–0.42; AL(lectotype): 0.89; OL: 0.10–0.12; PL: 0.45–0.50; PW: 0.83–0.97; ESL: 1.03–1.16; EW: 1.12–1.17; AW: 1.00–1.05; MTbL(lectotype): 0.50, MTrL(lectotype): 0.27 (MTrL 1–4: 0.17; MTrL 5: 0.10); AedL: 0.37; TL: 2.35–3.20 (lectotype: 3.18).</p><p>Body elongate, subconvex. Body yellow-brown to reddish brown, head and abdomen darker in some specimens; antennomeres 4–11 brown; mouthparts, antennomeres 1–3, lateral portions of pronotum, shoulders of elytra and legs yellow in some specimens. Middle part of vertex without, or with fine transverse microsculpture, middle part of head between eyes and posterior portions with coarse sculpture between punctures, more rugose on posterior portions of infraorbital ridges and less elevated in middle (some specimens with middle part of head without sculpture between punctures); middle part of neck and abdominal sternites with distinct isodiametric microsculpture. Head with irregular, large and deep punctation, distinctly larger and coarser on infraorbital ridges, with markedly elevated diagonal and transverse wrinkles between punctures; middle part of neck with indistinct to distinct, small and regular punctation; pronotum with irregular, moderately dense punctation, smaller on medioapical and mediobasal parts, slightly larger on lateral portions, sparser in middle, with small impunctate area between mediobasal impression in some specimens; visible part of scutellum with several small punctures, or without them; punctation of elytra as that on pronotum, moderately sparse, distinctly denser and coarser in parascutellar area, each elytron with six to eight vague and tangled longitudinal rows of punctures, with moderately wide impunctated areas between rows 3 and 4 in some specimens; abdominal tergites without visible punctation. Habitus as in Fig. 77.</p><p>Head 1.4–1.7 times as wide as long; middle portion slightly or distinctly convex, without or with indistinct transverse impression between peyes at posterior third, with short and deep grooves in front of ocelli, usually reaching middle length of eyes; postocular ridges smooth; anterior portion between antennal insertion and anterior margin of eye with wide and moderately deep notch. Ocelli large, situated at level of postocular ridges; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Length × width of antennomeres (lectotype): 1: 0.11 × 0.07; 2: 0.10 × 0.05; 3: 0.11 × 0.02; 4: 0.08 × 0.03; 5: 0.10 × 0.03; 6: 0.07 × 0.03; 7: 0.07 × 0.04; 8: 0.05 × 0.04; 9–10: 0.05 × 0.06; 11: 0.10 × 0.06.</p><p>Pronotum 1.8–1.9 times as wide as long, 1.5 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin distinctly narrower than posterior margin; anterior angles acute or obtuse; lateral edges of pronotum with irregular, small and smooth crenulation; pronotum with wide, slightly elevated middle portion, without or with indistinct to distinct, deep and wide longitudinal impression usually reaching middle area near distinct and markedly deep semioval mediobasal impressions.</p><p>Elytra about as long as wide, markedly more than twice as long as pronotum, moderately narrow, slightly widened apicad from middle, reaching apical margin of abdominal tergite IV or middle of tergite V; middle part of each elytron with indistinct longitudinal elevations between punctures, forming smooth and very indistinct three elevations on each elytron in some specimens.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII widely rounded. Aedeagus (Fig. 86) narrow, median lobe with wide subtruncate apex; parameres significantly exceeding apex of median lobe, widened apically, with two short apical and preapical setae; internal sac narrow, long. Aedeagus laterally as in Fig. 87.</p><p>Female. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Anthobium sociale may be distinguished from all remaining species of the fusculum group by elongate elytra more than twice as long as pronotum, and by the shape and internal structure of the aedeagus.</p><p>Distribution. The species is known from Kashmir, Uttarakhand and West Bengal states of India and central Nepal (Fig. 64).</p><p>Bionomics. Specimens were collected at elevations from 3100 to 3417 m a.s.l. Specimens from the Valley of Flowers in Uttarkhand were collected by sifting leaves and mosses in Rhododendron shrubs near small stream (Fig. 88). Specimens from Ghoropani Pass were taken by sifting moss, leaf litter and other debris along a creek, around bases of old trees and in old, wet Rhododendron forest with bushy undergrowth. Specimens from Meme Pokhari and Jaljala La were taken by sifting leaf litter in Rhododendron forest (J. Schmidt, pers. comm.), and were collected together with A. monticola (see above).</p><p>Remarks. Anthobium sociale is here recorded for the first time from Uttarakhand and West Bengal states of India and from Nepal.</p></div>	https://treatment.plazi.org/id/03EB87F8F155FF9CFF31F9B07D9749A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F15AFF99FF31FA4B7DCE4E6B.text	03EB87F8F15AFF99FF31FA4B7DCE4E6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium splendidulum Shavrin & Smetana 2018	<div><p>Anthobium splendidulum Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 78, 89–93)</p><p>Type material examined: Holotype ♂ [specimen dissected]: ‘ CHINA (Yunnan) | Nujiang Lisu Aut. Pref., | Gaoligong Shan, valley 18 km | W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.50361&amp;materialsCitation.latitude=27.798332" title="Search Plazi for locations around (long 98.50361/lat 27.798332)">Gongshan</a>, 3020 m | 27°47'54''N, 98°30'13''E | (mixed forest, litter, moss,’ &lt;rectangular label, printed&gt;, ‘wood sifted) | 7.VI.2007 D.W.Wrase [24]’ &lt;rectangular label, printed &gt;, HOLOTYPE | Anthobium | splendidulum sp.n. | Shavrin A. &amp; Smetana A. 2018 ’ &lt;red rectangular label, printed&gt; (CSC) .</p><p>Paratypes (10 specimens): 2 ♀♀: ‘P.R. CHINA, Yunnan, E | slope N Gaoligongshan, | N 27°59'01.0'' E 098°32' | 56.9'', 27.v.2010, 3018m, | sifting22, V.Grebennikov’ &lt;rectangular label, printed&gt; (NSMT) ; 1 ♀: ‘P.R. CHINA, Yunnan, E | slope N Gaoligongshan, | N 27°59'11.5'' E 098°33' | 25.3'', 28.v.2010, 3086m, | sifting24, V.Grebennikov’ &lt;rectangular label, printed&gt; (CNC) ; 1 ♂: ‘ CHINA: Yunnan [CH07-24], Nujiang | Lisu Aut. Pref., Gaoligong Shan, valley 18 | km W Gongshan, 3020 m 27°47'64'' N, | 98°30'13''E, mixed forest, litter, moss, | wood sifted, 7.VI.2007, M. Schülke’ &lt;rectangular label, printed&gt; (CS) ; 1 ♂ [specimen dissected]: ‘ CHINA: Yunnan [CH07-21], | Nujiang Lisu Aut. Pref., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.44722&amp;materialsCitation.latitude=27.774166" title="Search Plazi for locations around (long 98.44722/lat 27.774166)">Gaoligong Shan</a>, | pass 22 km W Gongshan, N slope 3350- | 3400 m, 27°46'27''N, 98°26'50''E, fern, | moss, litter, sifted, 6.VI.2007, M. Schülke’ &lt;rectangular label, printed&gt; (CS) ; 1 ♂: ‘ CHINA: Yunnan, Nujiang Lisu Pref., | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.68361&amp;materialsCitation.latitude=25.9725" title="Search Plazi for locations around (long 98.68361/lat 25.9725)">Gaoligong Shan</a>, “Cloud pass” | 21 km NW Liuku, 3150 m, | 25°58'21''N, 98°41'01''E, | shrubs &amp; bamboo, litter sifted, | 2.IX.2009, leg. M. Schülke [CH09-22]’ (CSC) ; 1 ♂ [specimen dissected], 2 ♀♀: ‘ CHINA: N-Yunnan [C 2005-06] | Diqing Tibet. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.09116&amp;materialsCitation.latitude=28.323" title="Search Plazi for locations around (long 99.09116/lat 28.323)">Aut. Pref.</a>, | Deqin Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.09116&amp;materialsCitation.latitude=28.323" title="Search Plazi for locations around (long 99.09116/lat 28.323)">Baima Shan</a>, pass | 25 km SE Deqin, 4225 m,’ &lt;rectangular label, printed&gt;, ‘ 28°19.38'N, 99°05.47'E, small | creek valley, Rhododendron, Salix, | leaf litter, moss, dead wood, sifted, | 8.VI.2005, M. Schülke [C 2005-06]’ &lt;rectangular lable, printed&gt; (CSC) ; 1 ♂ [specimen dissected]: ‘ CHINA: N-Yunnan [C 2005-09] | Diqing Tibet. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.7725&amp;materialsCitation.latitude=28.457834" title="Search Plazi for locations around (long 98.7725/lat 28.457834)">Aut. Pref.</a>, | Deqin Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.7725&amp;materialsCitation.latitude=28.457834" title="Search Plazi for locations around (long 98.7725/lat 28.457834)">Meili Xue Shan</a>, | E-side, 14 km W Deqin, 2580 m,’ &lt;rectangular label, printed&gt;, ‘ 28°27.47'N, 98°46.35'E, creek | valley below glacier, mixed, | leaf litter, moss, dead wood, sifted, | 11.VI.2005, M. Schülke [C 2005-09]’ &lt;rectangular label, printed&gt; (CSC) . All paratypes with additional red rectangular printed label : ‘ PARATYPE | Anthobium | splendidulum sp.n. | Shavrin A. &amp; Smetana A. des. 2018’.</p><p>Description. Measurements (n=9): HW: 0.62–0.70, HL: 0.38–0.45; AL(holotype): 1.11; OL: 0.17–0.20; PL: 0.50–0.52; PW: 1.07–1.17; ESL: 1.32–1.45; EW: 1.43–1.55; AW: 1.28–1.42; MTbL(holotype): 0.62, MTrL(holotype): 0.25 (MTrL 1–4: 0.18; MTrL 5: 0.07); AedL: 0.53–0.60; TL: 2.49–3.34 (holotype: 3.22).</p><p>Body wide, subconvex. Body yellow-brown to reddish brown, some specimens with darker head and abdomen (immature specimens with reddish middle part of head and longitudinal narrow spot on middle of pronotum); antennomeres 4–11 reddish-brown to brown; lateral portions of pronotum and some specimens with shoulders of elytra yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Posterior portions of infraorbital ridges with indistinct transverse andrugose microsculpture (some paratypes without sculpture between punctures there) and abdominal tergites with very indistinct, fine isodiametric microsculpture sculpture or without it. Head with irregular, moderately large and deep punctation, distinctly sparser in middle part (some paratypes only with several punctures) and denser on infraorbital ridges, punctures on posterior part of infraorbital ridges smaller; middle part of neck with irregular, small and sparse punctation; pronotum with irregular dense punctation, larger and deeper than that on head (some paratypes with interspaces between punctures on medioapical part of pronotum as long as puncture diameter), sparser on lateral portions, some specimens with impunctate small area between mediobasal impression; visible part of scutellum with several small punctures or without them; punctation of elytra as that on pronotum, denser and smaller on parascutellar area and along suture, each elytron with indistinct, five to seven vague and tangled longitudinal rows of punctures, usually with moderately wide impunctate longitudinal area between rows 3 and 4; abdominal tergites with very indistinct small punctation. Habitus as in Fig. 78.</p><p>Head 1.5–1.6 times as wide as long; middle portion somewhat flattened or slightly elevated, with very indistinct or distinct and moderately deep transverse impression between middle and posterior third of eyes, with very short and some with indistinct small grooves in front of ocelli usually as long as diameter of ocellus or slightly longer; posterior ridges smooth; anterior portion between antennal insertion and anterior margin of eye with superficial wide notch. Ocelli large, situated at level of postocular ridges; distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp slightly less than three times as long as preceding segment. Length × width of antennomeres (holotype): 1: 0.13 × 0.06; 2: 0.10 × 0.05; 3–4: 0.11 × 0.04; 5: 0.10 × 0.05; 6–8: 0.09 × 0.05; 9–10: 0.07 × 0.06; 11: 0.15 × 0.06.</p><p>Pronotum transverse, distinctly more than twice as wide as long, 1.6–1.7 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin slightly narrower than posterior margin; anterior angles widely rounded and strongly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with irregular and very smooth crenulation; pronotum with very wide markedly elevated middle portion, without or with indistinct, small longitudinal impression on medioapical part and with a pair of indistinct or distinct and deep semioval impressions on mediobasal third.</p><p>Elytra slightly broader than long, more than two and half times as long as pronotum, distinctly widened apicad, reaching apical margin of abdominal tergite IV to VII; surface between punctures on medioapical and middle portions of each elytron with irregular longitudinal elevations in some specimens.</p><p>Male. Apical margin of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 89) bulbose basally, with median lobe markedly, gradually narrowing toward widely rounded apex; parameres narrow, significantly exceeding apex of median lobe, with two small apical and preapical setae and with two additional setae in some specimens (Fig. 91); internal sac moderately wide, elongate, spiralled in basal part. Aedeagus laterally as in Fig. 90.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Anthobium splendidulum Shavrin &amp; Smetana, sp.n. may be distinguished from all remaining species of the fusculum group by the very short grooves in front of ocelli and by parameres significantly exceeding apex of median lobe.</p><p>Distribution. The species is known from several locations (Fig. 92) in Gaoligong Shan, Meili Xue Shan and Baima Shan ranges in Yunnan, China.</p><p>Bionomics. Specimens were collected at elevations from 2580 to 4225 m a.s.l. by sifting leaf litter and mosses in mixed forests; one specimen from Gaoligong Shan was taken by sifting litter under shrubs and bamboo (locality: CH 09-22). The locality of one paratype which was collected in Gaoligong Shan (locality: CH 0721) is shown in Fig. 93.</p><p>Etymology. The specific epithet is the Latin adjective splendidus, - a, - um (splendid). It refers to the external attractiveness of the species.</p></div>	https://treatment.plazi.org/id/03EB87F8F15AFF99FF31FA4B7DCE4E6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F15FFF9AFF31F91278664BAA.text	03EB87F8F15FFF9AFF31F91278664BAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium alticola (Coiffait 1977) Shavrin & Smetana 2018	<div><p>Anthobium alticola (Coiffait, 1977) comb.nov.</p><p>(Figs. 94, 96–97, 100)</p><p>Deliphrum alticola Coiffait, 1977: 265</p><p>Deliphrum alticola: Herman, 2001: 281; Smetana, 2004: 242; Schülke &amp; Smetana, 2015: 312</p><p>Type material examined: Holotype by monotypy of Deliphrum alticola Coiffait, 1977 ♀: ‘Umg. Mahidoela-Pass |</p><p>5000m,b.Maharigaon’ &lt;rectangular label, printed&gt;, ‘Gebiet von Jumla | Westnepal, lg.H. Franz’ &lt;rectangular label, printed&gt;, ‘TYPE’ &lt;red rectangular label, printed&gt;, ‘ Deliphrum | alticola Cam | H.Coiffait det. 1976’ &lt;rectangular label, printed, except handwritten “76”&gt; (NHMW; Franz collection).</p><p>Additional material: 1 ♂: ‘NEPAL-HIMALAYA | SE-Annapurna mts. | leg. O. Jäger 1997’, ‘ Telbrung Danda | 3700m, 9.VI’ (SNSD) .</p><p>Redescription. Measurements (n=2): HW: 0.67–0.68; HL: 0.40; AL(holotype): 1.11; OL: 0.17–0.19; PL: 0.60; PW: 1.20; ESL: 1.52–1.55; EW: 1.42–1.48; AW: 1.15–1.45; MTbL(holotype): 0.75, MTrL(holotype): 0.40 (MTrL 1–4: 0.25; MTrL 5: 0.15); AedL: 0.70; TL: 3.50.</p><p>Body moderately elongate, convex. Body and antennomeres 4–11 brown (holotype paler, with yellow-brown elytra); lateral, basal and apical parts of pronotum yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Body very shiny, forebody with bronze sheen; anterior portion of head between antennal insertion and anterior margin of eye with fine transverse meshes, posterior parts of head between ocelli and posterior margins of eyes with traces of fine transverse sculpture; abdomen with distinct isodiametric microsculpture. Head with irregular small punctation, slightly denser on infraorbital ridges; middle part of neck with irregular small punctation; pronotum with punctation markedly denser, larger and deeper than that of head, sparser on basal third and lateral portions; scutellum without or with several small punctures; punctation of elytra markedly larger and deeper than that on pronotum, dense, distinctly smaller on parascutellar area, each elytron with six, vague and tangled longitudinal rows of punctures. Habitus as in Fig. 94.</p><p>Head 1.6 times as wide as long; middle portion of head somewhat flattened, without transverse impression in middle; occipital furrow between ocelli wide and deep; grooves in front of ocelli long and deep, reaching apical third of length of eye; postocular ridges distinct, smooth; anterior portion between antennal insertion and anterior margin of eye with wide and moderately deep semicircular notch. Ocelli large and very convex, situated at level of postocular ridges, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Length × width of antennomeres (holotype): 1: 0.15 × 0.05; 2: 0.10 × 0.05; 3: 0.12 × 0.04; 4: 0.07 × 0.04; 5: 0.10 × 0.05; 6: 0.10 × 0.06; 7–10: 0.08 × 0.07; 11: 0.15 × 0.07.</p><p>Pronotum twice as wide as long, 1.7 times as wide as head, widest in middle, evenly narrowed both posteriad and anteriad; anterior angles widely rounded, not protruded anteriad; posterior angles obtuse or rounded; lateral edges of pronotum with irregular smooth crenulation; disc of pronotum with wide and markedly convex middle elevation, with pair of indistinct transverse impression on mediobasal third; lateral portions wide and slightly explanate, each with depp oval pit in middle.</p><p>Elytra slightly longer than wide, slightly widened apicad, reaching apical margin of abdominal tergite V, about two and half times as long as pronotum, with widely rounded apical margins truncated at suture; lateral portions moderately narrow, slightly explanate; latero-apical margins with small flattened crenulation.</p><p>Abdomen distinctly narrower than elytra, with a pair of small rounded tomentose wing-folding spots in middle of tergite V.</p><p>Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 96) with median lobe wide basally, gradually narrowed toward truncate apex; parameres markedly exceeding apex of median lobe, with two pairs of short apical and preapical setae. Aedeagus laterally as in Fig. 97.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Based on the shape of the convex body, the flattened head and the sparse punctation of the forebody, A. alticola is most similar to the Himalayan A. ivani Shavrin &amp; Smetana, 2017, from which it differs by shorter body, paler coloration, longer elytra and by the shape and internal structure of the aedeagus.</p><p>Distribution. The species is at present known from two locations in central and eastern Nepal (Fig. 100).</p><p>Bionomics. Specimens were collected at elevations from 3700 and 5000 m a.s.l. The detailed ecological data are unknown.</p><p>Remarks. The study of the holotype and the additional male from central Nepal revealed that Deliphrum alticola belongs to the nigrum group of the genus Anthobium (Shavrin &amp; Smetana 2017) .</p></div>	https://treatment.plazi.org/id/03EB87F8F15FFF9AFF31F91278664BAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F15CFF84FF31FC1B7D0448DB.text	03EB87F8F15CFF84FF31FC1B7D0448DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium flaveolum Shavrin & Smetana 2018	<div><p>Anthobium flaveolum Shavrin &amp; Smetana, sp.n.</p><p>(Figs. 95, 98–100)</p><p>Type material examined: Holotype ♂ [specimen dissected]: ‘ NEPAL:nr. Sanghu [underlined by yellow] | Middle oak for. [handwritten] | 27°19[handwritten]'N,87°31[handwritten]' | 26.x. [handwritten]1961 [1-handwritten] | c. 9350 ft. [handwritten]’ &lt;rectangular label, printed&gt;, ‘ Funnel sample | no. 155[handwritten]’ &lt;rectangular label, printed&gt;, ‘BMNH Nepal [underlined by yellow] | Exp. 1961-1962 | B.M. 1975-287’ &lt;rectangular label, printed &gt;, ‘ HOLOTYPE | Anthobium | flaveolum sp.n. | Shavrin A. &amp; Smetana A. 2018 ’ &lt;red rectangular label, printed&gt; (BMNH) .</p><p>Paratype ♀: same data as the holotype, ‘ PARATYPE | Anthobium | flaveolum sp.n. | Shavrin A. &amp; Smetana A. 2018 ’ &lt;red rectangular label, printed&gt; (BMNH) .</p><p>Description. Measurements (n=2): HW: 0.67–0.71, HL: 0.40–0.44; AL(holotype): 1.09; OL: 0.17–0.19; PL: 0.50–0.54; PW: 1.07–1.15; ESL: 1.47–1.48; EW: 1.47–1.48; AW: 1.17–1.20; MTbL(holotype): 0.70, MTrL(holotype): 0.35 (MTrL 1–4: 0.25; MTrL 5: 0.10); AedL: 0.40; TL: 3.15(holotype)–3.64.</p><p>Body elongate, very convex. Head reddish brown; antennomeres 4–11, pronotum, elytra and abdomen yellowbrown; mouthparts, antennomeres 1–3, lateral portions of pronotum and elytra, and legs yellow. Body very shiny, without microsculpture except for anterior portion of head between antennal insertion, and anterior margin of eye and posterior portions of head behind eyes with traces of transverse fine microsculpture; abdomen with distinct isodiametric microsculpture. Head with irregular small punctation, with moderately wide impunctate areas in middle and on infraorbital ridges; middle part of neck with several small punctures; pronotum with irregular and sparse punctation, markedly deeper than that on head, denser on medioapical portions, with wide impunctate area between mediobasal impression; scutellum with several small punctures; punctation of elytra larger and deeper than that on pronotum, denser and smaller on parascutellar area and along suture, each elytron with indistinct six, vague and tangled longitudinal rows of punctures, with moderately wide impunctate longitudinal areas between rows 3 and 4; abdominal tergites without visible punctation. Habitus as in Fig. 95.</p><p>Head 1.6 times as wide as long; middle portion of head somewhat flattened or very gently elevated, without or with (holotype) very indistinct transverse impression in middle; occipital furrow between ocelli wide and deep; grooves in front of ocelli narrow and deep, reaching posterior third of length of eye; postocular ridges smooth; anterior portion between antennal insertion and anterior margin of eye with wide semicircular notch. Ocelli large and convex, situated at level of posterior margins of eyes, distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Length × width of antennomeres (holotype): 1: 0.13 × 0.05; 2: 0.10 × 0.04; 3: 0.11 × 0.04; 4: 0.08 × 0.04; 5: 0.08 × 0.05; 6–8: 0.09 × 0.05; 9: 0.09 × 0.06; 10: 0.08 × 0.06; 11: 0.15 × 0.06.</p><p>Pronotum slightly more than twice as wide as long, 1.5–1.6 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; anterior angles widely rounded, slightly protruded anteriad; posterior angles more or less rectangular; lateral edges of pronotum with very indistinct, irregular smooth crenulation; disc of pronotum with wide, convex middle elevation with distinct and wide, moderately deep semicircular impression on mediobasal third; lateral portions wide and distinctly explanate, each with very deep oval pit slightly in front of middle.</p><p>Elytra as wide as long, slightly widened to apical third and widely rounded apically, reaching basal margin of abdominal tergite V, slightly less than three times as long as pronotum; lateral portions narrow, slightly elevated; latero-apical margins with five relatively large, acute teeth of crenulation becoming flattened toward middle.</p><p>Abdomen very narrow, with indistinctly elevated paratergites, with a pair of small, rounded tomentose wingfolding spots near apical margin of tergite V.</p><p>Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (Fig. 98) with median lobe wide basally, gradually narrowing toward rounded apex; parameres rather short, somewhat exceeding apex of median lobe, with two apical and preapical setae; internal sac simple, very narrow. Aedeagus laterally as in Fig. 99.</p><p>Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</p><p>Comparative notes. Anthobium flaveolum Shavrin &amp; Smetana, sp.n. may be distinguished from all remaining species of the nigrum group by the pale coloration of the body, the very indistinct lateral crenulation of the pronotum, the moderately large teeth on latero-apical margins of elytra, and by the shape and internal structure of the aedeagus.</p><p>Distribution. The species is known only from the type locality in eastern Nepal (Fig. 100).</p><p>Bionomics. Specimens were collected at elevation about 2800 m a.s.l. The detailed ecological data are unknown.</p><p>Etymology. The specific epithet is the Latin adjective flaveolus, - a, - um (yellowish). It refers to the coloration of the body.</p></div>	https://treatment.plazi.org/id/03EB87F8F15CFF84FF31FC1B7D0448DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
03EB87F8F142FF84FF31FEC479844C07.text	03EB87F8F142FF84FF31FEC479844C07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthobium nigrum (Cameron 1924)	<div><p>Key to species of Anthobium nigrum group (modified Shavrin &amp; Smetana 2017)</p><p>1 Lateral edges of pronotum with very indistinct smooth crenulation, latero-apical margins of elytra with moderately large teeth. Head reddish-brown, pronotum, elytra and abdomen yellow-brown. Aedeagus as in Fig. 98. Habitus as in Fig. 95. Body length: 3.15–3.64 mm. Nepal ................................................... A. flaveolum Shavrin &amp; Smetana, sp.n.</p><p>- Lateral edges of pronotum with distinct crenulation, latero-apical margins of elytra with small acute teeth. Body darker..... 2</p><p>2 Anterior angles of pronotum strongly protruded anteriad. Elytra markedly wider than long. Aedeagus as in Fig. 77 (Shavrin &amp; Smetana 2017). Habitus as in Fig. 67 (Shavrin &amp; Smetana 2017). Body length: 3.90–4.20 mm. China: Yunnan ................................................................................. A. conjunctum Shavrin &amp; Smetana, 2017</p><p>- Anterior angles of pronotum slightly protruded anteriad. Elytra as wide as long or slightly longer than wide.............. 3</p><p>3 Punctation of pronotum very dense. Aedeagus as in Fig. 80 (Shavrin &amp; Smetana 2017). Habitus as in Fig. 68 (Shavrin &amp; Smetana 2017). Body length: 3.50–4.27 mm. China: Yunnan ............... A. densepunctatum Shavrin &amp; Smetana, 2017</p><p>- Punctation of pronotum sparse.......................................................................... 4</p><p>4 Pronotum slightly flattened in cross section, with indistinct mediobasal impression. Aedeagus as in Fig. 72 (Shavrin &amp; Smetana 2017). Habitus as in Fig. 66 (Shavrin &amp; Smetana 2017). Body length: 4.15–4.37 mm. China: Gansu .................................................................................A. anishchenkoi Shavrin &amp; Smetana, 2017</p><p>- Pronotum very convex, with deep mediobasal impression...................................................... 5</p><p>5 Middle portion of head distinctly convex, with indistinct transverse impression. Aedeagus as in Fig. 70 (Shavrin &amp; Smetana 2017). Habitus as in Fig. 65 (Shavrin &amp; Smetana 2017). Body length: 3.44–4.75 mm. India ..... A. nigrum (Cameron, 1924)</p><p>- Middle portion of head flattened, without or with very indistinct transverse impression. Aedeagus different.............. 6</p><p>6 Elytra as wide as long. Body reddish-brown to black. Median lobe of aedeagus narrow, apical portions of parameres widened (Shavrin &amp; Smetana 2017: Fig. 82). Habitus as in Fig. 69 (Shavrin &amp; Smetana 2017). Body larger: 4.00– 4.71 mm. Nepal, Bhutan ...................................................................... A. ivani Shavrin &amp; Smetana, 2017</p><p>- Elytra slightly longer than wide. Body brown. Median lobe of aedeagus moderately wide, apical portions of parameres narrow (Fig. 96). Habitus as in Fig. 94. Body smaller: 3.50 mm. Nepal ............................ A. alticola (Coiffait, 1977)</p></div>	https://treatment.plazi.org/id/03EB87F8F142FF84FF31FEC479844C07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shavrin, Alexey V.;Smetana, Aleš	Shavrin, Alexey V., Smetana, Aleš (2018): A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group. Zootaxa 4508 (4): 451-506, DOI: 10.11646/zootaxa.4508.4.1
