taxonID	type	description	language	source
03ED65146305F97CBB81FD17FD47FDE0.taxon	materials_examined	Type species. Phanolinus discedens Sharp, 1884, fixed by original designation and monotypy.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146305F97CBB81FD17FD47FDE0.taxon	diagnosis	Diagnosis. Phanolinopsis stands out among other xanthopygines because it is one of few genera that has an impunctate or nearly impunctate pronotum (Figs. 7 – 12); with the exception of punctures around the margins of pronotum, species have at most four (or on rare occasions five) punctures at the center of the pronotum, each one delimiting the corner of a square. The only other xanthopygines with similar punctation patterns on the pronotum to Phanolinopsis are Ocyolinus amethystinus Sharp (Chatzimanolis & Ashe 2009; Chatzimanolis In press), and some species of Torobus Herman. A second key diagnostic characteristic of Phanolinopsis is the presence of an extended and punctate postmandibular ridge, visible both in dorsal (Figs. 7 – 12) and lateral (Figs. 13 – 18) views of the head that makes the posterolateral corners of the head appears ‘ angular’. This feature easily distinguishes Phanolinopsis from Ocyolinus amethystinus (in addition to the lack of long mandibles present in Ocyolinus amethystinus). Unfortunately, some species of Torobus have both a pronotum punctation pattern and a postmandibular ridge similar to Phanolinopsis. Thankfully, these taxa in Torobus can be easily distinguished from Phanolinopsis by the presence of a postcoxal process (absent in Phanolinopsis), and the presence of a porose structure on abdominal sternum VII in males (absent in Phanolinopsis). Other diagnostic features of Phanolinopsis include the following: head transverse (not subquadrate or trapezoid); antennomeres longer than wide or subquadrate but not transverse; mandibles short, blunt, with small tooth; neck prominent, with punctures; elytra shiny with medium to large punctures; and terga V – VI without subbasal (arch-like) carinae. Despite the name, Phanolinopsis is probably not closely related to Phanolinus. Besides the metallic coloration of the head and pronotum, these two genera share few other characteristics. Based on the structure of the antennae, and the morphology of the head and pronotum, Phanolinopsis is probably closely related to the genus Isanopus Sharp (Chatzimanolis 2008). The molecular phylogenetic analysis of Xanthopygina (Chatzimanolis 2014 b) did not include any specimens of Phanolinopsis, however, a forthcoming analysis (Chatzimanolis in prep.) that will incorporate morphological data (in addition to the molecular dataset) and a more comprehensive species list will provide more definitive answers.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146305F97CBB81FD17FD47FDE0.taxon	description	Description. Habitus as in Figs. 1 – 6, body medium sized, 9.5 – 15.7 mm in total length. Color of head and pronotum metallic green-brown, purple-brown, brown, blue, green or red; elytra metallic blue-green, blue-purple, purple-brown, brown-green, blue or green; mouthparts, antennae, mesoscutellum, ventral surface of body and legs, reddish brown to brown; abdomen reddish brown to brown except posterior part of VII and VIII orange. Head transverse, with medium-sized to large setose punctures around margin of head and microsculpture; with two large punctures anteriorly, each adjacent to antenna; surface of epicranium matte due to micropunctures and microsculpture. Clypeus slightly emarginate; anteclypeus not expanded. Eyes medium to large, prominent, occupying 1 / 2 to 3 / 4 of lateral margins of head. Ventral surface of head with transverse microsculpture and large sparse punctures; postoccipital suture and ventral basal ridge present; infraorbital ridge pronounced posteriorly; postmandibular ridge present, prominent, extending from near mandible to lateral side of head, delineated by multiple setose punctures (Figs. 13 – 18); gular sutures separated throughout length with narrowest point between them medio-posteriorly; nuchal depression prominent forming well defined neck; neck with microsculpture, micropunctures and multiple small punctures. Antenna (Fig. 21) with antennomeres 1 – 3 with multiple rows of macrosetae; antennomeres 4 – 11 with few macrosetae but covered with microtrichiae; antennomeres 1 – 9, 11 longer than wide; antennomere 10 longer than wide or subquadrate; antennomere 1 twice as long as antennomere 2; antennomere 3 longer than 2; antennomeres 4 – 7 subequal in size; antennomeres 8 – 10 subequal in size, typically shorter than antennomeres 4 – 7. Mouthparts with labrum medially emarginate to its base. Mandibles as in Fig. 19; small curved, blunt; left mandible with small bicuspid molar; right mandible with small tooth medially; mandibles with dorsolateral groove extending from condyle to just above tooth; prostheca setose. Maxilla as in Fig. 20; galea and lacinia densely setose; maxillary palpi 4 - segmented; P 1 small, about 1 / 3 as long as P 2; P 2 curved, elongate, longer than P 3; P 2 – P 3 with large setae apically; P 4 elongate, subequal in length to P 3. Hypopharynx as in Fig. 22. Submentum with one long and one shorter anterolateral setae in each end; labial palpi 3 - segmented; with transverse microsculpture; P 1 subequal in length to P 2; both P 1 and P 2 with several long setae; P 3 with distal end slightly dilated, not securiform. Pronotum subquadrate to slightly longer than wide; lateral margins of pronotum concave in dorsal aspect; pronotum broadest in apical 1 / 3 and narrower at basal angles. Hypomeron expanded, with transverse microsculpture and few micropunctures; superior and inferior marginal lines of hypomeron separate throughout their lengths; superior line fully visible from above, extending around anterolateral margin of pronotum and contacting inferior line at neck fossa; no portion of dorsum of pronotum visible from below. Surface of pronotum matte due to microsculpture and micropunctures; with scattered large setose punctures around margins but with at maximum 4 – 5 punctures on pronotum disc; margins of pronotum with several large setae. Postcoxal process absent. Mesoscutellum prominent, with dense polygon-shaped microsculpture and micropunctures; with multiple rows of small punctures. Basisternum with dense polygon-shaped microsculpture and carina; anterior marginal depression present; furcasternum with medial carina pointed vertically; furcasternum without polygon-shaped microsculpture. Elytra slightly shorter than pronotum; with multiple rows of nearly confluent punctures and large setae; with sparse longitudinal microsculpture; elytra appearing shining due to lack of extensive microsculpture. Hind wings fully developed. Mesoventrite with anterior margin forming “ lip ”; with dense polygon-shaped microsculpture and few punctures along edges; without median carina. Metaventrite with dense uniform medium-sized punctures; metaventral process small, rounded, with v-shaped emargination. Legs with tarsal segmentation 5 - 5 - 5; meso- and metatibia with multiple rows of spurs; protibia without multiple rows of spurs but with single row of spurs apically. Protarsus enlarged in both sexes, with spatulate setae ventrally; meso- and metatarsus not enlarged. Empodium with two small setae. Abdomen with paired prototergal glands present; abdomen expanding from segment III to segment V (widest) and then becoming narrower towards segment VIII. Abdominal terga III – V with tergal basal and without subbasal (arch-like) carina. Segments with distinctive microsculpture (Figs. 23 – 24) anteriorly and on anterolateral corners; sterna with deep elongate uniform punctation in multiple rows; terga V – VII with elongate punctures containing microsculpture. Males with secondary sexual structures of sternites VIII – IX; without porose structure on abdominal sternum VII; sternum VIII with emargination posteriorly; sternite IX with U or V-shaped emargination. Lateral tergal sclerites of the abdominal segment IX long and straight, covered with long macrosetae. Male genitalia with aedeagus typical of Xanthopygina (Figs. 25 – 42); with median lobe longer and wider than paramere; median lobe with two dorsal teeth; paramere long, not divided into lobes. Paramere with peg setae and short apical setae. Spermatheca not sclerotized.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146301F970BB81FADAFE33FA83.taxon	materials_examined	Type material. Holotype, female, on paper card mount in Sharp’s handwriting: “ ♀ Phanolinus discedens, Type, D. S. ” / “ Type ” / “ Bugaba, Panama. Champion. ” / “ B. C. A. Col. I. 2. Phanolinus discedens, Sharp ” / “ Sharp Coll. 1905 - 313. ” / “ HOLOTYPE Phanolinus discedens Sharp, 1884 det. R. G. Booth 2011 ”. In the collection of BMNH. Sharp (1884) mentioned that he had a single female specimen, which is the holotype. Additional Material. COSTA RICA: Puntarenas: Altamira Biological Station, 1510 – 1600 m, 9 ° 01.76 ’ N 83 ° 00.49 ’ W, 4 – 7. vi. 2004, J. S. Ashe, Z. Falin, I. Hinojosa, FIT, CR 1 AFH 04 144, SM 0606681, SM 0607497, SM 0607486 (3 ♂ SEMC); Las Alturas Biological Station, 1660 m, 8 ° 56.17 ’ N 82 ° 50.01 W, 31. v. – 3. vi. 2004, J. S. Ashe, Z. Falin, I. Hinojosa, FIT, CR 1 AFH 04 0 92, SM 0606932, SM 0606860 (2 ♂ SEMC); same locality, 1500 m, 27. v. 1993, J. S & A. K. Ashe, # 63, FIT, SM 0079878 (1 ♂ SEMC); PANAMA: Bocas del Toro: Cerro Pata de Macho Trail, W. of Cerro Horqueta, near Boquete, 8 ° 53 ’ N 82 ° 23 ’ W, 1780 m, 10 – 12. viii. 1987, D. M. Olson, # 754, lower montaine rainforest, pitfall trap, Field Museum (1 ♂ FMNH); Chiriquí: Cerro Pelota, 4 km N. Sta. Clara, 1500 m, vii. 1982, B. Gill (2 ♂ CNC); La Fortuna, Continental Divide Trail, 8 ° 46 ’ N 82 ° 12 ’ W, 1150 m, 23. v. – 9. vi. 1995, J. S. Ashe, R. Brooks, # 155, FIT, SM 0007031; SM 0079927 (1 ♂ SEMC, 1 ♂ UTCI); same locality, 9. vi. 1995, R. Anderson, PAN 2 A 95 10 F, Berlese forest litter, SM 0035769 (1 ♂ SEMC); La Fortuna, Hydrolog. trail, 8 ° 42 ’ N 82 ° 14 ’ W, 1150 m, 23. v. – 9. vi. 1995, J. S. Ashe, R. Brooks, # 156, FIT, SM 0003744 (1 ♂ SEMC); same locality and collectors, 1200 m, 9 – 12. vi. 1995, # 187, FIT, SM 0079928 (1 ♀ SEMC); Unknown Country: (1 ♂ FMNH).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146301F970BB81FADAFE33FA83.taxon	diagnosis	Diagnosis. Among all species of Phanolinopsis, P. discedens can be diagnosed based on the following characteristics: medium size (9.5 – 13.2 mm); disc of pronotum with four punctures at the center of the pronotum, each one delimiting the corner of a square (few specimens with less punctures); coloration of head and pronotum not red; posterolateral corners of head not extremely pointed; aedeagus as in Figs. 25 – 27. Phanolinopsis metaksenios is sympatric with P. discedens through much of their range, but these two species can be distinguished from each other based on their coloration (pronotum and elytra different color in P. discedens (Fig. 1); similar color in P. metaksenios (Fig. 5 )); the distance between the eye and the postmandibular ridge in lateral view (short in P. discedens (Fig. 13); long in P. metaksenios (Fig. 17 )), and the shape of the aedeagus (tip of median lobe pointed and apex of paramere narrow in P. discedens (Figs. 25 – 27); tip of median lobe wider and rounded and apex of paramere wide in P. metaksenios (Figs. 37 – 39 )).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146301F970BB81FADAFE33FA83.taxon	description	Description. Body length 9.5 – 13.2 mm. Head and pronotum metallic green-brown; elytra dark metallic bluegreen. Mouthparts, antennae, mesoscutellum, ventral surface of thorax and legs reddish brown. Abdominal terga and sterna reddish brown but with darker areas medially; posterior half of terga slightly lighter color; posterior 1 / 5 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.47. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin), but with no other punctures on epicranium. Eyes large, length of eyes / length of head ratio = 0.66, distance between eyes as wide as 1.44 times length of eye. Area between postmandibular ridge and eye (lateral side of head) narrow, with polygon-shaped microsculpture; posterolateral corner of head not pointed. Antennomeres 1 – 9, 11 longer than wide; antennomere 10 subquadrate. Neck with micropunctures, microsculpture, and with sparse small punctures. Pronotum subquadrate, width: length ratio = 0.95; surface of pronotum uniformly covered with sparse micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum with four large punctures at center, each one delimiting corner of square, but some specimens with three or fewer punctures (at least one puncture present). Elytra with large uniform punctures (about 10 punctures / elytron width); punctures almost confluent. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga sparsely punctate with small to medium-sized punctures. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 25 – 27; in dorsal view paramere converging to rounded tip; paramere shorter and narrower (apically) than median lobe; in lateral view paramere straight, narrower apically; paramere with peg setae along the lateral margins from tip to middle. Median lobe in dorsal view wide, converging to pointed apex, with two broad teeth apically; in lateral view becoming much narrower near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146301F970BB81FADAFE33FA83.taxon	distribution	Distribution. Known from the province of Puntarenas in Costa Rica and the provinces of Bocas del Toro and Chiriquí in Panama. Habitat. Collected with flight intercept traps, pitfall traps and in leaf litter from cloud forests between 1150 – 1780 m.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146301F970BB81FADAFE33FA83.taxon	discussion	Remarks. An additional specimen from FMNH can be tentatively included here. The specimen was labeled as “ Trigonopselaphus assmanni ” by Bierig as a manuscript type from San Isidro del Cotou, Costa Rica. However, I hesitate to include it above with the other “ additional specimens ” since the specimen is in bad condition and it is hard to see all the characters.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630CF972BB81FAF3FC4DFF78.taxon	materials_examined	Type material. Holotype, here designated, male, with labels: “ Guat. [emala], Chim. [altenango], 5 mi. S. Acatenago, 2400 m, 2. ix. [19] 72, J. Helava ” / “ Eugastus bicolor Shr. Smetana det. 1992 ” / “ New genus? not Eugastus det. A. Davies ” / “ Holotype Phanolinopsis erythros Chatzimanolis, des. Chatzimanolis 2017 ”. In the collection of CNC. Paratype. One, male, with labels: “ Panama, Veraguas Prov., 8 km W. Santa Fe, Cerro Tute, el. 3000 ft, 8 ° 30 ’ 26 ’’ N 81 ° 6 ’ 49 ’’ W, 24 – 26. vii. 1999, malaise, J. B. Woolley 99 / 057 ” / “ Duplicate ex series at TAMU (3), Field Museum Nat. His. ” / “? Eugastus det. Newton 2002 ” / “ Paratype Phanolinopsis erythros Chatzimanolis, des. Chatzimanolis 2017 ”. In the collection of FMNH.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630CF972BB81FAF3FC4DFF78.taxon	diagnosis	Diagnosis. Phanolinopsis erythros is easily distinguished from other species of Phanolinopsis based on the bright red coloration of the head, pronotum, mesoscutellum, legs, and ventral side of the body. Additionally, the tip of the paramere in dorsal view is emarginate (flat or rounded in other species). This species obviously does not belong in Philothalpus Kraatz (the current name for Eugastus Sharp), which belongs in a different subtribe, despite the references made above on the labels (see Chatzimanolis & Ashe 2005 and Chani-Posse et al. 2017 for more details on Philothalpus).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630CF972BB81FAF3FC4DFF78.taxon	description	Description. Body length 13.8 – 14.2 mm. Head and pronotum bright red; elytra dark metallic blue-purple or blue-green. Mesoscutellum red with brown border; mouthparts, ventral surface of thorax and legs red to reddish brown; antennomeres red but antennomeres 4 – 11 with darker setae. Abdominal terga and sterna III – IV red; V – VII reddish brown with brown area medially; posterior 1 / 3 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.38. Epicranium with mainly transverse and few polygon-shaped microsculpture; with sparse micropunctures; with few large to medium-sized punctures around margin of head; and numerous medium-sized punctures in 3 – 4 rows from lateral margins to center and from posterior margin to center; center of epicranium impunctate. Eyes medium-sized, length of eyes / length of head ratio = 0.48, distance between eyes as wide as twice length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, with transverse microsculpture; posterolateral corner of head not pointed. Antennomeres 1 – 11 longer than wide. Neck with micropunctures, microsculpture, and with many small punctures. Pronotum subquadrate, width: length ratio = 0.92; surface of pronotum uniformly covered with sparse micropunctures; appearing shiny. Pronotum with few large punctures around margin; disc of pronotum with four large punctures at center, each one delimiting corner of square. Elytra with large to medium-sized punctures (about 10 – 11 punctures / elytron width); distance punctures from almost confluent to 0.5 times width of puncture; punctures more clustered together near lateral margins. Elytra appearing shiny; with sparse polygon-shaped microsculpture. Abdominal terga with at least 3 – 4 rows of medium-sized punctures each. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Females unknown. Aedeagus as in Figs. 28 – 30; in dorsal view paramere wide, parallel-sided, with emarginate apex; paramere shorter and narrower than median lobe; in lateral view paramere slightly concave; paramere with peg setae as in Fig. 30. Median lobe in dorsal view narrow, converging to pointed apex, with two broad teeth apically; in lateral view becoming narrower and elongate near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630CF972BB81FAF3FC4DFF78.taxon	distribution	Distribution. Known from the department of Chimaltenango in Guatemala and the province of Veraguas in Panama. Habitat. Collected with malaise traps from elevation between 914 – 2400 m in cloud forests. Etymology. The specific epithet is derived from the Greek word ερΥΘρός (red) and refers to the coloration of the head and pronotum. The epithet is treated as a noun in apposition.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630EF973BB81FEE3FAA5FC25.taxon	materials_examined	Type material. Holotype, female, with labels: “ Columbia occ. Cali. Fassl ” / “ Cañon del Monte Tolima, 1700 m, iiiii. 1910 ” / “ fassli Bern. Typus ” / “ Chicago NHMus, M. Bernhauer Collection ” / HOLOTYPE teste A. Westricht 2015 GDI imaging Project ” / “ PHOTOGRAPHED Kelsey Keaton 2015 Emu Catalog ” / “ FMNHINS 3047816 Field Museum Pinned ” / “ Phanolinopsis fassli (Bernhauer) det. Chatzimanolis 2017 ”. In the collection of FMNH. Bernhauer (1917) added the following to the locality label on the specimen: “ Umgebung von Cali am Cauca ”, and mentioned that he had only a single female specimen, which is the holotype. Additional Material. Unknown Country: Coll. Kraatz (1 ♂ SDEI).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630EF973BB81FEE3FAA5FC25.taxon	diagnosis	Diagnosis. Among all species of Phanolinopsis, P. fassli and P. norahae are easily distinguished from other species due to the lack of punctures on the disc of the pronotum in these two species. Phanolinopsis fassli can be distinguished from P. norahae based on the following: head and pronotum with weak micropunctation (Fig. 9; much starker in P. norahae, Fig. 12); reduced microsculpture between eyes and postmandibular ridge (Fig. 15; not reduced in P. norahae, Fig. 18); tip of paramere flat (Figs. 31, 33; more rounded in P. norahae, Figs. 40, 42). The two species also differ in coloration (Figs. 3, 6).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630EF973BB81FEE3FAA5FC25.taxon	description	Description. Body length 12.6 – 14.0 mm. Head, pronotum and elytra metallic purple-brown with green overtones. Mouthparts, antennae, mesoscutellum, ventral surface of thorax, legs and abdomen brown. Posterior half of abdominal terga slightly lighter color; posterior 1 / 4 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.32. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin), but with no other punctures on epicranium. Eyes medium-sized, length of eyes / length of head ratio = 0.39, distance between eyes as wide as 2.67 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, without polygon-shaped microsculpture but with sparse transverse microsculpture; posterolateral corner of head not pointed. Antennomeres 1 – 8, longer than wide; antennomeres 9 – 11 missing. Neck with micropunctures, microsculpture, and with dense small punctures. Pronotum subquadrate, width: length ratio = 0.98; surface of pronotum uniformly covered with sparse micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum impunctate. Elytra with medium-sized punctures (about 12 – 13 punctures / elytron width); punctures almost confluent. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga with 1 – 2 rows of small punctures. Male secondary sexual structures with sternum VIII having shallow V-shaped emargination medially; sternum IX with deep V-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 31 – 33; in dorsal view paramere converging to flat tip; paramere shorter and narrower (apically) than median lobe; in lateral view paramere concave apically; paramere with peg setae as in Fig. 33. Median lobe in dorsal view wide, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrower near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630EF973BB81FEE3FAA5FC25.taxon	distribution	Distribution. Known from the type locality of west Colombia in the mountainous surrounding of Cali. Habitat. Unknown, but given the high altitude (1700 m), the species is probably found in cloud forests.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630FF974BB81FC52FBAEF82A.taxon	materials_examined	Type material. Holotype, here designated, male, with labels: “ Costa Rica: Puntarenas, Monteverde, Estacion Biol. Monteverde, 1730 m, 10 ° 19 ’ 10 ’’ N 84 ° 48 ’ 57 ’’ W, 12. vi. 2001, montane forest litter, R. Anderson, CR 1 A 01 107 ” / “ [barcode label] SM 0516461 ” / “ Holotype Phanolinopsis goniakos Chatzimanolis, des. Chatzimanolis 2017 ”. In the collection of SEMC. Paratypes. 45: same label as holotype, SM 0516460 (1 ♂ SEMC); “ Costa Rica: Puntarenas, Monteverde, Biol. Station, 1540 m, 10 ° 19.672 ’ N 84 ° 49.141 ’ W, 10 – 17. vi. 2001, S & J. Peck, 01 - 09, FIT, cloud forest, CR 1 P 01 001 ”, barcode label SM 0459582 (1 ♂ SEMC); “ Costa Rica: Puntarenas, Monteverde, 1570 m, 15. v. 1989, J. S. Ashe, R. Brooks, R Leschen, pitfall traps ” / “ Snow Entomol. Mus. Costa Rica Exped. # 205 ”, barcode labels SM 0079890, SM 0079896, SM 0079886, SM 0079905, SM 0079897, SM 0079906, SM 0079885, SM 0079887, SM 0079889, SM 0079899, SM 0079900, SM 0079901, SM 0079895, SM 0079888, SM 0079893, SM 0079894, SM 0079891, SM 0079892, SM 0079903, SM 0079904, SM 0079902, SM 0079898 (1 ♂, 1 ♀ UTCI; 17 ♂, 3 ♀ SEMC); same locality and collectors, 10. v. 1989, # 101, barcode labels SM 0079919, SM 0079924, SM 0079922, SM 0079921, SM 0079925, SM 0079923, SM 0079920 (1 ♂ UTCI; 5 ♂, 1 ♀ SEMC); same locality and collectors, 12. v. 1989, # 129, barcode labels SM 0079910, SM 0079911, SM 0079909 (3 ♂ SEMC); same locality and collectors, 9. v. 1989, # 074, barcode labels SM 0079908, SM 0079907 (1 ♂, 1 ♀ SEMC); same locality and collectors, 23. v. 1989, # 412, barcode labels SM 0079918, SM 0079917 (2 ♀ SEMC); same locality and collectors, Chomogo Trail, 1690 m, 23. v. 1989, # 410, barcode labels SM 0079915, SM 0079912, SM 0079914, SM 0079913 (2 ♂, 2 ♀ SEMC); same locality, 1520 m, 30. iv. 1986, J. S. Ashe, leaf litter, barcode label SM 0079926 (1 ♂ SEMC); same locality, 5000 ft, 23 – 24. v. 1979, J. M. & B. A. Campbell (1 ♂ CNC); “ Costa Rica: Guanacaste, Santa Elena, Santa Elena Cloud For. Reserve, 1650 m, 10 ° 20 ’ 42 ’’ N 84 ° 47 ’ 53 ’’ W, 11. vi. 2001, cloud for. litter, R. Anderson, CR 1 A 01 104 ”, barcode label SM 0516591 (1 ♂ SEMC). All paratypes with label “ Paratype Phanolinopsis goniakos Chatzimanolis, des. Chatzimanolis 2017 ”.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630FF974BB81FC52FBAEF82A.taxon	diagnosis	Diagnosis. This species can be easily distinguished from all other taxa of Phanolinopsis based on the morphology of the head. In P. goniakos the posterolateral corners of the head are pointed (Figs. 4, 10, 16) more so than any other known species.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630FF974BB81FC52FBAEF82A.taxon	description	Description. Body length 10.5 – 13.0 mm. Head and pronotum metallic brown, sometimes with green overtones; elytra dark metallic brown-green. Mouthparts, mesoscutellum, ventral surface of thorax and legs brown. Antennomeres 1 – 6 brown; antennomeres 7 – 11 dark yellow. Abdominal terga and sterna brown; posterior 1 / 5 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.45. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin); with 3 – 4 other large puncture near center but with no other punctures on epicranium. Eyes medium-sized, length of eyes / length of head ratio = 0.48, distance between eyes as wide as 2.07 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, with polygon-shaped microsculpture; posterolateral corner of head pointed. Antennomeres 1 – 9, 11 longer than wide; antennomere 10 subquadrate. Neck with micropunctures, microsculpture, and with sparse small punctures. Pronotum subquadrate, width: length ratio = 0.98; surface of pronotum uniformly covered with sparse micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum with four large punctures at center, each one delimiting corner of square. Elytra with medium-sized punctures (about 8 – 9 punctures / elytron width); distance between punctures 0.5 – 1 width of puncture. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga sparsely punctate with small to medium-sized punctures. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 34 – 36; in dorsal view paramere converging to rounded tip; paramere about as long as and narrower than median lobe; in lateral view paramere slightly concave apically; paramere with peg setae as in Fig. 36. Median lobe in dorsal view wide, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrower near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630FF974BB81FC52FBAEF82A.taxon	distribution	Distribution. Known from the provinces of Guanacaste and Puntarenas in Costa Rica. Habitat. Collected with pitfall traps, flight intercept traps and in leaf litter in cloud forests at elevations between 1520 – 1690 m.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630FF974BB81FC52FBAEF82A.taxon	etymology	Etymology. The specific epithet is derived from the Greek word γωνία (angle) and refers to the sharp angle of the posterolateral corners of the head. The epithet is treated as a noun in apposition.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146309F977BB81FF32FBA8FF55.taxon	materials_examined	Type material. Holotype, here designated, male, with labels: “ Costa Rica: Puntarenas, Las Alturas Stanford Bio. [logical] Stn. [Station], 29 km NE San Vito, 1500 m, 27. v. 1993, J. S. & A. K. Ashe, # 063, ex: flight intercept trap ” / “ [barcode label] SM 0079937 ” / Holotype Phanolinopsis metaksenios Chatzimanolis, des. Chatzimanolis 2017 ”. In the collection of SEMC. Paratypes. 35: “ Costa Rica: Heredia, La Selva, 3.2 km SE Puerto Viejo, 100 m, 21. iii. 1992, W. Bell. ex: flight intercept trap ”, barcode label SM 0079988 (1 ♂ SEMC); same locality and collector, 24. ii. 1992, barcode label SM 0079939 (1 ♂ SEMC); same locality and collector, 6. iii. 1992, barcode label SM 0079940 (1 ♂ SEMC); “ Costa Rica: Puntarenas Prov., Las Alturas Biol. Sta., 1660 m, 8 ° 56.17 N 82 ° 50.01 W, 31. v. – 3. vi. 2004, J. S. Ashe, Z. Falin, I. Hinojosa, ex. flight intercept trap, CR 1 AFH 04 092 ”, barcode labels SM 0606976, SM 0606985, SM 0606873, SM 0606871, SM 0606931 (2 ♂, 3 ♀ SEMC); “ Costa Rica: Puntarenas, San Vito, Estac. Biol. Las Alturas, Alturas, 2 km NE, 8 ° 56 ’ 56 ’’ N 82 ° 50 ’ 1 ’’ W, 1520 m, 20. vi. 1998, R. Anderson, CR 1 A 98 104 ex: Berlese leaf litter ”, barcode label SM 0115543 (1 ♂ SEMC); same locality, x. 1991, P. Hanson, barcode label SM 0079936 (1 ♂ SEMC); same locality, xii. 1991, malaise trap (1 ♂ MAIC); “ Costa Rica: Puntarenas Prov., Altamira Biol. Sta., 1510 – 1600 m, 9 ° 01.76 ’ N 83 ° 00.49 W, 4 – 7. vi. 2004, J. S. Ashe, Z. Falin, I. Hinojosa, ex: flight intercept trap, CR 1 AFH 04 144 ”, barcode labels SM 0607495, SM 0607496 (1 ♂ SEMC; 1 ♂ UTCI); “ Costa Rica: Puntarenas, 24 km W Piedras Blances, R. F. Golfo Dulce, 200 m, 8 ° 46 ’ 0 ’’ N 83 ° 24 ’ 0 ’’ W, 1. viii. – 30. ix. 1993, CR 1 H 95 - 96 18, P. Hanson, ex: Malaise trap ”, barcode labels SM 0134806, SM 0134813 (2 ♂ SEMC); same locality and collector, xi. 1990, barcode label SM 0079877 (1 ♂ SEMC); same locality and collector, xii. 1990, barcode label SM 0063457 (1 ♀ SEMC); “ Costa Rica: Puntarenas Prov., Las Cruces Biol. Sta., 1330 m, 8 ° 47.14 N 82 ° 57.58 ’ W, 28 – 31. v. 2004, J. S. Ashe, Z. Falin, I. Hinojosa, ex: flight intercept trap, CR 1 AFH 04 060 ”, barcode label SM 0606584 (1 ♂ SEMC); “ Costa Rica: [Puntarenas Prov.] San Vito de C. B. [Coto Brus], Las Cruces, vii. 1982, 1200 m, B. Gill, FIT ” (2 ♂ CNC); “ Costa Rica, Prov. Puntarenas, P. N. Corcovado, Sector La Leona, Cerro Puma, 100 – 300 m, 17. ix. – 5. x. 2003, K. Caballero, Tp. Intersección, # 6 L _ S _ 267000 _ 518900, # 75587 ”, barcode label INB 0003780485 (1 ♂ ZMUC); same locality, 19. vi. – 8. vii. 2003, M. Moraga, A. Azofeifa, K. Caballero, # 74479, barcode labels INB 0003736774, INB 0003736776 (1 ♂, 1 ♀ ZMUC); “ Costa Rica, Prov. Puntarenas, Fca. Cafrosa, Est. Las Mellizas, P. N. Amistad, 1300 m, J. C. Saborio, vi – vii. 1990, L _ S _ 316100 _ 596100 ”, barcode label INBIOCRI 000673323 (1 ♂ ZMUC); same locality, M. Ramirez, G. Mora, barcode label INBIOCRI 000162656 (1 ♀ ZMUC); “ Costa Rica: Puntarenas Prov., Hacienda La Amistad, 8 ° 56.395 ’ N 82 ° 47.465 ’ W, 1500 m, premont. moist forest, FIT, 9 – 11. vi. 2012, Solodovnikov, Brunke, Puliafico, Selvantharan ” / “ Chatzimanolis DNA Voucher, extraction: SC- 406, species: Phanolinopsis, extraction date: 27. iii. 2015 ” (1 ♂ ZMUC); “ Costa Rica, Prov. Puntarenas, Peninsula de Osa, Rancho Quemado, 200 m, 12 – 24. v. 1993, A. Gutiérrez, L _ S _ 292500 _ 511000 ”, barcode label INBIOCRI 000188826 (1 ♂ ZMUC); same locality, xi. 1992, F. Quesada, barcode label INBIOCRI 000917070 (1 ♀ ZMUC); “ Costa Rica [Cartago], Carpintera, 6. viii. 1941 ” / “ Field Mus. Nat. Hist. 1966 A. Bierig. Collen. Acc. Z- 13812 (1 ♀ FMNH); same labels except 29. i. 1939 (1 ♀ FMNH); same labels except 19. iv. 1940 (1 ♀ FMNH); same labels except 8. iv. [19] 39, “ Trigonopselaphus gymnus Brg [manuscript name by A. Bierig] (1 ♂ FMNH); “ Panama: Chiriqui Prov., 27.7 km W. Volcan, Hartmann’s Finca, 8 ° 45 ’ N 82 ° 48 ’ W, 1450 m, 16. vi. 1995, J. Ashe, R. Brooks, # 230 ”, barcode label SM 0058244 (1 ♀ SEMC); “ Panama: Chiriqui, Cerro Pelota, 4 km N. Sta. Clara, 1500 m, vii. 1982, B. Gill (1 ♂, 1 ♀ CNC). All paratypes with label “ Paratype Phanolinopsis metaksenios Chatzimanolis, des. Chatzimanolis 2017 ”.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146309F977BB81FF32FBA8FF55.taxon	diagnosis	Diagnosis. Among all species of Phanolinopsis, P. metaksenios can be identified based on the following characteristics: large size (11.0 – 15.5 mm); disc of pronotum with four punctures at the center of the pronotum, each one delimiting the corner of a square (few specimens with five punctures); coloration of head and pronotum not red; posterolateral corners of head not extremely pointed; aedeagus as in Figs. 37 – 39. Phanolinopsis metaksenios is sympatric with P. discedens through much of their range, but these two species can be distinguished from each other based on their coloration (pronotum and elytra different color in P. discedens (Fig. 1); similar color in P. metaksenios (Fig. 5 )); the distance between the eye and the postmandibular ridge in lateral view (short in P. discedens (Fig. 13); long in P. metaksenios (Fig. 17 )) and the shape of the aedeagus (tip of median lobe pointed and apex of paramere narrow in P. discedens (Figs. 25 – 27); tip of median lobe wider and rounded and apex of paramere wide in P. metaksenios (Figs. 37 – 39 )).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146309F977BB81FF32FBA8FF55.taxon	description	Description. Body length 11.0 – 15.5 mm. Head and pronotum metallic green-blue; elytra dark metallic blue with purple overtones. Mouthparts, antennae, mesoscutellum, ventral surface of thorax and legs brown and most of abdomen brown. Abdominal segment VII with variable coloration, either completely dark orange or brown with posterior 1 / 3 orange; segment VIII orange. Head transverse, width: length ratio = 1.44. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin); with numerous medium-sized punctures in 3 – 4 rows from lateral margins to center; center of epicranium impunctate. Eyes large, length of eyes / length of head ratio = 0.59, distance between eyes as wide as 1.65 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, with polygon-shaped microsculpture; posterolateral corner of head not pointed. Antennomeres 1 – 9, 11 longer than wide; antennomere 10 subquadrate. Neck with micropunctures, microsculpture, and with sparse small punctures. Pronotum slightly longer than wide, width: length ratio = 0.88; surface of pronotum uniformly covered with sparse micropunctures and dense rectangular-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum with four large punctures at center, each one delimiting corner of square; few specimens with five punctures. Elytra with mediumsized punctures (about 10 – 11 punctures / elytron width); distance between punctures from confluent to 0.5 times width of puncture. Elytra appearing shiny; with sparse longitudinal and polygon-shaped microsculpture. Abdominal terga with 2 – 3 rows of small punctures. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 37 – 39; in dorsal view paramere converging to rounded tip; paramere slightly shorter and narrower than median lobe; in lateral view paramere slightly concave apically; paramere with peg setae as in Fig. 39. Median lobe in dorsal view wide, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrow and elongate near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146309F977BB81FF32FBA8FF55.taxon	distribution	Distribution. Known from the provinces of Heredia and Puntarenas in Costa Rica and the province of Chiriquí in Panama. Habitat. Collected with flight intercept traps, malaise traps and in leaf litter at elevations between 100 – 1600 m at lowland and cloud rainforests.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED65146309F977BB81FF32FBA8FF55.taxon	etymology	Etymology. The specific epithet is derived from the Greek word µετάξι (silk) and refers to the silky appearance of the head and pronotum. The epithet is treated as a noun in apposition.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630BF968BB81FE87FF02FA8D.taxon	materials_examined	Type material. Holotype, here designated, male with labels: “ Colombia, Nariño, R. N. La Planada vía Hondón, 1 ° 15 ’ N 78 ° 15 ’ W, 1930 m, pitfall, 16 – 18. viii. 2000, G. Oliva leg., M 1025 ” / “ [barcode label] SM 0548892 ” / “ Holotype Phanolinopsis norahae Chatzimanolis, des. Chatzimanolis 2017 ”. In the collection of SEMC. Paratypes. Seven: “ Colombia, Nariño, R. N. La Planada Parcela Permanente, 1 ° 15 ’ N 78 ° 15 ’ W, 1885 m, malaise, 14 – 29. ii. 2004, G. Oliva leg., M 4353 ” / “ [barcode label] SM 0349446 ” (1 ♂ SEMC); “ Colombia, Nariño, R. N. La Planada Centro Administrativo, 1 ° 15 ’ N 78 ° 15 ’ W, 1700 m, red, 9 – 12. viii. 2004, D. Arias leg., M 4901 ” / “ [barcode label] SM 0349447 ” (1 ♂ SEMC); “ Colombia, Nariño, R. N. La Planada Parcela Olga, 1 ° 15 ’ N 78 ° 15 ’ W, 1850 m, malaise, 29. i. – 14. ii. 2004, G. Oliva leg., M 4356 ” / “ [barcode label] UTCI 000006014 ” (1 ♂ UTCI); “ Ecuador, Pichincha, 45 km NWW Quito, Macquipucuna Station, 1600 – 1650 m, 3 – 18. iv. 1996, ECU 1 H 96 0 12, P. Hibbs, ex: flight intercept trap ” / “ [barcode label] SM 0089851 ” (1 ♂ SEMC); same locality and collector, 18. v. – 5. vi. 1996, ECU 1 H 96 0 22, barcode label SM 0092764 (1 ♀ SEMC); “ Ecuador, Pichincha, Macquipucuna For. Res., 50 km NW Quito, 1750 m, 23. xii. 1991, C. Carlton, R. Leschen. # 62, ex: flight intercept trap ” / “ [barcode label] SM 0079941 ” (1 ♂ SEMC); “ Ecuador, Pichincha, Macquipucuna Biological Station, 0 ° 5 ’ 34 ’’ N 78 ° 37 ’ 37 ’’ W, 1600 m, 29. x. 1999, R. Anderson, ECU 1 A 99 214 D, ex: ridge-top montane forest litter, mixed Cecropia / cloud forest ” / “ [barcode label] SM 0366942 ” (1 ♀ SEMC). All paratypes with label “ Paratype Phanolinopsis norahae Chatzimanolis, des. Chatzimanolis 2017 ”.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630BF968BB81FE87FF02FA8D.taxon	diagnosis	Diagnosis. Among all species of Phanolinopsis, P. fassli and P. norahae are easily distinguished from other species due to the lack of punctures on the disc of the pronotum. Phanolinopsis norahae can be distinguished from P. fassli based on the following: head and pronotum with stark micropunctation (Fig. 12; much weaker in P. fassli, Fig. 9); dense microsculpture between eyes and postmandibular ridge (Fig. 18; reduced in P. fassli, Fig. 15); tip of paramere rounded (Figs. 40, 42; flatter in P. fassli, Figs. 31, 33). The two species also differ in coloration (Figs. 3, 6).	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630BF968BB81FE87FF02FA8D.taxon	description	Description. Body length 13.0 – 15.7 mm. Head, pronotum and elytra dark metallic blue or green with purple overtones. Mouthparts, mesoscutellum, ventral surface of thorax and legs brown. Antennomeres 1 – 8 brown; antennomeres 9 – 11 light brown. Abdominal terga and sterna brown; posterior 1 / 5 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.37. Epicranium with transverse and polygon-shaped microsculpture and dense micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin); with numerous medium-sized punctures in 2 – 3 rows from lateral margins to center; center of epicranium impunctate. Eyes medium-sized, length of eyes / length of head ratio = 0.48, distance between eyes as wide as 1.44 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, with stark polygon-shaped microsculpture; posterolateral corner of head not pointed. Antennomeres 1 – 9, 11 longer than wide; antennomere 10 subquadrate. Neck with micropunctures, microsculpture, and with dense small punctures. Pronotum subquadrate, width: length ratio = 0.94; surface of pronotum uniformly covered with dense micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum impunctate. Elytra with medium-sized punctures (about 11 – 12 punctures / elytron width); distance between punctures from confluent to 0.5 times width of puncture. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga with 1 – 2 rows of small punctures. Male secondary sexual structures with sternum VIII having shallow emargination medially; sternum IX with deep U-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 40 – 42; in dorsal view paramere parallel-sided, converging to rounded tip; paramere slightly shorter and narrower than median lobe; in lateral view paramere slightly concave apically; paramere with peg setae as in Fig. 42. Median lobe in dorsal view narrow, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrow and elongate near apex.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630BF968BB81FE87FF02FA8D.taxon	distribution	Distribution. Known from the department of Nariño in Colombia and the province of Pichincha in Ecuador. Habitat. Collected with flight intercept, malaise and pitfall traps at elevations between 1600 – 1930 m in cloud forests.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
03ED6514630BF968BB81FE87FF02FA8D.taxon	etymology	Etymology. The specific epithet is in honor of my daughter, Norah Chatzimanolis, who really likes blue beetles.	en	Chatzimanolis, Stylianos (2017): And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4323 (1): 49-67, DOI: 10.11646/zootaxa.4323.1.4
