taxonID	type	description	language	source
03EDDF2BFFC7F21FFF0CF9C21C52FB3A.taxon	description	(Figs 1 A – C, 4 A, 5 A, B, D, E, 7)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC7F21FFF0CF9C21C52FB3A.taxon	materials_examined	Type locality. Peru, Junín, Acolla near Jauja, 3,460 m, - 11.743 ° - 75.549 °. Type material. 1 Paratype female in coll. Guignot (MNHN) Perou, Acolla près, Jauja, 3460 m, III- 1955 / ♀ / Paratypes. (According to Guignot, 1955, collected in “ I- 1954 ”, by F. Blancas). Other material studied (526 exs.). 5 exs.: “ Peru: Junín, Jauja, Lagoon Tragadero, 3459 m, 04. viii. 2018, - 11.767 ° - 75.532 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 03) ”; 28 exs.: “ Peru: Junín, La Oroya, Nueva Ciudad Morococha, 4244 m, 06. viii. 2018, - 11.579 ° - 76.07 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 04) ”; 165 exs.: “ Peru: Junín, Junín, Lagoon Huacra, 4461 m, 14. viii. 2018, - 11.289 ° - 76.007 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 13) ”; 7 exs.: “ Peru: Junín, Junín, Lagoon Estancia Azulmito, 4452 m, 14. viii. 2018, - 11.303 ° - 76.989 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 14) ”; 1 ex.: “ Peru: Junín, Junín, Lagoon Yanamyacu, 4327 m, 14. viii. 2018 ,, - 11.314 ° - 75.956 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 16) ”; 25 exs.: “ Peru: Junín, Junín, Shoqui, 4324 m, 15. viii. 2018, - 11.046 ° - 75.897 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 18) ”; 5 exs.: “ Peru: Junín, Junín, Shoqui, 4330 m, 15. viii. 2018, - 11.049 ° - 75.9 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 19) ”; 1 ex.: “ Peru: Junín, Jauja, Lagoon Vinso, Patacancha, 3911 m, 15. viii. 2018, - 11.921 ° - 75.505 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 20) ”; 37 exs.: “ Peru: Cusco, Quispicanchi, Marcapata, Yanatturo, 4458 m, 23. viii. 2018, - 13.634 ° - 71.143 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 24) ”; 11 exs.: “ Peru: Cusco, Canchis, Checacupe, Cayena, 4613 m, 25. viii. 2018, 14.027 ° - 71.114 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 29) ”; 21 exs.: “ Peru: Cusco, Canas, Layo District, 4122 m, 26. viii. 2018, - 14.507 ° - 71.064 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 30) ”; 62 exs.: “ Peru: Cusco, Canas, Langui District, 32. Road 34 G, 3960 m, 26. viii. 2018, - 14.434 ° - 71.279 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 31) ”; 70 exs.: “ Peru: Cusco, Acomayo, Acopia District, Lagoon Pomacanchi, 3675 m, 27. viii. 2018, - 14.039 ° - 71.496 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 33) ”; 18 exs.: “ Peru: Cusco, Acomayo, Acopia District, Lagoon P′osqoqocha, 3846 m, 27. viii. 2018, - 14.085 ° - 71.481 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 34) ”. Notes. This is a moderately large, oval species with yellowish to brownish dorsal surface; the pronotum is generally rather dark in the middle with paler sides (Figs 1 A – C). Head with cervical line that can be faint in some specimens; pronotum with basal plicae; elytron with short basal plicae. TL 2.1 – 2.2 mm, TL-H 1.8 – 1.9 mm; TW 1.0 – 1.1 mm. Male genitalia as in Fig. 4 B, median lobe in lateral view with apical portion thin, paramere with apical piece moderately long and narrow. Dorsal surface of male shiny, female dull due to fine mircoreticulation. Metathoracic wings fully developed. BOLD platform. We provided 45 entries in the “ COLLI ” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC7F21FFF0CF9C21C52FB3A.taxon	distribution	Distribution. High Andes, central and southeastern Peru and Lake Titicaca of Bolivia (Fig. 7), ca. 3,400 – 4,600 m. Habitat. Shallow and exposed puddles, collected with strainer out of mats of vegetation (Figs 5 A, B, D, E, F; 6 C). Other Dytiscidae associated with this species were species of Rhantus Dejean, 1833 and Lancetes Sharp, 1882, as well as Liodessus alpinus sp. nov., L. rhigos sp. nov. and L. thespesios sp. nov.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC4F219FF0CFB061D57FF63.taxon	description	(Figs 1 D, E, 4 B, 6 D – F, 7)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC4F219FF0CFB061D57FF63.taxon	materials_examined	Type locality. Bolivia, La Paz Dept., Lake Titicaca, Huatajata, ca. 3,900 m, - 16.203 ° - 68.694 ° (our alt / lat / long data inferred from GoogleEarth). Type material. Holotype female (ZSM). “ Titicacasee, Huatajata ca. 4000 m, 6. - 17.7.1954 / Bolivia 1954, leg. W. Förster / Type ♀ / F. Guignot, 1955, Liodessus andinus n. sp., Type ♀ ”. 3 female paratypes (ZSM) from the same sampling event. According to Guignot, 1957, there is a male “ Allotype ” from Peru, Chucuito, Lake Titicaca, Chucuito, ca. 3,900 m, - 15.905 ° - 69.888 °, not studied here but possibly stored in the Guignot collection at MNHN. Other material studied (130 exs.). 2 exs., “ Peru: Cusco, Acomayo, Acopia District, Lagoon Misk′iqocha, 3787 m, 27. viii. 2018, - 14.123 ° - 71.460 °, Y. S. Megna & N. Zenteno (MUSM, ZSM); (PER _ YSM _ 2018 _ 35) ”; 128 exs., “ Peru: Cusco, Canchis, San Pedro District, 3486 m, 27. viii. 2018, - 14.189 ° - 71.339 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 36) ” (MUSM, ZSM). Notes. This is a moderately large and (in many but not all specimens) appearing more broadly oval species with characteristic coloration of darker spots on otherwise mainly yellowish dorsal surface (Figs 1 D, E). TL 2.3 – 2.6 mm, TL-H 2.2 – 2.3 mm; TW 1.2 – 1.4 mm. Head with cervical line; pronotum with basal plicae; elytron with short basal plicae. Male genitalia as in Fig. 4 B, median lobe in lateral view thickened before apex, paramere with apical piece rather long and narrow. Dorsal surface of male shiny, female dull due to fine mircoreticulation. Metathoracic wings fully developed. BOLD platform. We provided eight entries in the “ COLLI ” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC4F219FF0CFB061D57FF63.taxon	distribution	Distribution. High Andes in southeastern Peru and Lake Titicaca of Bolivia (Fig. 7), ca. 3,400 – 4,000 m. Habitat. Shallow and exposed puddles, collected with strainer out of mats of vegetation (Figs 5 C; 6 D – F). Other Dytiscidae associated with this species were species of Rhantus Dejean, 1833 and Lancetes Sharp, 1882.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC2F219FF0CFEBE19F8F8D7.taxon	description	(Figs 2 A, B, 4 C, 5 C, E, 7)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC2F219FF0CFEBE19F8F8D7.taxon	materials_examined	Type locality. Peru: Junín, Junín, Lagoon Estancia Azulmito, 4,452 m, - 11.303 ° - 75.989 °. Type material. Holotype: Male (MUSM): “ Peru: Junín, Junín, Lagoon Estancia Azulmito, 4452 m, 14. viii. 2018, - 11.303 ° - 75.989 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 14) ”; “ Holotype Liodessus alpinus sp. nov. Balke, Megna, Hendrich des. 2020 ” [red printed label]. 85 Paratypes (MUSM, ZSM): 5 exs.: same data as holotype; 55 exs.: “ Peru: Junín, Junín, Shalacancha, 4600 m, 11. viii. 2018, - 11.09 ° - 75.879 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 09) ”; 2 exs.: “ Peru: Junín, Junín, Shalacancha, 4320 m, 11. viii. 2018, - 11.075 ° - 75.863 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 10) ”; 15 exs.: “ Peru: Junín, Junín, Junín, 4099 m, 12. viii. 2018, - 11.164 ° - 76.008 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 11) ”; 18 exs.: “ Peru: Junín, Junín, Shoqui, 4324 m, 15. viii. 2018, - 11.046 ° - 75.897 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 18) ”; 7 exs.: “ Peru: Cusco, Canchis, Checacupe, Cayena, 4906 m, 25. viii. 2018, - 14.009 ° - 71.067 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 28) ”. All paratypes are provided with our red printed paratype labels.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC2F219FF0CFEBE19F8F8D7.taxon	description	Description of holotype. Habitus with distinct discontinuity between pronotum and elytra; broadest at about 1 / 3 of elytral length and body therefore appearing slightly pear shaped (as in Figs 2 A, B). TL: 2.0 mm; TL-H: 1.7 mm; TW: 0.9 mm. Colouration. Mostly blackish dorsally and ventrally (as in Figs 2 A, B). Surface sculpture. Head with faint microreticulation at base and along hind angles of eyes, frons and clypeus more or less smooth and with few fine setiferous punctures; pronotum shiny with with sparser and fine setiferous punctation; elytra shiny with dense and coarser setiferous punctation. Structures. Antenna stout. Head without cervical line but few serial punctures instead. Pronotum on each lateral side (or margin) with distinct bead and with distinct and deep basal plicae. Elytron with short and faint basal plicae only visible using diffuse lateral lighting. Metathoracic wings short, about half the length of elytron. Genitalia. Median lobe of aedeagus thick, bulky in lateral view, tip with short nose which is also prominent in ventral view, dorsally on both sides at about mid-length with shallow lateral keels; lateral lobes (parameres) with distal part longish (Fig. 4 C). Variation. TL: 2.0 – 2.2 mm; TL-H: 1.7 – 1.8 mm; TW: 0.9 – 1.0 mm. Few specimens show an extremely faint, interrupted cervical line. We assessed the length of the metathoracic wings in 10 specimens, which all had the same short wing length. While these specimens are certainly not able to fly actively, there might be few fully winged individuals that are capable of flight. Female. Dorsal surface as in male, few specimens with more or less extensive microreticulation on clypeus and along eyes, pronotal disc and laterally as well as on elytron apically only or from about basal 1 / 3 up to apex (dullest form depicted in Fig. 2 B). BOLD platform. We provided 17 entries in the “ COLLI ” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC2F219FF0CFEBE19F8F8D7.taxon	etymology	Etymology. The specific name is the Latin adjective alpinus (of high mountains).	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC2F219FF0CFEBE19F8F8D7.taxon	distribution	Distribution. High Andes of central and southeastern Peru (Fig. 7), ca. 4,300 – 4,900 m. Habitat. Shallow and exposed puddles, collected with strainer out of mats of vegetation (Fig. 5 C, E). Other Dytiscidae associated with this species were species of Rhantus Dejean, 1833 and Lancetes Sharp, 1882, as well as Liodessus acollensis.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F218FF0CFF6318B6FBE6.taxon	description	(Figs 3 C, 7)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F218FF0CFF6318B6FBE6.taxon	materials_examined	Type locality. Peru, Huánuco, Churubamba, Unchog, 3,600 m, - 9.741 ° - 76.169 °. Type material. Holotype: Female (MUSM): “ Peru, Huánuco, Churubamba, Unchog, 3600 m, - 9.740642 ° - 76.169581 °, swampy area ”; “ Holotype Liodessus hauthi sp. nov. Balke, Megna, Hendrich des. 2020 ” [red printed label]. Note on generic affiliation. In the absence of male specimens and DNA sequence data, we can only tentatively assign this species to the genus Liodessus. We opted for this approach as Liodessus is the most widespread Bidessini genus in the high Andes. The morphological diversity of other Andean Liodessus suggests that L. hauthi indeed fits into that genus. We thus chose a conservative approach rather than introducing a new genus.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F218FF0CFF6318B6FBE6.taxon	description	Description of holotype. Large species. Habitus with slight discontinuity between pronotum and elytra (Fig. 3 C). TL: 3.2 mm; TL-H: 2.9 mm; TW: 1.5 mm. Colouration. Orange with darker pattern on pronotal base and on elytra (Fig. 3 C). Surface sculpture. Head dull, with distinct microreticulation and with moderately dense, coarse setiferous punctation; pronotum dull with distinct microreticulation and with sparser and a coarse setiferous punctation; elytra dull with distinct microreticulation and with denser and a coarse setiferous punctation. Structures. Antenna stout. Head with thin cervical occipital line. Pronotum on each lateral side (or margin) with distinct bead; in place of the basal plicae with rough, coarse punctures. Elytron without basal plicae but in place of striae with indistinct imprint. Metathoracic wings not investigated. Notes. This is a highly characteristic species due to its size, the orange coloration as well as being distinctly microreticulate and coarsely punctate on head, pronotum and elytra. BOLD platform. No data available.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F218FF0CFF6318B6FBE6.taxon	etymology	Etymology. Named after our colleague David Hauth, Munich. The specific epithet is a substantive in the genitive case.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F218FF0CFF6318B6FBE6.taxon	distribution	Distribution. Only known from the type locality in the high Andes of central Peru (Fig. 7), 3600 m. Habitat. Collected from a swampy area according to the label, the habitat is most likely like in the other species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F21BFF0CFB3B19F8FDFB.taxon	description	(Figs 2 C, D, 4 D, 5 D, F)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F21BFF0CFB3B19F8FDFB.taxon	materials_examined	Type locality. Peru: Junín, Junín, Lagoon Huacra, 4,461 m, - 11.289 ° - 76.007 °. Type material. Holotype: Male (MUSM): “ Peru: Junín, Junín, Lagoon Huacra, 4461 m, 14. viii. 2018, - 11.289 ° - 76.007 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 13) ”; “ Holotype Liodessus rhigos sp. nov. Balke, Megna, Hendrich des. 2020 ” [red printed label]. 24 Paratypes (MUSM, ZSM): 22 exs.: same data as holotype; 1 exs.: “ Peru: Junín, Junín, Lagoon Yanamyacu, 4327 m, 14. viii. 2018, - 11.314 ° - 75.956 °, Y. S. Megna & N. Zenteno. (PER _ YSM _ 2018 _ 16) ”. All paratypes are provided with our red printed paratype labels.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F21BFF0CFB3B19F8FDFB.taxon	description	Description of holotype. Habitus with indistinct discontinuity between pronotum and elytra; appearing less pear shaped than other species (as in Figs 2 C, D). TL: 2.1 mm; TL-H: 1.8 mm; TW: 0.9 mm. Colouration. Mostly blackish dorsally and ventrally (as in Figs 2 C, D). Surface sculpture. Head with faint microreticulation except on frons and with few fine setiferous punctures; pronotum shiny with with sparser and fine setiferous punctation; elytra shiny with moderately dense and fine setiferous punctation (as in Fig. 2 C). Structures. Antenna stout. Head with faint cervical line. Pronotum on each lateral side (or margin) with distinct bead and with distinct and deep basal plicae. Elytron without basal striae. Metathoracic wings short, slightly longer than half the length of elytron. Genitalia. Median lobe of aedeagus thin and evenly curved in lateral view, apex moderately thin, apex in ventral view more strongly narrowing and pointed; lateral lobes (parameres) with distal part longish (Fig. 4 D). Variation. TL: 2.0 – 2.1 mm; TL-H: 1.6 – 1.8 mm; TW: 0.9 – 1.0 mm. Few specimens with cervical line extremely faint and interrupted. We assessed the length of the metathoracic wings in 10 specimens, which all had the same short wing length. While these specimens are certainly not able to fly actively, there might be few fully winged individuals that are capable of flight. Female. Dorsal surface dull due to presence of distinct microreticulation (Fig. 2 D). BOLD platform. We provided 3 entries in the “ COLLI ” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F21BFF0CFB3B19F8FDFB.taxon	etymology	Etymology. From the Greek rhigos (cold) referring to the chilly collecting experience. The name is a noun standing in apposition.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC3F21BFF0CFB3B19F8FDFB.taxon	distribution	Distribution. Known from the type locality in the high Andes of central Peru (Fig. 7), ca. 4,300 – 4,500 m. Habitat. Shallow and exposed puddles, collected with strainer out of mats of vegetation (Fig. 5 D). Other Dytiscidae associated with this species were species of Rhantus Dejean, 1833 and Lancetes Sharp, 1882, as well as Liodessus acollensis.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC0F216FF0CFD4619F8FEDB.taxon	description	(Figs 3 A, B, 4 E, 6 A, B, 7)	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC0F216FF0CFD4619F8FEDB.taxon	materials_examined	Type locality. Peru, Cusco, Calca, Lares District, Abra Lares, 4,353 m, - 13.190 ° - 71.940 °. Type material. Holotype: Male (MUSM): “ Peru: Cusco, Calca, Lares District, Abra Lares, 4353 m, 2. ix. 2018, - 13.190 ° - 71.940 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 42) ”; “ Holotype Liodessus thespesios sp. nov. Balke, Megna, Hendrich des. 2020 ” [red printed label]. 420 Paratypes (MUSM, ZSM): 50 exs.: same data as holotype; 54 exs.: “ Peru: Cusco, Quispicanchi, Marcapata, Yanatturo, 4433 m, 23. viii. 2018, - 13.633 ° - 71.151 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 23) ”; 35 exs.: “ Peru: Cusco, Quispicanchi, Marcapata, Yanatturo, 4458 m, 23. viii. 2018, - 13.634 ° - 71.143 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 24) ”; 37 exs.: “ Peru: Cusco, Quispicanchi, Pirhuayani, 4662 m, 23. viii. 2018, - 13.641 ° - 71.107 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 25) ”; 19 exs.: “ Peru: Cusco, Quispicanchi, Pirhuayani, 4702 m, 23. viii. 2018, - 13.641 ° - 71.104 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 26) ”; 35 exs.: “ Peru: Cusco, Calca, Calca District, Chaypa, 4129 m, 2. ix. 2018, - 13.202 ° - 71.92 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 40) ”; 171 exs.: “ Peru: Cusco, Calca, Calca District, Pitociray, 4315 m, 2. ix. 2018, - 13.203 ° - 71.94 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 41) ”; 50 exs.: “ Peru: Cusco, Calca, Lares District, Abra Lares, 4353 m, 2. ix. 2018, - 13.19 ° - 71.94 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 42) ”; 19 exs.: “ Peru: Cajamarca, Cajamarca, Lares District, Pampacorral, Mapacocha, 3948 m, 2. ix. 2018, - 13.178 ° - 72.004 °, Y. S. Megna & N. Zenteno (PER _ YSM _ 2018 _ 43) ”. All paratypes are provided with our red printed paratype labels	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC0F216FF0CFD4619F8FEDB.taxon	description	Description of holotype. Habitus with distinct discontinuity between pronotum and elytra (as in Figs 3 A, B). TL: 2.2 mm; TL-H: 2.0 mm; TW: 1.0 mm. Colouration. Mostly dark dorsally and ventrally (as in Figs 3 A, B). Surface sculpture. Head with faint microreticulation at base and along hind angles of eyes, frons and clypeus more or less smooth and with few fine setiferous punctures; pronotum shiny with sparser and fine setiferous punctation; elytra shiny with dense and coarser setiferous punctation. Structures. Antenna stout. Head without cervical line but few serial punctures instead. Pronotum on each lateral side (or margin) with distinct bead; with distinct and deep basal plicae. Elytron without basal plicae. Metathoracic wings short, about half the length of elytron. Genitalia. Median lobe of aedeagus thick in lateral view, tip with short nose; lateral lobes (parameres) with distal part appearing comparably broad (Fig. 4 E). Variation. TL: 2.2 – 2.3 mm; TL-H: 1.8 – 2.0 mm; TW: 1.0 – 1.0 mm. Very few specimens show a short depression at the elytral base in place of the elytral plicae. Additionally, very few specimens show an extremely faint, interrupted occipital line. We assessed the length of the metathoracic wings in 10 specimens, which all had the same short wing length. While these specimens are certainly not able to fly actively, there might be few fully winged individuals that are capable of flight. Female. Dorsal surface dull due to distinct microreticulation (Fig. 3 B). BOLD platform. We provided 30 entries in the “ COLLI ” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
03EDDF2BFFC0F216FF0CFD4619F8FEDB.taxon	etymology	Etymology. From the Greek thespesios (divine, wondrous). The name is a noun standing in apposition. Distribution. High Andes of southeastern Peru (Fig. 7), ca. 3,900 – 4,700 m. Habitat. Shallow and exposed puddles, collected with strainer out of mats of vegetation (Fig. 6 A, B). Other Dytiscidae associated with this species were species of Rhantus Dejean, 1833 and Lancetes Sharp, 1882, as well as Liodessus acollensis.	en	Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis, Hendrich, Lars (2020): New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini). Zootaxa 4852 (2): 151-165, DOI: 10.11646/zootaxa.4852.2.1
