taxonID	type	description	language	source
03EC292BFFAAFFA6FD9C5893DD09FDE7.taxon	description	Figs 1 – 24; Supp. file 1, Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFAAFFA6FD9C5893DD09FDE7.taxon	type_taxon	Type species Neolindus religans (Sharp, 1876), designated by monotypy.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFAAFFA6FD9C5893DD09FDE7.taxon	diagnosis	Diagnosis (modified from Herman 1991) Neolindus is distinguished from all Paederinae genera due to the following combination of characters: head with 1 (Fig. 2 A – B) or 2 pairs of trichobothria (Fig. 2 C – D); maxillary palpomere 4 nipple-shaped (Supp. file 1, Supp. file 2); mesothoracic peritreme small; 1 pair of paratergites on segment III present; paratergites absent on segments IV – VIII; tergite and sternite IV – VI fused into cylinder (Figs 4 – 24 A).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFAAFFA6FD9C5893DD09FDE7.taxon	description	Redescription COLOURATION. Head, pronotum and abdomen from light or dark brown to black, abdominal intersegmental membrane light brown, only in some species darker (e. g., Neolindus maya Guzman, Tokareva & Żyła sp. nov. and N. parahermani Guzman, Tokareva & Żyła sp. nov.); appendages brown, sometimes lighter than body. HEAD. Head capsule in most species wider than long, shape square to trapezoidal; mandibular articulation rounded and prominent; posterior margin of head rounded to straight, slightly emarginate in front of neck; posterior angles of head rounded to straight, often with distinct dorsal ridge with setae; neck about ¼ to ⅓ width of head, dorsal surface with transverse groove, foramen magnum width up to ½ or ⅔ of occiput, with 2 round, anteriorly joined, lateral pits in ventral view. Eye without setae between ommatidia, shorter than or equal to temple length, covering majority of lateral side of head in some species. Epicranium shiny, convex, without microsculpture; setation with 1 pair of SAS and IOS arranged in square, with or without OS and FS, with 1 – 3 PFS arranged in parallel row to PMS in head lateral part (Fig. 1). Gena with lateral margin expanded (Figs 1, 2 A – F), setation with MS, POS, PTOS, and 1 or 2 pairs of trichobothria (OT, or OT and POT); if 2 pairs of trichobothria, then sockets of POT more oval and elongate (Fig. 2 C – D). Ventral side of head with IFOS and POCS; anterior part of gula with 1 or 2 setae in species with small body (up to and including 10 mm) size or 1 row of 6 – 8 irregular setae in species with large body size (more than 10 mm); gular sutures wide apart; postgena with 1 row of 2 – 5 setae in line from eye to gular suture (Supp. file 2). Antenna with antennomere 1 either as long as combined length of antennomeres 2 and 3, or extending as far as combined length of antenomeres 2 – 4. Labrum bilobed or with multiple lobes, with 3 pairs of setae on anterior margin. Mandibula long, slender, sickle-shaped, adentate, prostheca as membranous lobe. Maxilla (Supp. file 1) with row of 4 dark setae on outer margin of stipes; cardo with 1 seta; galea with thick and dark setae on outer margin; lacinia with additional sclerotisation on outer margin. Maxillary palpus with palpomere 1 slightly broadened at tip, glabrous; palpomere 2 elongate, shorter or as long as 3, slightly broadened at tip, with scattered setae; palpomere 3 rounded, expanding near middle, then narrowing toward tip, with denser setation than palpomere 2; palpomere 4 minute, nipple-like and small, glabrous. Ligula bilobed, lobes separate; mentum and submentum each with 1 pair of setae. Labial palpus with palpomere 1 shorter than 2, palpomere 2 longer than 3; palpomere 1 and 2 almost same width; palpomere 2 wider than 3. THORAX. Pronotum nearly quadrate in most species, widest at middle, without microsculpture; anterior margin slightly convex, angles round; lateral margin smooth, without stout setae; posterior margin with rounded angles, slightly emarginate. Prosternum with superior marginal line parallel to anterior margin, not deflexed; pronotosternal suture present, poorly developed near coxal cavity and absent anteriorly (Supp. file 2); basisternum triangular with longitudinal ridge; furcasternum broad, well separated from hypomeron, triangular, with longitudinal ridge in middle. Hypomeron with postcoxal process with additional parallel carina along its margin. Procoxal cavity opened posteriorly. Prepectus longitudinally elongate, with umbilicate punctures on lateral side. Mesosternum, basisternum pentagonal, with acute angle producing between coxa; tuberculate micropunctuation sparse; longitudinal ridge not extended to anterior margin; transversal carina and sternopleural sutures elevated; furcasternum with longitudinal ridge; mesanepisternum smooth, transversely convex. Metasternum with scattered umbilicate micropunctuation; metanepisternum with 2 elevated longitudinal ridges. Mesothoracic peritremes small. Scutellum with 2 emarginate transversal ridges, impunctate. Elytron longer than wide; epipleural and additional ridge present; apical margin emarginate; apical angle rounded; dorsal surface with highly to moderately dense umbilicate micropunctures forming 5 – 8 longitudinal rows. LEGS. Procoxa longer than wide; profemur 3 times as long as wide; protibia with 3 – 5 well-developed combs of setae on ventral side; protarsomeres 1 – 4 wider than long, wider than apex of tibia, with long adhesive setae on ventral side and short dense setae between each other; mesocoxa longer than wide; mesofemur 3 times as long as wide; mesotibia with apical ctenidium on both sides in most species, and metatibia with apical ctenidium on both sides in all species; metacoxa longer than wide; metafemur 3 times as long as wide; mesotarsomeres and metatarsomeres 1, 2, and 5 longer than 3 and 4; mesotrasomeres and metatarsomeres 3 and 4 subequal. ABDOMEN. Surface without microsculpture, micropunctures varying in density from high to low, arranged randomly. Sternite II narrow and fused to sternite III; posterior margin of sternite II with row of setae. Sternite III with central longitudinal keel. Segment III with 1 pair of paratergites. Segments III – VI with straight posterior margin. Segments IV – VI with fused tergite and sternite forming cylinder. Abdominal segments III – IX sometimes with internal canals at base of sternite and tergite. Tergite IX with straight apices, fused or divided medially. No glandular openings in intersegmental membranes. Males: segment VII and VIII with tergite and sternite separate, posterior margin of various form, varies between species; aedeagus, median lobe oval to triangular, basal foramen acentric, near to base of median lobe, with asymmetrical appendages; some species with complex of movable sclerites at apex around opening of median foramen. Females: usually lack of emargination on abdominal segments VII and VIII, sometimes present, but shallow, always shallower than in respective male.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFAAFFA6FD9C5893DD09FDE7.taxon	distribution	Distribution and habitat Neolindus can be found in various localities across South and Central America. Central America ranges from south of Mexico in Chiapas, to Panama, including records from Honduras, Costa Rica, and Nicaragua. In South America, Neolindus is known from Venezuela, Colombia, Ecuador, Peru, Bolivia, Brazil, Suriname, Guyana, and French Guiana, with the Bolivian department Pando being the southernmost occurrence (Fig. 3). Species of Neolindus are typically found in forest leaf litter at altitudes ranging from 2 (N. utriensis Guzman, Tokareva & Żyła sp. nov.) to 2900 m a. s. l. (N. campbelli Herman, 1991).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	description	urn: lsid: zoobank. org: act: A 3 AE 98 B 8 - 192 C- 498 F- 82 DA-F 147 AF 44 B 503 Fig. 4; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	diagnosis	Diagnosis This species is similar to N. apiculus and N. campbelli in having a broad emargination of sternite VIII (Herman 1991: figs 168, 173; Fig. 4 E). However, the structure of the aedeagus in N. bicornis Guzman, Tokareva & Żyła sp. nov. is different (Herman 1991: figs 170, 174; Fig. 4 F – I). It resembles the aedeagus of N. triangularis Guzman, Tokareva & Żyła sp. nov., but it differs by having a rounded base of the median lobe and a narrower pLS in their middle part (the base of the median lobe acute in N. triangularis) (Figs 4 F – I, 20 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	etymology	Etymology The name is a combination of the Latin prefix ‘ bi ’ meaning ‘ two’ and the Latin word ‘ cornum ’, which means ‘ a horn’. It describes the characteristic form of apical aedeagal sclerites. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus bicornis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Roura, 18.4 Km SSE, 240 m [m a. s. l.]; 4 ° 36 ” 38 ” N, 52 ° 13 ’ 25 ’’ W; 29 May – 10 Jun 1997; J. Ashe, R Brooks Leg.; FG 1 AB 97 180; ex: flight intercept trap \\ SM 0100688 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes FRENCH GUIANA • 2 ♀♀, 2 ♂♂; “ Neolindus bicornis Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Cayenne, Roura, 27.4 Km SSE; 280 m [m a. s. l.]; 4 ° 44 ’ 20 ” N, 52 ° 13 ’ 25 ’’ W; 10 Jun 1997; J. Ashe, R Brooks, FG 1 AB 97 177 ex: flight intercept trap \\ [barcode] SM 0133909 KUNHM-ENT ”; KUNHM-ENT • 1 ♂; same data as for preceding; “ SM 0133998 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ 18.4 Km SSE; 240 m; 4 ° 36 ” 38 ” N, 52 ° 13 ’ 25 ’’ W; 25 – 29 May 1997; FG 1 AB 97 081; ex: flight intercept trap \\ SM 0098969 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	description	Description MEASUREMENTS. BL (9), H (0.49, 1.04), A (1.71), a 1 (0.34, 0.11), a 2 (0.14, 0.07), a 3 (0.26, 0.08), a 4 (0.12, 0.09), a 5 (0.13, 0.09), a 6 (0.12, 0.08), a 7 (0.12, 0.09), a 8 (0.11, 0.09), a 9 (0.11, 0.08), a 10 (0.11, 0.09), a 11 (0.14, 0.1), NKW (0.49), GL (0.21), P (1.09, 1.29), E (1.59, 1.39), PC (0.5, 0.29), PF (0.86, 0.39), PT (0.65, 0.15), MSC (0.4, 0.46), MSF (1.13, 0.29), MST (0.82, 0.11), MTC (0.46, 0.25), MTF (1.15, 0.28), MTT (0.9, 0.13). COLOURATION. Head dark brown; pronotum brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin straight with emargination in front of neck; posterior angles straight; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 1 pair of PFS, without OS. Gena with depression from mandibular base to middle of eye, setation with 1 OT, MS, POS and PTOS; sockets of MS and POS umbilicate. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 3, shorter than head and pronotum combined; antennomeres longer than wide, tomentose pubescence starting from antennomere 3; antennomere 1 shorter than antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, emargination deep, U-shaped, with 2 dark, thick seta in median line of labrum lobes (Fig. 4 A, Supp. file 2). THORAX. Pronotum slightly wider than long, with randomly distributed umbilicate micropunctuatoin and slightly convex anterior margin. Basisternum of prosternum with tuberculate micropunctuation. Metasternal intercoxal process with 1 pair of acute processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 8 – 10 longitudinal rows (Fig. 4 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae, apical comb longitudinal, composed of widely separated, thick setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as mesotarsomere 2, mesotarsomere 3 shorter than mesotarsomere 1, but as long as mesotarsomere 4, mesotarsomere 5 as long as mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomere 2 longer than metatarsomere 3, metatarsomere 4 as long as metatarsomere 3, metatarsomere 5 shorter than 1 but longer than 2. ABDOMEN. Male: posterior margin of sternite VII sinuate, with midlongitudinal emargination and rounded lateral angles (Fig. 4 C). Posterior margin of tergite VIII with elongate rounded apex (Fig. 4 D). Posterior margin of sternite VIII with median V-shaped emargination and rounded lateral angles (Fig. 4 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length, wider at margin and narrow towards apex; aedeagus, median lobe rounded at base, widening apically and well-sclerotised; pLS fused into flattened arcuate sclerite of almost even width with 2 acute processes on each side protruding distad from median foramen and ending bent to antiparameral side. APS flattened, arcuate, smaller than pLS, with 2 acute processes on each side protruding distad from median foramen and ending bent to parameral side, shorter than processes of pLS. pPMS not visible (Fig. 4 F – I). Female: lack of emargination on abdominal segments.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFA1FFB8FE0C5E53DCFBF9EE.taxon	distribution	Distribution This species is known from the type locality in French Guyana (Cayenne, Roura) and the surrounding area. It was collected by a flight interception trap at an altitude of 240 – 280 m a. s. l.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	description	urn: lsid: zoobank. org: act: F 14030 CB- 549 D- 48 BA-B 857 - E 961 C 078 B 08 D Fig. 5; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	diagnosis	Diagnosis This species is similar to N. milleri in shape of sternite and tergite VIII. However, in N. elegans Guzman, Tokareva & Żyła sp. nov., the posterior margin of sternite VII has a wide middle emargination with a straight middle part, delimited on sides by 2 long acute processes (Fig. 5 C). In contrast, N. milleri has a shallower emargination lacking acute processes on the sides (Herman 1991). Furthermore, the aedeagus of N. elegans displays a complex composition of apical sclerites surrounding the median foramen of the median lobe (Fig. 5 F – I), while the aedeagus of N. milleri has a simpler arrangement of sclerites (Herman 1991: fig. 189).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	etymology	Etymology The name is derived from the Latin word ‘ elegans ’, which means ‘ elegant’ or ‘ tasteful’. It refers to the symmetry of the aedeagus in parameral view, particularly the straight angle between the median lobe apex and some apical sclerites. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus elegans Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Saül, 7 km N, 3 km NW, Les Eaux Claires, Mt. La Fumée trail, 230 - 400 m [m a. s. l.]; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 1 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 103; ex: fungusy log \\ SM 0094312 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	description	Description MEASUREMENTS. BL (5.2), H (0.42, 0.68), A (1.06), a 1 (0.24, 0.08), a 2 (0.06, 0.06), a 3 (0.09, 0.06), a 4 (0.07, 0.06), a 5 (0.09, 0.6), a 6 (0.08, 0.08), a 7 (0.09, 0.08), a 8 (0.08, 0.07), a 9 (0.08, 0.08), a 10 (0.08, 0.07), a 11 (0.1, 0.08), NKW (0.31), GL (0.2), P (0.7, 0.8), E (0.92, 0.9), PC (0.3, 0.2), PF (0.5, 0.2), PT (0.2, 0.1), MSC (0.26, 0.2), MSF (0.7, 0.12), MST (0.54, 0.07), MTC (0.2, 0.25), MTF (0.6, 0.16), MTT (0.42, 0.08). COLOURATION. Head and pronotum dark brown; legs brown; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded; posterior angles rounded; setation with 1 pair of PCS, 1 row of PMS, and 2 long setae at neck junction. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 1 pair of PFS, without OS. Gena with depression around trichobothria, setation with 1 OT, MS, POS and PTOS; sockets of POS concave. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae close to gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 3, shorter than head and pronotum combined; antennomeres longer than wide; antennomeres 4 – 11 with tomentose pubescence; antennomere 1 as long as antenomeres 2 and 3 combined; antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination, with 2 dark thick seta in median line of labrum lobes (Fig. 5 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Metasternum with umbilicate micropunctuation in longitudinal rows, metasternal intercoxal process with pair of barely visible rounded processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 10 – 12 longitudinal rows (Fig. 5 A; Supp. file 2). LEGS. Protibia with 3 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than 2, mesotarsomeres 3 and 4 as long as mesotarsomere 2, mesotarsomere 5 longer than 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 as long as 2, metatarsomere 3 shorter than 1 and as long as metatarsomere 4, metatarsomere 5 longer than metatarsomere 1. ABDOMEN. Male: tergites with low-density random micropunctuation. Posterior margin of tergite VII slightly convex, almost straight (Fig. 5 B). Posterior margin of sternite VII with wide midpoint emargination with straight angles, delimited by 2 long teeth-like processes with acute ends (Fig. 5 C). Posterior margin of tergite VIII with elongate rounded apex (Fig. 5 D). Posterior margin of sternite VIII with midlongitudinal V-shaped emargination, lateral angles rounded (Fig. 5 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, median lobe vase-shaped, rounded at base, broadening towards apex; apex notched on parameral side with elongate median hole or cavity not reaching basal foramen elevation, apical part of median lobe with 2 pairs of processes near middle, as part of joint with movable pPMS. pPMS robust, of complex form, protruding upwards, shorter than APS, upper part curved on parameral side, with multiple small denticles on curved top. pLS small, oval, obscure, shorter than other apical sclerites. APS spoon-shaped, longer than pLS and pPMS, with central emargination having acute tip and 2 acute narrow short processes on sides (Fig. 5 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBFFFBBFE035A55DC31FD2C.taxon	distribution	Distribution The species is known only from the type locality in French Guyana (Saint-Laurent-du-Maroni, Saül). It was collected on Monts La Fumée between 230 – 400 m a. s. l. from fungus log.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	description	urn: lsid: zoobank. org: act: 39 F 8 C 257 - B 035 - 4 DC 9 - 8952 - 55 AE 96 FC 0 E 1 D Fig. 6; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	diagnosis	Diagnosis Neolindus longithorax Guzman, Tokareva & Żyła sp. nov. resembles N. bidens in the structure of tergite VIII as well as the broader median lobe of the aedeagus. However, the posterior margin of tergite VIII in N. longithorax differs from that of N. bidens with a more prominent central lobe and less developed lateral lobes (Herman 1991: fig. 221; Fig. 6 D). Additionally, the aedeagus of N. longithorax has narrower and more elongate median lobe without median hole, contrasting with robust and medially widened median lobe with distinct median hole on parameral side of aedeagus in N. bidens (Herman 1991: fig. 222; Fig. 6 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	etymology	Etymology The specific epithet is a combination of Latin words ‘ longus ’ meaning ‘ long’ and ‘ thorax ’ as an anatomical part of an insect and refers to relatively long thorax of this species. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus longithorax Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Saül, Galbao summit, 740 m [m a. s. l.]; 3 ° 37 ’ 18 ’’ N, 53 ° 16 ’ 42 ’’ W, 5 – 7 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 154; ex: flight intercept trap \\ SM 0099826 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	description	Description MEASUREMENTS. BL (9.4), H (0.57, 0.9), A (1.95), a 1 (0.42, 0.1), a 2 (0.15, 0.07), a 3 (0.23, 0.07), a 4 (0.15, 0.09), a 5 (0.14, 0.09), a 6 (0.15, 0.08), a 7 (0.15, 0.07), a 8 (0.14, 0.09), a 9 (0.14, 0.09), a 10 (0.13, 0.09), a 11 (0.15, 0.09), NKW (0.43), GL (0.24), P (1.01, 1.02), E (1.48, 1.24), PC (0.38, 0.37), PF (0.8, 0.5), PT (0.63, 0.2), MSC (0.54, 0.32), MSF (0.9, 0.23), MST (0.66, 0.1), MTC (0.43, 0.37), MTF (1.04, 0.26), MTT (0.1, 1.08). COLOURATION. Head brown; pronotum light brown; legs brown; elytra dark brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded, with emargination in front of neck; posterior angles rounded; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 1 pair of PFS, without OS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, MS, POS, and PTOS; sockets of all setae umbilicate. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae near gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 4, longer than head and pronotum combined (Fig. 6 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with V-shaped emargination (Fig. 6 A; Supp. file 2). THORAX. Pronotum quadrate, with umbilicate randomly distributed micropunctures absent in medial longitudinal line. Metasternal intercoxal process with 1 pair of minute rounded processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 8 – 10 longitudinal rows (Fig. 6 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thick setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than 2, mesotarsomeres 3 and 4 as long as 2, mesotarsomere 5 longer than 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomeres 3 and 4 as long as metatarsomere 2, metatarsomere 5 longer than metatarsomere 1. ABDOMEN. Male: tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite and sternite VII straight (Fig. 6 B – C). Posterior margin of tergite VIII trilobed, with slightly triangular central lobe, lateral lobes less developed (Fig. 6 D). Posterior margin of sternite VIII with midlongitudinal U-shape emargination; 2 lobes delimiting emargination with row of 5 dark setae; lateral angles slightly elevated (Fig. 6 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ½ of segment length; aedeagus, in parameral view, median lobe of uniform width along sides, slightly narrowing at base; apex with median emargination with elevated thinner disrupted edges laterally. pPMS of complex form, quadrate, flattened, shorter than APS. pLS not visible. APS spoon-like, flat, protruding upwards, with apex divided into 2 oval lobes (Fig. 6 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBCFFBDFE1A5F17DC86F9D1.taxon	distribution	Distribution This species is known only from the type locality in French Guiana (Saint-Laurent-du-Maroni, Saül, Mont Galbao). It was collected at a higher altitude, 740 m a. s. l., by a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	description	urn: lsid: zoobank. org: act: B 0810 A 98 - 899 B- 46 D 2 - 8654 - 16 AFB 54371 E 1 Fig. 7; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	diagnosis	Diagnosis Neolindus luxipenis Guzman, Tokareva & Żyła sp. nov. is most similar to N. lirellus and N. brachiatus. However, N. luxipenis differs from N. lirellus in sternite VII by the more pronounced central process of posterior margin, and tergite VIII, where the posterior margin is rounded (Fig. 7 C – D; Herman 1991: figs 108 – 109, 111). In comparison to N. brachiatus, sternite VII of N. luxipenis has a central narrower emargination, with 2 protruded processes on each side and invaginated surface around emargination; in N. brachiatus central emargination is wider and the processes are less protruded (Fig. 7 C; Herman 1991: fig. 101).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	etymology	Etymology The name combines the Latin word ‘ luxus ’ for ‘ luxury’ or ‘ extravagant’ and ‘ penis ’ which addresses the aedeagus here. It refers to an elaborate structure of aedeagus in the species. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	materials_examined	Type material Holotype BOLIVIA • ♂; “ Neolindus luxipenis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Cochabamba, Cochabamba, 67.5 km NE, Est. Biol. Valle de Sajita, Univ. de San Simon; 300 m [m a. s. l.]; 17 ° 6 ’ 33 ’’ S, 64 ° 47 ’ 52 ’’ W; 9 – 13 Feb 1999; F. Genier Leg.; BOL 1 G 99 069; ex: light intercept trap \\ SM 0173627 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratype BOLIVIA • 1 ♂; “ Neolindus luxipenis Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ same data as for holotype \\ 7 Feb 1999; R. Anderson Leg.; BOL 1 A 99 027; ex: rain forest litter \\ SM 0169264 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	description	Description MEASUREMENTS. BL (8.08), H (0.46, 0.7), A (1.08), a 1 (0.25, 0.09), a 2 (0.09, 0.06), a 3 (0.14, 0.07), a 4 (0.07, 0.07), a 5 (0.01, 0.08), a 6 (0.08, 0.09), a 7 (0.09, 0.08), a 8 (0.09, 0.08), a 9 (0.08, 0.08), a 10 (0.06, 0.09), a 11 (0.12, 0.09), NKW (0.33), GL (0.21), P (0.78, 0.83), E (0.94, 0.91), PC (0.52, 0.27), PF (0.7, 0.34), PT (0.44, 0.12), MSC (0.34, 0.22), MSF (0.75, 0.17), MST (0.48, 0.08), MTC (0.36, 0.26), MTF (0.8, 0.22), MTT (0.69, 0.1). COLOURATION. Head and pronotum brown; coxa, femur and tibia brown, tarsomeres light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded; setation with 1 pair of PCS, 1 row of PMS, and 2 long setae at neck articulation. Epicranial setation with 1 pair of FS and 2 pairs of PFS, without OS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, 1 pair of MS, POS and PTOS; sockets of MS umbilicate; sockets of POS concave. Postgena with row of 3 setae in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 1 seta on each side of gular sutures. Neck with foramen magnum occupying ⅔ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 7 A), antennomeres 1 – 4, 6 – 8, and 11 longer than wide, antennomeres 5, 9, 10 wider than long; tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, emargination U-shaped, with 2 dark thick seta in median line of labrum lobes (Fig. 7 A; Supp. file 2). THORAX. Pronotum nearly quadrate, with umbilicate micropunctures and 2 distinct paramedial rows of setae; anterior margin slightly convex. Metasternal intercoxal process with 1 pair of obtuse processes. Elytron longer than wide, surface of elytra with umbilicate micropunctures in 5 – 7 moderately dense longitudinal rows (Fig. 7 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb composed of widely separated, thick setae; mesotibial apical ctenidium only on outer side; mesotarsomere 1 longer than 2, mesotarsomere 3 as long as 2, mesotarsomere 4 as long as 2, mesotarsomere 5 longer than 1; metatarsomere 1 longer than 2, metatarsomere 2 longer than 3, metatarsomere 3 as long as 4, metatarsomere 5 equal to metatarsomere 1. ABDOMEN. Male: segments V – VII wider than III, IV, and VIII. Tergites III – VI with high density of micropunctuation; tergites VII and VIII with low density of micropunctuation, punctures randomly distributed. Posterior margin of tergite VII straight (Fig. 7 B). Posterior margin of sternite VII with median tooth-like processes (Fig. 7 C). Tergite VIII with 4 internal canals with carinated margins; 2 median canals forming quadrate area. Posterior margin of tergite VIII apically extended and rounded (Fig. 7 D). Sternite VIII with broad midlongitudinal emargination covering ⅓ of segment length, starting as narrow and widening towards rounded lateral edges (Fig. 7 E). Posterior margin of tergite IX with midlongitudinal deep emargination ½ of segment length, fused; aedeagus, median lobe with uniform width from base to apex, apex emarginate; pLS flat with rounded edges, expanded laterally and folded inward; APS complex with 2 processes, each process with 2 arms, apical arm acute, frontal arms bend downward and subdivided to long tooth and group of several small teeth at top of acute process; pPMS absent (Fig. 7 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFBAFFB0FE0A5A38DD2EFCE5.taxon	distribution	Distribution This species is known only from the type locality in Bolivia (Cochabamba, Cochabamba, Valle de Sajta Biological Station). It was collected at a lower altitude (300 m a. s. l.) using a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	description	urn: lsid: zoobank. org: act: C 0 B 837 AA- 1976 - 4 F 78 - 80 D 3 - C 0 DD 7 AC 72 E 69 Fig. 8; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	diagnosis	Diagnosis Habitus of N. maya sp. nov. resembles that of N. punctogularis, but these species are separated by characters of abdominal sclerites and the aedeagus. N. maya differs from N. punctogularis in the shape of the posterior margin of sternite VII, which narrows towards the apex with a moderately deep concave middle emargination in N. maya and oblique rest of the posterior margin on the sides of the emargination (Fig. 8 C). In contrast, in N. punctogularis, the margin is straight and the emargination is bordered by wide rounded processes (Herman 1991: fig. 207). Moreover, N. maya differs in the shape of the posterior margin of sternite VIII with V-shaped emargination and a narrow elevated midlongitudinal invagination on the sternite (Fig. 8 E), while N. punctogularis lacks the invagination (Herman 1991: fig. 205); in the shape of the posterior margin of tergite VIII, with the middle acute lobe longer than the 2 lateral acute lobes (Fig. 8 D), in contrast to all lobes almost equal in length in N. punctogularis (Herman 1991: fig. 206); and in a different shape of the median lobe of the aedeagus, as well as apical arms of pPMS (Herman 1991: fig. 204; Fig. 8 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	etymology	Etymology The name honours the Maya communities that inhabit the type locality in Mexico. A noun in apposition.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	materials_examined	Type material Holotype MEXICO • ♂; “ Neolindus maya Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Chiapas, Najá; 16 ° 58.450 ’ N, 91 ° 35.155 ’ W; 950 m [m a. s. l.]; 14 Jul 2007; J. Longino Leg.; ex. winkler, montane wet forest leaf litter; LLAMA 07 JTL 6047 - s \\ SM 0787486 KUNHM-ENT [barcode] \\ Oedichirus M. K. Thayer det. 2017 ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	description	Description MEASUREMENTS. BL (10.07), H (0.75, 1.38), A (2.62), A 1 (0.53, 0.14), A 2 (0.16, 0.09), A 3 (0.27, 0.97), A 4 (0.24, 0.09), A 5 (0.23, 0.09), A 6 (0.22, 0.09), A 7 (0.21, 0.12), A 8 (0.22, 0.08), A 9 (0.2, 0.1), A 10 (0.18, 0.11), A 11 (0.18, 0.1), NKW (0.62), GL (0.4), P (1.3, 1.67), E (2.02, 2.18), PC (0.69, 0.36), PF (1.1, 0.58), PT (0.71, 0.2), MSC (0.56, 0.37), MSF (1.5, 0.34), MST (0.18, 0.12), MTC (0.62, 0.77), MTF (1.61, 0.35), MTT (1.61, 0.15). COLOURATION. Shiny black body and appendages; abdominal intersegmental membrane between segments III and IV dark brown, rest black. HEAD. Head capsule wider than long; anterior margin straight; posterior margin rounded, with emargination in front of neck; posterior angles straight; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 2 – 3 pairs of PFS in rows parallel to FS, without OS. Gena with irregular surface, with depression from mandibular base to posterior margin of head; setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 3 setae in line from eye to gular suture; gular sutures not reaching posterior margin of head; anterior part of gula with irregular row of 6 – 8 setae. Neck with foramen magnum occupying ¾ of occiput. Antenna thin, longer than head and pronotum combined (Fig. 8 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with V-shaped emargination (Fig. 8 A; Supp. file 2). THORAX. Pronotum wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Prosternum with superior marginal line parallel to anterior margin. Metasternal intercoxal process with 1 pair of rounded processes. Elytron wider than long, surface of elytra with moderately dense umbilicate punctures in 10 – 11 longitudinal rows (Fig. 8 A; Supp. file 2). LEGS. Protibia with 5 – 6 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; metatibial apical ctenidium on both sides, inner longer than outer; metatarsomere 1 longer than 2, metatarsomere 2 longer than 3, metatarsomere 3 longer than 4, metatarsomere 5 as long as 2. ABDOMEN. Male: tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite VII straight (Fig. 8 B). Posterior margin of sternite VII with moderate midline emargination bordered by rounded processes (Fig. 8 C). Posterior margin of tergite VIII trilobed, median lobe wide with triangle tip, lateral lobes narrow (Fig. 8 D). Posterior margin of sternite VIII with midlongitudinal V-shaped emargination ¼ of segment length, lateral angles rounded (Fig. 8 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, in parameral view, median lobe of even width, apex with moderately deep emargination. pPMS elongate, robust, with 2 acute apical arms bent inwards; arm closer to median foramen longer than arm closer to parameral side. pLS thin, with net-like microsculpture, forming elongate flat processes equal in length to pPMS and slightly longer than APS. APS spoon-shaped, with blunt top, wider than pLS (Fig. 8 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB7FFB2FE3C5F6DDC62F992.taxon	distribution	Distribution The species is known only from the type locality in Mexico (Chiapas, Laguna Naja). It sets the northernmost border of the distribution for the genus. It was collected at the altitude of 950 m a. s. l. in the montane wet forest by sifting leaf litter and using a Winkler extractor.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	description	urn: lsid: zoobank. org: act: D 5782 A 9 D- 41 EE- 4 C 22 - 93 EB- 31655 E 31 E 39 B Fig. 9; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	diagnosis	Diagnosis Neolindus minutus Guzman, Tokareva & Żyła sp. nov. and N. bullus are the smallest species in the genus with body length no longer than 4 mm. Both species have the same shape of the posterior margin of sternite VIII with U-shaped median emargination, but it is deeper in N. minutus (Herman 1991: fig. 154; Fig. 9 E). Other differences are in the shape of the posterior margin of tergite VIII, which is rounded in N. minutus (Fig. 9 D), but emarginated in N. bullus (Herman 1991: fig. 157). Furthermore, aedeagus in N. minutus has a different form of median lobe as well as structure and composition of apical sclerites (Herman 1991: fig. 156; Fig. 9 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	etymology	Etymology The Latin epithet ‘ minutus ’ means ‘ very small, little, minute’ and describes the noticeably smaller size of this species of Neolindus compared with others. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	materials_examined	Type material Holotype GUYANA • ♂; “ Neolindus minutus Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Region 8, Iwokrama Forest, 1 km W Kurupukari, Iwokrama Field Stn [Station]; 60 m [m a. s. l.]; 4 ° 40 ’ 19 ’’ N, 58 ° 41 ’ 4 ’’ W; 21 MAY 2001, R. Brooks, Z. Falin, GUY 1 BF 01 005 ex: Acromyrmex hystrix refuse pile \\ SMO 568781 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes ECUADOR • 1 ♀; “ Neolindus minutus Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Napo, Yuturi Longe, Rio Napo; 270 m; 0 ° 32 ’ 54 ’’ S, 76 ° 02 ’ 18 ’’ W; 20 – 21 Mar 1999; R. Brooks, D. Brzoska Leg.; ECU 1 B 99 010; ex: flight interception trap \\ SM 0153469 KUNHM-ENT [barcode] ”; KUNHM-ENT. FRENCH GUIANA • 1 ♀; “ Neolindus minutus Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Saül, 7 km N, 0.5 km ESE, Les Eaux Claires, Mt. La Fumée; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 300 m; 4 – 8 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 164; ex: flight interception trap \\ SM 0134044 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; “ Roura, 39.4 km SSE; 270 m; 4 ° 32 ’ 43 ’’ N, 52 ° 8 ’ 26 ’’ W; 25 – 29 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 076; ex: flight interception trap \\ SM 0131319 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; “ 27.4 km SSE; 280 m; 4 ° 44 ’ 20 ’’ N, 52 ° 13 ’ 25 ’’ W; 25 – 29 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 079; ex: flight interception trap \\ SM 0101388 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ FG 1 AB 97 177; ex: \\ SM 0133995 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Cayenne, 33.5 km S and 8.4 km NW of HWY N 2 on HWY D 5; 30 m; 4 ° 48 ’ 18 ’’ N, 52 ° 28 ’ 41 ’’ W; 25 – 29 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 171; ex: flight interception trap \\ SM 0131037 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Saül, 7 km N, Les Eaux Claires; 220 m; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 30 May – 4 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 143 B; ex: flight interception trap \\ SM 0099466 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Roura, 39.4 km SSE; 270 m; 4 ° 32 ’ 43 ’’ N, 52 ° 8 ’ 26 ’’ W; 29. May – 10 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 172; ex: flight interception trap \\ SM 0099394 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; same data as for preceding; “ FG 1 AB 97 076 \\ SM 0131322 KUNHM-ENT [barcode] ”; KUNHM-ENT. GUYANA • 1 ♀; same data as for holotype; “ PARATYPE [yellow label] \\ GUY 1 BF 01 034 \\ SM 0566025 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 0567438 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 05661128 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 0566308 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 0565913 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 0567446 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 146 \\ SM 0567615 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for holotype; “ GUY 1 BF 01 097 \\ SM 0570780 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 0566193 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; same data as for holotype; “ GUY 1 BF 01 034 \\ SM 05661153 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; same data as for holotype; GUY 1 BF 01 034 \\ SM 056663 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Kabocalli Field Stn [Station]; 3 – 5 Jun 2001; GUY 1 BF 01 146 \\ SM 0567596 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Turtle Mt. [Mountains] Summit; 290 m; 4 ° 43 ’ 57 ’’ N, 58 ° 44 ’ 1 ’’ W; 30 May – 1 Jun 2001; GUY 1 BF 01 09 \\ SM 0570865 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♂; “ Kabocalli Field Stn; 60 m; 21 May 2001; GUY 1 BF 01 005; ex: Acromyrmex sp. \\ SM 0568401 KUNHM-ENT [barcode] ”; KUNHM-ENT. NICARAGUA • 1 ♀; “ Rio San Juan Dept. [Departamento], 60 km SE San Carlos, Refugio Bartola; 100 m; 10 ° 58 ’ 52.5 ” N, 84 ° 20 ’ 08.9 ” W; 25 – 28 May 2002; R. Brooks, Z. Falin, S. Chatzimanolis Leg.; ex: flight interception trap; NIC 1 BFC 02 112 \\ SM 0555340 KUNHM-ENT [barcode] ” • 1 ♀; same data as for preceding “ NIC 1 BFC 02 112 \\ SM 0540900 KUNHM-ENT [barcode] ” • 1 ♀; same data as for preceding; “ NIC 1 BFC 02112 \\ SM 0540901 KUNHM-ENT [barcode] ”; KUNHM-ENT. PERU • 1 ♀; “ Madre de Dios, Concha Cashu Bio. Stn [Biological Station], Manu National Park; 350 m; 11 ° 53 ’ 45 ’’ S, 71 ° 24 ’ 24 ’’ W; 17 – 19 Out 2000; R. Brooks Leg.; PERU 1 B 00 042; ex: flight interception trap \\ SM 0210599 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; “ Concha Salvador, Reserved Zone Manu National Park; 310 m; 12 ° 0 ’ 13 ’’ S, 71 ° 31 ’ 36 ’’ W; 20 – 21 Out 2000; R. Brooks Leg.; PERU 1 B 00 070; ex: flight interception trap \\ SM 0210838 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; “ Pantiacolla Lodge, Alto Madre de Dios River; 400 m; 12 ° 39 ’ 22 ’’ S, 71 ° 13 ’ 55 ’’ W; 23 – 26 Out 2000; R. Brooks Leg.; PERU 1 B 00 099; ex: flight interception trap \\ SM 0210824 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	description	Description MEASUREMENTS. BL (2.65), H (0.3, 0.3), A (0.65), A 1 (0.09, 0.03), A 2 (0.31, 0.02), A 3 (0.03, 0.03), A 4 (0.03, 0.04), A 5 (0.04, 0.05), A 6 (0.04, 0.05), A 7 (0.04, 0.06), A 8 (0.04, 0.06), A 9 (0.04, 0.06), NKW (0.20), GL (0.1), P (0.43, 0.44), E (0.54, 0.46), PC (0.25, 0.13), PF (0.29, 0.1), PT (0.13, 0.08), MSC (0.17, 0.09), MSF (0.26, 0.07), MST (0.21, 0.06), MTC (0.13, 0.1), MTF (0.27, 0.09), MTT (0.21, 0.06). COLOURATION. Head and pronotum brown, legs light brown, abdomen brown. HEAD. Head capsule nearly as wide as long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded, with emargination in front of neck; posterior angles rounded; setation with 1 pair of PCS and 2 pairs of PMS, central and distal. Epicranium shiny, without micropunctuation, setation with 1 pair of FS, without OS and PFS. Gena with irregular surface, with depression from mandibular base to posterior margin of head, setation with 1 OT, MS, and POS. Sockets of MS and POS umbilicate, socket of OT oval, narrowed anteriorly. Postgena with 1 seta in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae near gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 9 A); antennomeres 1 and 2 longer than wide; antennomeres 3 – 9 wider than long; antennomeres 10 – 11 longer than wide, tomentose pubescence starting from antennomere 5; antennomere 1 shorter than antennomeres 2 and 3 combined. Labrum slightly bilobed (Fig. 9 A, Supp. file 2). THORAX. Pronotum quadrate, with umbilicate micropunctures, setation with 1 pair of setae in middle of dorsal side and 1 pair of setae in posterior third part of dorsal side; anterior margin slightly convex with anterior angles rounded; posterior margin rounded and slightly emarginate, posterior angles rounded. Metasternal intercoxal process with 1 pair of acute processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 4 – 5 longitudinal rows (Fig. 9 A; Supp. file 2). LEGS. Protibia with 2 well-developed combs of setae; mesotibial apical ctenidium with spines; mesotarsomere 1 longer than 2, mesotarsomeres 3 and 4 as long as 2, mesotarsomere 5 longer than 1; metatarsomere 1 as long as 2, metatarsomeres 3 and 4 as long as metatarsomere 2, metatarsomere 5 longer than 1. ABDOMEN. Tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite and sternite VII straight (Fig. 9 B – C). Posterior margin of tergite VIII apically rounded (Fig. 9 D). Male: posterior margin of sternite VIII with midlongitudinal U-shaped emargination, ⅓ of segment length, lateral angles rounded (Fig. 9 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, median lobe widened in first half, narrowed to apex; with wide oval median invagination above basal foramen; apex of lobe with 2 spoon-shaped flattened lobes with acute tips and rounded lateral edges, joined with U-shape emargination. pPMS fused in conical structure folding inside median foramen. pLS long, narrow, acute, curved. APS absent, antiparameral side membranous, with bilobed cover folding pLS (Fig. 9 F – I). Female: lack of emargination on abdominal segments VIII and IX.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB5FFB6FE0F5A7CDD3FFC3A.taxon	distribution	Distribution The species is known from the type locality in Guyana (Region 8, Potaro-Siparuni, Iwokrama Field Station), and several other countries, i. e., Ecuador (Napo, El Eden), French Guiana (Cayenne, MontsineryTonnegrande; Regina-Kaw; Roura, Roura; Saül, Cambrouze), Nicaragua (Rio San Juan, El Bambu), Peru (Madre de Dios, Fitzcarrald District and Itahuania). The holotype has a note that it was collected from the Acromyrmex hystrix (Latreille, 1802) refuse pile. One paratype specimen was also collected near ants from the same genus (Acromyrmex Mayr, 1865), while the others by using a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	description	urn: lsid: zoobank. org: act: 43 C 4 CAA 7 - 3 DC 0 - 4835 - 92 DA- 252113 D 9 E 6 AF Fig. 10; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	diagnosis	Diagnosis Similar to N. apiculus and N. campbelli in the shape of the posterior margin of sternite VIII and tergite VIII. However, N. napo Guzman, Tokareva & Żyła sp. nov. differs from these species by the different structure of apical sclerites of aedeagus, especially, by much larger bifid protruding APS (Herman 1991: figs 170, 174; Fig. 10 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	etymology	Etymology The epithet ‘ napo’ is derived from the toponym of the type locality near the Napo River basin in Ecuador. A noun in apposition.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	materials_examined	Type material Holotype ECUADOR • ♂; “ Neolindus napo Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ ECUADOR: Sucumbíos, Sacha Lodge; 0.5 ° S, 76.5 ° W; 270 m [m a. s. l.]; 13 – 25 Jul 1994; Hibbs Leg.; ex: malaise \\ SM 0019856 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes ECUADOR • 1 ♂; “ Neolindus napo Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ same data as for holotype \\ SM 0024346 KUNHM-ENT [barcode] \\ Neolindus spp. Det. J. S. Ashe 1996 ”; KUNHM-ENT. SURINAME • 1 ♂; “ Neolindus napo Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Saramacca, West Suriname Road, 178 km WSW, Zaderij Airport; 25 m; 4 ° 59 ’ 6 ’’ N, 56 ° 18 ’ 48 ’’ W; 12 – 14 Jun 1999; Z. H. Falin, B. DeDijin Leg.; SUR 1 F 99 073; ex: flight intercept trap \\ SM 0175295 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	description	Description MEASUREMENTS. BL (5.67), H (0.3, 0.64), A (1.73), A 1 (0.24, 0.07), A 2 (0.1, 0.06), A 3 (0.13, 0.06), A 4 (0.09, 0.06), A 5 (0.08, 0.06), A 6 (0.63, 0.09), A 7 (0.09, 0.07), A 8 (0.09, 0.07), A 9 (0.08, 0.06), A 10 (0.08, 0.06), A 11 (0.11, 0.07), NKW (0.30), GL (0.2), P (0.74, 0.81), E (1.08, 0.84), PC (0.44, 0.21), PF (0.54, 0.24), PT (0.27, 0.11), MSC (0.38, 0.22), MSF (0.68, 0.18), MST (0.54, 0.07), MTC (0.39, 0.24), MTF (0.54, 0.2), MTT (0.56, 0.08). COLOURATION. Head and pronotum brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded, with emargination in front of neck; posterior angles rounded; setation with 1 pair of PCS and 1 row of PMS. Epicranium shiny, without micropunctuation, setation with 1 pair of FS and 2 pairs of PFS, without OS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, IOS, MS, and POS. Sockets of MS and POS umbilicate. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae near gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, longer than head and pronotum combined (Fig. 10 A); antennomeres longer than wide, antennomeres 3 – 11 with tomentose pubescence; antennomere 1 shorter than antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination (Fig. 10 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Prosternum with superior marginal line parallel to anterior margin. Metasternal intercoxal process with 1 pair of acute processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 8 – 9 longitudinal rows (Fig. 10 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thick setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than 2, metatarsomeres 3 and 4 as long as 2, mesotarsomere 5 longer than 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than 2, metatarsomeres 2, 3 and 4 equal, metatarsomere 5 as long as metatarsomere 1. ABDOMEN. Male: tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite VII straight (Fig. 10 B). Posterior margin of sternite VII sinuate (Fig. 10 C). Posterior margin of tergite VIII apically rounded (Fig. 10 D). Posterior margin of sternite VIII with midlongitudinal glabrous depression and midpoint U-shaped emargination, 1 / 5 of segment length, lateral angles rounded (Fig. 10 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, in parameral view, median lobe widest at middle, with pair of oval invaginations on sides from basal foramen to apex of lobe, apex of median lobe with 2 flattened processes. pPMS of complex form, with 2 pairs of acute triangular arms protruding towards parameral side. pLS longer than pPMS and APS, fused into plate on parameral side, with 2 acute flat triangular elongate arms bent towards parameral side. APS flat, with 2 long arms with rounded tips aligned with pLS (Fig. 10 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFB1FFB7FE335806D9A6F87F.taxon	distribution	Distribution This species is known from the type locality in Ecuador (Sucumbíos, Shushufindi), and Suriname (Sipaliwini, Kabalebo). It was collected at low altitudes between 25 to 270 m a. s. l. using Malaise and flight interception traps.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	description	urn: lsid: zoobank. org: act: 574 C 159 C- 8 F 90 - 43 B 7 - 981 D- 66215377 F 07 B Figs 2 B, 11; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	diagnosis	Diagnosis The unusual head chaetotaxy of this species including 2 pairs of PCS, the morphology of abdominal apical sclerites, and the aedeagus make this species similar to N. punctogularis. Neolindus niger Guzman, Tokareva & Żyła sp. nov. differs from it by the shiny black colouration of the head and pronotum (brown in N. punctogularis) and the presence of setae and paired long midlongitudinal invagination anteriorly to the emargination of abdominal sternite VIII (absent in N. punctogularis) (Herman 1991: fig. 205; Fig. 11 E).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	etymology	Etymology The name is a Latin adjective ‘ niger ’ which means ‘ shiny black’ and refers to the colouration of specimens.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	materials_examined	Type material Holotype COSTA RICA • ♂; “ Neolindus niger Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Punta. Prov. Rincon de Osa, 50 m [m a. s. l.]; 8 ° 41.141 ’ N, 83 ° 31.117 ’ W; 23 – 26 Jun 2001; S. & J. Peck Leg.; 01 - 13; ex FIT; CR 1 P 01 005 \\ SM 0551715 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratype COSTA RICA • 1 ♂; “ Neolindus niger Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Puntarenas, Corcovado National Park, Sirena Stn. Corcovado Trail, 150 m [m a. s. l.]; 8 ° 29 ’ 7 ’’ N, 83 ° 34 ’ 39 ’’ W; 28 Jun – 1 Jul 2000; Z. H. Falin Leg.; CR 1 ABF 00 059; ex flight intercept trap \\ SM 0203548 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	description	Description MEASUREMENTS. BL (11.76), H (0.95, 1.24), A (2.91), a 1 (0.56, 0.13), a 2 (0.17, 0.09), a 3 (0.3, 0.12), a 4 (0.25, 0.08), a 5 (0.27, 0.09), a 6 (0.28, 0.11), a 7 (0.27, 0.07), a 8 (0.25, 0.71), a 9 (0.19, 0.08), a 10 (0.18, 0.1), a 11 (0.19, 0.08), NKW (0.60), GL (0.4), P (1.36, 1.65), E (2.01, 1.67), PC (0.73, 0.26), PF (1.28, 0.49), PT (0.9, 0.23), MSC (0.63, 0.39), MSF (1.27, 0.37), MST (1.12, 0.14), MTC (0.53, 0.26), MTF (1.28, 0.29), MTT (1.64, 0.14). COLOURATION. Head and pronotum dark brown; coxa and femur dark brown, tibia and tarsomeres light brown; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin straight with emargination in front of neck; posterior angles straight; setation with 2 pairs of PCS, PMS setae in single row. Epicranium with dense micropunctuation, setation with 1 pair of FS, 1 pair of OS and 2 pairs of PFS. Gena with irregular surface, with depression from mandibular base to middle of eye, with 1 OT; socket of mandibular seta umbilicate, sockets of POS concave. Postgena with row of 3 setae in line from eye to gular suture and 1 seta parallel to PTOS on each side; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula with irregular row of 6 – 8 setae. Neck with foramen magnum occupying ½ of occiput. Antenna thin and elongate, longer than head and pronotum combined (Fig. 11 A); antennomeres longer than wide; tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with V-shaped emargination and 2 dark thick setae in median line of lobes (Fig. 11 A; Supp. file 2). THORAX. Pronotum slightly wider than long. Basisternum of prosternum with row of 8 setae on anterior margin. Metasternal intercoxal process with 1 pair of rounded processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 8 – 10 longitudinal rows (Fig. 11 A; Supp. file 2). LEGS. Protibia with 5 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 equal to mesotarsomere 2, mesotarsomere 3 shorter than 1 and as long as 4, mesotarsomere 5 longer than 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than 2, metatarsomere 2 longer than 3, metatarsomere 4 equals 3, metatarsomere 5 equals 2. ABDOMEN. Male: tergites III – VI with high density of micropunctuation, tergites VII and VIII with low density of micropunctuation. Tergite VII with posterior margin straight (Fig. 11 B). Sternite VII with posterior margin moderately emarginate, bordered by obtuse distinct processes (Fig. 11 C). Tergite VIII with posterior margin trilobed, central lobe triangular, apically acute; lateral lobes slightly acute (Fig. 11 D). Sternite VIII posterior margin with broad emargination; with pair of midlongitudinal invaginations on sides of flat elevation in middle (Fig. 11 E). Tergite IX with posterior margin deeply emarginated to ¾ of segment length (Fig. 11 A; as in Herman 1991: fig. 137); aedeagus, median lobe medially widened, apex with deep emargination bordered by rounded processes; pLS thin, with net-like microsculpture, forming elongate flat processes; APS spoon-shaped, with rounded apical edge; pPMS sickle-like, wider than long in lateral view; apical arm closer to parameral side, bent towards it, apical arm closer to antiparameral side, straight (Fig. 11 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8EFF8BFE3D5D05D88CFBB2.taxon	distribution	Distribution This species is known from the type locality in Costa Rica (Puntarenas, Rincon de Osa) and Corcovado National Park (Puntarenas). It was collected in a cloud forest at a low altitude using flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	description	urn: lsid: zoobank. org: act: C 9 A 162 F 2 - 2 A 4 E- 490 A- 842 A- 36 D 00 DE 02441 Figs 2 A, 12; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	diagnosis	Diagnosis Neolindus ornatus Guzman, Tokareva & Żyła sp. nov. is similar to N. cuneatus in sternite and tergite VIII (Fig. 12 C, D and figs 177, 178 in Herman 1991, respectively). However, N. ornatus has longer pPMS and longer, bifid pLS of aedeagus than N. cuneatus (Herman 1991: fig. 179; Fig. 12 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	etymology	Etymology The name is derived from the Latin word ‘ ornatus ’ meaning ‘ adorned’ or ‘ equipped’, referring to the complex structure of the aedeagus. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus ornatus Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Matoury, 41.5 km SSW on HWY N 2; 4 ° 37 ’ 22 ’’ N, 52 ° 22 ’ 35 ’’ W; 50 m [m a. s. l.]; 29 May – 9 Jun 1997, J. Ashe, R. Brooks Leg.; FG 1 AB 97 170; ex: light intercept trap \\ SM 0100511 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes FRENCH GUIANA • 1 ♂; “ Neolindus ornatus Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Matoury, 41.5 km SSW on HWY N 2; 4 ° 37 ’ 22 ’’ N, 52 ° 22 ’ 35 ’’ W, 50 m; 29 May – 9 Jun 1997, J. Ashe, R. Brooks Leg.; FG 1 AB 97 170 ex: flight intercept trap \\ [barcode] SM 0100507 KUNHM-ENT ”; KUNHM-ENT • 1 ♂; “ Cayenne, 33.5 km S and 8.4 km NW of Hwy N 2 on Hwy D 5; 4 ° 48 ’ 18 ’’ N, 52 ° 28 ’ 41 ’’ W; 30 m; 26 – 28 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 059 ex: flight intercept trap \\ [barcode] SM 0099850 KUNHM-ENT ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	description	Description MEASUREMENTS. BL (7.3), H (0.48, 0.79), A (1.31), A 1 (0.3, 0.09), A 2 (0.05, 0.06), A 3 (0.13, 0.06), A 4 (0.09, 0.06), A 5 (0.1, 0.06), A 6 (0.01, 0.07), A 7 (0.12, 0.07), A 8 (0.11, 0.07), A 9 (0.11, 0.08), A 10 (0.11, 0.09), A 11 (0.17, 0.09), NKW (0.40), GL (0.2), P (0.89, 0.91), E (1.16, 0.96), PC (0.41, 0.26), PF (0.69, 0.26), PT (0.32, 0.12), MSC (0.44, 0.24), MSF (0.82, 0.19), MST (0.47, 0.09), MTC (0.46, 0.33), MTF (0.75, 0.22), MTT (0.65, 0.11). COLOURATION. Whole body and appendices brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded, with emargination in front of neck; posterior angles rounded, setation with 1 pair of PCS, 1 row of PMS, and longer setae near posterior margin before neck. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 2 pairs of PFS, without OS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 1 or 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae near gular sutures. Neck with foramen magnum occupying ⅔ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 12 A); antennomeres longer than wide, tomentose pubescence from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination (Fig. 12 A; Supp. file 2). THORAX. Pronotum quadrate, with umbilicate randomly distributed micropunctures. Metasternal intercoxal process with 1 pair of acute processes, joining medially in U-like invagination. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 10 – 11 longitudinal rows (Fig. 12 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thick setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as 2, mesotarsomere 3 equals to 4, shorter than 2, mesotarsomere 5 longer than 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than 2, metatarsomeres 3 and 4 as long as metatarsomere, metatarsomere 5 as long as metatarsomere 1. ABDOMEN. Male: tergites with low-density micropunctuation. Posterior margin of tergite VII straight (Fig. 12 B). Posterior margin of sternite VII slightly emarginated, with small midpart invagination (Fig. 12 C). Posterior margin of tergite VIII truncate (Fig. 12 D). Posterior margin of sternite VIII with midlongitudinal V-shape emargination, with elevated area in midpart (Fig. 12 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length, divided; aedeagus, in parameral view, median lobe basally narrower, and wider in middle, apex deeply invaginated, with prominent sclerotised rounded processes on each side; foramen triangular, with round angles; APS not visible; pLS flattened with 2 arms, apical arm, curving towards non-parameral side, ending in acute tip, basal arm projecting to parameral side ending as acute tip; pPMS flattened with 2 arms, apical arm tapering to end as acute tip, basal arm projects in parameral side with rounded end (Fig. 12 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF8CFF8EFE02589EDC0BFEAA.taxon	distribution	Distribution The species is known from the type locality in French Guyana (Cayenne, Roura, near Matoury) and the surrounding area. It was collected at low altitudes using a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	description	urn: lsid: zoobank. org: act: 2 D 2 F 08 A 4 - E 884 - 4 C 1 C- 8 AED-B 289223 FDC 9 B Fig. 13; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	diagnosis	Diagnosis Neolindus parahermani sp. nov. resembles N. hermani in habitus and genitalia morphology. However, they can be distinguished by the structure of pPMS of aedeagus. Antiparameral apical arms of pPMS in N. parahermani are bifid with two acute processes, while in N. hermani, they end with a rounded tip (Asenjo 2011: figs 9 – 10; Fig. 13 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	etymology	Etymology The specific epithet is the combination of ‘ para ’, a Latin prefix that means ‘ similar’ or ‘ equal’, and ‘ hermani ’ as the name of the species described by Asenjo (2011), N. hermani. It indicates the similarity of morphology between aedeagi of the two species. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus parahermani Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Réserve Trésor, cca 225 m: 4 ° 36 ’ 37.0 ” N, 52 ° 16 ’ 42.5 ” W; Dec 2009; Stephane Brule Leg.; ex. Window trap \\ BMNH {E} 2010 - 62 ”; NHMUK.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	description	Description MEASUREMENTS. BL (13.63), H (0.96, 2.03), A (3.56), A 1 (0.78, 0.16), A 2 (0.19, 0.14), A 3 (0.37, 0.13), A 4 (0.27, 0.1), A 5 (0.28, 0.11), A 6 (0.33, 0.1), A 7 (0.34, 0.1), A 8 (0.31, 0.1), A 9 (0.26, 0.1), A 10 (0.2, 0.11), A 11 (0.21, 0.1), NKW (0.85), GL (0.4), P (1.87, 2.3), E (2.63, 2.49), PC (0.7, 0.42), PF (1.35, 0.76), PT (0.81, 0.35), MSC (0.72, 0.59), MSF (1.58, 0.47), MST (1.14, 0.15), MTC (0.71, 0.64), MTF (1.72, 0.49), MTT (1.7, 0.22). COLOURATION. Head and pronotum black; legs dark brown; abdomen black; intersegmental membrane dark brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin rounded; with emargination in front of neck; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, high density micropunctuation, setation with 2 pairs of FS and 1 pair of PFS, without OS. Gena with irregular surface, with depression from mandibular base to posterior margin of head, setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 3 – 5 setae in line from eye to gular suture; gular sutures not reaching posterior margin of head; anterior part of gula with 6 – 8 setae. Neck with foramen magnum occupying ¾ of occiput. Antenna thin, longer than head and pronotum combined (Fig. 13 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination (Fig. 13 A; Supp. file 2). THORAX. Pronotum wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Metasternal intercoxal process with pair of acute processes light brown at tips. Elytron longer than wide, surface of elytra with moderately dense umbilicate micropunctures in 8 – 10 longitudinal rows (Fig. 13 A; Supp. file 2). LEGS. Protibia with 5 – 6 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as 2, mesotarsomere 3 shorter than 1 and as long as 4, mesotarsomere 5 as long as mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomere 3 shorter than metatarsomere 2, metatarsomere 4 as long metatarsomere 3, metatarsomere 5 as long as metatarsome 1. ABDOMEN. Male: tergites with low-density random micropunctuation. Posterior margin of tergite VII straight (Fig. 13 B). Posterior margin of sternite VII with wide midline emargination delimited with 2 long teeth-like processes with acute ends (Fig. 13 C). Posterior margin of tergite VIII trilobed, with rounded middle lobe and triangle acute lateral lobes (Fig. 13 D). Posterior margin of sternite VIII with broad midline emargination about ½ of segment length and 1 pair of longitudinal depressions on sides of midline emargination (Fig. 13 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ½ of segment length; aedeagus, in parameral view, median lobe slightly widened at apical part, apex with deep V-shaped emargination and with narrow flattened lateral processes adjacent to pair of lateral elongated ‘ windows’ in membrane. pPMS elongated, complex, curved around longitudinal axis, with 2 apical arms almost fully fused and pointing distad and 1 apical arm short and bent laterad; bases of pPMS triangular, anchored in median foramen with internal sclerotised ring. pLS thin, slightly longer than pPMS and APS, with net-like microsculpture, forming elongate flat processes with lateral rounded lobes. APS spoon-shaped, with slightly emarginate top, shorter than pPMS (Fig. 13 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF89FF80FE625D96DC4AFB6C.taxon	distribution	Distribution This species is only known from the type locality in French Guiana (Roura, Réserve Naturelle Régionale Trésor). It was collected at a low altitude of 225 m a. s. l. using window trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	description	urn: lsid: zoobank. org: act: 1 C 892 CF 5 - 7 D 91 - 49 E 1 - A 867 - 02 FC 75 A 3 D 1 D 8 Fig. 14; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	diagnosis	Diagnosis Neolindus paraplectrus Guzman, Tokareva & Żyła sp. nov. is most similar to N. plectrus by the morphology. However, N. paraplectrus can be distinguished by the straight posterior margin of abdominal sternite VII, which is widely emarginate in N. plectrus (Fig. 14 C; Herman 1991: fig. 83) and the median lobe of aedeagus with tapering triangular apex with additional ridge outlining the triangle, absent in N. plectrus (Herman 1991: fig. 81; Fig. 14 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	etymology	Etymology The specific epithet is the combination of ‘ para ’, a Latin prefix that means ‘ similar’ or ‘ equal’, and ‘ plectrus ’ as the name of the species described by Herman (1991), N. plectrus. It indicates the similarity of morphology between the aedeagi of these species. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	materials_examined	Type material Holotype VENEZUELA • ♂; “ Neolindus paraplectrus Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Merida, Tabay, 7 km E., La Mucuy Station, Sierra Nevada Natl Park; 2300 – 2700 mm; 8 ° 37 ’ 44 ’’ N, 71 ° 2 ’ 26 ’’ W; 24 May 1998, R. Anderson Leg.; VEN 1 A 98 034 B; ex: cloud forest litter \\ SM 0110682 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	description	Description MEASUREMENTS. BL (7.4), H (0.55, 0.86), A (1), a 1 (0.21, 0.07), a 2 (0.09, 0.07), a 3 (0.08, 0.06), a 4 (0.07, 0.08), a 5 (0.07, 0.09), a 6 (0.08, 0.1), a 7 (0.08, 0.09), a 8 (0.09, 0.11), a 9 (0.08, 0.1), a 10 (0.08, 0.1), a 11 (0.08, 0.1), NKW (0.50), GL (0.22), P (1.12, 0.97), E (0.8, 0.88), PC (0.35, 0.22), PF (0.7, 0.42), PT (0.48, 0.16), MSC (0.42, 0.24), MSF (0.59, 0.22), MST (0.51, 0.13), MTC (0.25, 0.38), MTF (0.73, 0.38), MTT (0.78, 0.12). COLOURATION. Head and pronotum dark brown; legs brown; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded with emargination in front of neck; posterior angles rounded; setation with 1 pair of PCS and 1 row of PMS clustered in middle. Epicranium convex, elevated above gena, with low-density micropunctuation in first part of head, setation with 1 pair of OS, without FS and PFS. Gena with smooth surface, with depression from mandibular base to posterior margin of head, setation with 2 trichobothria, MS close to antennal incisions, sockets of POS and PTOS umbilicate. Postgena with row of 3 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula without setae, with lateral invaginations; midlength of gula with 2 setae; posterior margin of head with 2 setae close to gular sutures. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 14 A); antennomeres 1 – 4 longer than wide, antennomeres 5 – 11 wider than long, tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 2 longer than 3. Labrum with multiple lobes and 1 wide quadrate emargination, marginal clypeal setae of equal lengths (Fig. 14 A; Supp. file 2). THORAX. Pronotum slightly longer than wide, with umbilicate randomly distributed micropunctures and irregular setation. Prosternal basisternum with longitudinal carina and smooth surface. Mesosternum with scutiform basisternum with scarce tuberculate micropunctuation. Mesanepisternum smooth, transversely convex. Metasternal intercoxal process with 1 pair of rounded processes. Elytron wider than long, shorter than pronotum; surface of elytra with umbilicate micropunctures in 8 – 9 moderately dense longitudinal rows (Fig. 14 A; Supp. file 2). LEGS. Protibia with 3 well-developed combs of setae, one of combs with thickened spines-like setae; mesotibial apical ctenidium on both sides, spines-like; mesotarsomeres 1, 2 and 3 as long as mesotarsomere 4, mesotarsomere 5 as long as mesotarsomeres 1 – 4 combined; inner ctenidium of metatibia longer than outer; metatarsomeres 1 to 4 subequal, metatarsomere 5 as long as metatarsomeres 1 – 4 combined. ABDOMEN. Male: abdomen gradually widened towards segment VII. Tergites with low density of micropunctuation. Posterior margin of tergite and sternite VII straight (Fig. 14 B, C). Posterior margin of tergite VIII elongate, rounded (Fig. 14 D). Posterior margin of sternite VIII with midlongitudinal V-shaped emargination, and longitudinal glabrous invagination with carinated margin in middle part of emargination, lateral angles rounded (Fig. 14 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length, fused; aedeagus, in parameral view, median lobe elongated, gradually narrowed towards apex; apex of lobe with triangular tapering process with additional ridge outlining triangular area. pPMS and pLS absent. APS long, spoon-like, flattened, shorter than median lobe, partially covering median foramen (Fig. 14 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF87FF83FE6158D4DCA2FEAB.taxon	distribution	Distribution This species is known only from the type locality in Venezuela (Merida, Tabay). It was collected in a cloud forest in Sierra Nevada National Park at a high altitude between 2300 – 2700 m a. s. l.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	description	urn: lsid: zoobank. org: act: FAD 92 BD 2 - 8 A 53 - 4 DBC- 88 F 1 - 3 BD 907287 FD 2 Fig. 15; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	diagnosis	Diagnosis This species resembles N. sinuatus and N. basisinuatus, but can be distinguished by a combination of characters. Unlike in N. sinuatus, the posterior margin of male sternite VIII in N. parasinuatus Guzman, Tokareva & Żyła sp. nov. has a wide emargination with a concave middle part surrounded by invagination on the sclerite (Herman 1991: fig. 199; Fig. 15 E). In contrast to N. basisinuatus, the posterior margin of tergite VIII in N. parasinuatus is rounded (Herman 1991: fig. 194; Fig. 15 D). In comparison to both species, the aedeagus of N. parasinuatus lacks a pair of setal brushes near the median invagination on the parameral side (Herman 1991: figs 193, 198; Fig. 15 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	etymology	Etymology The specific epithet is a combination of ‘ para ’, a Latin prefix that means ‘ similar to’ or ‘ equal’, and ‘ sinuatus ’ as the name of the species described by Herman (1991), N. sinuatus. It indicates the similarity in the morphology of the aedeagi of the two species. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	materials_examined	Type material Holotype COLOMBIA • ♂; “ Neolindus parasinuatus Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Amazonas, PNN [National Natural Park] Amacayacu, San Martin; 3 ° 46 ’ S, 70 ° 18 ’ W; Malaise; 19 – 27 Out 1997; 150 m [m. a. s. l]; B. Amado Leg.; M. 839 \\ SM 0549284 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes COLOMBIA • 1 ♂; “ Neolindus parasinuatus Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Amazonas, PNN Amacayacu, San Martin; 3 ° 46 ’ S, 70 ° 18 ’ W; 150 m; Malaise; 11 – 29 Nov 2000; B. Amado Leg.; M. 1318. \\ SM 0549282 KUNHM-ENT [barcode] ”: KUNHM-ENT • 1 ♂; same data as for preceding; “ 19 – 27 Nov 2000 \\ SM 549297 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	description	Description MEASUREMENTS. BL (7.9), H (0.85, 0.69), A (1.67), a 1 (0.37, 0.08), a 2 (0.13, 0.05), a 3 (0.15, 0.07), a 4 (0.13, 0.07), a 5 (0.11, 0.08), a 6 (0.14, 0.09), a 7 (0.14, 0.09), a 8 (0.13, 0.86), a 9 (0.12, 0.89), a 10 (0.1, 0.1), a 11 (0.14, 0.09), NKW (0.37), GL (0.23), P (0.86, 0.92), E (1.27, 1.2), PC (0.63, 0.38), PF (0.71, 0.29), PT (0.43, 0.14), MSC (0.52, 0.3), MSF (0.1, 0.2), MST (0.67, 0.12), MTC (0.39, 0.33), MTF (1.16, 2.28), MTT (1, 0.1). COLOURATION. Whole body and appendages brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin slightly convex anteriad with emargination in front of neck; posterior angles rounded; setation with 1 PCS, PMS in moderately dense row. Epicranium with low-density micropunctuation, setation with 1 pair of FS, 1 pair of OS, and 3 pairs of PFS. Gena with irregular surface, with depression from mandibular base to middle of eye, with 1 OT; socket of mandibular seta umbilicate, sockets of POS with raised edges. Postgena with row of 3 setae in line from eye to gular suture; posterior margin close to gular sutures with 2 setae; gular sutures straight, extended to posterior margin of head; anterior part of gula with 2 rows of setae. Antenna shorter than head and pronotum combined (Fig. 15 A), moniliform starting from antennomere 5; antennomeres 1 – 7, 10 and 11 longer than wide, antennomeres 8 and 9 wider than long; tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum tetralobed, with 2 dark thick setae in median line of lobes, central lobes more developed than lateral lobes, emargination deep, V-shaped (Fig. 15 A; Supp. file 2). THORAX. Pronotum quadrate, with umbilicate micropunctures, 2 paramedial and 2 lateral distinct rows of setae. Basisternum of prosternum triangular, with row of 8 setae on anterior margin, longitudinal ridge, and wrinkled surface. Metasternal intercoxal process with 1 pair of acute long processes. Elytron longer than wide, surface of elytra with punctures in 7 – 8 longitudinal rows (Fig. 15 A; Supp. file 2). LEGS. Protibia with 3 well-developed combs of setae, 1 anterior row covering ⅔ of tibia length; mesotibial apical ctenidium only on inner side; outer margin with row of spines; metatarsomere 1 as long as metatarsomere 2, metatarsomere 3 as long as metatarsomere 4 and shorter than 1, metatarsomere 5 as long as metatarsomere 1. ABDOMEN. Male: tergites with sparse micropunctures. Anterior margin of tergite VII with 4 basal canals with elevated margin, posterior margin straight (Fig. 15 B). Posterior margin of sternite VII straight (Fig. 15 C). Anterior margin of tergite VIII with 4 internal canals, central canals fused, forming inverted pentagon; posterior margin rounded (Fig. 15 D). Posterior margin of sternite VIII with wide and shallow midlongitudinal emargination with straight middle part and rounded lateral angles (Fig. 15 E). Posterior margin of tergite IX with midlongitudinal deep emargination of ¾ of segment length (Herman 1991: fig. 192); aedeagus, median lobe rounded at base, with parallel lateral sides, apex on parameral side deeply notched with elongate, blunt-end median hole or cavity. pPMS shell-shaped, triangular, robust, immersed into opening of median foramen. pLS absent. APS flattened, wider than long, shorter than pPMS; 1 pair of inner sclerites attached to pPMS and single, symmetrical, flattened sclerite attached to paired ones (Fig. 15 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF84FF85FE625D96DB94FA5E.taxon	distribution	Distribution The species is known only from the type locality in Colombia (Amazonas, Leticia, Parque Nacional Natural Amacayacu). It was collected by Malaise trap at low altitude.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	description	urn: lsid: zoobank. org: act: 89840 CD 7 - 5 CE 1 - 4759 - 8 B 2 A- 305 CAB 96681 D Fig. 16; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	diagnosis	Diagnosis This species resembles N. utriensis sp. nov. and N. maya sp. nov. in habitus, but can be distinguished from them by the characters of the apical sclerites of the aedeagus. pPMS of N. parautriensis Guzman, Tokareva & Żyła sp. nov. are crossed with each other, narrower than in N. utriensis and N. maya with apical arms of equal lengths, slightly bent inwards, while in N. utriensis and N. maya, pPMS are more robust, not crossing each other, with apical arms of different lengths (Fig. 16 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	etymology	Etymology The name is a combination of ‘ para ’, a Latin prefix that means ‘ similar to’ or ‘ equal’, and ‘ utriensis ’, which refers to one of the species described in this paper. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	materials_examined	Type material Holotype PANAMA • ♂; “ Neolindus parautriensis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Colón, 14 km N jct Escobal & Piña Rds.; 02 – 11 Jun 1996; J. Ashe & R. Brooks Leg.; PAN 1 AB 96 181 B; ex: flight interception trap \\ SM 0016476 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes PANAMA • 1 ♀; “ Neolindus parautriensis Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Colón, 14 km N jct Escobal & Piña Rds.; 02 – 11 Jun 1996; J. Ashe & R. Brooks Leg.; PAN 1 AB 96 181 B; ex: flight interception trap \\ SM 0016473 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ SM 0016472 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ SM 0016477 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	description	Description MEASUREMENTS. BL (11.48), H (1.01, 1.68), A (3.15), a 1 (0.63, 0.14), a 2 (0.23, 0.12), a 3 (0.3, 0.12), a 4 (0.26, 0.11), a 5 (0.27, 0.12), a 6 (0.26, 0.09), a 7 (0.24, 0.11), a 8 (0.25, 0.1), a 9 (0.27, 0.10), a 10 (0.21, 0.10), a 11 (0.22, 0.10), NKW (0.60), GL (0.38), P (1.38, 1.98), E (2.24, 1.96), PC (0.94, 0.52), PF (1.46, 0.46), PT (0.06, 0.36), MSC (0.44, 0.5), MSF (1.32, 0.42), MST (0.95, 0.18), MTC (0.72, 0.98), MTF (1.93, 0.40), MTT (1.74, 0.18). COLOURATION. Head and pronotum dark brown; antennae and maxillary palpi light brown; legs brown and partially light brown; meso- and metatibia lighter than other parts of respective legs; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin straight with emargination in front of neck; posterior angles straight; setation with 1 pair of PCS, PMS setae in dense straight row. Epicranium with dense micropunctuation, setation with 1 pair of FS, 1 OS, 2 – 3 pairs of PFS in row parallel to PMS, and without OS. Gena with irregular surface, setation with 1 POS, 2 PTOS, 1 MS with umbilicate socket, and 1 OT, sockets of POS concave. Postgena with row of 3 – 5 setae in line from eye to gular suture, gular sutures not reaching posterior margin of head, anterior part of gula with irregular row of 6 setae. Antenna thin, longer than head and pronotum combined (Fig. 16 A); antennomeres longer than wide; tomentose pubescence starting from antennomere 3; antennomere 1 as long as antennomeres 2 and 3 combined; antennomere 3 longer than 2. Labrum bilobed, with shallow V-shaped emargination (Fig. 16 A; Supp. file 2). THORAX. Pronotum wider than long, almost quadrate (ratio 0.87), with umbilicate punctures and 2 paramedial rows of setae. Basisternum of prosternum triangular, with row of 8 setae on anterior margin, well-developed longitudinal ridge, and wrinkled surface. Metasternal intercoxal process with 1 pair of acute spine-like processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 6 – 8 longitudinal rows (Fig. 16 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; mesotibial apical ctenidia on both sides, inner longer than outer; mesotarsomere 1 longer than mesotarsomere 2; metatarsomere 1 as long as metatarsomere 2, metatarsomere 3 as long as metatarsomere 4 but shorter than metatarsomere 2, metatarsomere 5 as long as metatarsomere 1. ABDOMEN. Tergites with sparse micropunctures. Posterior margin of tergite VII straight (Fig. 16 B). Anterior margin of tergite VIII with 4 internal canals, central canals fused, forming inverted pentagon. Anterior margin of sternite VIII with 4 internal canals. Male: posterior margin of sternite VII with moderate emargination at middle, with acute processes on each side (Fig. 16 C). Posterior margin of tergite VIII with 3 acute processes, wide 1 in middle and 2 narrow ones on sides (Fig. 16 D). Posterior margin of sternite VIII bilobed, with narrow median emargination reaching up to ¼ of segment length, lobes obtuse (Fig. 16 E). Posterior margin of tergite IX with midlongitudinal deep emargination, reaching up to ¾ of segment length (Fig. 16 A; as in Asenjo 2011: fig. 12); aedeagus, median lobe broadened at base, apex with moderately deep emargination and rounded processes on each side. pLS thin, with net-like microsculpture, forming elongate flat processes equal in length to pPMS. APS spoon-shaped with widened distal third, wider than pLS, with slightly emarginate apical margin. pPMS elongate, thin, crossed with each other, with 2 acute apical arms of same length, slightly bent inwards (Fig. 16 F – I). Female: lack of emargination on abdominal segments VIII and IX.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF82FF98FE6D59A5DD4CFD0F.taxon	distribution	Distribution This species is known from the type locality in Panama (Colón, Chagres). Based on the label information, it was collected in Bosque Protector San Lorenzo at a low altitude using a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	description	urn: lsid: zoobank. org: act: 1 F 0 AFABD-E 3 B 6 - 4886 - 88 AA- 1 E 909 C 32 AC 13 Figs 2 E – G, 17; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	diagnosis	Diagnosis Neolindus pseudosensillaris Guzman, Tokareva & Żyła sp. nov. resembles N. elegans sp. nov in the shape of posterior margin of sternites VII, VIII, and tergite VIII, as well as structure of aedeagus with denticles on pPMS. However, N. pseudosensillaris has a unique character among all known species of Neolindus: 3 pairs of tapered, fusiform, smooth structures resembling sensilla on the epicranium (Fig. 2 E – G). Moreover, in N. pseudosensillaris, posterior margin of sternite VII has narrower midline emargination without acute protruding processes on its sides, posterior margin of sternite VIII has narrower V-shape emargination, posterior margin of tergite VIII is slightly truncate (Figs 5 C – E, 17 C – E), and the shape of median lobe as well as morphology of APS is different from those of N. elegans (Figs 5 F – I, 17 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	etymology	Etymology The name combines ‘ pseudo ’, the Latin prefix for ‘ false’, with the New Latin ‘ sensillum ’, a diminutive of Latin ‘ sensus ’ (‘ perception, feeling’). It refers to the three sensilla-like structures on the head of unknown origin. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	materials_examined	Type material Holotype Brazil • ♂; “ Neolindus pseudosensillaris Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Amazonas, Manaus, INPA / Smithsonian Resv; 2 ° 25 ’ 00 ’’ S, 59 ° 50 ’ 00 ’’ W; 93 m [m a. s. l.]; Jun 1994; R. Didham Leg.; 631 | 5 ”; NHMUK. Paratypes BRAZIL • 1 ♂; “ Neolindus pseudosensillaris Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ same data as for holotype \\ 901 | 3 (BMNH) ”; NHMUK • 1 ♀; same data as for preceding; “ 21 | 3 ’ (BMNH) ”; NHMUK • 1 ♀; same data as for preceding; “ 197 | 10 (BMNH) ”; NHMUK • 1 ♀; same data as for preceding; “ 501 | 1 (BMNH) ”; NHMUK • 1 ♀; “ Reserva Florestal Adolpho Ducke; 2 ° 57 ’ 47.4 ” S, 59 ° 55 ’ 21.2 ” W; 109 m; Feb 1995, Barbosa Leg., M. G. V; G 26.30 (BMNH) ”; NHMUK.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	description	Description MEASUREMENTS. BL (5.3), H (0.43, 0.62), A (1.15), A 1 (0.25, 0.07), A 2 (0.1, 0.06), A 3 (0.1, 0.07), A 4 (0.07, 0.06), A 5 (0.08, 0.06), A 6 (0.08, 0.06), A 7 (0.09, 0.08), A 8 (0.07, 0.07), A 9 (0.09 0.08), A 10 (0.08, 0.08), A 11 (0.15, 0.07), NKW (0.25), GL (0.2), P (0.66, 0.73), E (0.98, 0.87), PC (0.4, 0.1), PF (0.36, 0.17), PT (0.61, 0.25), MSC (0.31, 0.14), MSF (0.66, 0.16), MST (0.48, 0.07), MTC (0.29, 0.22), MTF (0.66, 0.16), MTT (0.68, 0.08). COLOURATION. Head and pronotum dark brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin rounded, with emargination in front of neck; posterior angles straight; setation with 1 pair of PCS and 1 row of PMS. Epicranium smooth, without micropunctuation, setation with 1 pair of FS and 2 pairs of PFS, without OS; 3 pairs in line of tapered fusiform smooth structures resembling sensilla with sockets posterior to antennal incisions. Gena with irregular surface, with depression from mandibular base to middle of eye; setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 3 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining with neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 pairs of setae. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 17 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with U-shaped emargination (Fig. 17 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Prosternum with superior marginal line parallel to anterior margin. Metasternal intercoxal process with 1 pair of rounded processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 8 – 10 longitudinal rows (Fig. 17 A; Supp. file 2). LEGS. Protibia with 5 – 6 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thick setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than 2, mesotarsomeres 3 and 4 as long as 2, mesotarsomere 5 longer than mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 as long as metatarsomere 2, metatarsomere 3 shorter than metatarsomere 1 and as long as metatarsomere 4, metatarsomere 5 longer than metatarsomere 1. ABDOMEN. Tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite VII straight (Fig. 17 B). Posterior margin of tergite VIII elongate, rounded, with lateral edges slightly curved (Fig. 17 D). Male: posterior margin of sternite VII with moderately deep midline emargination bordered by acute processes, with internal margin of emargination slightly bent posteriad in middle; surface adjacent to emargination with shallow, median, glabrous, and quadrate invagination (Fig. 17 C). Posterior margin of sternite VIII with midlongitudinal V-shaped emargination, lateral angles rounded (Fig. 17 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, in parameral view, median lobe slightly widened at base, apex with narrow emargination not exposing margin of median foramen and with flattened lateral processes folded towards emargination. Apical sclerites with more than 3 pairs of elements. pPMS robust, flattened, curved around longitudinal axis, with multiple denticles on top. pLS absent. APS flattened, spoon-like, with 3 apical projections; middle projection wide, flat, enfolding pPMS, longer than pPMS, with elevated angles; lateral projections acute, bent towards parameral side, shorter than pPMS (16 F – I). Female: lack of emargination on abdominal segments VII and VIII.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9FFF99FE455EFAD92CF812.taxon	distribution	Distribution The species is known from the type locality in Brazil (Amazonas, Manaus, Projeto Dinâmica Biológica de Fragmentos Florestais reserve) and nearby Reserva Florestal Adolpho Ducke. It was collected at a low altitude.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	description	urn: lsid: zoobank. org: act: D 5349 E 92 - F 31 A- 4 FBA-B 904 - 7 D 7326937 DC 4 Fig. 18; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	diagnosis	Diagnosis The habitus of this species, as well as the broadly rounded posterior margin of tergite VIII, resembles that of N. paralellus. However, N. sauron Guzman, Tokareva & Żyła sp. nov. differs from N. paralellus in the morphology of sternite VIII with a concave, broad emargination of the posterior margin and a glabrous midlongitudinal invagination on the rest of the sclerite (Fig. 18 E), while it is deeply and narrowly emarginated in N. paralellus (Herman 1991: fig. 88). Additionally, N. sauron differs in the shape of the apical complex of sclerites of the aedeagus (Herman 1991: fig. 85; Fig. 18 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	etymology	Etymology The name is inspired by the structure of antiparameral sclerite in the aedeagus resembling the ‘ Eye of Sauron’ in the movie trilogy ‘ The Lord of the Rings’ based on J. R. R. Tolkien’s book series. A noun in apposition.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	materials_examined	Type material Holotype GUYANA • ♂; “ Neolindus sauron Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Region 8, Iwokrama Forest, Turtle Mt. Summit; 4 ° 43 ’ 57.0 ” N, 58 ° 44 ’ 01.0 ” W; 290 m [m a. s. l.]; 30 May – 1 Jun 2001; R. Brooks, Z. Falin Leg.; GUY 18 F 01 097; ex. flight interception trap \\ SM 0253302 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes GUYANA • 1 ♂; “ Neolindus sauron Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Region 8, Iwokrama Forest, 26 km SW Kurupukari, Iwokrama Mt. Basecamp; 4 ° 20 ’ 17.0 ” N, 58 ° 48 ’ 38.0 ” W, 300 m; 23 – 25 May 2002; R. Brooks, Z. Falin Leg.; GUY 18 F 01 031; ex. flight intercept trap \\ SM 0253198 KUNHM-ENT [barcode] ” • 1 ♂; same data as for preceding; “ SM 0253197 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	description	Description MEASUREMENTS. BL (6.3), H (0.38, 0.78), A (1.23), A 1 (0.28, 0.07), A 2 (0.08, 0.06), A 3 (0.07, 0.14), A 4 (0.09, 0.06), A 5 (0.1, 0.06), A 6 (0.12, 0.07), A 7 (0.11, 0.07), A 8 (0.10, 0.07), A 9 (0.1, 0.07), A 10 (0.09, 0.07), A 11 (0.09, 0.09), NKW (0.40), GL (0.21), P (0.92, 0.94), E (1.23, 1.1), PC (0.5, 0.26), PF (0.78, 0.34), PT (0.43, 0.12), MSC (0.38, 0.25), MSF (0.77, 0.21), MST (0.38, 0.1), MTC (0.28, 0.26), MTF (0.911, 0.16), MTT (0.64, 0.07). COLOURATION. Head and pronotum brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded with emargination at middle; posterior angles rounded; setation with 1 pair of PCS, 1 row of PMS, and 2 thick and dark setae at neck articulation. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 1 pair of PFS, without OS. Gena with depression from mandibular base to middle of eye, setation with 1 OT, MS, and POS; sockets of POS umbilicate. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head, joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae close to gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 18 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 2 longer than 3. Labrum bilobed, with shallow, V-shaped emargination (Fig. 18 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate micropunctures, 2 paramedial and 2 lateral rows of setae. Metasternal intercoxal process with 1 pair of rounded processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 10 – 11 longitudinal rows (Fig. 18 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thin setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than mesotarsomere 2, mesotarsomeres 3 and 4 as long as mesotarsomere 2, mesotarsomere 5 as long as mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomere 2 longer than metatarsomere 3, metatarsomere 3 as long as metatarsomere 4, metatarsomere 5 as long as metatarsomere 1. ABDOMEN. Male: tergites with low-density micropunctuation. Posterior margin of tergite VII slightly convex, almost straight (Fig. 18 B). Posterior margin of sternite VII sinuate, with small midlongitudinal depression (Fig. 18 C). Posterior margin of tergite VIII broad, apically rounded (Fig. 18 D). Posterior margin of sternite VIII with median broad, concave emargination, lateral angles rounded; disc of sternite with longitudinal glabrous invagination (Fig. 18 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length, fused; aedeagus, median lobe of same width from parameral side, rounded at base, apex notched on parameral side with elongate median hole or cavity reaching basal foramen elevation, apex of median lobe on parameral side with sclerotised processes on each sides of median foramen; pPMS elongate, thin, bifid at apex, longer apex pointing laterad. pLS shorter than pPMS and APM, pocket-shaped, enfolding pPMS. APS long, spoon-shaped, apically bifid; apical processes flattened, with distinct angle protruding laterad; emargination dividing 2 apical processes rounded (17 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9CFF9DFE075D05D9F8FA8E.taxon	distribution	Distribution This species is known from the type locality in Guyana (Upper Demerara-Berbice, Iwokrama Forest, Turtle Mountain) and the nearby Iwokrama Mountains. It was collected at a lower altitude with a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	description	urn: lsid: zoobank. org: act: 66 D 4 DDA 9 - 79 E 6 - 49 AF-BCCB- 2 DC 7 F 98 E 33 D 3 Figs 2 C – D, 19; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	diagnosis	Diagnosis Neolindus sibyllae Guzman, Tokareva & Żyła sp. nov. is similar to N. apiculus, but can be distinguished by the more broadly rounded posterior margin of tergite VIII and short, flat APS of aedeagus (pointed in N. apiculus) (Fig. 19 F – I; Herman 1991: figs 174 – 175).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	etymology	Etymology The name is given in honour of Maria Sibylla Merian, German entomologist, naturalist and scientific illustrator. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	materials_examined	Type material Holotype VENEZUELA • ♂; “ Neolindus sibyllae Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Táchira, San Cristobal, 8 km SE, P. N. [National Park] Chorro El Indio; 1425 m [m a. s. l.]; 7 ° 44 ’ 15 ’’ N, 72 ° 11 ’ 37 ’’ W; 29 May 1998; R. Anderson Leg.; VEN 1 A 98 050 D; ex: cloud forest litter \\ SM 0113752 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes VENEZUELA • 1 ♂; “ Neolindus sibyllae Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Táchira, San Cristobal, 8 km SE, P. N. Chorro El Indio, 1425 m; 7 ° 44 ’ 15 ’’ N, 72 ° 11 ’ 37 ’’ W; 28 May 1998; R. Anderson Leg.; VEN 1 A 98 047 A; ex: cloud forest litter \\ SM 0111014 KUNHM-ENT [barcode] ”; KUNHM-ENT • ♂; same data as for preceding; “ 29 May 1998; VEN 1 A 98 050 C \\ SM 0111081 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	description	Description MEASUREMENTS. BL (6.02), H (0.5, 0.7), A (1.41), a 1 (0.24, 0.06), a 2 (0.07, 0.06), a 3 (0.05, 0.05), a 4 (0.07, 0.05), a 5 (0.6, 0.06), a 6 (0.07, 0.07), a 7 (0.08, 0.08), a 8 (0.07, 0.06), a 9 (0.08, 0.08), a 10 (0.08, 0.07), NKW (0.42), GL (0.20), P (0.9, 0.9), E (0.7, 0.76), PC (0.54, 0.24), PF (0.66, 0.33), PT (0.36, 0.13), MSC (0.25, 0.16), MSF (0.62, 0.17), MST (0.5, 0.1), MTC (0.3, 0.2), MTF (0.5, 0.2), MTT (0.73, 0.1). COLOURATION. Head and pronotum brown, legs light brown, abdomen brown. HEAD. Head capsule wider than long; anterior and posterior margins rounded, posterior angles rather rounded; setation with 2 pairs of equal-sized PCS and PMS arranged in 2 pairs, directed upright. Epicranium with umbilicate and sparse micropunctuation, setation with 1 pair of OS, without FS and PFS. Gena smooth, with 2 OT; sockets of MS and POS umbilicate. Postgena with gular sutures reaching posterior margin of head joining neck pits; anterior part of gula with 1 seta near midline of gular suture and 2 close to posterior margin of gular sutures. Antenna widened towards tip, shorter than head and pronotum combined (Fig. 19 A); antennomeres 1 – 11 longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 equals to antennomeres 2 and 3 combined, antennomere 2 longer than 3. Labrum bilobed, with U-shaped emargination and 2 darker, thicker setae along ⅔ line of lobes; marginal clypeal seta light, nearly same size (Fig. 19 A; Supp. file 2). THORAX. Pronotum quadrate with umbilicate micropunctures and 2 paramedial and lateral longitudinal rows of setae. Basisternum of prosternum with well-developed longitudinal carina and smooth surface. Elytron wider than long, surface of elytra with umbilicate micropunctures in 10 – 11 moderately dense longitudinal rows (Fig. 19 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than mesotarsomere 2, mesotarsomeres 3 and 4 equal to mesotarsomere 2, mesotarsomere 5 longer than mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 as long as metatarsomere 2, metatarsomere 3 shorter than metatarsomere 1 and equal to metatarsomere 4, metatarsomere 5 longer than metatarsomere 1. ABDOMEN. Male: tergites III – VI with high density of micropunctuation; tergites VII and VIII with low density of micropunctuation, punctures randomly distributed. Posterior margin of tergite and sternite VII straight (Fig. 19 B – C). Posterior margin of tergite VIII truncate, with lateral angles rounded (Fig. 19 D). Posterior margin of sternite VIII with midlongitudinal emargination, lateral angles rounded (Fig. 19 E). Posterior margin of tergite IX divided; aedeagus, median lobe uniformly wide, apex of median lobe with deep median cavity with 2 pairs of elongate processes on each side; bottom of median lobe cavity sclerotised with thin spoon-like lamella tapering upwards from top of parameral side. pLS and pPMS difficult to identify. APS rather small, oval, shorter than elongate processes of median lobe (Fig. 19 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF9AFF90FE03597ADCDBFEAC.taxon	distribution	Distribution The species is known only from the type locality in Venezuela (Tachira, San Cristobal, Parque Nacional Chorro El Indio). It was collected in the cloud forest at higher altitude – 1425 m a. s. l.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	description	urn: lsid: zoobank. org: act: 040 BF 5 AB-B 5 AC- 4 EFB-BDA 4 - 1163885 E 5865 Fig. 20; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	diagnosis	Diagnosis Neolindus triangularis sp. nov. is similar to N. milleri and N. bicornis. nov. in the shape of the median lobe of the aedeagus. However, the aedeagus of N. triangularis differs from that of N. milleri in the structure and composition of apical sclerites (Herman 1991; fig. 189; Fig. 20 F – I) and has an acute base of the median lobe with a short median ridge on the parameral side, in contrast to the median lobe of the aedeagus in N. bicornis. Additionally, N. triangularis has noticeably wider fused pLS in the middle part, and a longitudinal invagination in sternite VIII continuing from the middle part of the emargination, limited by 2 ridges on the sides, which is absent in N. bicornis (Figs 4 E – I, 20 E – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	etymology	Etymology The name is derived from the Latin noun ‘ triangulus ’ meaning ‘ a triangle ”, and refers to the form of the median lobe of aedeagus. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus triangularis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Saül, 7 km N, 0.5 km ESE, Les Eaux Claires, Monts La Fumée; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W, 300 m [m a. s. l.]; 4 – 8 Jun 1997; J. Ashe, R. Brooks Leg.; FGAB 97 164; ex: flight interception trap \\ SMO 134030 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes FRENCH GUIANA • 1 ♀; “ Neolindus triangularis Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Matoury, 41.5 km SSW on Hwy N 2; 4 ° 37 ’ 22 ’’ N, 52 ° 22 ’ 35 ’’ W; 50 m; 26 - 28 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 060; ex: flight intercept trap \\ SM 0099449 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; “ Saül, 7 km N, Les Eaux Claires; 220 m; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 30 May – 04 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 144 \\ SM 0131297. KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	description	Description MEASUREMENTS. BL (7.9), H (0.43,1.06), A (1.43), A 1 (0.4, 0.1), A 2 (0.11, 0.07), A 3 (0.22, 0.08), A 4 (0.15, 0.09), A 5 (0.15, 0.09), A 6 (0.13, 0.09), A 7 (0.12, 0.09), A 8 (0.12, 0.1), NKW (0.42), GL (0.23), P (0.99, 1.22), E (1.56, 1.47), PC (0.63, 0.31), PF (0.96, 0.28), PT (0.67, 0.12), MSC (0.44, 0.41), MSF (1.1, 0.2), MST (0.76, 0.09), MTC (0.36, 0.31), MTF (1.24, 0.23), MTT (0.83, 0.1). COLOURATION. Head and pronotum brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded; posterior angles straight; setation with 1 pair of PCS and 1 row of PMS. Epicranium with umbilicate, low-density micropunctuation, setation with 1 pair of FS and 2 pairs of PFS, without OS. Gena with irregular depressions; setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 2 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula with 2 setae; posterior margin of head with 2 setae near gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 20 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 3 longer than 2. Labrum tetralobed; central lobes well-developed; central emargination shallow, V-shaped (Fig. 20 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate randomly distributed micropunctures. Metasternal intercoxal process with 1 pair of acute processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 10 – 11 longitudinal rows (Fig. 20 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; apical comb longitudinal, composed of widely separated, thin setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 longer than mesotarsomere 2, mesotarsomere 2 longer than mesotarsomere 3, mesotarsomere 3 as long as mesotarsomere 4, mesotarsomere 5 as long as mesotarsomere 2; inner ctenidium of metatibia longer than outer. Metatarsomere 1 longer than metatarsomere 2, metatarsomere 2 longer than metatarsomere 3, metatarsomere 3 as long as metatarsomere 4, metatarsomere 5 as long as 2. ABDOMEN. Tergites with low-density randomly distributed micropunctuation. Posterior margin of tergite VII straight (Fig. 20 B). Male: posterior margin of sternite VII with small short midside emargination (Fig. 20 C). Posterior margin of sternite VIII with V-shaped midlongitudinal emargination, lateral angles rounded (Fig. 20 D). Posterior margin of tergite VIII with median acute elongation, lateral angles slightly lobed (Fig. 20 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length; aedeagus, median lobe acute at base, widening apically and well-sclerotised; with small suture connecting basal acute tip of median lobe with elevated area around basal foramen. pLS fused into flattened arcuate sclerite widening in middle with 2 acute processes on each side protruding distad from median foramen and ending bent to anti-parameral side. APS flattened, arcuate, smaller than pLS, with 2 acute processes on each side protruding distad from median foramen and ending bent to parameral side shorter than processes of pLS. pPMS not visible (Fig. 20 F – I). Female: lack of emargination on posterior of sternite VII; posterior margin of tergite VIII sligthtly lobulated.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF97FF92FE655D97DB89FA7A.taxon	distribution	Distribution The species is known from the type locality in French Guyana (Saint-Laurent-du-Maroni, Saül, Monts La Fumée), the surrounding forest, and around 130 km SE from the type locality (Cayenne, Matoury). It was collected between 50 – 300 m a. s. l. using flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	description	urn: lsid: zoobank. org: act: 2 EEFAB 58 - D 70 D- 45 C 4 - AF 11 - 381017182864 Fig. 21; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	diagnosis	Diagnosis Neolindus tropicalis Guzman, Tokareva & Żyła sp. nov. has a very close structure of aedeagus to those of N. lodhii, but can be distinguished by the combination of the following characters: posterior margin of sternite VII with acute processes on each side of emargination (absent in N. lodhii), lobes of posterior margin of sternite VIII without longitudinal straight rows of short black setae (present in N. lodhii) and without short median carina between the lobes of sternite VIII (present in N. lodhii) (Herman 1991: figs 182, 187; Fig. 21 C, E).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	etymology	Etymology The name is derived from the Latin word ‘ tropicus ’, which refers to the tropics. It indicates that the species type locality is in tropical Brazil. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	materials_examined	Type material Holotype BRAZIL • ♂ (specimen lacks hind tarsi); “ Neolindus tropicalis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Amazonas, Manaus, Reserva Florestal Adolpho Ducke; 2 ° 57 ’ 47.4 ” S, 59 ° 55 ’ 21.2 ” W; 109 m [m a. s. l.]; 1995 – 1996 ”; NHMUK.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	description	Description MEASUREMENTS. BL (6), H (0.43, 0.71), A (1.19), A 1 (0.27, 0.71), A 2 (0.07, 0.06), A 3 (0.1, 0.06), A 4 (0.08, 0.06), A 5 (0.09, 0.06), A 6 (0.08, 0.07), A 7 (0.08, 0.07), A 8 (0.09, 0.08), A 9 (0.09, 0.07), A 10 (0.12, 0.08), A 11 (0.12, 0.08), NKW (0.32), GL (0.2), P (0.76, 0.82), E (1.05, 0.96), PC (0.33, 0.24), PF (0.4, 0.33), PT (0.47, 0.11), MSC (0.35, 0.25), MSF (0.75, 0.18), MST (0.49, 0.09), MTC (0.28, 0.23), MTF (0.801, 0.2), MTT (0.62, 0.09). COLOURATION. Head and pronotum dark brown; legs light brown; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin sinuate, bent dorsoventrally, slightly elevated; posterior margin rounded with emargination in front of neck; posterior angles straight; setation with 1 pair of PCS and 1 row of PMS. Epicranium shiny and smooth, without micropunctuation, setation with 1 pair of OS, and 1 pair of PFS, without FS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, MS, and POS; sockets of MS and POS umbilicate. Postgena with row of 3 setae in line from eye to gular suture; gular sutures not reaching posterior margin of head; anterior part of gula with 2 setae; posterior margin of head with 2 setae close to gular sutures. Neck with foramen magnum occupying ¾ of occiput. Antenna moniliform from antennomere 5, shorter than head and pronotum combined (Fig. 21 A); antennomeres longer than wide, tomentose pubescence starting from antennomere 4; antennomere 1 as long as 2 and 3 combined, antennomere 3 longer than 2. Labrum bilobed, with V-shaped emargination (Fig. 21 A; Supp. file 2). THORAX. Pronotum slightly wider than long, with umbilicate micropunctures and 3 longitudinal rows of seta, 1 median and 2 lateral. Metasternal intercoxal process with 1 pair of acute processes with light brown tips. Elytron longer than wide; surface of elytra with umbilicate micropunctures in 6 – 8 moderately dense longitudinal rows (Fig. 21 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae, apical comb with wider spaced and thicker longitudinal setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as 2, mesotarsomere 3 shorter than 1 and as long as 4, mesotarsomere 5 as long as 1; inner ctenidium of metatibia longer than outer. ABDOMEN. Male: tergites with low density of micropunctuation. Posterior margin of tergite VII straight (Fig. 21 B). Posterior margin of sternite VII with moderate deep emargination flanked by acute processes on each side (Fig. 21 C). Posterior margin of tergite VIII with single broad, triangular lobe with rounded tip (Fig. 21 D). Posterior margin of sternite VIII with midline broad emargination, of about ¼ of segment length, lateral angles obtuse (Fig. 21 E). Posterior margin of tergite IX with midlongitudinal deep emargination in ¾ of segment length (Fig. 21 A); aedeagus, in parameral view, median lobe with consistent width throughout, apex emarginate with rounded sclerotised processes on each side; pPMS conical, sclerotised, slightly bent towards each other, shorter than pLS and APS. pLS partially sclerotised, narrow, flattened; APS longer than pLS and pPMS, apically divided in 2 partially membranous arrowlike processes (Fig. 21 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF95FF93FE1559C6DB6DF809.taxon	distribution	Distribution The species is known only from the type locality in Brazil (Amazonas, Manaus, Reserva Florestal Adolpho Ducke). It was collected at a low altitude.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	description	urn: lsid: zoobank. org: act: D 8 A 5 DF 7 F- 1903 - 4 F 28 - B 5 AD- 49 E 6 F 04 A 16 E 6 Fig. 22; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	diagnosis	Diagnosis This species resembles N. parautriensis sp. nov. and N. maya sp. nov. in terms of habitus, but can be distinguished from them by the characters of the apical sclerites of the aedeagus: pPMS of N. utriensi s sp. nov. is straight, narrower than in N. maya and wider than in N. parautriensis, bearing apical arms of uneven lengths, bent inwards (Fig. 22 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	etymology	Etymology The species name is derived from the name of the Utría National Natural Park in Chocó, Colombia, which is the type locality. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	materials_examined	Type material Holotype COLOMBIA • ♂; “ Neolindus utriensis Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Chocó, Bahía Solano, PNN [National Natural Park] Utría Centro de Visitantes; 6 ° 1 ’ 0.00 ’’ N, 77 ° 20 ’ 0.00 ’’ W; 2 m [m. a. s. l]; Malaise; 15 – 30 Out 2000; J. Perez Leg.; M. 816 \\ SM 0549296 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes COLOMBIA • 1 ♀; “ Neolindus utriensis Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Chocó, Bahía Solano, PNN Utría Centro de Visitantes; 6 ° 1 ’ 0.00 ’’ N, 77 ° 20 ’ 0.00 ’’ W; 2 m; Malaise; 26 Dec 2000 – 01 Feb 2001; J. Perez Leg.; M. 1465 \\ SM 0549286 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ M. 1342 \\ SM 0549283 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	description	Description MEASUREMENTS. BL (12.50), H (0.75, 1.53), A (3.25), a 1 (0.53, 0.11), a 2 (0.20, 0.08), a 3 (0.42, 0.09), a 4 (0.28, 0.08), a 5 (0.29, 0.08), a 6 (0.29, 0.07), a 7 (0.28, 0.06), a 8 (0.28, 0.07), a 9 (0.25, 0.08), a 10 (0.21, 0.06), a 11 (0.21, 0.08), NKW (0.70), GL (0.35), P (1.45, 1.86), E (2.30, 1.80), PC (1.35, 0.65), PF (1.65, 0.75), PT (1.1, 0.3), MSC (0.9, 0.6); MSF (1.76, 0.5), MST (1.41, 0.15), MTC (0.73, 1.0), MTF (1.89, 0.37), MTT (1.83, 0.2). COLOURATION. Head and pronotum dark brown, legs brown, abdomen dark brown. HEAD. Head capsule wider than long; anterior margin straight; posterior margin straight with emargination in front of neck; posterior angles straight; setation with 1 PCS, PMS in dense straight row. Epicranium with dense micropunctuation, setation with 1 pair of FS, 2 pairs of PFS in row parallel to PMS, and without OS. Gena with irregular surface, with depression from mandibular base to posterior eye margin, and with 1 OT. Postgena with row of 3 setae in line from eye to gular suture, gular sutures not reaching posterior margin of head, anterior part of gula with irregular row of 6 – 8 setae. Antenna thin, longer than head and pronotum combined; antennomeres longer than wide; tomentose pubescence starting from antennomere 3; antennomere 1 shorter than antennomeres 2 and 3 combined; antennomere 3 longer than 2. Labrum bilobed, with shallow U-shaped emargination (Fig. 22 A; Supp. file 2). THORAX. Pronotum wider than long, almost quadrate (ratio 0.87), with umbilicate punctures and 2 paramedial rows of setae. Basisternum of prosternum triangular, with row of 8 setae on anterior margin, longitudinal ridge present, and surface wrinkled. Metasternal intercoxal process with 1 pair of acute spine-like processes. Elytron longer than wide, surface of elytra with moderately dense umbilicate punctures in 6 – 8 longitudinal rows (Fig. 22 A; Supp. file 2). LEGS. Protibia with 4 well-developed combs of setae; mesotibial apical ctenidia on both sides, inner longer than outer; mesotarsomere 1 as long as mesotarsomere 2; mesotarsomeres 3 and 4 shorter than mesotarsomere 1; mesotarsomere 5 longest; metacoxa wider than long; metatarsomere 1 as long as metatarsomeres 2 or 5; metatarsomeres 3 and 4 combined shorter than 5. ABDOMEN. Tergites with sparse micropunctures. Anterior margin of tergite VIII (base of tergite) with 4 internal canals, elevated, central canals forming inverted pentagon. Anterior margin of sternite VIII with 4 internal canals. Male: posterior margin of tergite VII straight (Fig. 22 B). Posterior margin of sternite VII with moderate emargination at middle, with acute processes on each side (Fig. 22 C). Posterior margin of tergite VIII with wide median process with narrow invaginations on sides and acute angles (Fig. 22 D). Posterior margin of sternite VIII bilobed, with narrow median emargination reaching up to ¼ of segment length, lobes rounded (Fig. 22 E); posterior margin of tergite IX with midlongitudinal deep invagination at ¾ of segment length (Asenjo 2011: fig. 12); aedeagus, median lobe broadened at base, apex with moderately deep emargination and rounded processes on each side. pLS thin, with net-like microsculpture, forming elongate flat processes equal in length to pPMS and slightly longer than APS. APS spoon-shaped, wider than pLS, with slightly emarginate apical edge. pPMS elongate, thin, with 2 acute apical arms bent inwards; arm closer to median foramen longer than arm closer to parameral side (Fig. 4 A – B). Female: posterior margin of sternite VII sinuate; sternite VIII bilobed, slightly emarginate (Fig. 22 F – I).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF92FF97FE095D05DBD0FBA5.taxon	distribution	Distribution This species is known only from the type locality in Colombia (Choco, Bahía Solano, Utría National Natural Park). It was collected at a low altitude by Malaise trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	description	urn: lsid: zoobank. org: act: 2 DDCFAA 0 - 665 A- 41 E 8 - 8 F 11 - 6 B 69 B 9 E 06 A 7 A Fig. 23; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	diagnosis	Diagnosis Neolindus volkeri Guzman, Tokareva & Żyła sp. nov. resembles N. schubarti and N. irmleri in aedeagus morphology. However, in N. volkeri, the median invagination on the parameral side of the median lobe is wider than in the other two species (Herman 1991: fig. 210; Asenjo 2011: fig. 2; Fig. 23 F – G). Unlike the two similar species, N. volkeri has a deep midline emargination with a convex median part on the posterior margin of sternite VII, with acute protruding processes on both sides (emargination shallow and without processes in both N. schubarti and N. irmleri) (Herman 1991; fig. 208; Asenjo 2011: fig. 4; Fig. 23 C). Additionally, the posterior margin of tergite VIII has deeper pair of emarginations surrounded by invaginated areas (shallower and without invagination in N. schubarti and N. irmleri) (Fig. 23 D; Herman 1991: fig. 206; Asenjo 2011: fig. 6).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	etymology	Etymology This species is dedicated to the great taxonomist, Volker Assing, who passed away prematurely. He was a renowned specialist in Palaearctic and Oriental Paederinae and described two species of Neolindus.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	materials_examined	Type material Holotype FRENCH GUIANA • ♂; “ Neolindus volkeri Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Roura, 8.4 Km SSE; 200 m [m a. s. l.]; 4 ° 40 ’ 41 ’’ N, 52 ° 13 ’ 25 ’’ W; 25 – 29 May 1997; J. Ashe, R Brooks Leg.; FG 1 AB 97 O 88; ex: flight intercept trap \\ SM 0133899 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes FRENCH GUIANA • 1 ♂; “ Neolindus volkeri Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ Roura, 8.4 lm SSE; 200 m; 4 ° 40 ’ 41 ’’ N, 52 ° 13 ’ 25 ’’ W; 25 – 29 May 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 O 88; ex: flight intercept trap \\ SM 0133900 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ SM 0131259 KUNHM-ENT [barcode] ”; KUNHM-ENT • 1 ♀; same data as for preceding; “ 26 May – 10 Jun 1997; FG 1 AB 97 182 \\ SM 0330895 KUNHM-ENT [barcode] \\ Neolindus sp. det. J. S. Ashe 1998 ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	description	Description MEASUREMENTS. BL (12.95), H (1.46, 1.82), A (4.04), a 1 (0.63, 0.12), a 2 (0.13, 0.09), a 3 (0.37, 0.09), a 4 (0.22, 0.07), a 5 (0.28, 0.09), a 6 (0.29, 0.08), a 7 (0.28, 0.08), a 8 (0.27, 0.07), a 9 (0.23, 0.08), a 10 (0.18, 0.08), a 11 (1.16, 0.82), NKW (0.62), GL (0.4), P (1.46, 1.82), E (2.22, 1.99), PC (0.6, 0.36), PF (1.01, 0.42), PT (0.62, 0.26), MSC (0.56, 0.54), MSF (1.54, 0.27), MST (0.86, 0.15), MTC (0.53, 0.44), MTF (1.69, 0.36), MTT (1.24, 0.17). COLOURATION. Head and pronotum dark brown; legs brown; abdomen dark brown. HEAD. Head capsule wider than long; anterior margin straight, posterior margin rounded, posterior angles rather rounded; setation with 1 pair of PCS and multiple PMS arranged in 1 row. Epicranium with umbilicate, highly dense micropunctuation, setation with 1 pair of FS and 1 pair of PFS, without OS. Gena with irregular surface, with depression from mandibular base to middle of eye, setation with 1 OT, 1 pair of MS, POS and PTOS; sockets of MS and POS concave. Postgena with row of 3 setae in line from eye to gular suture; gular sutures not reaching head posterior margin; anterior part of gula with irregular row of 6 – 8 setae. Neck with foramen magnum occupying ½ of occiput. Antenna thin and elongate, longer than head and pronotum combined, all antennomeres longer than wide, tomentose pubescence starting from antennomere 3, antennomere 1 as long as antennomere 2 and 3 combined, antennomere 3 longer than 2. Labrum tetralobed, central lobes well-developed, central emargination shallow, V-shaped, with 2 dark thick seta in median line of labrum lobes (Fig. 23 A; Supp. file 2). THORAX. Pronotum nearly quadrate, with umbilicate micropunctures and 2 distinct paramedial and lateral rows of setae. Basisternum of prosternum with 4 anterior setae and 1 posterior pair of setae. Metasternal intercoxal process with 1 pair of acute processes. Elytron longer than wide, surface of elytra with umbilicate micropunctures in 8 – 9 moderately dense longitudinal rows (Fig. 23 A; Supp. file 2). LEGS. Protibia with 5 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer; mesotarsomere 1 as long as mesotarsomere 2, mesotarsomere 3 shorter than mesotarsomere 1 and as long as mesotarsomere 4, mesotarsomere 5 as long as mesotarsomere 1; inner ctenidium of metatibia longer than outer; metatarsomere 1 longer than metatarsomere 2, metatarsomere 2 as long as metatarsomeres 3 and 4 combined, metatarsomere 3 longer than metatarsomere 4, metatarsomere 5 as long as metatarsomere 2. ABDOMEN. Tergites with low density, randomly placed micropunctures. Posterior margin of tergite VII straight (Fig. 23 B). Male: posterior margin of sternite VII with moderate emargination and acute processes on each side (Fig. 23 C). Posterior margin of tergite VIII trilobed, acute triangular lobe in middle and slightly acute-angled lateral lobes, each as long as central lobe (Fig. 23 D). Posterior margin of sternite VIII with shallow midlongitudinal emargination and rounded lateral angles (Fig. 23 E). Posterior margin of tergite IX with midlongitudinal deep emargination about ¾ of segment length; aedeagus, median lobe robust, strongly sclerotised, medially widened, apex with 2 sclerotised processes, deep midpoint emargination reaching middle of lobe, edges of emargination with brush of 5 – 7 short, dark setae; pLS small, oval, hidden under APS; APS oval, spoon-shape, covering pLS and pPMS. pPMS of complicated form, with several lamellas and arms, anchored with sclerotised ring under membrane (Fig. 23 F – I). Female: lack of emargination on posterior margin of sternite VII; posterior margion of tergite VIII slightly sinuated.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFF90FFEAFE075892DA94FE22.taxon	distribution	Distribution The species is only known from the type locality in French Guiana, (Cayenne, Roura). It was collected at a low altitude using a flight interception trap.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	description	urn: lsid: zoobank. org: act: F 2 E 11144 - A 406 - 413 E- 9 DBF- 465 E 1 B 0252 DB Fig. 24; Supp. file 2	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	diagnosis	Diagnosis Among species with two pairs of head trichobothria, N. yotokae Guzman, Tokareva & Żyła sp. nov. resembles N. paralellus with a wide midlongitudinal invagination in sternite VIII and the structure of aedeagus. However, in contrast to N. paralellus, this invagination is glabrous (without setation), and the concave deep emargination of the posterior margin of sternite VIII is covering the whole width and has acute lateral ends (Herman 1991: fig. 88; Fig. 24 E). Moreover, tergite VIII has a rounded posterior margin, compared to straight margin in N. paralellus (Herman 1991: fig. 87; Fig. 24 D), and the aedeagus of N. yotokae has shorter apical invagination on the parameral side compared to these structures in N. paralellus (Herman 1991: fig. 85; Fig. 24 F – G).	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	etymology	Etymology The name is dedicated to Dr Karla Yotoko, evolutionary biologist, and Camilo Guzman’s PhD advisor. An adjective.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	materials_examined	Type material Holotype VENEZUELA • ♂; “ Neolindus yotokae Guzman, Tokareva & Żyła 2024 HOLOTYPE [red label] \\ Aragua, [Mario Briceño Iragorry, Henri Pitttier National Park] Rancho Grande Biological Station; 10 ° 21 ’ 38 ’’ N, 67 ° 41 ’ 38 ’’ W; 1550 m [m a. s. l.]; 14 May 1998; R. Anderson Leg.; VEN 1 A 98 008 A; ex: cloud forest litter \\ SM 0129897 KUNHM-ENT [barcode] ”; KUNHM-ENT. Paratypes VENEZUELA • 1 ♂; “ Neolindus. yotokae Guzman, Tokareva & Żyła 2024 PARATYPE [yellow label] \\ same data as for holotype \\ 1550 m; 10 ° 21 ’ 38 ’’ N, 67 ° 41 ’ 38 ’’ W; 14 May 1998; VEN 1 A 98 007 E \\ [barcode] SM 0112571 KUNHM-ENT ”; KUNHM-ENT • 1 ♂; same data as for preceding; “ 1100 – 1300 m; 10 ° 21 ’ 32 ’’ N, 67 ° 40 ’ 46 ’’ W; 14 MAY 1998; J. Ashe, R. Anderson Leg.; VEN 1 ABH 98 009; ex: fungus log \\ SM 0334139 KUNHM-ENT [barcode] ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	description	Description MEASUREMENTS. BL (5.7), H (0.34, 0.5), A (0.77), a 1 (0.2, 0.06), a 2 (0.09, 0.05), a 3 (0.07, 0.05), a 4 (0.06, 0.05), a 5 (0.05, 0.05), a 6 (0.05, 0.06), a 7 (0.06, 0.06), a 8 (0.06, 0.07), a 9 (0.06, 0.07), a 10 (0.06, 0.08), a 11 (0.01, 0.08), NKW (0.33), GL (0.20), P (0.8, 0.7), E (0.06, 0.62), PC (0.32, 0.13), PF (0.57, 0.3), PT (0.3, 0.1), MSC (0.4, 0.25), MSF (0.6, 0.2), MST (0.4, 0.1), MTC (0.26, 0.21), MTF (0.38, 0.22), MTT (0.4, 0.07). COLOURATION. Head and pronotum brown; legs light brown; abdomen brown. HEAD. Head capsule wider than long; anterior margin sinuate, dorsoventrally deflexed, slightly elevated; posterior margin rounded with emargination in front of neck; posterior angles round; setation with 2 pairs of PCS and 1 row with 8 setae of PMS, central PMS wide apart, setae near neck longer than others. Epicranium with low-density micropunctuation, setation with 1 pair of OS; IOS and SAS displaced towards second and third parts of head. Gena with smooth surface, with depression from mandibular base to posterior margin of head, setation with 2 trichobothria, MS, 1 POS, and 1 PTOS between margin of eye and POT. Postgena with row of 3 setae in line from eye to gular suture; gular sutures reaching posterior margin of head joining neck pits; anterior part of gula without setae, with lateral depressions; midlength of gula with 2 setae; posterior margin of head with 2 setae close to gular sutures. Antenna moniliform from antennomere 3, shorter than head and pronotum combined; antennomeres 1 – 7 longer than wide, antennomeres 8 – 11 wider than long, tomentose pubescence starting from antennomere 5; antennomere 1 as long as antennomeres 2 and 3 combined, antennomere 2 longer than 3. Labrum bilobed, with wide quadrate emargination (Fig. 24 A, Supp. file 2). THORAX. Pronotum slightly longer than wide, with umbilicate randomly distributed micropunctures. Prosternal basisternum with longitudinal carina and smooth surface. Mesosternum with scutiform basisternum with scarce tuberculate micropunctuation. Metasternal intercoxal process with 1 pair of rounded processes. Elytron wider than long, shorter than pronotum; surface of elytra with umbilicate micropunctures in 4 – 6 moderately dense longitudinal rows (Fig. 24 A, Supp. file 2). LEGS. Protibia with 3 well-developed combs of setae; mesotibial apical ctenidium on both sides, inner longer than outer, outer spine-like; mesotarsomeres 1, 2 and 3 as long as mesotarsomere 5 as longer than mesotarsomeres 1, 2, 3 and 4 combined; inner ctenidium of metatibia longer than outer; metatarsomeres 1, 2, 3 and 4 subequal, metatarsomere 5 as long as metatarsomeres 1 – 3 combined. ABDOMEN. Male: tergites with moderate density of micropunctuation. Posterior margin of tergite VII straight (Fig. 24 B). Posterior margin of sternite VII sinuate, with moderately shallow midline emargination, area around emargination invaginated (Fig. 24 C, Supp. file 2). Posterior margin of tergite VIII elongate, rounded (Fig. 24 D). Posterior margin of sternite VIII with midline concave emargination as long as posterior margin width, with wide longitudinal glabrous stripe throughout entire segment, lateral angles rounded (Fig. 24 E, Supp. file 2). Posterior margin of tergite IX with midlongitudinal deep emargination in ⅓ of segment length, fused; aedeagus, in parameral view, median lobe of uniform width along sides, apex of median lobe with 2 lateral flattened rounded processes. pPMS and pLS absent. APS short, triangular, flattened, no longer than median lobe, partially covering median foramen (Figs. 24 F – I). Female: unknown.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEDFFEBFE035E2EDCD7F98B.taxon	distribution	Distribution The species is known from the type locality in Venezuela (Aragua, Rancho Grande Biological Station). It was collected in the cloud forest at high altitude (1550 m a. s. l.) by sifting leaf litter.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEAFFEDFDBA5D05DB6CFE74.taxon	materials_examined	BOLIVIA • 1 spec.; “ Cochabamba, Cochabamba, 67.5 km NE, Est. Biol. [Biological Estation] Valle del Sajita, Universidad de San Simon; 300 m; 17 ° 6 ’ 33 ’’ S, 64 ° 47 ’ 52 ’’ W; 7 FEB 1999; R. Anderson Leg.; BOL 1 A 99 027; Rain forest litter ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEAFFEDFD975DDFDC23FD69.taxon	materials_examined	COLOMBIA • 1 spec.; “ Cauca, PNN [Natural National Park] Gorgona, El Roble; 130 m; 2 ° 58 ’ 0.00 ’’ N, 78 ° 11 ’ 0.0 ’’ W; 9 - 27. VIII. 2001; H. Torres Leg.; M. 2124 ”; KUNHM-ENT. PANAMA • 1 spec.; “ Chiriqui, Prov. [Province] La Fortuna, Cont. Divide Trail; 1080 m; 08 ° 46 ’ 0.00 ’’ N, 82 ° 12 ’ 0.00 ’’ W; 21 May 1995; J. Ashe, R. Brooks Leg.; # 033 ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEAFFEDFDEF5ED1DA88FA0F.taxon	materials_examined	COSTA RICA • 1 spec.; “ Heredia, Puerto Viejo, Estacion Biologica La Selva - OET; 100 m; [10 ° 25 ’ 52.0 ” N, 84 ° 00 ’ 13.0 ” W]; I-II. 1993; P. Hanson Leg.; ex: Malaise ”; KUNHM-ENT • 1 spec.; “ Punta. [Puntarenas]; Prov. Rincon de Osa; 150 m; 08 ° 41 ’ 141 ’’ N ,, 83 ° 31 ’ 117 ’’ W; 23 - 26. VI. 2001; S. Peck, J. Peck Leg.; CR 1 P 01 006; 1 - 13, Flight Intercept Trap ”; KUNHM-ENT • 2 specs; same data as for preceding; “ CR 1 P 01 006; 1 - 14 ”; KUNHM-ENT • 2 specs; “ Puntarenas, Prov. Rincon de Osa; 50 m; 08 ° 41 ’ 141 ’’ N, 83 ° 31 ’ 117 ’’ W; 23 - 26. VI. 2001; S. Peck, J. Peck Leg.; CR 1 P 01 005; 1 - 13, Flight Intercept Trap ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ CR 1 P 01 005; 1 - 14 ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ 150 m, CR 1 P 01 006; 1 - 13 ”; KUNHM-ENT. [8 specimens.] PANAMA • 1 spec.; “ Panama, Colon, Parque Nac. Soberania, Pipeline Rd. Km 6; 40 m; 09 ° 07 ’ 0.00 ’’ N, 79 ° 45 ’ 0.00 ’’ W; 7 - 21. VI. 1995; J. Ashe, R. Brooks, Z. Fallin; # 265; ex: FIT [Flight Intercept Trap] ”; KUNHM-ENT • 1 spec.; “ Panama, Prov. Barro Colorado Island; 200 m; 9 ° 11 ’ 0.0 ” N, 79 ° 51 ’ 0.0 ” W; 7 July 2000; S. Chatzimanolis; PAN 1 C 00 046; ex: FIT ”; KUNHM-ENT • 3 specs; same data as for preceding; KUNHM-ENT • 1 spec.; same data as for preceding; “ 8 July 2000; PAN 1 C 00 052 ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ 17 – 23 July 2000 July 2000; PAN 1 C 00 087 ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ 40 m; 14 - 18 July 2000; PAN 1 C 00 07 ”; KUNHM-ENT • 2 specs; same data as for preceding; “ 100 m; 9 ° 11 ’ 0.0 ” N, 79 ° 51 ’ 0.0 ” W; 6. VIII. 1994; D. Banks Leg.; ex: Flight Intercept Trap ”; KUNHM-ENT • 1 spec.; “ Colon, 14 km N jct Escobal & Piña Rds; [100] m; [9 ° 14 ’ 33.2 ” N, 80 ° 02 ’ 11.0 ” W]; 02 - 11. VII. 1996; J. Ashe, R. Brooks Leg.; PAN 1 AB 96 181 B; ex: Flight Intercept Trap ”; KUNHM-ENT. [11 specimens.]	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFEAFFEEFD9C59FADC16FDE7.taxon	materials_examined	COSTA RICA • 2 specs; “ Alajuela, E. B. San Ramon, R. B. San Ramon, 27 km N, 8 km W; 850 - 950 m; 10 ° 13 ’ 30 ’’ N, 84 ° 35 ’ 30 ’’ W; 29 Jun - 6 Jul 1999; R. Anderson Leg.; CR 1 A 99 108 A; ex: Wet premontane forest ”; KUNHM-ENT • 1 spec.; “ Guanacaste, Guanacaste, Conservación Area, Maritza Field Station; 950 m; [10 ° 57 ’ 25.5 ” N, 85 ° 29 ’ 41.4 ” W]; 13 Feb 1996; R. Anderson Leg.; CR 1 A 96 010 A; ex: Drytropical wet forest trans, litter ”; KUNHM-ENT • 2 specs; “ Guanacaste, Patilla Biological Station; 610 m; 10 ° 59 ’ 22 ’’ N, 85 ° 25 ’ 33 ’’ W; 13 – 15 Jul 2000; J. Ashe, R. Brooks; CR 1 ABF 00 135; ex: FIT [Flight Intercept Trap] ”; KUNHM-ENT • 1 spec.; “ Guanacaste, Parque Nacional Guanacaste, Maritza Field Station; 950 m; 10 ° 57 ’ 25.5 ” N, 85 ° 29 ’ 41.4 ” W; 13 Feb 1996; R. Anderson Leg.; CR 1 A 96 010 A; Ex: Drytropical wet forest trans, litter ”; KUNHM-ENT • 1 spec.; “ Guanacaste, Santa Cecilia, Patilla Biological Station; 610 m; 10 ° 59 ’ 22 ’’ N, 85 ° 25 ’ 33 ’’ W; 13 - 15 Jun 2000; J. Ashe, R. Brooks Leg.; CR 1 ABF 00 135; ex: FIT ”; KUNHM-ENT • 3 specs; “ Punta. [Puntarenas], Rincon de Osa; 150 m; 08 ° 41 ’ 41 ’’ N, 83 ° 31 ’ 17 ’’ W; 23 - 26 Jun 2001; S. Peck, J. Peck Leg.; CR 1 P 01 006; 1 - 14, ex: FIT ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Las Cruces Biol. Sta., San Vito; 1425 m; 8 ° 46 ’ 59 ’’ N, 82 ° 59 ’ 18 ’’ W; 22 Jun 1998; R. Anderson Leg.; CR 1 A 98 108; Berlese leaf litter ”; KUNHM-ENT • 1 spec.; “ Prov. Rincon de Osa; 150 m; 08 ° 41 ’ 141 ’’ N, 83 ° 31 ’ 117 ’’ W; 23 – 26 Jun 2001; S. Peck, J. Peck; CR 1 P 01 006; 1 - 14, ex: FIT ”; KUNHM-ENT. [12 specimens.] PANAMA • 2 specs; “ Barro Colorado Island; 40 m; 9 ° 09 ’ 06.2 ” N, 79 ° 50 ’ 44.7 ” W; 25 – 30 Jun 2000; S. Chatzimanolis Leg.; PAN 1 C 00 033; ex: FIT ” • same data as for preceding; “ 30 Jun – 5 Jul 2000; PAN 1 C 00 039 ”; KUNHM-ENT • 1 spec.; “ Panama, Cerro Campa, nr Capira; 790 m; 08 ° 44 ’ 0.00 ’’ N, 79 ° 57 ’ 0.00 ’’ W; 18 May 1995; C. Chaboo Leg. ”; KUNHM-ENT. [3 specimens.]	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE9FFEEFD905E53DC40FC9C.taxon	materials_examined	COSTA RICA • 1 spec.; “ Puntarenas, Osa Penn., Fundacion Neotrop. 10 km W Rincon; 20 m; 8 ° 42 ’ 30 ’’ N, 83 ° 31 ’ 30 ’’ W; 23 Jun 1997; R. Anderson Leg.; CR 1 A 97 028 B; ex: Berlese forest litter ”; KUNHM-ENT. PANAMA • 1 spec.; “ Chiriqui ,, 20.4 km N San Felix; 950 m; 8 ° 22 ’ 0.00 ’’ N, 81 ° 46 ’ 0.00 ’’ W; 8 Jun 1995; R. Anderson Leg.; PAN 2 A 95 09 A; ex: Berlese forest litter ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE9FFEEFD8E5F64DA58FB4C.taxon	materials_examined	COSTA RICA • 1 spec.; “ Puntarenas, Rancho Quemado, Rincon de la Osa; 150 m; 08 ° 41 ’ 41 ’’ N, 83 ° 31 ’ 17 ’’ W; 23 – 26 Jun 2001; S. Peck, J. Peck Leg.; CR 1 P 01 006; 1 - 14, esx: FIT ”; KUNHM-ENT. FRENCH GUIANA • 1 spec.; “ Saint-Laurent-du-Maroni, Saul, 7 km N, 0.5 km ESE Les Eaux Claires, Mt. La Fumee; 220 m; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 31 May – 3 Jun 1997; J. Ashe, R. Brooks; FG 1 AB 97 124; Malaise • 1 spec.; same data as for preceding; “ 4 – 8 Jun 1997; FG 1 AB 97 164 ”; flight intercept trap; KUNHM-ENT. [2 specimens.]	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE9FFEEFD8458B7DAAFFA5F.taxon	materials_examined	FRENCH GUIANA • 1 spec.; “ Saûl, 7 km N, 0.5 km ES Les Eaux Claires, Mt. La Fumee, 3 ° 39 ’ 46 ” N, 53 ° 13 ’ 19 ” W, 300 m 4 – 8 JUN 1997; J. Ashe, R. Brooks Leg.: FG 1 AB 97 164 ex: flight intercept trap ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ 220 m 31 MAY- 3 JUN 1997; FG 1 AB 97 124; ex: malaise trap ”; KUNHM-ENT. [2 specimens.]	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE9FFEFFD8759AADB2CFE99.taxon	materials_examined	FRENCH GUIANA • 1 spec.; “ Roura, 18.4 Km SSE; 240 m; 4 ° 36 ’ 38 ’’ N, 52 ° 13 ’ 25 ’’ W; 29 May – 10 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 180; ex: FIT [Flight Intercept Trap] ”; KUNHM-ENT • 1 spec.; “ Cayenne, 33.5 km S and 8.4 km NW of Hwy N 2 On Hwy D 5; 30 m; 4 ° 48 ’ 18 ’’ N, 52 ° 28 ’ 41 ’’ W; 29 May – 9 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 A 97 171; ex: FIT. ”; KUNHM-ENT • 1 spec.; “ Cayenne, Matoury, 41.5 km SSW or HWY N 2; 50 m; 4 ° 37 ’ 22 ’’ N, 52 ° 22 ’ 35 ’’ W; 29 May – 9 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 170; ex: FIT ”; KUNHM-ENT • 2 specs; “ Saul, 7 km N, 0.5 km ESE Les Eaux Claires, Mt. La Fumee; 300 m; 3 ° 39 ’ 46 ’’ N, 53 ° 13 ’ 19 ’’ W; 4 - 8. VI. 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 164; ex: FIT ”; KUNHM-ENT. [5 specimens.] SURINAME • 2 specs; “ Brokopondo, Brownsberg Nature [Reserve], Witi Creek Trail; 420 m; 4 ° 56 ’ 55 ’’ N, 55 ° 10 ’ 53 ’’ W; 22 – 23 Jun 1999; Z. H. Falin, H. Hiwat Leg.; SUR 1 F 99 097; ex: Flight Intercept Trap ”; KUNHM-ENT • 1 spec.; “ Marowijne, Paloemeu; 160 m; 3 ° 20 ’ 56 ’’ N, 55 ° 26 ’ 18 W; 5 – 9 Jun 1999; D. Konoe Leg.; SUR 1 F 99 185; ex: FIT ”; KUNHM-ENT • 1 spec.; “ Saramacca, West Suriname Road; 30 m; 5 ° 13 ’ 37 ’’ N, 55 ° 52 ’ 54 ’’ W; 12 – 14 Jun 1999; Z. H. FaliN, B. DeDijin Leg.; ex: FIT ”; KUNHM-ENT • 2 specs; “ Marowijne, Paloemeu; 160 m; 3 ° 20 ’ 56 ’’ N, 55 ° 26 ’ 18 W; 5 – 9 Jun 1999; D. Konoe Leg.; SUR 1 F 99 185; ex: FIT ”; KUNHM-ENT. [6 specimens.]	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE8FFEFFDE95D60DCB4FDAA.taxon	materials_examined	COSTA RICA • 1 spec.; Heredia, Puerto Viejo, La Selva Biol. Res. Sta. [Biological Research Station] 3 km S; 100 m; 10 ° 25 ’ 52.0 ” N, 84 ° 00 ’ 13.0 ” W; 2 – 15. Jun 1996; R. Hanley Leg.; CR 1 H 96 016; ex: flight intercept trap ”; KUNHM-ENT. FRENCH GUIANA • 1 spec.; “ Roura, 18.4 km SSE; 240 m; 4 ° 36 ’ 38 ’’ N, 52 ° 13 ’ 25 ’’ W; 29 May 10 Jun 1997; J. Ashe, R. Brooks Leg.; FG 1 AB 97 180; ex: flight intercept trap ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE8FFEFFDF75E96DC42F981.taxon	materials_examined	COSTA RICA • 1 spec.; “ Heredia, Puerto Viejo, OTS-La Selva; 100 m; 10 ° 25 ’ 52.0 ” N, 84 ° 00 ’ 13.0 ” W; II-III. 1993; P. Hanson Leg.; ex: Malaise ”; KUNHM-ENT • 1 spec.; “ Heredia, 3 km S Puerto Viejo, OTS-La Selva; 100 m; [10 ° 25 ’ 52.0 ” N, 84 ° 00 ’ 13.0 ” W]; II-III. 1993; P. Hanson Leg.; ex: Malaise ”; KUNHM-ENT • 1 spec.; “ Heredia, Puerto Viejo, La Selva Biol. Res. Sta. 3 km S; [100 m]; 10 ° 26 ’ 00.0 ” N, 84 ° 01 ’ 00.0 ” W; 21. I- 3. II. 1991; J. S. Noyes Leg.; ex: Malaise ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Alturas, 2 km NE, Estac. Biol. Las Alturas; 1520 m; 8 ° 56 ’ 56 ” N, 82 ° 50 ’ 1 ” W; 20 Jun 1998; R. Anderson Leg.; CR 1 A 98 104; ex: Berlese leaf litter ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Sirena, Corcovado National Park, Sirena Stn, Upper Ollas Trail; 140 m; 8 ° 29 ’ 7 ’’ N, 83 ° 34 ’ 39 ’’ W; 24 – 28 Jun 2000; Z. H. Falin Leg.; CR 1 ABF 00 036; ex: Flight Intercept Trap ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Prov. Rincon de Osa; 50 m; 08 ° 41 ’ 141 ’’ N, 83 ° 31 ’ 117 ’’ W; 23 – 26 Jun 2001; S. Peck, J. Peck Leg.; CR 1 P 01 005; ex: 1 - 13, Flight Intercept Trap ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Estacion Biologica Monteverde; 1540 m; 10 ° 19 ’ 40.0 ” N, 84 ° 49 ’ 08.0 ” W; 12 Jun 2001; R. Anderson Leg.; CR 1 A 01 108; ex: Montane Forest Litter ”; KUNHM-ENT • 1 spec.; “ Puntarenas, Bahia Chal, Reserva Forestal Golfo Dulce, 5 km W Piedras Blancas; 100 m; [8 ° 45 ’ 34.7 ” N, 83 ° 28 ’ 54.8 ” W]; Feb – Mar. 1993; Leg. P. Hanson; ex: Malaise ”; KUNHM-ENT • 1 spec.; “ San Jose, San Antonio de Escazu; 1300 m; 9 ° 39 ’ 16 ’’ N, 84 ° 08 ’ 16 ’’ W; 1 - 30. IX. 1998; W. Eberhard, P. Hanson Leg.; CR 1 EH 99 04 ”; KUNHM-ENT • 1 spec.; same data as for preceding; “ CR 1 EH 99 02 ”; KUNHM-ENT • 1 spec.; “ Guanacaste, Parque Nacional Guanacaste, Maritza Field Station; 950 m; [10 ° 57 ’ 25.5 ” N, 85 ° 29 ’ 41.4 ” W]; 13 Feb 1996; R. Anderson Leg.; CR 1 A 96 010 A; ex: Dry tropical - Wet forest trans. litter ”; KUNHM-ENT. [11 specimens.] NICARAGUA • 1 spec.; “ Rio San Juan Dept., 8 km SE, El Castillo, Refugio Bartola; 80 m; 10 ° 58 ’ 06.0 ” N, 84 ° 20 ’ 04.0 ” W; 23 – 31 May 2002; S. Peck Leg.; NIC 1 P 02 002; ex: Flight Intercept Trap ”; KUNHM-ENT. PANAMA • 1 spec.; “ Chiriqui, La Fortuna; 1200 m; 8 ° 46 ’ 0.00 ’’ N, 82 ° 12 ’ 0.00 ’’ W; 9 Jun 1995; R. Anderson Leg.; PAN 2 A 95 10 C; ex: Berlese forest litter ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE8FFEFFD8F5A89D882F870.taxon	materials_examined	BOLIVIA • 1 spec.; “ Cochabamba, Cochabamba, 67.5 km NE Estacion Biologica Valle del Sajita, Universidad de San Simon; 300 m; 17 ° 6 ’ 33 ’’ S, 64 ° 47 ’ 52 ’’ W; 7 – 9 Feb 1999; F. Genier Leg.; BOL 1 G 99 044; ex: Dung Trap 2 ”; KUNHM-ENT. COSTA RICA • 1 spec.; “ Heredia, Puerto Viejo, La Selva Biol. Res. Sta. 3 km S; 80 m; 10 ° 25 ’ 0.00 ’’ N, N, 84 ° 0 ’ 0.00 ’’ W; 2 – 15 Jun 1996; R. Hanley Leg.; CR 1 H 96 016; ex: Flight Intercept Trap ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
03EC292BFFE7FFE0FD975D05DA4BFE74.taxon	materials_examined	PERU • 1 spec.; “ Madre de Dios, Parque Nacional del Manu, Pakitza Bio. Stn. Casianal Trail, Reserved Zone; 317 m; 11 ° 56 ’ 41 ’’ S, 71 ° 17 ’ 0.00 ’’ W; 15 – 16 Out 2000; R. Brooks Leg.; PERU 1 B 00 013; ex: Flight Intercept Trap ”; KUNHM-ENT.	en	Guzman, Yoan Camilo, Tokareva, Alexandra, Koszela, Katarzyna, Żyła, Dagmara (2024): Twenty-one new species of the Neotropical rove beetle genus Neolindus Scheerpeltz (Coleoptera, Staphylinidae, Paederinae). European Journal of Taxonomy 942: 1-76, DOI: 10.5852/ejt.2024.942.2581, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2581/11745
