taxonID	type	description	language	source
03EF6F19E120B3751B5EFE7FFBB1FC11.taxon	description	(FIG. 9 C, I, S) Type species: Selasia rhipiceroides Laporte, 1838; by monotypy. Diagnosis: Selasia as defined here can be recognized by the following combination of characters: frontoclypeal region short and wide; labrum transverse, more than four times wider than long, sometimes membranous; antenna flabellate; pronotum transverse, 1.60 – 1.90 times as wide as long (about 1.35 times in S. socotrana), with posterior margin shallowly and widely arcuately emarginate (Fig. 9 C), lateral carina almost complete, sublateral carina usually missing; prosternum (Fig. 9 C) transverse, with prosternal process reduced or short and narrow; scutellar shield longer than wide; mesoventrite (Fig. 9 I) with anterior margin slightly concave, without distinct margin; seven free abdominal ventrites (Fig. 9 S); and sternite IX basally emarginate. Species included: Altogether 64 species. Selasia apicalis Pic, 1914, S. arabica Geisthardt, 2003, S. atriventris Pic, 1914, S. auberti Pic, 1921, S. basalis Gorham, 1895, S. boruckae Kundrata, 2012, S. decipiens Guérin-Méneville, 1829, S. dembickyi Kundrata & Sormova, 2018, S. erlangeri Geisthardt, 2007, S. etoshaensis Wittmer, 1989, S. fulva Gorham, 1883, S. grandis Pic, 1914, S. incognita Geisthardt, 2007, S. jenisi Kundrata & Sormova, 2018, S. kilimana Kolbe, 1897, S. lata Pic, 1954, S. laticeps Pascoë, 1887, S. merkli Kundrata, 2012, S. minuta Kolbe, 1897, S. nigrobrunnea Kundrata, 2017, S. oberprieleri Wittmer, 1989, S. obscura Wittmer, 1962, S. pectoralis Pic, 1946, S. pulchella Gestro, 1878, S. raffrayi Pic, 1921, S. rhipiceroides Laporte, 1836, S. robecchii Gestro, 1892, S. sabatinellii Kundrata, 2017, S. socotrana Kundrata, 2012, S. striata (Pic, 1929), S. variabilis Wittmer, 1989, S. venusta Olivier, 1914, S. widenmanni Kolbe, 1897. The type material of S. asperulipennis Fairmaire, 1893 and S. isabellae Bourgeois, 1909 is most probably lost and their identity cannot be confirmed from the brief original descriptions. The following species currently placed in this genus differ morphologically from the here defined Selasia, but are not yet placed elsewhere: Selasia angustior Pic, 1954, S. atricornis Pic, 1946, S. basipennis Pic, 1913, S. bicolor Wittmer, 1953, S. castanea Wittmer, 1989, S. chaboti Pic, 1921, S. curtipennis Pic, 1918, S. diversicornis Pic, 1929, S. dumosa Geisthardt, 2007, S. endroedyi Wittmer, 1989, S. fuscula (Boheman, 1851), S. homhilia Geisthardt, 2003, S. incostata Wittmer, 1989, S. intermedia Wittmer, 1997, S. maynei Pic, 1931, S. murtulai Pic, 1930, S. nigricollis Wittmer, 1989, S. nigripennis Pic, 1925, S. pectinata Wittmer, 1989, S. posticalis Pic, 1931, S. pseudopectinata Geisthardt, 2007, S. ruficolor Pic, 1946, S. sibutensis Pic, 1914, S. testaceicolor Pic, 1914, S. transkeiensis Wittmer, 1989, S. transvaalensis Wittmer, 1989; S. unicolor Guérin-Méneville, 1829, S. wittmeri Geisthardt, 2007, S. zernyi Hicker, 1944. The systematic placement of these species will be examined in detail in future studies. Relationships: Selasia arabica from Yemen and Selasia sp. from Nigeria form an unsupported clade sister to S. erlangeri from Kenya in all but one analysis (Fig. 1; Supporting Information, Fig. S 1). Distribution: Afrotropical Region, southern part of the Arabian Peninsula, Socotra, Pakistan, Nepal, India, Sri Lanka, Thailand.	en	Kundrata, Robin, Bocak, Ladislav (2019): Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413-452
03EF6F19E123B3761979FD68FBE3FCB6.taxon	materials_examined	Type species: Kupeselasia minuta Kundrata & Bocak, 2017; by original designation.	en	Kundrata, Robin, Bocak, Ladislav (2019): Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413-452
03EF6F19E123B3761AFCF9B4FB28FD07.taxon	materials_examined	Type species: Lolosia transversalis Kundrata & Bocak, 2017; by original designation. Species included: Three described species: Lolosia gajduskovae Kundrata, 2018, L. smetkovae Kundrata, 2018, L. transversalis. Diagnosis: Lolosia is recognized by the following combination of characters: body minute, with smooth, glabrous surface; frontal carina incomplete; frontoclypeal region wide; eyes large, their frontal separation 1.20 times eye diameter; terminal maxillary and labial palpomere narrowed, subacute apically; antenna serrate; pronotum transverse, 1.85 times wider than long, lateral carina almost complete, sublateral carinae distinct, sinuate, almost reaching anterior margin; scutellar shield about as long as wide; abdominal ventrites free; sternite IX elongate, 2.1 times longer than wide; phallobase V-shaped; paramere relatively long, apically rounded (for more details see: Kundrata & Bocak, 2017). Distribution: Cameroon, Central African Republic, Democratic Republic of the Congo.	en	Kundrata, Robin, Bocak, Ladislav (2019): Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413-452
03EF6F19E122B37A19E6FE21FD55FCF6.taxon	description	(FIG. 9 E, K, U) Type species: Drilus flavescens (Geoffroy, 1785); by monotypy. Species included: Altogether 45 described and several undescribed species: Drilus adustus Chevrolat, 1854, D. akbesianus (Fairmaire, 1895), D. amabilis Schaufuss, 1867, D. attenuatus Pic, 1914, D. badius Kobieluszova & Kundrata, 2015, D. baenai Kundrata et al., 2015, D. bleusei (Olivier, 1913), D. bicolor Schaufuss, 1867, D. concolor Ahrens, 1812, D. creticus Pic, 1905, D. distincticollis Pic, 1907, D. flavescens (Geoffroy, 1785), D. frontalis Schaufuss, 1867, D. fulvicollis Audouin, 1824, D. fulvicornis Kiesenwetter, 1859, D. fulvitarsis Baudi di Selve, 1872, D. funebris Reitter, 1884, D. horasfakionus Kundrata et al. 2015, D. huijbregtsi Kobieluszova & Kundrata, 2015, D. humeralis Pic, 1931, D. iljini Barovskij, 1922, D. iranicus Wittmer, 1967, D. latithorax Pic, 1902, D. longulus Kiesenwetter, 1859, D. mauritanicus Lucas, 1842, D. mertliki Kobieluszova & Kundrata, 2015, D. nemethi Kundrata et al., 2014, D. novoathonius Sumakow, 1903, D. obscuricornis Pic, 1899, D. posticus Schaufuss, 1867, D. rectus Schaufuss, 1867, D. rittneri Petrzelkova & Kundrata, 2015, D. robustus Kobieluszova & Kundrata, 2015, D. rufipes (Baudi di Selve, 1872), D. sanliurfensis Kobieluszova & Kundrata, 2015, D. schwarzi Reitter, 1891, D. subparallelus Pic, 1934, D. testaceipes Pic, 1933, D. teunisseni Kobieluszova & Kundrata, 2015, D. turcicus Kobieluszova & Kundrata, 2015. The following five species described from the Afrotropical Region have different morphology from the here defined Drilus and will be revised in the near future: D. atripennis Pic, 1934, D. basilewskyi Wittmer, 1962, D. impressiceps Pic, 1913, D. ramosus Fairmaire, 1883, D. testaceipennis Pic, 1918. Diagnosis: Drilus is recognized by the following combination of characters: dorsal body surface uneven, usually matt; eyes relatively small to medium-sized, their frontal separation 1.60 – 3.00 times eye diameter; antenna slightly serrate to weakly flabellate; pronotum transverse, 1.20 – 1.70 times as wide as long, with lateral sides sinuate or rounded, lateral carina almost complete, and posterior margin usually only shallowly arcuately emarginate; prosternum (Fig. 9 E) with reduced prosternal process; mesoventrite (Fig. 9 K) V-shaped, with only indistinctly defined or shallow mesoventral cavity; surface of elytra usually without distinct striae or lines of punctures; abdominal ventrites free (Fig. 9 U), sternite IX and tergite X not apparently elongate, with more or less rounded sides, usually not more than 1.80 times as long as wide; paramere without latero-apical projection. Relationships: The phylogenetic relationships among the Drilus lineages are only partly resolved based on the six-marker phylogeny (Fig. 1; Supporting Information, Fig. S 1). The presence of the monophyletic group informally known as the ‘ Greek clade’, which is formed by the elongate, usually tiny species with the serrate antennae that are specialized predators of Albinaria snails and are distributed in the Peloponnese, the Ionian islands and Crete, is consistent with the results of the cox 1 analysis by Kundrata et al. (2015 a). Its statistically strongly supported sister group relationship with D. rufipes (Baudi di Selve) from Cyprus agrees with the results of the three-marker analysis by Sormova et al. (2018). The remaining species, which include usually more robust and hairy representatives with weakly flabellate antennae and a strongly transverse pronotum, formed a gradual branching in previous studies (Kundrata et al., 2015 a; Sormova et al., 2018), but they are found to be monophyletic in this study, although with low support (Fig. 1; Supporting Information, Fig. S 1). This clade included two branches, i. e. D. fulvicornis + (D. flavescens + D. mauritanicus) and D. concolor + (D. rectus + D. cf. mertliki). More species in the analysis in combination with a detailed morphological investigation of this genus are needed for a better understanding of Drilus phylogenetic interrelationships. Distribution: Northern Africa (Algeria, Morocco), Europe, Asia Minor, Levant, Caucasus, Iran. Species from tropical Africa do not belong to the genus (see comment above). GENUS MALACODRILUS KUNDRATA & BOCAK,	en	Kundrata, Robin, Bocak, Ladislav (2019): Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413-452
03EF6F19E129B37C1A9FFCD1FC44FE5C.taxon	description	(FIG. 9 F, L, W) Type species: Malacogaster passerinii Bassi, 1834; by monotypy. Species included: Eleven species: Malacogaster bassii Lucas, 1870, M. holomelas Peyerimhoff, 1949, M. maculiventris Reitter, 1894, M. nigripes Schaufuss, 1867, M. olcesei Pic, 1951, M. parallelocollis Reitter, 1894, M. passerinii Bassi, 1833, M. rubripes Peyerimhoff, 1949, M. rutllanti Alcaide, 1946, M. theryi Pic, 1951, M. tilloides Wollaston, 1864. Diagnosis: Malacogaster is recognized by the combination of the following characters: frontoclypeal region short and wide; eyes small, their frontal separation 2.20 – 2.65 times eye diameter; labrum large, subtrapezoidal; mandible with only small tooth medially at incisor; terminal maxillary and labial palpomeres short, gradually narrowed toward apex, apically obliquely truncate; antenna serrate; pronotum subquadrate to subtrapezoidal, about 1.10 – 1.40 times as wide as long, with lateral sides almost straight, usually gradually widened towards posterior margin, posterior margin with small arcuate median emargination; lateral carina short, reaching usually no more than half of pronotal length; prosternum (Fig. 9 F) with reduced prosternal process; mesoventrite (Fig. 9 L) V-shaped, with only indistinctly defined or shallow mesoventral cavity; elytra usually relatively short, dehiscent, with abdomen surpassing the tip of elytra; surface of elytra uneven, without striae or lines of punctures; abdominal ventrites free (Fig. 9 W); sternite IX and tergite X typically elongate, about or more than twice as long as wide; paramere with latero-apical projection on inner side. D i s t r i b u t i o n: S p a i n (C a n a r y I s l a n d s, I b e r i a n Peninsula, Balearic Islands), Italy (Sardinia, Sicily, Italian Peninsula), northern Africa (Morocco, Algeria, Tunisia, Libya).	en	Kundrata, Robin, Bocak, Ladislav (2019): Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413-452
