taxonID	type	description	language	source
B6CFF4826AFBD9EDBC0CB9FF5F44FA83.taxon	diagnosis	Diagnosis. Length 6.3 - 6.5 mm, width 2.5 mm. Head and prothorax red; elytra black or sometimes bluish black. Labrum (Fig. 1) semicircular in front, shortly depressed towards base, front pair of setae small and closely placed along the front margin, intermediate one at a distance from margin; ligula rectangular, with a wide and deep median impression, a pair of setae at a distance form apex depressed at base; apex truncate, outer apical angles rounded; mentum (Fig. 9) glabrous at base, both tooth and lobes elongate, slender and sharp at apex; tooth almost as long as lobes, two pairs of setae on base; lobes sinuate at a distance from apex along outer margin; maxillary palpi (Fig. 17) not dilated. Elytra without spots. Female genitalia. Gonocoxite short, stout, inner margin arcuate and not sinuate, apex short and sharp (Fig. 37).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
B6CFF4826AFBD9EDBC0CB9FF5F44FA83.taxon	distribution	Distribution. India.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
18642328DCEE29508CC7DA5469BADE1E.taxon	diagnosis	Diagnosis. Length 14.5 - 15.0 mm, width 5.0 - 5.5 mm. Black. Labrum (Fig. 2) with front margin rounded and bisinuate, three pairs of setae close to margin, and front two pairs on sinuated area; mandibles slightly obtuse at apex; mentum (Fig. 10) irregularly setose in basal half, both tooth and lobes elongate and sharp at apex, tooth slightly shorter than lobes; maxillary palpomere 4 (Fig. 18) cylindrical and gradually dilated toward apex, rounded and obliquely truncate at apex. Elytra without spots. Male genitalia. Median lobe dilated on dorsal side in middle partion, sinuate on ventral side; apical lamella elongate, not sinuate near apex, rounded at apex (Figs 25 - 26). Female genitalia. Gonocoxite slender, five setae on dorsal surface, sharp at apex (Fig. 38).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
18642328DCEE29508CC7DA5469BADE1E.taxon	distribution	Distribution. India (East India) and Indonesia (Java).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
D2B1C261B647E4CE3DE1A1272ADBDA99.taxon	diagnosis	Diagnosis. Length 24.7 mm, width 8.3 mm. Labrum (Fig. 3) elongate, three pairs of equidistant setae on upper surface near margin; ligula thickened, apex slightly narrowed, slightly emarginate in middle of front margin, deeply depressed near apex to form a median channel, with five pairs of setae along sides, hollowed out above with a median carina; mentum (Fig. 11) with both tooth and lobes stout, tooth shorter than lobes, contracted after middle, irregularly setose on ventral surface on basal half; maxillary palpomere 4 (Fig. 19) dilated, densely setose; labial palpomere 4 dilated, palpomere 3 not dilated inwards. Elytal spots nearly square, large, near the middle, cover intervals 2 - 7. Male genitalia. Median lobe of aedeagus strongly dilated and stout, strongly sinuate near paramere, straight towards apex, apical lamera elongate and narrowed at apex (Figs 27 - 28).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
D2B1C261B647E4CE3DE1A1272ADBDA99.taxon	distribution	Distribution. Myanmar.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
7E0E3BFCD8260C9B7D3852414329D2DC.taxon	diagnosis	Diagnosis. Length 16.0 - 16.3 mm, width 6.0 - 6.2 mm. Labrum (Fig. 4) convex, with a short furrow on each side of base, narrowed and pointed in front, with three pairs of setae away from the margin; ligula thickened, apex truncate towards sides, apical margin slightly emarginate, deeply depressed towards base, with a small median impression near apex; mentum (Fig. 12) glabrous at base; median tooth shorter than lobes, rather narrow and with apex rather pointed, two pairs of setae at basal area; lateral lobes obtuse at apex. Palpi (Fig. 20) not dilated. Elytral spots transverse, close to the middle of interval 5, covering intervals 3 - 7. Male genitalia. Median lobe stout, apical lamella short and round at apex (Figs 29 - 30). Female genitalia. Gonocoxite subapically dilated, three setae on dorsal surface, sharp at apex (Fig. 39).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
7E0E3BFCD8260C9B7D3852414329D2DC.taxon	distribution	Distribution. India (Nilgiri Hills, Malabar, Madras and Bombay) and Sri Lanka.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
91B688B2A97938D74EA03FE7D3B12274.taxon	distribution	Distribution. China, Vietnam, Laos, Myanmar, Indonesia and India.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
BA06BBD2C9B8E8B9ABEEB6CB0537FDE8.taxon	diagnosis	Diagnosis. Length 14.0 mm, width 4.8 mm. Labrum (Fig. 5) with apex rounded and pointed, front setae on apical margin; mandibles covered by the labrum, sharp at apex; mentum (Fig. 13) glabrous at base, median tooth nearly as long as lobes, about five setae at basal half; maxillary palpomere 4 (Fig. 21) cylindrically dilated, labial palpomere 4 flat and dilated, labial palpomere 3 dilated inwards; ligula wide and rectangular, apical margin straight, with a wide median impression beneath apex, a seta on either side near apex. Elytral spots large and oblong, covering intervals 3 - 7 in the middle. Female genitalia. Gonocoxite elongate, three setae on dorsal margin, sparsely setose on ventral surface; apex sharp (Fig. 40).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
BA06BBD2C9B8E8B9ABEEB6CB0537FDE8.taxon	distribution	Distribution. Indonesia (Sumatra).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
F56D815FA88F58569CC027DB91EF1BAD.taxon	diagnosis	Diagnosis. Length 10.0 mm, width 3.0 mm. Labrum large, semicircular anteriorly, with a flat depression on each sides, setae running along the sides in two small furrows; palpi short and stout, truncate at apex; mentum with a long, narrow, sharp median tooth, lateral lobes a little longer. Elytral spots oblong and covers intervals 2 - 7. The above description is after Louwerens (1949).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
F56D815FA88F58569CC027DB91EF1BAD.taxon	distribution	Distribution. Indonesia (Java).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
708FF3F174B0C919051F96B142E12AB9.taxon	distribution	Distribution. China, Laos, Vietnam, Borneo.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
E4F0FBCF0645EC39F995CEF60DEF93F7.taxon	distribution	Distribution. China (Guangdong: Zhanjiang).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
A5A7CE6D1EBA8B60E07D3CF5A368B189.taxon	distribution	Distribution. China (Guangxi).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
0595EE08D3E17E35D6633206457BC09F.taxon	diagnosis	Diagnosis. Length 12.0 - 12.5 mm, width 4.0 - 4.3 mm. Labrum (Fig. 6) strongly convex on anterior part, rounded and pointed in front, front seta on apical margin, intermediate one on upper surface close to margin; mandibles sharp at apex; mentum (Fig. 14) irregular and densely setose, both tooth and lobes stout, tooth shorter than lobes, densely setose at basal half; palpi slender, palpomere 4 (Fig. 22) not dilated, densely setose, rounded and truncate at apex, labial palpomere 3 not dilated, bisetose inside; ligula strongly thickened and dilated, deeply depressed at sides, apex rounded, slightly emarginate in middle with a seta on either side. Elytra with two pairs of spots, front spot rounded, covering intervals 3 - 7 in the middle, hind spot close to apical inner angles, nearly rectangular, covering intervals 1 - 5; apex roundly truncate. Male genitalia. Median lobe dilated in the middle ventrally, apical lamella long, wide and round at apex (Figs 31 - 32). Female genitalia: Gonocoxite slender, setose on ventral surface, three setae on dorsal side, apex sharp (Fig. 41).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
0595EE08D3E17E35D6633206457BC09F.taxon	distribution	Distribution. Myanmar and Indonesia (Java).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
84803ECE8D96A939D5D352C4460680EE.taxon	distribution	Distribution. China (Macao), Cambodia, Laos, Vietnam, the Philippines, Indonesia (Sumatra) and India.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
AE934238C6D1F5615603991692AE19C4.taxon	distribution	Distribution. China (Fujian, Jiangxi, Guangdong, Guangxi, Guizhou, Yunnan, Hongkong, Hainan), Laos, Vietnam, Myanmar, India and Sri Lanka.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
23063E13676E64838CF5800205CEC101.taxon	diagnosis	Diagnosis. Length 8.0 - 10.1 mm, width 3.0 - 3.5 mm. Labrum (Fig. 7) convex, arcuate at apex, front setae beneath apex, intermediate one on apical margin, hind one close to middle margin; mandibles sharp at apex; mentum (Fig. 15) setose at base; median tooth triangular and shorter than lobes, with two pairs of setae at base; lobes obtuse at apex; maxillary palpomere 4 (Fig. 23) roundly dilated, labial palpomere 4 triangular in shape and dilated, 3 not dilated and bisetose on inner sides; ligula thickened, apical margin arcuate inwards, deeply depressed at sides, with a median impression close to apex, outer apical angles rounded, with a seta on either of median impression close to apex. Elytra with front spot large, just before middle, almost rounded, covering intervals 3 - 7 and extended to small part of 2 and 8; hind spot on inner angles, more or less triangular, covering intervals 1 - 5. Male genitalia. Median lobe slender, apical lamella round (Figs 33 - 34). Female genitalia. Gonocoxite slender, arcuate, with three or four setae on dorsal side, apex slightly obtuse (Fig. 42).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
23063E13676E64838CF5800205CEC101.taxon	distribution	Distribution. Indonesia (Sumatra, Malaysia and Sulawesi).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
60883FA732714C9131457ED71CD2646F.taxon	description	Description. Length 9.5 mm, width 3.5 mm. Black; ligula, antennomeres 1 - 4, a spot on vertex, lateral margin of pronotum and legs reddish brown; sides of ligula, palpi, antennomeres 5 - 11 and elytral spots brown. Head convex; neck short and punctate on dorsal surface; frontoclypeal sulcus faint, frontal foveae short and shallow; clypeus with apical margin truncate, two setae on each side of apical outer angels, a row of 6 setae along apical margin, irregularly setose basally on each side; labrum (Fig. 8) convex anteriorly, apical margin rounded, front setae beneath near apex, middle one just on apical margin, hind one close to middle margin; mandibles covered by labrum, sharp at apex; mentum (Fig. 16) irregularly setose and punctate at base, median tooth triangular and shorter than lobes, with a pair of setae at base, lobes obtuse at apex; maxillary palpi (Fig. 24) not dilated, labial palpomere 3 with two setae on inner side; ligula thickened, apex deeply and widely emarginated in middle. Pronotum flat; widest before middle; faint median line, median and apical impression distinct, basal foveae deep; lateral margin round in front, strongly sinuate near base; hind angles nearly rectangular, with a small obtuse tooth and an emargination before tooth. Elytra flat, striae with large, close punctures and setae; intervals convex, with two rows of regular punctures and setae, interval 8 wider than others and densely and irregularly punctate and setose; spots small, front spot rounded, just before middle, covering intervals 3 - 6, hind spot rhombic, on inner apical angles, covering intervals 1 - 5. Male genitalia. Median lobe dilated in middle on ventral margin; apical lamella elongated, rounded at apex (Figs 35 - 36).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
60883FA732714C9131457ED71CD2646F.taxon	etymology	Etymology. This species is named in honor of Dr. Thierry Deuve (MNHN), a well known carabidologist.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
60883FA732714C9131457ED71CD2646F.taxon	distribution	Distribution. The Philippines.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
5F0E7FD7DC76B6E296FA45571E915D44.taxon	distribution	Distribution. China (Guangxi: Napo).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
35FD78E7AA4CCBEA7AE703CEB03914F1.taxon	distribution	Distribution. China (Guangxi: Dibei).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
A36059524D4F4F0838C308E53E2DD9A4.taxon	distribution	Distribution. China (Guangxi: Tengxian).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
88B506F05938593A97862D3087A42903.taxon	distribution	Distribution. China (Guangxi: Cangwu).	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
EDEB1DEFF5B4B45BC08B56194B887A00.taxon	distribution	Distribution. China (Guangxi: Fenghuangcheng). Distribution of Macrocheilus in the Oriental Region The zoogeographical pattern of Macrocheilus is illustrated in Figure 62. Most species are distributed in limited small areas, but three are widespread, occurring in many countries: Macrocheilus asteriscus in China, Vietnam, Laos, Myanmar, Indonesia and India; Macrocheilus chaudoiri in China, Cambodia, Laos, Vietnam, the Philippines, Sumatra and India; and Macrocheilus bensoni in China, Laos, Vietnam, Myanmar, India and Sri Lanka. At present the Macrocheilus faunas of Thailand and Nepal remain unknown.	en	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
