identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5A9D0E601DCC69CDF67C63886E0DCC13.text	5A9D0E601DCC69CDF67C63886E0DCC13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus Hope 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Genus  Macrocheilus Hope, 1838</p>
            <p> Macrocheilus Hope 1838: 166. Type species:  Macrocheilus bensoni Hope, 1838. Jeannel 1949: 1041. Zhao and Tian 2010:4. </p>
            <p> Acanthogenius Reiche 1842: 334. Type species:  Helluo impictus Wiedemann, 1823. </p>
            <p> Generic diagnosis. </p>
            <p>Medium sized (length from 8.0 to 17.0 mm), elongate, whole body coarsely punctuate and pubescent, except for labrum and middle region of ventral side of head. Head with two supraorbital setae on each side; ligula fairly wide, deeply impressed beneath at sides of base; mentum deeply emarginated, with a long and slender tooth; palpi varied in form; labrum extraordinarily or well developed; mandibles dorsally covered by labrum or not covered; antennae stout and flat, densely pubescent from antennomere 5. Pronotum truncate-cordiform or quadrate, basal margin more or less produced backwards in middle; lateral margin with two setae, one just before middle, and the other at hind angle. Elytra with deep striae and setiferous pores; intervals slightly or rather convex, interval 8 usually wide. Metepisterna elongate and very narrow in all species. Tarsomere 4 emarginate. Wings fully developed. No externally visible sexual dimorphic characters present. Median lobe of aedeagus varied in form. Parameres of aedeagus quite similar, elongate, the left one larger than the right one.</p>
            <p> Differences with other genera of  Helluonini in the Oriental Region. </p>
            <p> Macrocheilus spp., with larger body size, simple 4th tarsomeres, and non-securiform labial palpomere are distinctly different from  Colfax spp. (smaller body size, securiform labial palpomere) and  Creagris spp. (bilobed 4th tarsomere; smaller body size). Functional wings and the occurrence of a long spiniform median tooth of mentum of  Macrocheilus distinguish them from  Omphra spp. (which are brachypterous and have a short median tooth of the mentum). </p>
            <p>Generic range.</p>
            <p>Oriental Region (China, Vietnam, Laos, Cambodia, Myanmar, India, Sri Lanka, the Philippines, Malaysia and Indonesia), Palaearctic Region (Pakistan, Syria, Asia Minor), Ethiopian Region (Ethiopia, Tanzania, Uganda, Senegal, Guinea, Guinea-Bissau, Sierra Leone, Togo, Nigeria, Central Africa, Cameroon, Gabon, Congo, Zimbabwe, Angola, South Africa, Madagascar).</p>
            <p> Key to species of Oriental  Macrocheilus</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/5A9D0E601DCC69CDF67C63886E0DCC13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
B6CFF4826AFBD9EDBC0CB9FF5F44FA83.text	B6CFF4826AFBD9EDBC0CB9FF5F44FA83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus bicolor Andrewes 1920	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus bicolor Andrewes, 1920 Figs 1, 9, 17, 37, 43 </p>
            <p> Macrocheilus bicolor Andrewes 1920: 503; Andrewes 1930: 206; Csiki 1932: 1573; Lorenz 2005: 512. Type locality: India (Bombay: Belgaum), deposited in NHML. </p>
            <p>Diagnosis.</p>
            <p> Length 6.3-6.5 mm, width 2.5 mm. Head and prothorax red; elytra black or sometimes bluish black. Labrum (Fig. 1) semicircular in front, shortly depressed towards base, front pair of setae small and closely placed along the front margin, intermediate one at a distance from margin; ligula rectangular, with a wide and deep median impression, a pair of setae at a distance form apex depressed at base; apex truncate, outer apical angles rounded; mentum (Fig. 9) glabrous at base,  both tooth and lobes elongate, slender and sharp at apex; tooth almost as long as lobes, two pairs of setae on base; lobes sinuate at a distance from apex along outer margin; maxillary palpi (Fig. 17) not dilated. Elytra without spots. </p>
            <p>Female genitalia. Gonocoxite short, stout, inner margin arcuate and not sinuate, apex short and sharp (Fig. 37).</p>
            <p>Remarks.</p>
            <p> This species is allied to the next species,  Macrocheilus impictus (Wiedemann). Both differ from other species in the absence of elytral spot. </p>
            <p>Materials examined.</p>
            <p> Holotype: 1 female, labeled "Kaoeqaoe, 740, Type,  Macrochilus bicolor Type Andr., H. E. Andrewes det.; H. E. Andrewes Coll. B. M. 1945-97.;  Macrochilus
bicolor
 Andrewes, 1920", deposited in NHML. 1 ex, sex unknown (the abdomen missing), labeled "Bangalore, Chikkangalur, Taboourel, 1900;  Macrocheilus bicolor Andrewes, H. E. Andrewes det.", deposited in MNHN. </p>
            <p>Distribution.</p>
            <p>India.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/B6CFF4826AFBD9EDBC0CB9FF5F44FA83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
18642328DCEE29508CC7DA5469BADE1E.text	18642328DCEE29508CC7DA5469BADE1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus impictus (Wiedemann 1823) Wiedemann 1823	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus impictus (Wiedemann, 1823) Figs 2, 10, 18, 25, 26, 38, 44 </p>
            <p> Helluo impictus Wiedemann 1823: 49. Dejean 1825: 287; Reiche 1842: 335; Heller 1900: 3; Andrewes 1920: 503; 1921: 168; Csiki 1932: 1574; Lorenz 2005: 512. Type locality: India, deposited in ZMUC. </p>
            <p>Diagnosis.</p>
            <p>Length 14.5-15.0 mm, width 5.0-5.5 mm. Black. Labrum (Fig. 2) with front margin rounded and bisinuate, three pairs of setae close to margin, and front two pairs on sinuated area; mandibles slightly obtuse at apex; mentum (Fig. 10) irregularly setose in basal half, both tooth and lobes elongate and sharp at apex, tooth slightly shorter than lobes; maxillary palpomere 4 (Fig. 18) cylindrical and gradually dilated toward apex, rounded and obliquely truncate at apex. Elytra without spots.</p>
            <p>Male genitalia. Median lobe dilated on dorsal side in middle partion, sinuate on ventral side; apical lamella elongate, not sinuate near apex, rounded at apex (Figs 25-26).</p>
            <p>Female genitalia. Gonocoxite slender, five setae on dorsal surface, sharp at apex (Fig. 38).</p>
            <p>Remarks.</p>
            <p> Similar to  Macrocheilus bicolor without elytral spots, distinctly differs by larger size, body black, sinuate front margin of labrum and plurisetose mental tooth. </p>
            <p>Materials examined.</p>
            <p> 1 male, labeled "Indes Orientales, Mts Kodeicanel, J. Castets 1886"; 1 male, "India, Bangalore. P.S. Nathan, 1936"; 1 male, "Punjab Baddia (Indes Angl.); G. Babault Avril 1914"; 1 male, "S. India, Medungadu, P.S. Nathan. 1936"; 3 males,  “Java” , "Ex-Musaeo Chaudoir from Coll. Dejean"; 1 female,  “Java” , "Ex-Musaeo Mniszech from Coll. Dejean". All deposited in MNHN. </p>
            <p>Distribution.</p>
            <p>India (East India) and Indonesia (Java).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/18642328DCEE29508CC7DA5469BADE1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
D2B1C261B647E4CE3DE1A1272ADBDA99.text	D2B1C261B647E4CE3DE1A1272ADBDA99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus immanis Andrewes 1920	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus immanis Andrewes, 1920 Figs 3, 11, 19, 27, 28, 45 </p>
            <p> Macrocheilus immanis Andrewes 1920: 498. Csiki 1932: 1574;  Jedlička 1963: 469; Lorenz 2005: 512. Type locality: Myanmar (Taung-ngu), deposited in NHML. </p>
            <p>Diagnosis.</p>
            <p> Length 24.7 mm, width 8.3 mm. Labrum (Fig. 3) elongate, three pairs of equidistant setae on upper surface near margin; ligula thickened, apex slightly  narrowed , slightly emarginate in middle of front margin, deeply depressed near apex to form a median channel, with five pairs of setae along sides, hollowed out above with a median carina; mentum (Fig. 11) with both tooth and lobes stout, tooth shorter than lobes, contracted after middle, irregularly setose on ventral surface on basal half; maxillary palpomere 4 (Fig. 19) dilated, densely setose; labial  palpomere 4 dilated, palpomere 3 not dilated inwards. Elytal spots nearly square, large, near the middle, cover intervals 2-7. </p>
            <p>Male genitalia. Median lobe of aedeagus strongly dilated and stout, strongly sinuate near paramere, straight towards apex, apical lamera elongate and narrowed at apex (Figs 27-28).</p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus niger and  Macrocheilus asteriscus in having anterior setae of labrum obviously on upper surface, but differs from the other two species by the large and almost square elytral spots and stout median tooth of mentum. </p>
            <p>Materials examined.</p>
            <p> 1 male, the holotype, labeled "Toungoo; Type;  Macrochilus immanis , Type, Andr., H. E. Andrewes det.;  Macrochilus immanis Andrewes, 1920", deposited in NHML. </p>
            <p>Distribution.</p>
            <p>Myanmar.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/D2B1C261B647E4CE3DE1A1272ADBDA99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
7E0E3BFCD8260C9B7D3852414329D2DC.text	7E0E3BFCD8260C9B7D3852414329D2DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus niger Andrewes 1920	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus niger Andrewes, 1920 Figs 4, 12, 20, 29, 30, 39, 46 </p>
            <p> Macrocheilus niger Andrewes 1920: 499. Csiki 1932: 1574; Lorenz 2005: 512. Type locality: India (Nilgiri Hills), deposited in NHML. </p>
            <p>Diagnosis.</p>
            <p>Length 16.0-16.3 mm, width 6.0-6.2 mm. Labrum (Fig. 4) convex, with a short furrow on each side of base, narrowed and pointed in front, with three pairs of setae away from the margin; ligula thickened, apex truncate towards sides, apical margin slightly emarginate, deeply depressed towards base, with a small median impression near apex; mentum (Fig. 12) glabrous at base; median tooth shorter than lobes, rather narrow and with apex rather pointed, two pairs of setae at basal area; lateral lobes obtuse at apex. Palpi (Fig. 20) not dilated. Elytral spots transverse, close to the middle of interval 5, covering intervals 3-7.</p>
            <p>Male genitalia. Median lobe stout, apical lamella short and round at apex (Figs 29-30).</p>
            <p>Female genitalia. Gonocoxite subapically dilated, three setae on dorsal surface, sharp at apex (Fig. 39).</p>
            <p>Remarks.</p>
            <p> Similar to  Macrocheilus asteriscus , but differs with the tooth of mentum plurisetose and elytral spot not cruciform. </p>
            <p>Materials examined.</p>
            <p> 1 female, the holotype, labeled "type, 11, H. L. Andrewes, Nilgiri Hills, H. E. Andrewes Coll., B.M. 1945-97.;  Macrochilus niger Type-, Andr., H. E. Andrewes det.;  Macrochilus niger Andrewes, 1920", deposited in NHML. 1 male and 1 female, "Dehra-Dun 1940-45, Kumaon Himalaya, Liesenfeldt leg.", deposited in MNHN. </p>
            <p>Distribution.</p>
            <p> India (Nilgiri Hills, Malabar, Madras and Bombay) and Sri Lanka  .</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/7E0E3BFCD8260C9B7D3852414329D2DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
91B688B2A97938D74EA03FE7D3B12274.text	91B688B2A97938D74EA03FE7D3B12274.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus asteriscus White 1844	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus asteriscus White, 1844 Fig. 47 </p>
            <p> Macrocheilus asteriscus White 1844: 422; Bates 1892: 389; Andrewes 1919: 180; 1920: 500; 1924: 470; 1930: 206; Csiki 1932: 1573; Wu 1937: 188;  Jedlička 1963: 470;  Hůrka 2003:407; Lorenz 2005: 512; Zhao and Tian 2010: 6. Type locality: China (Hongkong), deposited in NHML. </p>
            <p> Planetes crucifer Redtenbacher 1867: 4. Type locality: China (Hongkong), deposited in NHML. </p>
            <p>Remarks.</p>
            <p> Macrocheilus asteriscus differs from other species in having cruciform elytral spots, the slender median tooth of mentum and the narrow apex of ligula. </p>
            <p>Materials examined.</p>
            <p> 2 males, 1 female,  “Hongkong” (MNHN); 2 males, "Hainan, Oct. 1979, Shaoming Zhuo leg." (SCAU); 2 males, "Guangdong: Zhanjiang, Jul. 1982" (SCAU); 1 male, 1 female, "Guangdong: Zhanjiang, May, 1983" (SCAU). 1 male, 1 female, "Annam, Phuc-Son, Nov. to Dec., H, Fruhstorfer", Central Vietnam (MNHN); 1 male, "Tonkin, P.  Lemée , 1903  –1906” , North Vietnam (MNHN); 1 male, "Laos. Mouhot" and "Janson Acq. 1884" (MNHN); 2 males, "Java, Preanger", Indonesia (MNHN); 1 female,  “Nilgherries” and "Ex. Musaeo H. W. Bates, 1892", India (MNHN). </p>
            <p>Distribution.</p>
            <p>China, Vietnam, Laos, Myanmar, Indonesia and India.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/91B688B2A97938D74EA03FE7D3B12274	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
BA06BBD2C9B8E8B9ABEEB6CB0537FDE8.text	BA06BBD2C9B8E8B9ABEEB6CB0537FDE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus binotatus Andrewes 1931	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus binotatus Andrewes, 1931 Figs 5, 13, 21, 40, 49 </p>
            <p> Macrocheilus binotutats Andrewes 1931: 68. Csiki 1932: 1573; Lorenz 2005: 512. Type locality: Indonesia (Sumatra), deposited in LMN. </p>
            <p>Diagnosis.</p>
            <p>Length 14.0 mm, width 4.8 mm. Labrum (Fig. 5) with apex rounded and pointed, front setae on apical margin; mandibles covered by the labrum, sharp at apex; mentum (Fig. 13) glabrous at base, median tooth nearly as long as lobes, about five setae at basal half; maxillary palpomere 4 (Fig. 21) cylindrically dilated, labial palpomere 4 flat and dilated, labial palpomere 3 dilated inwards; ligula wide and rectangular, apical margin straight, with a wide median impression beneath apex, a seta on either side near apex. Elytral spots large and oblong, covering intervals 3-7 in the middle.</p>
            <p>Female genitalia. Gonocoxite elongate, three setae on dorsal margin, sparsely setose on ventral surface; apex sharp (Fig. 40).</p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus macromaculatus having elytral spots oblong; and it differs from  Macrocheilus macromaculatus in having the anterior setae of labrum on the apical margin. </p>
            <p>Materials examined.</p>
            <p>1 female, "Paggar Alam, Sumatra, J. Bouchaud", deposited in MNHN.</p>
            <p>Distribution.</p>
            <p>Indonesia (Sumatra).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/BA06BBD2C9B8E8B9ABEEB6CB0537FDE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
F56D815FA88F58569CC027DB91EF1BAD.text	F56D815FA88F58569CC027DB91EF1BAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus macromaculatus Louwerens 1949	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus macromaculatus Louwerens, 1949</p>
            <p> Macrocheilus macromaculatus Louwerens 1949: 51. Lorenz 2005: 512. Type Locality: Indonesia (Java), deposited in LMN. </p>
            <p>Diagnosis.</p>
            <p>Length 10.0 mm, width 3.0 mm. Labrum large, semicircular anteriorly, with a flat depression on each sides, setae running along the sides in two small furrows; palpi short and stout, truncate at apex; mentum with a long, narrow, sharp median tooth, lateral lobes a little longer. Elytral spots oblong and covers intervals 2-7.</p>
            <p>The above description is after Louwerens (1949).</p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus binotatus , but the former is smaller in size and has the sides of elytra gently rounded behind, in contrast to the larger size and more sharply rounded hind region of elytra in  Macrocheilus macromaculatus . </p>
            <p>Distribution.</p>
            <p>Indonesia (Java).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/F56D815FA88F58569CC027DB91EF1BAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
708FF3F174B0C919051F96B142E12AB9.text	708FF3F174B0C919051F96B142E12AB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus vitalisi Andrewes 1920	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus vitalisi Andrewes, 1920 Fig. 48 </p>
            <p> Macrocheilus vitalisi Andrewes 1920: 500; Andrewes 1930: 208; Csiki 1932: 1575; Wu 1937: 188;  Jedlička 1963: 470;  Hůrka 2003: 407; Lorenz 2005: 512; Zhao and Tian 2010: 7. </p>
            <p>Remarks.</p>
            <p> Macrocheilus vitalisi is similar to  Macrocheilus binotatus from Sumatra, but as stated by Andrewes (1931),  Macrocheilus binotatus differs from  Macrocheilus vitalisi by "the dark colour, the elytral spot oblong and red; the upper surface generally is more coarsely and less densely punctuate, the genae are contracted more sharply to the neck, the sides of the prothorax are less sinuate behind, the elytral intervals more convex, with punctuation along the side more widely spaced and coarser". In addition, according to our examination, the anterior seta of the labrum is closer to the apical margin in  Macrocheilus vitalisi than in  Macrocheilus binotatus and the 4th maxillary palpomere is cylindrically dilated in  Macrocheilus vitalisi . </p>
            <p>Materials examined.</p>
            <p> 1 female, the holotype, "China, Bowring 63·47*, 986 27/2/53" (NHML); 1 female, "Tonkin,  Région de Hoa-Binh",  “Muséum Paris, 1932, A. de Cooman" (MNHN). </p>
            <p>Distribution.</p>
            <p>China, Laos, Vietnam, Borneo.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/708FF3F174B0C919051F96B142E12AB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
E4F0FBCF0645EC39F995CEF60DEF93F7.text	E4F0FBCF0645EC39F995CEF60DEF93F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus gigas Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus
gigas Zhao &amp; Tian, 2010
 Fig. 50 </p>
            <p> Macrocheilus gigas Zhao and Tian 2010: 8. Type Locality: China (Guangdong), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> Macrocheilus gigas is similar to  Macrocheilus parvimaculatus and  Macrocheilus tripustulatus in having anterior seta of the labrum on the apical margin and the ligula thickened at apex which distinguishes these three species from other species with two spots on each elytron. But  Macrocheilus gigas differs from the other two species by wide apex of labrum and having median tooth of mentum sinuate near apex on lateral margin. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, Guangdong: Zhanjiang, Jun.1983, deposited in SCAU; 1 male, Guangdong: Zhanjiang, Oct.1982, Zhichang Tan leg., deposited in SCAU; 2 males, Guangdong: Zhanjiang, July 1982, deposited in SCAU and MNHN.</p>
            <p>Distribution.</p>
            <p>China (Guangdong: Zhanjiang).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/E4F0FBCF0645EC39F995CEF60DEF93F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
A5A7CE6D1EBA8B60E07D3CF5A368B189.text	A5A7CE6D1EBA8B60E07D3CF5A368B189.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus parvimaculatus Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus parvimaculatus Zhao &amp; Tian, 2010 Fig. 51 </p>
            <p> Macrocheilus parvimaculatus Zhao and Tian 2010: 9. Type Locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> This species is easily distinguished from other species of  Macrocheilus by its strongly convex labrum, maxillary palpomere 4 dilated in apical half and with small spots on elytra. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, Guangxi: Liuzhou: Luzhai, 26 May 1980, Shaozhou Ruan leg.; 1 male, same data as holotype. All specimens deposited in SCAU.</p>
            <p>Distribution.</p>
            <p>China (Guangxi).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/A5A7CE6D1EBA8B60E07D3CF5A368B189	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
0595EE08D3E17E35D6633206457BC09F.text	0595EE08D3E17E35D6633206457BC09F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus tripustulatus (Dejean 1825) Dejean 1825	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus tripustulatus (Dejean, 1825) Figs 6, 14, 22, 31, 32, 41, 52 </p>
            <p> Helluo tripustulatus Dejean 1825: 286 (syn. in Reiche 1842: 334). Reiche 1842: 334; Chaudoir 1872: 212; Heller 1900: 3; Andrewes 1919: 124; Andrewes 1920: 501; Csiki 1932: 1575; Lorenz 2005: 512. Type locality: Indonesia (Java), deposited in MNHN. </p>
            <p>Diagnosis.</p>
            <p> Length 12.0-12.5 mm, width 4.0-4.3 mm. Labrum (Fig. 6) strongly convex on anterior part, rounded and pointed in front, front seta on apical margin, intermediate one on upper surface close to margin; mandibles sharp at apex; mentum (Fig. 14)  irregular and densely setose, both tooth and lobes stout, tooth shorter than lobes, densely setose at basal half; palpi slender, palpomere 4 (Fig. 22) not dilated, densely setose, rounded and truncate at apex, labial palpomere 3 not dilated, bisetose inside; ligula strongly thickened and dilated, deeply depressed at sides, apex rounded, slightly emarginate in middle with a seta on either side. Elytra with two pairs of spots, front spot rounded, covering intervals 3-7 in the middle, hind spot close to apical inner angles, nearly rectangular, covering intervals 1-5; apex roundly truncate. </p>
            <p>Male genitalia. Median lobe dilated in the middle ventrally, apical lamella long, wide and round at apex (Figs 31-32).</p>
            <p>Female genitalia: Gonocoxite slender, setose on ventral surface, three setae on dorsal side, apex sharp (Fig. 41).</p>
            <p>Remarks.</p>
            <p> Macrocheilus tripustulatus can be distinguished from  Macrocheilus parvimaculatus by the slender maxillary palpomere 4, greater convexity of labrum and larger elytral spots. </p>
            <p>Materials examined.</p>
            <p> 1 female, the holotype, "tripustulatus Dejean (non Fabr.), Java, Coll. Dejean, 3-pustulatus Wiedemann (  Helluo ). Fabr. (Brachinus), Western, Ex-Musaeo Chaudoir"; 1 male, "Java merid., 1500, 1891, H. Fruhstorfer."; 1 male, " Java", "Ex-Musaeo Mniszech"; 1 female, "Carin  Chebà , 900-1100 m, L. Fea, V Xii-88; Ex-Musaeo H.W. Bates 1892 ". All specimens deposited in MNHN. </p>
            <p>Distribution.</p>
            <p>Myanmar and Indonesia (Java).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/0595EE08D3E17E35D6633206457BC09F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
84803ECE8D96A939D5D352C4460680EE.text	84803ECE8D96A939D5D352C4460680EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus chaudoiri Andrewes 1919	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus chaudoiri Andrewes, 1919 Fig. 53 </p>
            <p> Macrocheilus chaudoiri Andrewes 1919: 130; 1920: 502; 1924: 470; 1930: 207; Csiki 1932: 1573;  Jedlička 1963: 470; Lorenz 2005: 512; Zhao and Tian 2010: 11. Type Locality: India, deposited in MNHN. </p>
            <p> Acanthogenius trimaculatus sensu Chaudoir 1872: 171 (non Oliver). Type Locality: India (Decan), deposited in MNHN. </p>
            <p> Macrocheilus ruficollis Heller 1923: 296; Andrewes 1926: 258. Type Locality: Philippines (Luzon), deposited in MDSG. </p>
            <p>Remarks.</p>
            <p> Macrocheilus chaudoiri ,  Macrocheilus nigrotibialis ,  Macrocheilus bensoni and  Macrocheilus deuvie are similar in having the anterior seta of the labrum beneath the apex and two pairs of elytral spots. This species is distinct from the other three species in having the head and pronotum reddish brown and maxillary palpomere 4 strongly dilated. </p>
            <p>Materials examined.</p>
            <p> 1 female, the holotype, "Ex-Musaeo Chaudoir; Macrocheilus Chaudoiri Andr., H.S. Andrewes det.; 3-maculatus Chaud., Deccan, Coll. Jeakes" (MNHN); 1 male, "Maissour, Sakrabail, IX 1897" (MNHN); 1 female, "Museum Paris, Cochinchine, Harmand 1872" (MNHN); 1 female,  “Java” and "Museum Paris, Lakhon, Harmand 1878" (MNHN); 1 female, "Insl. Phiip." and  “Thorey” (MNHN). </p>
            <p>Distribution.</p>
            <p>China (Macao), Cambodia, Laos, Vietnam, the Philippines, Indonesia (Sumatra) and India.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/84803ECE8D96A939D5D352C4460680EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
AE934238C6D1F5615603991692AE19C4.text	AE934238C6D1F5615603991692AE19C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus bensoni Hope 1838	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus bensoni Hope, 1838 Fig. 55 </p>
            <p> Macrocheilus bensoni Hope 1838: 166; Chaudoir 1872: 212; Bates 1892: 389; Heller 1900: 3; Andrewes 1919: 176, 202;  Hůrka 2003: 407; Lorenz 2005: 512: Zhao and Tian 2010: 10. Type locality: India, deposited in NHML. </p>
            <p> Carabus trimaculatus Olivier 1790: 347 (non Villers 1789); Andrewes 1919: 129, 176; 1920: 502; 1930, 208; Csiki 1932: 1574; Wu 1937: 188;  Jedlička 1963: 470. Type locality: China, deposited in NHML. </p>
            <p> Helluo quadrimaculata Guérin-Méneville 1840: 38; Chaudoir 1872: 212. Type locality: India, deposited in MNHN. </p>
            <p> Helluo tripustulata sensu  Guérin-Méneville 1843: 34 (non Dejean 1825); Andrewes 1923: 460. Type locality: China, deposited in MNHN. </p>
            <p> Macrocheilus quadripustulatus Schmidt-Göbel 1846: 65. Type locality: Myanmar, deposited in NMP. </p>
            <p> Acanthogenius infuscatus Bates 1892, 389; Andrewes 1920: 493. Type locality: Myanmar (Bhamo), deposited in MGI. </p>
            <p>Remarks.</p>
            <p> This species can be distinguished from  Macrocheilus chaudoiri by the black head and pronotum and from  Macrocheilus nigrobibialis by the slender maxillary palpomere 4. </p>
            <p>Materials examined.</p>
            <p>1 male, "North China, 1884, Janson" (MNHN); 1 male, "North India, Coll. Benson, Ex-Musaeo H.W. Bates, 1892" (MNHN); 1 male, "Ind. Angl., Coimbatore Dt, Siruveni, VI. 1937" (MNHN); 1 female, "Ind. Angl., Mysore" (MNHN); 1 male, "Guangdong: Zhanjiang, Oct.1982, Zhichang Tan leg." (SCAU); 2 males, "Guangdong: Yingde, 27 Mar.2003, Danyang Zhao leg." (SCAU and MNHN); 1 male, Guangxi, Dec.1983" (SCAU); 1 male, "Yunnan: Jinghong, Jul. 1985" (SCAU); 1 female, "Hainan: Diaoluoshan, 26 Nov.1963" (SCAU).</p>
            <p>Distribution.</p>
            <p>China (Fujian, Jiangxi, Guangdong, Guangxi, Guizhou, Yunnan, Hongkong, Hainan), Laos, Vietnam, Myanmar, India and Sri Lanka.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/AE934238C6D1F5615603991692AE19C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
23063E13676E64838CF5800205CEC101.text	23063E13676E64838CF5800205CEC101.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus nigrotibialis Heller 1900	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus
nigrotibialis Heller, 1900
 Figs 7, 15, 23, 33, 34, 42, 54 </p>
            <p> Macrochilus nigrotibialis Heller 1900: 2. Andrewes 1920: 497; Csiki 1932: 1574; Lorenz 2005: 512. Type locality: Indonesia (Sulawesi), deposited in MDSG. </p>
            <p>Diagnosis.</p>
            <p>Length 8.0-10.1 mm, width 3.0-3.5 mm. Labrum (Fig. 7) convex, arcuate at apex, front setae beneath apex, intermediate one on apical margin, hind one close to middle margin; mandibles sharp at apex; mentum (Fig. 15) setose at base; median tooth triangular and shorter than lobes, with two pairs of setae at base; lobes obtuse at apex; maxillary palpomere 4 (Fig. 23) roundly dilated, labial palpomere 4 triangular in shape and dilated, 3 not dilated and bisetose on inner sides; ligula thickened, apical margin arcuate inwards, deeply depressed at sides, with a median impression close to apex, outer apical angles rounded, with a seta on either of median impression close to apex. Elytra with front spot large, just before middle, almost rounded, covering intervals 3-7 and extended to small part of 2 and 8; hind spot on inner angles, more or less triangular, covering intervals 1-5.</p>
            <p>Male genitalia. Median lobe slender, apical lamella round (Figs 33-34).</p>
            <p>Female genitalia. Gonocoxite slender, arcuate, with three or four setae on dorsal side, apex slightly obtuse (Fig. 42).</p>
            <p>Remarks.</p>
            <p>This species differs from following two species in having the tibiae black, maxillary palpomere 4 more dilated, the median lobe of male genitalia stouter and the apical lamella more rounded.</p>
            <p>Materials examined.</p>
            <p> 1 male, the holotype,  “typus!” , "Drs. Sarasin N. Celebes Panot-Maimang",  “12623” , "Staatl. Museum fur Tierkunde. Dresden", deposited in SNSD. 1 female, labeled "Nord Borneo Mont Kina Balu 5-8, 1903, John Waterstradt;  Macrochilus nigrotibialis Heller, det. Andrewes", deposited in MNHN. </p>
            <p>Distribution.</p>
            <p>Indonesia (Sumatra, Malaysia and Sulawesi).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/23063E13676E64838CF5800205CEC101	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
60883FA732714C9131457ED71CD2646F.text	60883FA732714C9131457ED71CD2646F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus deuvie	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus deuvie sp. n. Figs 8, 16, 24, 35, 36, 56 </p>
            <p>Description.</p>
            <p>Length 9.5 mm, width 3.5 mm.</p>
            <p>Black; ligula, antennomeres 1-4, a spot on vertex, lateral margin of pronotum and legs reddish brown; sides of ligula, palpi, antennomeres 5-11 and elytral spots brown.</p>
            <p>Head convex; neck short and punctate on dorsal surface; frontoclypeal sulcus faint, frontal foveae short and shallow; clypeus with apical margin truncate, two setae on each side of apical outer angels, a row of 6 setae along apical margin, irregularly setose basally on each side; labrum (Fig. 8) convex anteriorly, apical margin rounded, front setae beneath near apex, middle one just on apical margin, hind one close to middle margin; mandibles covered by labrum, sharp at apex; mentum (Fig. 16) irregularly setose and punctate at base, median tooth triangular and shorter than lobes, with a pair of setae at base, lobes obtuse at apex; maxillary palpi (Fig. 24) not dilated, labial palpomere 3 with two setae on inner side; ligula thickened, apex deeply and widely emarginated in middle.</p>
            <p>Pronotum flat; widest before middle; faint median line, median and apical impression distinct, basal foveae deep; lateral margin round in front, strongly sinuate near base; hind angles nearly rectangular, with a small obtuse tooth and an emargination before tooth.</p>
            <p> Elytra flat, striae with large, close punctures and setae; intervals convex, with two rows of regular punctures and setae, interval 8 wider than others and densely and irregularly punctate and setose; spots small, front spot rounded, just before middle, covering intervals 3-6, hind spot rhombic, on inner apical angles, covering intervals 1-5. </p>
            <p>Male genitalia. Median lobe dilated in middle on ventral margin; apical lamella elongated, rounded at apex (Figs 35-36).</p>
            <p>Remarks.</p>
            <p> This species is very similar to  Macrocheilus bensoni , but differs by  it’s smaller size, curved labrum at anterior part, clypeus glabrous on middle, pronotum with lateral margin strongly sinuate near base, and male genitalia dilated on ventral margin. </p>
            <p>Type material.</p>
            <p>1 male, the holotype, "Philippines, Bohol. Ch. Semper", deposited in MNHN.</p>
            <p>Etymology.</p>
            <p>This species is named in honor of Dr. Thierry Deuve (MNHN), a well known carabidologist.</p>
            <p>Distribution.</p>
            <p>The Philippines.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/60883FA732714C9131457ED71CD2646F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
5F0E7FD7DC76B6E296FA45571E915D44.text	5F0E7FD7DC76B6E296FA45571E915D44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus fuscipennis Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus fuscipennis Zhao &amp; Tian, 2010 Fig. 57 </p>
            <p> Macrocheilus fuscipennis Zhao and Tian 2010: 12. Type locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus solidipalpis and  Macrocheilus cheni in having the larger elytral spots, but easily distinguishable from them by the labrum without additional setae, mandibles obtuse at apex, median tooth of mentum with lateral margin not sinuate and body brownish. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, "Guangxi: Napo, 10 Oct.1970, by light trap".</p>
            <p>Distribution.</p>
            <p>China (Guangxi: Napo).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/5F0E7FD7DC76B6E296FA45571E915D44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
35FD78E7AA4CCBEA7AE703CEB03914F1.text	35FD78E7AA4CCBEA7AE703CEB03914F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus solidipalpis Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus solidipalpis Zhao &amp; Tian, 2010 Fig. 58 </p>
            <p> Macrocheilus solidipalpis Zhao and Tian 2010: 13. Type locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus cheni but easily distinguishable from the latter by the presence of additional setae between the anterior and anterior setae, the median tooth of the mentum with sinuate lateral margins in middle (sinuate on api  cal one-third in  Macrocheilus cheni ), the median lobe larger, and the apical lamella long and narrowed towards apex. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, "Guangxi: Dibei, Oct.1980, Xiuzhen Mao leg., by light trap".</p>
            <p>Distribution.</p>
            <p>China (Guangxi: Dibei).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/35FD78E7AA4CCBEA7AE703CEB03914F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
A36059524D4F4F0838C308E53E2DD9A4.text	A36059524D4F4F0838C308E53E2DD9A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus cheni Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus cheni Zhao &amp; Tian, 2010 Fig. 59 </p>
            <p> Macrocheilus cheni Zhao and Tian 2010: 14. Type locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> This species is similar to  Macrocheilus solidipalpis and the differences from the latter were mentioned above. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype,"Guangxi: Tengxian, Oct.1980".</p>
            <p>Distribution.</p>
            <p>China (Guangxi: Tengxian).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/A36059524D4F4F0838C308E53E2DD9A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
88B506F05938593A97862D3087A42903.text	88B506F05938593A97862D3087A42903.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus quadratus Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus quadratus Zhao &amp; Tian, 2010 Fig. 60 </p>
            <p> Macrocheilus quadratus Zhao and Tian 2010: 16. Type locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p>This species is easily distinguished from other species by the shape of pronotum.</p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, labeled "Guangxi: Cangwu, 1980, in paddy field".</p>
            <p>Distribution.</p>
            <p>China (Guangxi: Cangwu).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/88B506F05938593A97862D3087A42903	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
EDEB1DEFF5B4B45BC08B56194B887A00.text	EDEB1DEFF5B4B45BC08B56194B887A00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrocheilus sinuatilabris Zhao & Tian 2010	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Macrocheilus sinuatilabris Zhao &amp; Tian, 2010 Fig. 61 </p>
            <p> Macrocheilus sinuatilabris Zhao and Tian 2010: 17. Type locality: China (Guangxi), deposited in SCAU. </p>
            <p>Remarks.</p>
            <p> Macrocheilus sinuatilabris is similar to  Macrocheilus quadratus in having three pairs of setae on the labrum and small elytral spots. However, it differs from  Macrocheilus quadratus in having  smaller elytral spots, the median tooth of the mentum obtuse at its apex, the rounded apex of the labrum, the close positioning of front and intermediate labral setae, and the slender median lobe of the aedeagus. </p>
            <p>Materials examined.</p>
            <p>1 male, the holotype, labeled "Guangxi: Fenghuangcheng, Sep.1981, by light trap".</p>
            <p>Distribution.</p>
            <p>China (Guangxi: Fenghuangcheng).</p>
            <p> Distribution of  Macrocheilus in the Oriental Region </p>
            <p> The zoogeographical pattern of  Macrocheilus is illustrated in Figure 62. Most species are distributed in limited small areas, but three are widespread, occurring in many countries:  Macrocheilus asteriscus in China, Vietnam, Laos, Myanmar, Indonesia and India;  Macrocheilus chaudoiri in China, Cambodia, Laos, Vietnam, the Philippines, Sumatra and India; and  Macrocheilus bensoni in China, Laos, Vietnam, Myanmar, India and Sri Lanka. At present the  Macrocheilus faunas of Thailand and Nepal remain unknown. </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/EDEB1DEFF5B4B45BC08B56194B887A00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhao, Danyang;Tian, Mingyi	Zhao, Danyang, Tian, Mingyi (2012): Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines. ZooKeys 228: 77-100, DOI: http://dx.doi.org/10.3897/zookeys.228.3401, URL: http://dx.doi.org/10.3897/zookeys.228.3401
