taxonID	type	description	language	source
03E1AF0CFFA4FFE8FF76FD9E570EC44F.taxon	materials_examined	Type material. Holotype male: “ S. Pedro (Vila Viçosa) \ Distrito Viseu, Portugal \ 6. XII. 2018 \ Col. A. Serrano ” \\ “ A. Serrano & \ C. Aguiar leg. ” \\ “ 41 º 1´10.35 ´´ N \ 8 º 13´36.49 ´´ W \ 220 m alt. ” \\ “ Holotype male \ Domene \ (Lathromene) \ boavidae sp. nov. \ A. Serrano det. 2024 ” [t] [h] [red card]. Paratypes, 2 females (1 female gold coated), same data as holotype; all paratypes with additional label “ Paratype female \ Domene \ (Lathromene) \ boavidae sp. nov. \ A. Serrano det. 2024 ” [t] [h] [red card].	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFA4FFE8FF76FD9E570EC44F.taxon	etymology	Etymology In modest homage to her memory, this new species is dedicated to the late colleague of the first author from the “ Departamento de Biologia Animal da Faculdade de Ciências de Lisboa ” (Portugal), Prof. Maria José Boavida, a friend and an eminent Portuguese limnologist which gave us scientific support and help to improve the English versions of some of our works.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFA4FFE8FF76FD9E570EC44F.taxon	description	Description. Microphthalmous, wingless and elongate body (Figs 1 b – c). Coloration: body reddish and abdomen dark-reddish. Body length: 8.5 mm (male), 6.3 ‒ 8.1 mm (females). Head (Figs 2 a – d, 3 a): orbicular, 1.1 ‒ 1.2 times wider than long (males and females) [length: 1.36 mm (male) and 1.07 ‒ 1.17 mm (females), width: 1.10 mm (male) and 0.86 ‒ 1.06 mm (females)], slightly wider than pronotum (1.1 times); eyes reduced, with about 15 ommatidia (Fig. 2 b); long and thin seta in supraocular small circular depression (supraocular trichobothrium); absence of small patch of minor pubescence above depression and behind eyes (Fig. 2 b); two dark spots in vertex marking insertions of dorsal arms of tentorium; gular sutures well defined, converging towards neck (Fig. 3 a); dorsal surface without any isodiametric microreticulation, with well-defined dense punctuation, similar to that of pronotum, on lateral sides and behind vertex; vertex and frons with sparse punctuation, almost smooth (Fig. 2 a); antennae filiform (Fig. 2 d), 2.5 mm long (male), 1.9 ‒ 2.5 mm (females), reaching base of pronotum when directed backwards; all antennomeres longer than wide, 1 st longest of all [length: 0.42 mm (male) and 0.32 ‒ 0.40 mm (females)]; 3 rd antennomere in average 1.2 – 1.3 times longer than the 4 th and 5 th antennomeres; the 6 th to 10 th antennomeres subequal and 11 th in average 1.3 times longer than preceding five; labrum deeply emarginated with 20 ‒ 23 large setae spread over disc and anterior margin (Fig. 2 c); mandibles symmetrical with four distinct teeth in inner edge, two of them more pronounced (Figs 2 c, 3 a); maxillary palpus with four palpomeres, second and third subequal, more than 3 times as long as broad, apical palpomere reduced and conical; internal lobe of maxilla with clusters of setae; labium with bilobed glossae, circumscribed by hairy paraglossae; labial palpus with three palpomeres, 2 nd longest and more robust, and 3 rd thinnest. Pronotum (Fig. 2 g): 1.2 times longer than wide (males and females) [length: 1.23 mm (male) and 0.99 ‒ 1.17 mm (females), width: 0.99 mm (male) and 0.80 ‒ 0.94 mm (females)], slightly narrower than head, widest anteriorly and distinctly tapering posteriad; anterior angles weakened, posterior angles marked but largely obtuse; basal margin well defined; punctuation similar to that of head, evenly distributed except for smooth midline ranging from anterior to posterior parts; absence of any middle sulcus, interstices without microsculpture. Elytra (Fig. 2 h): 1.2 ‒ 1.3 times wider than long [length: 0.78 mm (male) and 0.56 ‒ 0.74 mm (females), width: 0.96 mm (male) and 0.72 ‒ 0.93 mm (females)], approximately 0.6 times shorter than head or pronotum [elytra length / head length: 0.57 (male) and 0.52 ‒ 0.63 (females), elytra length / pronotum length: 0.63 (male) and 0.57 ‒ 0.63 (females)]; only slightly narrower than head (male: 0.9 times; females: 0.8 ‒ 0.9 times) and same width as pronotum (male: 1.0 times, females: 0.9 ‒ 1.0); flat in dorsal view; humeral angles obsolete; lateral margins slightly divergent posteriad; surface with coarse, rugose and wrinkled punctuation. Wingless. Abdomen (Figs 3 b ‒ f): Maximum width at segment VI (male: 1.17 mm; females: 0.96 – 1.18 mm), 1.2 – 1.3 times wider than elytra; tergites IV ‒ VI with fine and dense punctuation within microsculpture of transverse meshes, these more pronounced in intersegmental membranes; sternite III (male and females) without middle carina (intercoxal carina) in first two thirds; sternites III ‒ VI with fine and dense punctuation within microsculpture of transverse meshes, substituted by irregular rows of scale-shaped microtrichia anteriad near intersegmental membranes and by regular posteriad quadrate meshes [similar to Domene viriatoi Serrano & Boieiro, 2015, see fig. 5 b in Serrano et al. (2015)]. Male: sternite VII without any median impression posteriorly; sternite VIII slightly transverse, slightly and subtriangularly excised at posterior margin surrounded by small glabrous area with microsculpture of polygonal meshes (Fig. 3 b); sternite IX slightly arcuate apically; genital segment in ventral view as in Figure 3 d. Female: genital segment (dorsal view) with a large glabrous area in the center and in ventral view as in Figures 3 e ‒ f. Legs (Fig. 1 c): Long and slender; forelegs with antennal cleaning organ following the general pattern of the genus (Figs 2 e ‒ f). Aedeagus (Fig. 4): Length 1.3 mm; sclerotized ventral process symmetric, exceeding largely edge of dorsal plate (Fig. 4 a), with crested helmet shape apex, base of helmet with pair of small teeth in middle (Fig. 4 d); peduncular base of ventral process with expansion on each side directed inwards and upwards (Figs 4 a, 4 c); dorsal plate subparallel in dorsal view (Fig. 4 b) with fused lateral lobes (Fig. 4 c), presenting quadrate ‒ shaped sclerotized area in upper part (Figs 4 b – c) and ending by membranous process in apical edge (Fig. 4 a). Sexual dimorphism The sexual dimorphism is connected with the external morphological characters of the VIII and genital segments. The male sternite VIII is slightly and subtriangularly excised at posterior margin surrounded by small glabrous area (Fig. 3 b); sternite IX slightly arcuate apically; genital segment in dorsal and ventral views as in Figures 3 c and 3 d, respectively. The female sternite VIII unmodified; genital segment (dorsal view) with a large glabrous area in the center (Fig. 3 e) and in ventral view as in Figure 3 f.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFA4FFE8FF76FD9E570EC44F.taxon	discussion	Taxonomic, distribution and bionomic remarks The subgeneric classification of the genus Domene is at present artificial (e. g. see Assing & Feldmann 2014; Herman 2023) and in need of a thorough revision, which is out of the scope of the present work. So, using wellestablished taxonomic criteria (Coiffait 1982) (e. g. aedeagus with fused lateral lobes and the development of the lateral and ventral process) Domene boavidae sp. nov. should be included in the subgenus Lathromene. Among other morphological characteristics, the species shows a distinctive shape of the aedeagus. Taking into account the pieces of this structure, the new species can be easily segregated from D. scopaeella Fauvel, 1873, D. subiasi (Outerelo, 1977), D. gallaeciana Feldmann & Hernando, 2005, D. barraganensis Outerelo & Gamarra, 2012 and D. pedroi sp. nov., by the symmetry type of ventral process (symmetric in the new species vs. asymmetric in the previous five species). Among the species with symmetric ventral process, the closest species to D. boavidae sp. nov. seem to be D. caurelensis Outerelo, Gamarra & Salgado, 2000, D. orensis Struyve, 2018, D. eufemia Struyve, 2018 and D. darinkae Magrini & Carotti, 2019. However, unlike the new species, the ventral process of D. caurelensis only exceeds slightly the edge of the aedeagus and the apex is strongly bifurcated. In the other three species, the shape of the ventral process is that of a very elongated plough in lateral view and like a spearhead with the apical region converging to the apex (D. orensis) or more parallel (D. eufemia and D. darinkae) in ventral view [see figs 26 N – Q in Struyve (2018) and figs 4 – 7 in Magrini & Carotti (2019)]. Still within the symmetric ventral process group, D. lusitanica Reboleira & Oromi, 2011 and D. viriatoi Serrano & Boieiro, 2015, are two other species closely related to D. boavidae sp. nov. However, in D. lusitanica this structure does not exceed the edge of the aedeagus and presents a different shape of the pointed apex, also lacking the minor middle pair of teeth in the body base of apex (cf. Figs 4 a, 4 c – d and fig. 6 in Reboleira et al. 2011 b). On the other hand, the ventral process apex in D. viriatoi is bifurcated and its median region presents minor middle lateral hook-like expansions that are absent in the new species (cf. Fig. 4 a and fig. 7 a in Serrano et al. 2015). Furthermore, while in D. lusitanica the male sternite VIII shape and the setae on either side of the apex are similar to the new species, in D. viriatoi there are a deeply inverted U ‒ shaped incision with a cluster of 15 ‒ 21 modified, short and stout black striated setae [cf. Fig. 3 b and fig. 4 in Reboleira et al. (2011 b) and figs 5 c, 5 e – f in Serrano et al. (2015)]. The congeners with populations closer to the new species are D. darinkae (from Serra do Marão, Vila Real, Portugal), D. scopaeella (from Serra do Gerez, Portugal and Spain and Serra do Marão, Portugal), D. eufemia (from Torneiros, Orense, Spain) and D. sistelensis Struyve, 2018 (from Sistelo, Viana do Castelo, Portugal) (see also fig. 10 in Magrini & Carotti 2019). The latter species can be segregated from D. boavidae sp. nov. by the wide ventral process of aedeagus with two lateral teeth bent upwards, a big apical lobe with rounded incision and the dorsal side asymmetric (see figs 26 K – M in Struyve 2018). Although the male of D. hispanica Outerelo, 1985 is unknown, the species can easily be separated from the new species through several characters of the external morphology (e. g. shape and punctuation of the head and pronotum, size and absence of a median sulcus on the pronotum, femoral tooth of front leg) (see description in Outerelo 1985). The holotype and paratypes specimens of D. boavidae sp. nov. were sampled by hand directly among plates of schistose rocks below clayey brown-reddish soil on the side slopes of a country road (Fig. 1 a). Interestingly, all the specimens were found in chambers with organic remains within a Camponotus cruentatus (Latreille, 1802) (Hymenoptera, Formicidae) colony. The ant colony extended over a wide area among the schistose plates. The locality habitat was dominated by Pinus and Eucalyptus trees with brushwood, rock-roses, brooms, lavender and rosemary shrubs [Rubus fruticosus (Rosaceae), Cistus ladanifer (Cistaceae), Cistus salvifolius (Cistaceae), Cytisus spp. (Fabaceae), Lavandula sp. (Lamiaceae) and Rosmarinus officinalis (Lamiaceae)] (Fig. 1 a). We visited the same site during February 2019 and March 2024 trying to find more individuals, but without success. With no more data on its distribution, it is provisional accepted that the species is endemic of Portugal. According to the assumption of Reboleira et al. (2011 b) by their bigger size it was expected that this new species would be more connected with the hypogean environment. However, D. boavidae sp. nov. was found in a typical endogean environment (Fig. 1 a). Taking into account the reduced eyes, absence of hind wings, elongated body shape and long legs it is assumed that this species is a subterranean dwelling.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFABFFECFF76FB725050C503.taxon	materials_examined	Type material. Holotype male: “ Aldeia Cimeira (Pampilhosa da Serra) \ Distrito Coimbra, Portugal \ 3. IV. 2008 \ Col. A. Serrano ” \\ “ A. Serrano & \ C. Aguiar leg. ” \\ “ 40 º 1´38.01 ´´ N \ 7 º 58´22.61 ´´ W \ 635 m alt. ” \\ “ Holotype male \ Domene \ (Lathromene) \ pedroi sp. nov. \ A. Serrano det. 2024 ” [t] [h] [red card].	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFABFFECFF76FB725050C503.taxon	etymology	Etymology This new species is dedicated to Pedro Martins da Silva, a master’s and doctoral student of the first author and a renowned ecologist, who would like to have been a staphylinid taxonomist, but whose scientific career path led him to study insects encoded in numbers.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFABFFECFF76FB725050C503.taxon	description	Description. Microphthalmous, wingless and elongate body (Fig. 1 d). Coloration: body yellowish and abdomen slightly darker. Body length: 4.5 mm (male). Head (Figs 5 a – c): subquadrate / orbicular, 1.1 times longer than wide (length: 0.75 mm, width: 0.68 mm), slightly wider than pronotum (1.2 times); eyes strongly reduced, with 7 ommatidia (Fig. 5 c); long and thin seta in supraocular, small circular depression (supraocular trichobothrium); absence of patch of minor pubescence above depression and behind eyes (Fig. 5 c); two dark spots in vertex marking insertions of dorsal arms of tentorium; gular sutures well defined, diverging slightly towards neck (Fig. 5 b); dorsal surface without any isodiametric microreticulation, with well-defined coarse and dense punctuation, similar to that of pronotum, on lateral sides and vertex; frons with sparse punctuation, almost smooth (Fig. 5 a); antennae almost moniliform (Fig. 1 d), 1.3 mm long, not reaching the base of pronotum when directed backwards; all antennomeres slightly longer than wide, the 1 st longest of all (length: 0.21 mm); 2 nd antennomere slightly longer than the 3 rd to 10 th antennomeres (1.2 – 1.3 times); 3 rd to 10 th antennomeres subequal and the 11 th is 1.3 ‒ 1.4 times larger than the size of each of the previous eight antennomeres; labrum deeply emarginated with 17 large setae spread over the disk and the anterior margin (Fig. 5 a); mandibles symmetrical with four distinct teeth in the inner edge, two of them are more pronounced; maxillary palpus with four palpomeres, second and third subequal, more than 3 times as long as broad, apical palpomere reduced and conical; internal lobe of maxilla with clusters of setae; labium with bilobed glossae, circumscribed by hairy paraglossae; labial palpus with three palpomeres, the 2 nd being the longest and more robust, and the 3 rd the thinnest. Pronotum (Fig. 5 d): 1.2 times longer than wide (length: 0.72 mm, width: 0.59 mm), slightly narrower than head, widest anteriorly and distinctly tapering posteriad; anterior angles weakened, posterior angles marked but largely obtuse; basal margin well defined; punctuation similar to that of head, evenly distributed except for a smooth midline ranging from the anterior to the posterior parts; absence of any middle sulcus, interstices without microsculpture. Elytra (Fig. 5 e): 1.4 times wider than long (length: 0.43 mm, width: 0.59 mm), approximately 0.6 times shorter than head or pronotum [elytra length / head length: 0.57, elytra length / pronotum length: 0.60); only slightly narrower than head (0.9 times) and as wide as pronotum (0.59 mm); flat in dorsal view; humeral angles obsolete; lateral margins parallel; surface with coarse, rugose and wrinkled punctuation. Wingless. Abdomen (Figs 5 f – h): Maximum width at segment VI (0.69 mm), 1.2 times wider than elytra; tergites III ‒ VII with fine and dense punctuation within a polygonal microsculpture of transverse meshes, these more pronounced in the intersegmental membranes; sternite III with a middle carina (intercoxal carina) in the first half (Fig. 5 f) surrounded in both sides by a slight depression to accommodate the resting metacoxae; sternites III ‒ VI with fine and dense punctuation within a microsculpture of transverse meshes, substituted by irregular rows of scale-shaped microtrichia anteriad near the intersegmental membranes and by regular posteriad quadrate meshes. Sternite VII without any median impression posteriorly; sternite VIII slightly transverse, distinctly and circularly excised at posterior margin surrounded by a small glabrous area, surface with a microsculpture of polygonal meshes (Fig. 5 g); sternite IX slightly arcuate apically; genital segment in ventral view as in figure 5 h. Legs: Long and slender; forelegs with antennal cleaning organ following the general pattern of the genus (e. g. see Figs 2 e – f). Aedeagus (Fig. 6): Length 0.7 mm; sclerotized ventral process asymmetric, exceeding largely the edge of the dorsal plate (Figs 6 a – c), with a twisted leaf shape (Figs 6 a, 6 c – d); the base of the ventral process horn-shaped on each side directed inwards and upwards (Figs 6 a, 6 c); dorsal plate cylindrical ‒ shaped in dorsal view (Fig. 6 b), with fused lateral lobes (Fig. 6 c), an inverted V ‒ shaped sclerotized area towards the ventral process, supporting the upper part and the apical edge of the dorsal plate (Figs 6 b – c).	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFABFFECFF76FB725050C503.taxon	discussion	Taxonomic, distribution and bionomic remarks According to Coiffait (1982) taxonomic criteria (e. g. aedeagus with fused lateral lobes and the development of the lateral and ventral process) Domene pedroi sp. nov. should be included also in the subgenus Lathromene. Among other morphological characteristics the species shows a distinctive shape of the aedeagus. Taking into account the pieces of this structure, the new species can be easily segregated from all known Lathromene species with symmetric ventral process (D. bergidi, D. boavidae sp. nov., D. cantabrica, D. caurelensis, D. darinkae, D. eufemia, D. gridelliana, D. invernadeirensis, D. lusitanica, D. orensis, D. sistelensis, D. suidensis and D. viriatoi). Among the species with asymmetric ventral process, the closest species to D. pedroi sp. nov. seems to be D. subiasi (Outerelo, 1977). However, unlike the new species, the ventral process of D. subiasi presents a dilated axe-shaped apical tip in lateral view (Figs 44, 46 in Outerelo 1977) (leaf-shaped in the new species) and a small tooth directed laterally and obliquely backwards in the middle region on the left side (absent in the new species) (cf. Fig. 6 c and fig. 45 in Outerelo 1977). Furthermore, the apical tip is directed outward and in the new species it is directed slightly inward (cf. Fig. 6 a and fig. 46 in Outerelo 1977). Regarding morphological characteristics other than those of the aedeagus D. subiasi has a narrow and glabrous median stripe on the upper part of the head and a slight median longitudinal depression without setae in the VII sternite (see figs 40, 42 in Outerelo 1977, respectively), both characteristics are absent in the new species (e. g. see head in Fig. 5 a). The remaining three species with asymmetric ventral blade are D. scopaeella, D. gallaeciana and D. barraganensis. All can be segregated from the new species by some features of the aedeagus. In the D. scopaeella and D. barraganensis the ventral process has an apical dilated axe-shaped tip that bends inwards at a right angle [see figs 100 G – H in Coiffait (1982) and figs 10, 11 in Outerelo & Gamarra (2012), respectively]. Moreover, unlike the new species, the upper part and the apical edge of the dorsal plate in D. barraganensis is not directed towards the ventral process (see fig. 10 in Outerelo & Gamarra 2012). Concerning D. gallaeciana, the ventral process exceeds largely the edge of the aedeagus and from the middle to apex, gradually narrows, becoming filiform towards the tip (see figs 11 – 14 in Feldmann & Hernando 2005); median region presents also two contiguous lateral expansions that are absent in the new species (ventral view) (fig. 13 in Feldmann & Hernando 2005). Furthermore, in these three species there are no horn-shaped process on each side of the base of the ventral process (present in the new species and in D. subiasi). Some other morphological characteristics, like the ommatidia number, as well as male sternites VII and VIII features, allow to segregate the new species from D. scopaeella, D. gallaeciana and D. barraganensis too. For instance, D. gallaeciana and D. barraganensis are totally anophtalmous (7 ommatidae in the new species), the latter presents a middle depression on VII sternite and both have black modified setae in the sides of the posteriad excision of sternite VIII (not present in the new species). The congeners with populations closer to the new species are D. lusitanica (from Gruta da Cerâmica, Serra do Sicó, Portugal), D. viriatoi (from Buraco da Moura, Serra da Estrela, Portugal), D. hispanica (from El Cabaco, Salamanca, Spain) and D. subiasi (from Nuñomoral, Cáceres, Spain) (see also fig. 10 in Magrini & Carotti 2019). Domene pedroi sp. nov. can easily be separated from D. hispanica Outerelo, 1985 also through several characters of the external morphology (e. g. body size, shape and punctuation of the head and pronotum, absence of a median sulcus on the pronotum, femoral tooth of front leg) (see description in Outerelo 1985). The holotype specimen of D. pedroi sp. nov. was sampled by hand directly among plates of schistose rocks below thin layer of clayey brown-reddish soil on the side slopes of a country road. The locality habitat was dominated by Pinus trees with rock-roses, brooms, lavender and rosemary shrubs [Cistus ladanifer (Cistaceae), Cistus salvifolius (Cistaceae), Cytisus spp. (Fabaceae), Lavandula sp. (Lamiaceae) and Rosmarinus officinalis (Lamiaceae)]. With no more data on the distribution of the new species than that of the holotype it is provisional accepted that D. pedroi sp. nov. is endemic of Portugal. In the same region and date two endogean ground beetles were also found: Typhlocharis zaballozi Serrano & Aguiar, 2014 and Hypothyphlus lusitanicus Serrano & Aguiar, 2004 (see Serrano & Aguiar 2011, 2014). According to the assumption of Reboleira et al. (2011 b), this small size species would be more connected with the endogean environment. Indeed, D. pedroi sp. nov. was found in a typical endogean environment like D. boavidae sp. nov. (e. g. Fig. 1 a). Taking into account the reduced eyes, lack of hind wings, elongated body shape and long legs it is assumed that it is a subterranean dwelling species.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFAEFFEDFF76FA1E5028C7A2.taxon	materials_examined	Material examined. 1 male, 1 female: “ Sazes da Beira \ (Guarda — PT) \ 27. VI. 2009 \ Col. A. Serrano ” \\ “ 40 ° 20 ’ 32.44 ” N \ 7 ° 45 ’ 39.34 ” W \ 680 m alt. ” [h] \\ “ Domene \ (Lathromene) \ viriatoi \ Serrano & Boieiro \ A. Serrano det. 2018 ” [h] [t]; 1 male, 1 female: “ Vila Cova a Coelheira \ (Seia, Portugal) \ 27. IV. 2016 \ A. Serrano leg. ” \\ “ 40 ° 23 ’ 6.11 ” N \ 7 ° 43 ’ 54.99 ” W \ 402 m alt. ” \\ “ Domene \ (Lathromene) \ viriatoi \ Serrano & Boieiro \ A. Serrano det. 2018 ” [h] [t].	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
03E1AF0CFFAEFFEDFF76FA1E5028C7A2.taxon	discussion	Remarks. This species was originally described from a granitic natural cavity located in Serra da Estrela mountain foothills near Lapa dos Dinheiros (Seia, Portugal) (Serrano et al. 2015). It was postulated that D. viriatoi due to its bigger size and elongated antennae and legs would be more connected with the hypogean environment, following Reboleira et al. (2011 b). However, all the specimens here recorded were sampled among the plates of sunken schistose rocks in clayey brown-reddish soil on the side slopes of country roads close to the original locality. These additional records suggest that the species is not restricted to the deeper underground granitic cave, but lives also more superficially in the fissures of schistose rocks.	en	Serrano, Artur R. M., Aguiar, Carlos A. S. (2025): Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes. Zootaxa 5588 (3): 437-454, DOI: 10.11646/zootaxa.5588.3.3, URL: https://doi.org/10.11646/zootaxa.5588.3.3
