taxonID	type	description	language	source
03E0B947FF92FFBDFF4BFEFEFBB1F802.taxon	materials_examined	Type species: Alasia alpina Furth & Zhaurova, new species	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF92FFBDFF4BFEFEFBB1F802.taxon	description	Description. Body shape elongate narrow. Dorsal punctation generally medium-sized, sparse to moderately dense. Body length: 2.85 – 4.0 mm. Head: (Figs. 1 a – c; 4 a – b): Narrowed posteriorly (at base, in dorsal view); antennal calli distinctly delimited, especially posteriorly; antennae as long as entire body; frons in frontal view wide, broadly triangular, evidently expanded ventrally, in lateral view vertex and frons forming a 90 ° angle, frons strongly concave; vertex with medium-sized, sparse punctuation; eyes large, round, prominent, but total width (eye to eye) across head not apparently greater than pronotal anterior width, distance between inner eye margins broad (see measurements in species description); dorsal punctures sparse, medium-sized; labrum bearing four fine setiferous punctures on dorsal surface, two on each side. Pronotum: (Figs. 1 a; 4 c): Evidently narrowed at base (posteriorly); antero-lateral callosities (sensu Konstantinov 1998) not distinctly angled, only with slight sub-quadrate enlargement with setiferous pore; distinct pre-basal transverse impression, especially medially, not distinctly delimited laterally; dorsal punctation medium-sized, sparse, unevenly distributed. Elytra: (Figs. 1 a – b, 4 c): Parallel-sided; base distinctly wider than base of pronotum; humeri prominent; postbasal transverse depressed area creating basal, raised elytral bossae (calli); punctation confused medially but with some evident striae in middle and laterally, epipleura parallel-sided, equal width from near base to just before tapered apex, mostly smooth. Two baso-lateral patches of microspines on underside of each elytron adjacent to epipleuron (Fig. 4 d), i. e., binding patches (sensu Samuelson 1994). Venter: Procoxal cavities open (Fig. 1 d). [see description below of the type species for characters of the thoracic and abdominal sternites] Legs: Moderately long with spindle-shaped femora, metafemora swollen distinctly more so than pro or mesofemora, but not greatly swollen or fat. Metafemoral spring (Fig. 3 c) with dorsal lobe only gently arched - more or less straight, extended as extended arm significantly beyond ventral lobe, extended arm about 25 % of total spring length, extended arm gradually depressed apically, basal angle of ventral lobe, dorsal edge of ventral lobe descending only gradually towards apex, sclerotized recurve flange evident; spring belongs to the Phyllotreta Morpho-group (Furth & Suzuki 1998). Metatibiae with distinct sclerotised apical spine, pro and meso-tarsi with very small apical spur (almost not evident); Metatibial apex not unusually expanded or laterally compressed, with dorsal margins of extreme apex with stout spines, a distance subequal to length of tarsomere 2. First metatarsomere relatively long, almost as long as tarsomeres 2 – 5 combined; male first protarsal segment only slightly swollen; claws appendiculate.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF92FFBDFF4BFEFEFBB1F802.taxon	discussion	Remarks. Alasia is most similar to Pseudo s tenophyma Furth (described below) because of elongate somewhat parallel-sided elytra, shape of head in lateral view (90 ° angle of vertex / frons and frons concave), antennae as long as entire body, prominent, large round eyes, pronotal pre-basal transverse impression, procoxal cavities open, presence of frontal calli and elytral bossae (calli); male with first protarsal segment not distinctly swollen. However, Pseudo s tenophyma differs from Alasia in the following characters: frons narrow in shape, parallel-sided, ventrally tapered in frontal view; anterolateral callosities of pronotum distinctly angled; pronotal prebasal transverse impression evidently delimited laterally by short, longitudinal, sub-lateral carinae; pronotum quadrate or rectangular in shape, not evidently narrowed basally, with sides straight and parallel-sided; eyes larger, IOD smaller; gena laterally below eye narrower; dorsal punctation of head, pronotum more coarse, dense; elytral punctuation striate; antennal segment 3 significantly shorter than 4; pro and mesotibial apical spines very apparent; metafemoral spring shape differs (see descriptions); head shape broader basally / posteriorly. The genus Nasigona Jacoby is morphologically rather similar to Alasia but differs in having closed procoxal cavities, elytral pubescence, no elytral bossae, different pronotal shape, not narrowed basally and not having angled lateral margins, smaller eyes, considerably finer dorsal punctation throughout. Leptophysa Baly is also somewhat similar to Alasia, but differs by having pubescent elytra, no elytral bossae, pronotum laterally with straight parallel lateral margins not evidently narrowing at base, procoxal cavities closed, antennae not reaching to the apex of elytra, pronotal pre-basal transverse impression clearly delimited sublaterally by short longitudinal carina, head in lateral view with frons and vertex not at a sharp 90 ° angle and frons not strongly concave, eyes not bulging,	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF92FFBDFF4BFEFEFBB1F802.taxon	etymology	Etymology. Named for the Project ALAS (Arthropods of La Selva) at La Selva Biological Station, Heredia, Costa Rica, as a random combination of letters to be treated as feminine.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF90FFB7FF4BF8BEFC2CF937.taxon	description	Description. Body surface shiny, dark-brown to almost black, elongate, slender. Color of frons, antennal calli, antennal segments 1 – 3, most of legs lighter - yellow to light brown; extreme apex of elytra often lighter; apical 30 – 50 % of femora and all of tarsi evidently darkened (Figs. 1 a – b). Male (x 7): Body Length (Lb): Range: 2.85 – 3.45 [average = 3.17] mm. Elytral Length (Le): 1.98 – 2.45 [2.33] mm. Female (x 7): Lb: 3.5 – 4.0 [3.88] mm. Le: 2.90 – 2.97 [2.94] mm. Head: (Figs. 1 a – c, 4 a – b): Antennal calli sub-triangular, smooth, dorsally-oriented, as long as wide, small area immediately above / posterior to calli smooth; frons broad, subtriangular in frontal view, in lateral view slanted downwards sharply, sub-concave; anterofrontal and frontal ridges (sensu Konstantinov 1998) not apparent; surface of frons distinctly shagreened; genae in lateral view very broad, significantly broader than eye width in same view, genae in frontal view broad, almost as broad as head width, including eyes, in same view, widening towards mouth, in lateral view ca. one half eye width; lower edge of frons with 4 fine (two sub-lateral pair), setiferous punctures; frons dorsally prominent and protruding between antennal sockets; vertex with rather sparse, medium-sized punctation dorsally; supra-antennal sulcus well developed; several fine setiferous punctures along dorsal eye margin. Inter-ocular distance about 1.2 times the length of first antennal segment; inter-antennal distance approximately equal to distance between eye and antennal socket; eyes bulging, round; IOD (male) = 0.35 – 0.38 mm [0.368 mm], IOD (female) = 0.45 – 0.47 mm [0.46 mm]; penultimate segment of maxillary palpi swollen, sub-globose, last segment bearing a patch of five ingrown sensillae. Antennae long, filiform, usually equal to length of body; first segment swollen, 2.3 times longer than second; segment 2 shortest, swollen; 3 – 6 longest, narrow; segment length measurements (average): male: 30 - 13 - 35 - 42 - 39 - 34 - 33 - 26 - 25 - 22 - 27, female: 34 - 14 - 34 - 46 - 40 - 36 - 34 - 27 - 27 - 24 - 28. Pronotum: (Figs. 1 a, 4 c): Appearing basally (posteriorly) constricted, basal margin more or less straight, laterally rounded and thinly margined. Wp (male maximum / middle width) = 0.7 – 0.8 mm [0.764 mm – average]; Wp (female maximum / middle width) = 0.85 – 0.95 mm [0.904 mm]; Wpb (male basal width) = 0.6 – 0.7 mm [0.669 mm]; Wpb (female basal width) = 0.8 – 0.85 mm [0.813 mm]; Wpa (male anterior width) = 0.65 – 0.78 mm [0.726 mm]; Wpa (female anterior width) = 0.82 – 0.90 mm [0.87 mm]; Lp (male) = 0.45 – 0.55 mm [0.505 mm]; Lp (female) = 0.55 – 0.60 mm [0.576 mm]; antero-laterally somewhat thickened and more or less rounded, but not beveled (sensu Scherer 1962, 1983) or acutely angled; punctation coarse, dense, unevenly distributed, midline of basal half without punctures. sub-basal transverse impression evident, but weak, depressed especially in middle, ending sublaterally; postero-lateral longitudinal impressions not evident. Elytra: (Figs. 1 a – b, 4 c – d): Le (male) = 1.98 – 2.45 mm [2.33 mm]; Le (female): 2.90 – 2.97 mm [2.936 mm]. We (male) = 1.22 – 1.36 mm [1.293 mm]; We (female): 155 – 1.65 mm [1.61 mm]. Elytra glabrous; base of elytra 1.6 – 1.7 times broader than base of pronotum, extreme apex lighter brown, with punctures appearing less evident; dorsal punctures medium-dense, coarse, striate, but more confused medially near suture, more evident laterally; humeral calli strong, impunctate; basal bossae (calli) evident just postero-medial to humeri; epipleura long, originating at humeri, extending as parallel-sided, equal width, tapering just before apex; 8 – 10 setiferous punctures near apex. Ventral surface of elytra with two patches of sensillae near baso-lateral margin, the anterior patch larger and oval, posterior patch smaller and narrow-elongate. Venter: Prosternal process very narrow, short (Fig. 1 d); mesosternal process tapered, apically truncate but wider than prosternal process; metasternum somewhat inflated, with sparse longer pubescence except along median; abdominal sternites / ventrites with moderately dense pubescence; first abdominal sternite with broadly pointed process projecting anteriorly between the transverse metacoxae; ultimate abdominal sternite of male with black median line and with medial lobe. Legs: (Figs. 1 a – b): Femora long, slender, proximal portion light-brown to yellow, apically darkened; tibiae light-brown to yellow; tarsi evidently darkened. Genitalia: (Figs. 2 a – c, 3 a – b): Aedeagus with apex evenly rounded apically, four sclerotized groups of spines visible in internal sac; spermathecal ductus with apparent single coil; receptacle sub-parallel-sided with lateral margins slightly invaginated sub-basally giving appearance of base slightly enlarged, pump distinctly and sharply narrower; vaginal palpi distinctly bifurcate apically, darkened in apical one third, except lighter in extreme apex, with 3 – 4 apical and subapical setae on each side; tignum very slender, apically narrowed. Host plants. (Figs. 5 a – d, 6 a – c): Monochaetum spp., Miconia tonduzii Cogn. and Miconia sp. (Melastomataceae); Gunnera insignis (Oerst.) A. DC. (Gunneraceae) [identified by Jorge Gomez-Laurito, Escuela de Biologia, University of Costa Rica], and Macrocarpaea sp. (Gentianaceae) (K. Nishida, in litteris). All photos by K. Nishida, taken at the ALAS Project Vara Blanca site (ca. 2000 m), from 9 – 12 April 2002.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF90FFB7FF4BF8BEFC2CF937.taxon	materials_examined	Type locality. Refugio Vara Blanca in Braulio Carrillo National Park: COSTA RICA: Prov. Heredia: 6 km ENE Vara Blanca, 1950 – 2050 m, 10 ° 11 ' N 84 ° 07 ' W; INBio-OET-ALAS transect. Material examined. HOLOTYPE: Male – INBio barcode number INB 0003234047. COSTA RICA: Heredia ∶ 6 km • ENE Vara Blancaʼ 2 OOOmʼ ₁ Oο ₁₁ ʼNʼ 84 οO 7 ʼWʼ 22 Μarch 2 OO 2 ʼ leg • D • G • Γurthʼ sweep netʼ swept from Melastomataceae – deposited at INBio. PARATYPES {} = repository: COSTA RICA ∶ Heredia ∶ 6 km • ENE Vara Blancaʼ 2 OOOmʼ ₁ Oο ₁₁ ʼNʼ 84 οO 7 ʼWʼ leg • D • G • Γurthʼ sweep net ∶ 16 March 2002 [Males: INB 0003233640 {USNM}; + 3642 - 50 {+ 3642 - 46 USNM}, + 3647 - 50 INBio}; + 3652 - 61 {+ 3652 - 56 USNM, + 3657 - 61 INBio}; + 3664 - 77 {+ 3664 - 70 USNM, + 3671 - 77 INBio}; + 3682 (measured) {USNM}. Females: + 3644 {USNM}]; 18 March 2002 [Male: + 3684 (measured) {USNM}]; 19 March 2002, all sweep net except as noted [Male: + 3692 (measured), MV lite {USNM}; + 3693 (measured), MV lite {USNM}; + 3695 (measured), MV lite {USNM}; + 3725 (measured) {USNM}; + 3773 (measured) {USNM}; + 3718 {INBio}; + 3726 {INBio}; + 3729 - 31 {+ 3729 USNM, + 3730 - 31 INBio}; + 3735 {INBio}; + 3739 - 40 {+ 3739 USNM, + 3740 INBio}; + 3745 {USNM}; + 3747 {USNM}; + 3758 - 59 {+ 3758 USNM, + 3759 INBio}; + 3766 {INBio}; + 3772 {INBio}; + 3774 {INBio}; + 3776 {INBio}; + 3778 {INBio}; + 3791 - 92 {USNM}; + Female: + 3737 {USNM}; + 3741 {INBio}]; 20 March 2002 [Male: + 3905 - 06 {+ 3905 USNM, + 0906 INBio}; + 3910 {USNM}; + 3871 {USNM}; + 3896 {INBio}]; 21 March 2002 [Males: + 3971 {USNM}; + 4006 {USNM}], [Males: + 4016 - 20, leg. K. Nishida, tested negative on Gunnera {+ 4016 - 18 USNM, + 4019 - 20 INBio}]; 22 March 2002, swept from Melastomataceae [Males: + 4042 - 43 {USNM}; + 4044 - 46 {USNM}; + 4048 - 50 {USNM}; + 4052 - 78 {+ 4052 - 61 USNM, + 4062 - 78 INBio}. Females: + 4098 {USNM}]. There are 344 males and 24 females from 22 March 2002, in ethanol and, therefore, have not yet been individually assigned ALAS Project / INBio barcode labels; of these 172 males and 12 females will be deposited at INBio and 170 males and 10 females at USNM. A male and female each will be deposited at MIZA and NHM. 6 km ENE Vara Blanca, 1950 - 2050 m, 10 ° 11 ' N 84 ° 07 ' W, INBio-OET-ALAS transect - 10 Marzo 2002 [Males: IB 0003221255 {INBio}; + 1301 - 04 {+ 1301 USNM, + 1302 - 4 INBio}]; 14 Marzo 2002 [Males: + 5254 {INBio}; + 5482 {INBio}]; 16 Marzo [Males: + 4115 {INBio}]; 22 Marzo 2002 [Males: + 1418 {USNM}; + 1463 {INBio}; + 1468 - 70 {+ 1468 USNM, + 1469 - 70 INBio}; + 1493 {INBio}; + 1519 {INBio}; + 3438 {INBio}; + 3815 {USNM}. Female: + 1419 {INBio}]; 9 Abril 2002 [Males: + 0579 (mating pair) {INBio}; + 0826 {USNM}; + 0898 {USNM}; + 0907 {INBio}; + 0966 {INBio}; + 1045 (mating pair) {USNM}; + 1046 {INBio}; + 1565 - 66 {INBio}; + 1627 - 29 {INBio}; + 1706 {INBio}; + 1710 {INBio}; + 1781 - 82 {INBio}; + 1826 {USNM}. Females: + 0578 (measured) {USNM}; + 0823 {USNM}; + 0888 (measured) USNM}; + 0823 {USNM}; + 0906 {INBio}; + 0976 {INBio}; + 1047 (measured) (USNM}; + 1567 {INBio}; + 1626 {INBio}; + 1703 {INBio}; + 1705 {INBio}; + 1707 - 09 {+ 1707 USNM, + 1708 - 09 INBio}; + 1711 {USNM}; + 1827 {USNM}]; 13 Abril 2002 [Female: + 5278 {INBio}]; 16 Abril [Males: + 5295 {INBio}]; 21 Abril 2002 [Males: + 0663 {INBio}; + 0679 (mating pair) {USNM}; + 0702 {USNM}; + 1839 (mating pair) {INBio}; + 1844 {INBio}; + 1953 {INBio}; + 2002 {USNM}; + 3129 {INBio}; + 3181 {USNM}; + 3248 - 49 {INBio}; + 3541 {INBio}; + 3543 {INBio}; + 3547 {USNM}; + 3613 {INBio}; + 3621 {INBio}; + 3659 {INBio}; + 3661 {USNM}. Females: + 0635 {INBio}; + 1876 (measured) {USNM}; + 1952 {INBio}; + 1973 - 74 {INBio}; + 2001 {INBio}; + 2080 (measured) {USNM}; + 3184 (measured) {USNM}; + 3185 (measured) {USNM}; + 3542 {INBio}; + 3544 - 45 {INBio}; + 3730 {INBio}; + 3866 {USNM}].	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF90FFB7FF4BF8BEFC2CF937.taxon	distribution	Distribution. Costa Rica	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF90FFB7FF4BF8BEFC2CF937.taxon	discussion	Remarks. Pseudostenophyma modesta differs from Alasia alpina by the following characters: Color yellow to light-brown (Figs. 7 a – c). Area posterior to antennal calli punctate (Figs. 7 a, 11 a), with no median margin. Punctures on vertex more numerous and coarse (Figs. 7 a, 11 a). Frons narrower, subparallel-sided, not as broadly triangular; surface smooth (Fig. 7 c). Eyes larger; IOD distance smaller, 0.8 times the length of 1 st antennal segment (Figs. 7 a – c, 11 a). Base of pronotum wider, almost as wide as the base of elytra (Figs. 7 a, 11 b); surface slightly rugose; punctures more dense, coarse; anterolateral callosities distinctly angled (Figs. 7 a, 11 b). Elytral punctures coarser; distinctly striate, including near suture. Basal bossae (calli) more apparently convex; humeral calli slightly weaker. (Figs. 7 a – b, 11 b). Aedeagus shape different, narrowing basally in lateral view; apex of median lobe bifid in dorsal view; no spines visible on internal sac (Figs. 9 a – c). Spermathecal duct with 2 full coils (Fig. 10 a). Vaginal palpi wider posteriorly; pubescence very sparse or absent at apex (Fig. 10 b). Currently no other species are known from this new genus. Other similar genera are discussed above. A search of the USNM collections yielded two specimens from Monte Verde, Costa Rica that differ from Vara Blanca specimens only by elytral coloration. Indications from other museum specimens are that there may be additional new species from Ecuador and Colombia. Further study is required for positive identification.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF90FFB7FF4BF8BEFC2CF937.taxon	etymology	Etymology. The specific name “ alpina ” refers to the fact that all known specimens of this new species are found at rather high elevation and, therefore, are considered as alpine.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9BFFB5FF4BFF26FDAEFCDC.taxon	materials_examined	Type species. Stenophyma modesta Weise, 1921, 137: Male: Brasil: “ Rio Purus ”. “ Amazon, Roman ”, “ Hyutanahã, 2 / 215. Hõglãnd “ [hand-written]. Stenophyma modesta m. [hand-written] (Swedish Museum of Natural History) – Herein designated as the Lectotype. Male: Brasil: “ Rio Autaz ”, “ Amazon, Roman ” (Swedish Museum of Natural History) – Herein designated as the Paralectotype. These Lectotype and Paralectotype designations are being made because no primary type was mentioned in the original description by Weise (1921), thus they are syntypes. Therefore, the designation of a primary type specimen establishes a single specimen as the unique bearer of this name and the standard for its application in the future, thus avoiding any confusion.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9BFFB5FF4BFF26FDAEFCDC.taxon	description	Description. Body shape elongate, narrow. Dorsum with coarse, dense punctation. Body length: 2.7 – 3.05 mm. Head: (Figs. 7 a – c; 11 a – b): Broadly connected to pronotum at base (posteriorly); vertex with coarse, dense punctation; frons narrow, smooth, narrowing ventrally, concave in lateral view, ca. at a 90 ° angle to vertex in lateral view; antennal calli evident, delimited posteriorly; eyes large, round, prominent, but total eye to eye width not apparently greater than pronotal anterior width; IOD narrow; antennae almost as long or longer than body, antennal segments 2 and 3 shortest, 4 distinctly longest; labrum bearing four fine setiferous punctures on dorsal surface, two on each side. Pronotum: Quadrate / rectangular, not distinctly narrowed basally (posteriorly); laterally straight-sided, parallel; anterolateral callosities distinctly angled; punctation coarse, dense; pre-basal transverse impression deep, laterally delimited by apparent short, longitudinal, sublateral carinae. Elytra: Parallel-sided; base somewhat wider than base of pronotum; humeri prominent; elytral bossae distinct; punctation coarse, strongly striate throughout; epipleurae parallel-sided, equal width tapered subapically laterad of elytral apex. Two basolateral patches of microspines on underside of each elytron (Fig. 11 c). Ve n te r: Procoxal cavities open (Fig. 8 c). Legs: Metafemora distinctly swollen, much more so than pro- or mesofemora; metafemoral spring (Fig. 10 c) with dorsal lobe rather strongly arched dorsally at base with extended arm strongly depressed at apex, extended arm about 17 % of total spring length, basal angle of ventral lobe obtuse, dorsal edge of ventral lobe descending sharply towards apex, recurve flange evident, most similar in shape to the Phyllotreta spring Morpho-group (Furth & Suzuki 1998), but differs significantly from Alasia (above); tibiae with evident apical spines; metatibial apex not unusually expanded or laterally compressed, dorsal margins with only a short distance with stout spines (subequal to length of tarsomere 2); metatarsus with first tarsomere distinctly longer than in pro or mesotarsi, first metatarsomere relatively long, almost as long as tarsomeres 2 – 5 combined; first foretarsal segment not swollen in males; claws appendiculate.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9BFFB5FF4BFF26FDAEFCDC.taxon	discussion	Remarks. Pseudostenophyma is similar to Alasia in general body shape, antennal length, lateral view of frons, but differs in IOD; narrow gena laterally below eye; pronotal shape, pronotal anterolateral angles; distinctly striate elytral punctation throughout; male and female genitalia, and metafemoral spring form (see also Remarks for Alasia alpina above). It is also clearly very different from Stenophyma elegans (Fig. 12 a – c) [see Remarks section for S. elegans].	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF99FFB0FF4BFB8EFC0BFD32.taxon	description	Description. Color entirely yellow to light brown (Figs. 7 a – c), but rarely with darkening along elytral suture and even laterally on elytra and narrowly on pronotal lateral margins (Figs. 8 a – b). Elongate slender. Male (x 6): Lb: 2.70 – 2.85 [average = 2.77] mm; Le: 1.95 – 2.05 [1.99] mm. Female (x 6): Lb: 2.90 – 3.05 [2.96] mm. Le: 205 – 2.30 [2.16] mm. Head: (Figs. 7 a – c, 8 a – b, 11 a): Antennal calli apparent, subtriangular with smooth surface, area immediately posterior smooth, raised; antennae long, approximately as long as body; segment length measurements (average): male: 32 - 11 - 19 - 39 - 32 - 33 - 29 - 27 - 26 - 24 - 29, female: 32 - 11 - 18 - 34 - 32 - 31 - 28 - 26 - 25 - 24 - 29; segment 1 very long, segment 2 smallest, segment 3 considerably longer than 2, 4 and 5 subequal, very long, distinctly longer than others, each approximately equal to segments 1 – 3 combined; frons in lateral view somewhat concave, in frontal view flattened, surface smooth, anterofrontal and frontal ridges (sensu Konstantinov 1998) not apparent; genae in lateral view under eye dorso-ventrally considerably narrower (ca. one third) than eye width in same view and tapering ventrally near mouth; in frontal view frons and clypeuslabrum tapering ventrally towards mouth; eyes large, round, bulging; IOD (male) 0.25 – 0.30 mm, IOD (female) 0.26 – 0.29 mm; antennal calli apparent, triangular in shape, smooth; vertex with coarse, moderately dense punctation; maxillary palpi pointed apically with penultimate segment somewhat swollen but not more than basal segment; Pronotum: (Figs. 7 a – b, 8 a – b, 11 b): Sub-quadrate shape; anterolateral callosities strongly angled; lateral margins straight / parallel, not constricted basally; basal margin straight; transverse pre-basal impression distinct, delimited sub-laterally by short longitudinal carinae; punctation coarse, moderately dense; WP (male): 0.72 – 0.78 mm; WP (female): 0.73 – 0.85 mm. LP (male): 0.50 – 0.55 mm; LP (female): 0.52 – 0.55 mm. Elytra: (Figs. 7 a – b, 8 a – b, 11 b): Le (male): 1.95 – 2.05 mm; Le (female): 2.05 – 2.30 mm. We (male): 0.59 – 0.62 mm; We (female): 0.65 – 0.67 mm. Striate with coarse punctures, striae distinct across elytra; striae interstices smooth surface; humeri distinct; basal bossae (calli) evident just postero-medial to humeri; epipleura long, originating near humeri, extending as parallel-sided, equal width, tapering just before apex. Venter: prosternum without extension between coxae (Fig. 8 c); procoxae sub-conical; mesocoxae subspherical; metasternum slightly inflated in lateral view with sparse long setae laterally. abdominal sternites with sparse pubescence. Legs: (Fig. 7 a – b, 8 a – b): Femora long, slender metatibial apex not unusually expanded or laterally compressed; Genitalia: (Figs. 9 a – c, 10 a – b): Aedeagus in ventral view apically slightly tapered and rounded, in lateral view internal sac separating dorsum and venter, in dorsal view apex truncate with slight invagination of apical margin; spermathecal ductus with apparent double coils, receptacle slightly larger basally, gradually narrowing apically to pump; vaginal palpi unicolorous, apically tapered, not evidently bifurcate, with one subapical seta on each side; tignum very slender, apically narrowed. Host plants. Unknown. However, in Costa Rica, the ALAS Project Phases I – IV collected specimens through fogging of Pentaclethra macroloba (Willd.) O. Ktze. (Fabaceae), Virola koschnyi Warb. (Myristicaceae), as well as a few other trees. The ALAS Project Phase V (2001 – 2005) collected specimens using several techniques, including trapping by Malaise, flight intercept, sweep net, yellow pan, fogging from dates ranging from 9 February through 21 April and from elevations ranging from 300 – 1550 meters (Furth, unpublished).	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF99FFB0FF4BFB8EFC0BFD32.taxon	materials_examined	Type locality. Brazil: Amazonas, Rio Purus: Hyutanahã, 2 Feb. 1915; Rio Autáz, Oct.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF99FFB0FF4BFB8EFC0BFD32.taxon	distribution	Distribution. Brazil (Amazonas); Costa Rica; Heredia Province, La Selva Biological Station, 10 ° 26 ' N 84 ° 01 ' W (Furth et al., 2003)	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF99FFB0FF4BFB8EFC0BFD32.taxon	discussion	Remarks. Somewhat similar to Alasia alpina, but differs as mentioned in Remarks for that species. Differs from A. alpina, especially by head in frontal view with frons narrower and tapered ventrally toward mouth; eye size; pronotal anterolateral angles; pronotal lateral margins; pronotal pre-basal impression; prosternite projection; elytral punctation / striae; metafemoral spring shape.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF99FFB0FF4BFB8EFC0BFD32.taxon	etymology	Etymology. The new name for this genus just indicates that this is not the same as the true Stenophyma as discovered when examining the holotype of S. elegans Baly (see below).	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFB0FF4BFB3BFCB6FCFF.taxon	materials_examined	Type species: Stenophyma elegans Baly, 1877: 176, by monotypy	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFAEFF4BFBECFF19FDC2.taxon	materials_examined	Holotype. Male: Type, H. T. [circular label with red rim, typical NHM type label]; Bresil [hand-written on small rectangular green label]; 327 [printed? on square blue label]; Stenophyma elegans Baly, Brazil [handwritten on large rectangular green label, typical from Baly collection]. Type specimen was not removed from original card, i. e., observations and measurements were made from original position.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFAEFF4BFBECFF19FDC2.taxon	description	Description. Elongate narrow; overall color yellow to light brown; basal ca. 20 % of elytra and 75 % of lateral margins darker brown. Lb = 3.78 mm Head: Eyes round, bulging, very prominent, eye to eye width distinctly wider than pronotum; IOD = 0.42 mm; vertex smooth, impunctate; frons very short, not concave, not angled from vertex in lateral view, i. e., in same plane; antennal calli flat but evident, triangular; longitudinal frontal ridge raised, evident; gena below eye very narrow; maxillary palpomeres not evidently swollen; antennae distinctly longer than body; segments 1 – 3 yellow, 4 – 11 dark brown and somewhat thickened; segmental lengths: 30 - 12 - 12 - 60 - 58 - 55 - 55 - 55 - 56 - 56 - 67 [only left antenna complete; right antenna with only segments 1 – 6]. Pronotum: Rectangular; lateral margins more or less straight / parallel-sided, slightly narrower at base, distinctly narrower than elytral base; pre-basal transverse impression deep, straight, laterally fading into base before postero-lateral angles; antero-lateral callosities only slightly thickened at setal pore, not at all angled; surface smooth, impunctate. WP (middle) = 0.92 mm; WP (anterior) = 0.94 mm; WP (base) = 0.90; LP = 0.78 mm. Elytra: Parallel-sided; dorsally flattened, elytral bossae (calli) not apparent; distinctly striate throughout with fine, dark punctures, striae ending sub-apically leaving apex smooth; epipleura yellow / light brown, mostly equally wide / parallel-sided until sub-apex, except evidently widened below humerus, with weakly punctured, median, undulating, longitudinal stria. Le = 2.75 mm; We = 1.50 mm. Ve n te r: Procoxal cavities closed; apical sternite (male) typically concave with median, longitudinal, thin, dark line; metasternum rather flat, not inflated. Legs: Metafemur distinctly swollen; all tibiae with evident apical spine; first tarsomere of proleg slightly swollen (no female for comparison); all tibiae with evident apical spines; metatibiae in lateral view with apical 25 % expanded, laterally compressed, with row of stout spines on inner and outer margins; metatarsus with first tarsomere very long, distinctly longer than all remaining metatarsomeres combined, pro and meso first tarsomeres distinctly shorter; claws appendiculate. Host. Unknown. Genitalia: Not dissected.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFAEFF4BFBECFF19FDC2.taxon	materials_examined	Type locality. Brazil.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFAEFF4BFBECFF19FDC2.taxon	distribution	Distribution. Brazil.	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
03E0B947FF9CFFAEFF4BFBECFF19FDC2.taxon	discussion	Remarks. Stenophyma elegans is the type species of this now monotypic genus. The type specimen is housed in the Natural History Museum (London). It is distinctly different from Alasia and Pseudostenophyma by lateral view of head (vertex / frons angle); enhanced prominence of eyes; antennal segment proportions; lack of distinct elytral bossae (calli); pronotal surface; pronotal shape; pronotal prebasal transverse impression; metatibial apex shape / spines. Somewhat reminiscent also of Glenidion Clark and Huarinillasa Bechyné	en	Furth, David G., Zhaurova, Kira M. (2010): Two new flea beetle genera: Alasia alpina gen. et sp. nov. from a Costa Rican cloud forest and Pseudostenophyma gen. nov. from Brazil (Coleoptera: Chrysomelidae: Alticinae). Zootaxa 2679: 32-50, DOI: 10.5281/zenodo.199276
