identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E087FBFFF05975FED4A5D9FBE6F836.text	03E087FBFFF05975FED4A5D9FBE6F836.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maurocarpilius Ossó & Gagnaison & Bailleul 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  
Maurocarpilius 
n. gen
.
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            <p>urn:lsid:zoobank.org:act: 4A10F6F4-A9CA-4C7A-B679-E10D5266A77C</p>
            <p>  TYPE SPECIES. —  Maurocarpilius binodosus n. gen., n. sp. by monotypy  ETYMOLOGY. — From mauri, the name given to the people who </p>
            <p>lived in Mauritania Province, during the Roman Empire, which is the part of the current Morocco where the studied specimens were recovered. Gender: masculine.</p>
            <p>DIAGNOSIS. — Carapace transversely ovate, wider than long, L/W ratio about 0.72, smooth. Vaulted in both senses, mainly anteriorly. Maximum width at half-length of carapace. Front slightly subtriangular, bilobed, strongly downturned. Orbits small, rounded, supraorbital margin entire, slightly rimmed. Anterolateral margin strongly convex, acute, bearing two small nodes, one at half-length level and a second at the angle with the posterolateral margin. Posterolateral margin concave, entire, acute in anterior half, starting about 60° respect to the axis. Posterior margin slightly convex. Regions not defined. Branchiocardiac grooves defined by muscle scars. Gastric pits visible. Ventral features and appendages no preserved.</p>
            <p>REMARKS</p>
            <p> Arose in the early Eocene carpiliids are defined by a transversely ovate and broad carapace with well demarcated margins, a usually smooth surface of carapace, anterolateral margins entire or with blunt teeth or nodes, and from poorly to not defined regions (e.g. Karasawa &amp; Schweitzer 2006: 43; Davie et al. 2015: 1074; Schweitzer et al. 2018: 2). The inexpressiveness of the dorsal carapaces of members of  Carpiliidae , often leaves comparisons between their different genera and species reduced to subtle details, as the number of anterolateral teeth, if present, to length/width or fronto-orbital ratios, shape of them, or the angle degrees of posterior margins. Feldmann et al. (2011: t. 3) in the revision of Palaeocarpilius A. Milne- Edwards, 1862, summarized the aforementioned characters of all the genera placed within  Carpiliidae , which will be used in the following comparisons as well. Therefore, Maurocarpilius n. gen. is compared with all the known carpiliid genera as follow. </p>
            <p> Braggicarpilius Beschin, Busulini &amp; Tessier, 2015, from the Ypresian of northern Italy differs from Maurocarpilius n. gen. in having entire anterolateral margins, and convex posterolateral margins (see Beschin et al. 2015; Schweitzer et al. 2018: 3, fig. 3.1). An array of carpiliids, such as Bryocarpilius Feldmann, Schweitzer, Bennet, Frantescu, Resar &amp; Trudeau, 2011, from the Eocene;  Ocalina Rathbun, 1929 , from the Eocene; Palaeocarpilius, from the Eocene to Miocene; and  Paraocalina Beschin, Busulini, De Angeli &amp; Tessier 2007 , from Eocene, all of them with wide Tethysian and also Atlantic distribution; Proxicarpilius Collins &amp; Morris, 1978, from the Eocene of Pakistan; and Tethyscarpilius De Angeli &amp; Alberti, 2016, all those genera clearly differ from Maurocarpilius n. gen., in having the anterolateral margins armed from four, to up to nine nodes or teeth, instead of only two small nodes in the new genus and different ratios (see Feldmann et al. 2011: t. 3), and furthermore, in the case of Ocalina, in having a tuberculated dorsal surface (see Feldmann et al. 2011; 345-346, fig. 13; Rathbun 1929; A. Milne-Edwards 1862: 51-53; Beschin &amp; De Angeli 2006: 11-23, figs 2-5, pls 1-4; Beschin et al. 2007: 42-43, pl. 6, figs 1b, b; Collins &amp; Morris 1978: 972; Schweitzer et al. 2004: 100-102, fig. 6; Schweitzer et al. 2018; De Angeli &amp; Alberti 2016).  Carpilius Desmarest, 1823 , from the early Eocene to recent, differs from Maurocarpilius n. gen. in having only one anterolateral tooth instead of two, and a short but well-developed oblique carina that extends from that tooth (epibranchial) to the dorsal surface (see Desmarest 1823: 227-228; Schweitzer et al. 2018: 3, fig. 2a-b). Corallicarpilius De Angeli &amp; Ceccon, 2015, from the early Eocene of northern Italy, differs clearly from Maurocarpilius n. gen. in having a broader and more ovate carapace, without distinction between the posterolateral and posterior margin, and in having the dorsal surface ornated with irregular tubercles, instead of smooth in the new genus (see De Angeli &amp; Ceccon 2015: 125-126, fig. 4; Schweitzer et al. 2018: 5, fig. 3, 3a-b). Eocarpilius Blow &amp; Manning, 1996 from the Eocene of eastern North America and westernmost Tethys, differs from Maurocarpilius n. gen. in having a longer ovoid carapace, anterolateral margins entire with only a small protuberance at the corner with antero- and posterolateral margin, posterolateral margins straight, and by its punctulate surface (see Blow &amp; Manning 1996: 20-22, pl. 5, figs 2a, 2b; Feldmann et al. 1998: 11-12, figs 13-14; Schweitzer et al. 2018: 5, fig. 3, 3a-b). Holcocarcinus Withers, 1924, from the Eocene of Nigeria, differs clearly for the new genus in having the carapace with two marked transverse dorsal ridges (see Withers 1924: 94, pl. 5, figs 1-2; Schweitzer et al. 2018: 5, fig. 4.1). Laticarpilius Feldmann, Schweitzer, Bennet, Frantescu, Resar &amp; Trudeau, 2011, from the Eocene of Egypt, differs from Maurocarpilius n. gen. in having a broader carapace, and a less downturned and completely triangular front (see Feldmann et al. 2011: 342-344, fig. 12; Schweitzer et al. 2018: 5, figs 4, 2a, 2b). Liopsalis von Meyer, 1862 from the Eocene of western Tethys and India, differs from the new genus in having a completely entire margins, and a triangular front not lobed (see von Meyer 1862: 161- 163, pl. 17, figs 9-12; Stoliczka 1871 pl. 5, figs 6, 6a-b; Vía 1969: 231-237, fig. 28, t. 23, figs 3-4; Feldmann et al. 2011: 344-345; Schweitzer et al. 2018: 5, fig. 4, 3). Lovaracarpilius Beschin, De Angeli, Checchi &amp; Zarantonello, 2016 from the Eocene of northern Italy, differs from Maurocarpilius n. gen. in having shorter anterolateral margins, convex posterolateral margins, strongly punctulate surface and a triangular front (see Beschin et al. 2016: 55-56, fig. 47, pl. 9, fig. 4A-B; Schweitzer et al. 2018: 5, fig. 4, 4a-b). Oscacarpilius Artal &amp; Van Bakel, 2018 from the northern of Iberian Peninsula, differs from the new genus in having a completely entire anterolateral margin, rounded lateral corners and a densely pitted surface (Artal &amp; Van Bakel 2018: 23-29, figs 1-2). </p>
            <p> As mentioned above, the Moroccan specimens match perfectly with the general diagnosis of  Carpiliidae . However, as can be seen in the comparisons, they do not fit completely with the characters or ratios of any of the aforementioned genera. Therefore, we propose the new genus, Maurocarpilius n. gen., to accommodate them. </p>
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	https://treatment.plazi.org/id/03E087FBFFF05975FED4A5D9FBE6F836	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ossó, Àlex;Gagnaison, Cyril;Bailleul, Julien	Ossó, Àlex, Gagnaison, Cyril, Bailleul, Julien (2020): First report of Early Eocene Decapods in Morocco: description of a new genus and a new species of Carpiliidae (Decapoda: Brachyura) with remarks on its paleobiogeography. Geodiversitas 42 (4): 47-56, DOI: 10.5252/geodiversitas2020v42a4
03E087FBFFF2597AFF1FA4F9FD5BFC4A.text	03E087FBFFF2597AFF1FA4F9FD5BFC4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maurocarpilius binodosus Ossó & Gagnaison & Bailleul 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Maurocarpilius binodosus n. gen., n. sp. (Fig. 3) </p>
            <p>urn:lsid:zoobank.org:act: 647DD93A-97B4-4C0F-B651-BD01DCC0B193</p>
            <p> MATERIAL AND MEASUREMENTS (IN MM). — Holotype. ULB-IV- A(1a), L = 17; W = 23; FOW = 12.5; F = 7.</p>
            <p>Paratypes. ULB-IV-A(1b), L = 16; W = 23; FOW = 11; F = 6.5. — ULB-IV-A(1c) (deformed), L = 19.5; W = 25.5; FOW = 13; F = 7.5.</p>
            <p>TYPE LOCALITY AND HORIZON. — Tamalout section, northern face of the Jbel Talouit, North of Kelâa M’Gouna (East of Ouarzazate, Maroc). Ait Ouarhitane Formation, Ypresian (early Eocene).</p>
            <p>ETYMOLOGY. — In allusion to the two anterolateral nodes of the anterolateral margin.</p>
            <p>DIAGNOSIS. — As for the genus.</p>
            <p>DESCRIPTION</p>
            <p>Carapace transversely ovate, wider than long, L/W ratio about 0.72; surface of carapace smooth. Carapace vaulted transversely, strongly vaulted longitudinally in the anterior part of carapace. Maximum width at half-length of carapace, between the first and second (epibranchial) anterolateral teeth. Front strongly downturned, bilobed, slightly rimmed, forms a very flattened inverted isosceles triangle; the two median lobes, with the inner orbital lobes, gives a quadrilobate appearance; slightly arched and produced in dorsal view. Orbits small, rounded, supraorbital margin entire, slightly rimmed only at the inner half of the supraorbital margin; inner orbital tooth blunt, outer orbital tooth short, acute subtriangular; visible dorsally. Anterolateral margin largely convex, acute, bearing two small nodes, one at half-length level, weakly visible, and a second more prominent (epibranchial tooth) at posterolateral junction. Posterolateral margin concave, entire, acute rounded rim in anterior half, starting about 60° respect to the axis. Posterior margin slightly convex. Regions not defined; cardiac region bounded by the branchiocardiac grooves. Gastric pits visible. Branchiocardiac grooves defined by muscle scars. Ventral features and appendages no preserved.</p>
            <p>REMARKS</p>
            <p> The dorsal carapace and the general outline of  Maurocarpilius binodosus n. gen., n. sp. , greatly recall several species present in the early Eocene, which present unarmed anterolateral margins and a smooth carapace, but differing, however, in the absence or presence of subtle, but distinctive characters, that precludes a congeneric relationship. </p>
            <p> For instance, Braggicarpilius  marginatus Beschin, Busulini &amp; Tessier, 2015 , from the Ypresian (early Eocene) of northern Italy, differs from the new species by its more produced front, completely unarmed and rimmed anterolateral margin, and convex posterolateral margins. In contrary,  Maurocarpilius binodosus n. gen., n. sp. presents two anterolateral nodes and </p>
            <p>1 2 3 4 5 6 7 8 9 10</p>
            <p> their posterolateral margins are clearly concave (Beschin et al. 2015: 82, fig-text. 11, t. 5, fig. 1; Schweitzer et al. 2018: 3, fig. 3.1; Fig. 4A). Carpilius petreus Beschin, Busulini, De Angeli &amp; Tessier, 2007, from the middle-late Ypresian of northern Italy, differs from the new species because presents only one anterolateral tooth (epibranchial), instead of two in  Maurocarpilius binodosus n. gen., n. sp. , and by the presence of the short oblique carina, typical for the genus, that extends from that tooth to the dorsal surface, which is absent in the new species, and in having a shorter posterior margin (Beschin et al. 2007: 41-42, pl. 7, fig. 7a-b, 8a-b; Fig. 4B). Eocarpilius ortegai Artal &amp; Van Bakel, 2018, from the Ilerdian (early Ypresian) of northeastern of the Iberian Peninsula, presents a different L/W ratio, about 0.77/080 vs 0.73 in the new species. As well, their anterolateral margins are unarmed, with only a small node at the anterolateral/posterolateral angle, and its dorsal surface is densely covered by small granules, instead of the two clear anterolateral nodes and completely smooth dorsal surface of  Maurocarpilius binodosus n. gen., n. sp. , which makes the difference between the two species (Artal &amp; Van Bakel 2018: 30-34, figs 3-6; Fig. 4C, D). Oscacarpilius rotundus Artal &amp; Van Bakel, 2018, also from the Ilerdian of northeastern of the Iberian Peninsula, shows a very similar dorsal and outline pattern than  Maurocarpilius binodosus n. gen., n. sp. , however, differs from the latter in having a different L/W ratio, 0.80 vs 0.73, completely entire antero- and posterolateral margin, without nodes or sharp angles and by its dorsal surface pitted (Artal &amp; Van Bakel 2018: 23-29, figs 1-2; Fig. 4E, F). </p>
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	https://treatment.plazi.org/id/03E087FBFFF2597AFF1FA4F9FD5BFC4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ossó, Àlex;Gagnaison, Cyril;Bailleul, Julien	Ossó, Àlex, Gagnaison, Cyril, Bailleul, Julien (2020): First report of Early Eocene Decapods in Morocco: description of a new genus and a new species of Carpiliidae (Decapoda: Brachyura) with remarks on its paleobiogeography. Geodiversitas 42 (4): 47-56, DOI: 10.5252/geodiversitas2020v42a4
