taxonID	type	description	language	source
03E48782925CE978D48BFD9388EFFD2A.taxon	materials_examined	Additional material examined: 1 ex., Makhedonía, Vermion, above Seli, 1500 m, mixed forest, sifted, 11. IV. 1998, leg. Assing (cAss); 8 exs., Makedhonía, Kavála, Pangéo, peak region, 1900 m, 28. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782925CE978D48BFE2488E1FD8F.taxon	materials_examined	Additional material examined: 1 ex .., Thessalia, Ossa Oros, NE Spilia, near refuge, 39 ° 48 N, 22 ° 41 E, 1450 m, N-slope with scattered Abies, grass and ferns, 22. VII. 2004, leg. Assing (cAss); 11 exs., Makhedonía, Olympos, above Refuge A, 40 ° 05 N, 22 ° 22 E, 2460 m, grassy cirque, under stones, 25. VII. 2004, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782925CE978D48BFC258AFCFB8F.taxon	materials_examined	Additional material examined: 1 ex., Fthiotis, ca. 20 km SSE Lamia, Oros Kallidromo, 38 ° 44 N, 22 ° 32 E, 1250 m, N-slope, under stones on meadow, 7. IV. 2001, leg. Assing (cAss); 1 ex., NW- Evvoia, Oros Lichas, 38 ° 51 N, 22 ° 54 E, 525 m, 8. IV. 2001, leg. Assing (cAss); 3 exs., Evvoia, Oros Dirfys, N Kato Steni, SE peak, 38 ° 37 N, 23 ° 52 E, 1120 m, 8. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782925DE979D48BFC7C8A40FBBC.taxon	materials_examined	Additional material examined: 4 exs., Makedhonía, Kavála, Pangéo, road from Akrovouni to ski resort, oak forest, 500 m, with Lasius fuliginosus, 28. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782925DE979D48BFBFC8AB3FB3F.taxon	materials_examined	Additional material examined: 1 ex., Makedhonía, Sérrai, Vrontous, above Lailias, 1800 m, 25. V. 1999, leg. Assing (cAss); 3 exs., same data, but 1750 m, beech forest (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782925DE97AD48BFB638A86F98F.taxon	materials_examined	Holotype 3: GR. [1] Flórina, 20 km SSW Flórina, Oros Vitsi, N-slope, 1850 - 1900 m, 40 ° 38 ' 32 N, 21 ° 22 ' 46 E, 22. V. 2005, V. Assing / Holotypus 3 Cousya dimorpha sp. n. det. V. Assing 2005 (cAss). Paratypes: 733, 10 ♀♀: same data as holotype (OÖLL, cAss); 433, 4 ♀♀: same data, but leg. P. Wunderle (cWun). D e s c r i p t i o n: Very small species, 1.9 - 2.4 mm (abdomen fully extended). Coloration: forebody uniformly dark brown to blackish; abdomen blackish with the apex slightly paler; legs brown; antennae dark brown, with the basal two antennomeres often slightly paler. Habitus as in Figs 10 - 11. Head approximately as wide as long; eyes moderately large and protruding from lateral outline of head, slightly shorter than postocular region in dorsal view; microsculpure distinct; puncturation very fine and sparse, barely noticeable. Mouthparts as in Fig. 12. Antenna with antennomeres I and II of subequal length and about twice as long as wide; III much shorter than II and only slightly longer than wide; IV-X of increasing width and increasingly transverse; IV almost 1.5 times as wide as long; X approximately twice as wide as long; XI about 1.5 times as long as wide and slightly longer than the combined length of IX and X. Pronotum transverse, approximately 1.25 times as wide as long and 1.25 times as wide as head; maximal width approximately in the middle; posterior angles weakly marked; microsculpture and puncturation similar to those of head; pubescence along midline directed posteriad. Elytra with microsculpture and puncturation more pronounced than those of head and pronotum, surface matt. Width and length of elytra polymorphic. Macropterous specimens (Fig. 10): elytra 1.3 - 1.4 times as wide and at suture approximately 1.1 times as long as pronotum; hind wings fully developed. Brachypterous specimens (Fig. 11): elytra about 1.2 - 1.3 times as wide and at suture 1.0 - 1.1 times as long as pronotum. Hind wing development dimorphic, either fully developed or of reduced length and only slightly longer than elytra. Abdomen subparallel, in macropterous specimens distinctly (0.8 - 0.9 x) narrower than elytra, in brachypterous specimens approximately as wide as elytra; tergites III-V with shallow anterior transverse impression; tergite VI without such impression; microsculpture distinct; puncturation fine and sparse; posterior margin of tergite VII with palisade fringe in both morphs; tergite VIII without appreciable sexual dimorphism, its posterior margin weakly convex in both sexes. 3: posterior margin of sternite VIII strongly convex and with long thin marginal setae, especially in the middle (Fig. 13); median lobe of aedeagus and apical lobe of paramere as in Figs 14 - 17. ♀: posterior margin of sternite VIII broadly convex, sometimes with shallow concavity in the middle, and with row of moderately stout modified marginal setae (Fig. 18); spermatheca as in Figs 19 - 21. E t y m o l o g y: The name (adjective) refers to the dimorphism of the wings. C o m p a r a t i v e n o t e s a n d s y s t e m a t i c s: Cousya dimorpha is readily distinguished from its Western Palaearctic congeners by its small size, the distinctly transverse pronotum, the wing dimorphism, the relatively short antennae and maxillary palpi, as well as by the morphology of the genitalia. In body shape, as well as in the morphology of the mouthparts, the antennae, and the genitalia, C. dimorpha resembles species of the genus Zoosetha MULSANT & REY, from which it is distinguished by the much finer puncturation of head and pronotum. However, the character constellation of this species raises the question if Cousya MULSANT & REY and Zoosetha should really be considered distinct genera. D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from the peak of the Oros Vitsi, a mountain inhabited by several species with restricted distributions or even local endemics: Pareudectus vitsiensis ASSING & WUNDERLE, an undescribed species of Coryphiodes BERNHAUER, Lathrobium vitsiense, Quedius bernhaueri, Geostiba torisuturalis, and an undescribed species of Tectusa BERNHAUER. The fact that C. dimorpha was not found on any of the neighbouring mountains suggests that the species may have a restricted distribution, too. The types were collected by sifting grass roots and moss on a north slope at an altitude of approximately 1900 m. Seven type specimens (13, 6 ♀♀) are macropterous, 22 (1433, 8 ♀♀) are brachypterous.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829250E976D48BFF7C88D0FB18.taxon	materials_examined	Holotype 3: GR. [12] N-Pindos, ca. 40 km NNE Konitsa, Oros Arénes, 1900 m, 40 ° 17 ' 39 N, 20 ° 55 ' 22 E, 27. V. 2005, V. Assing / Holotypus 3 Tectusa recta sp. n. det. V. Assing 2005 (cAss). Paratypes: 233, 3 ♀♀: same data as holotype (cAss); 933, 2 ♀♀: same data, but leg. P. Wunderle (cWun). D e s c r i p t i o n: 2.5 - 3.0 mm (abdomen fully extended); habitus as in Fig. 22. Coloration: forebody uniformly blackish brown, with the abdominal apex slightly paler; legs reddish brown, with the femora slightly darker; antennae reddish brown, usually gradually darkened towards apex. Head approximately as wide as long; eyes small, composed of less than 20 ommatidia, and weakly protruding from lateral outline of head, less than half the length of postocular region in dorsal view; microreticulation of integument distinct; puncturation very fine and sparse, barely noticeable (Fig. 23). Antenna with antennomeres I and II of subequal length and about twice as long as wide; III somewhat shorter than II and about 1.5 times as long as wide; IV weakly transverse; IV-X of increasing width; IX almost twice as wide as long; X longer than IX and about 1.5 times as wide as long; XI of ovoid shape and shorter than the combined length of IX and X. Pronotum distinctly convex in cross-section, transverse, approximately 1.1 times as wide as long, and about 1.15 times as wide as head; maximal width approximately in the middle; posterior angles weakly marked; midline without impression; microsculpture and puncturation similar to those of head; pubescence along midline directed posteriad, in anterior 1 / 6 of midline sometimes anteriad, and on either side of midline laterad (Fig. 23). Elytra about as wide and at suture about 0.6 times as long as pronotum; microsculpture pronounced; puncturation fine, but somewhat more distinct than that of head and pronotum; pubescence short and more or less depressed (Fig. 23). Hind wings reduced. Abdomen slightly wider than elytra, widest at segments V / VI (Fig. 22). Anterior impressions of tergites III-V moderately shallow, that of tergite VI very shallow; microsculpture distinct; puncturation fine and sparse; posterior margin of tergite VII without palisade fringe; tergite VIII without appreciable sexual dimorphism. 3: posterior margin of sternite VIII strongly convex and with relatively short thin marginal setae (Fig. 24); median lobe of aedeagus at base of ventral process with pronounced folds (see arrows in Figs 25 - 27), ventral process in lateral view straight, not curved (Figs 25 - 28). ♀: posterior margin of sternite VIII broadly and weakly convex, marginal setae not distinctly modified (Fig. 29); spermatheca as in Figs 30 - 31. E t y m o l o g y: The name (Lat., adjective) refers to the straight ventral process of the median lobe of the aedeagus. C o m p a r a t i v e n o t e s: Greece is inhabited by numerous endemic species of Tectusa, most of which, however, are undescribed. Among the described species, the geographically closest congeners are T. vrontousensis ASSING & WUNDERLE from the Vrontous range near Séres and T. meschniggi (BERNHAUER) from the Olympos range. From all known species of Tectusa, the new species is separated especially by the distinctive shape of the median lobe of the aedeagus. From T. vrontousensis, it is additionally distinguished by its distinctly smaller eyes (in T. vrontousensis with more than 30 ommatidia), the lighter coloration of the legs and antennae, the much finer and more indistinct puncturation of the forebody, the absence of a median impression on the pronotum, the shallower anterior impression of the abdominal tergite VI, as well as by the much longer duct of the spermatheca. From T. meschniggi, it is also separated by the finer puncturation and the darker coloration of the forebody, by the darker femora, the more convex pronotum, by the posteriorly more strongly convex male sternite VIII, and by the much longer spermathecal duct. For illustrations of the genitalia of T. vrontousensis and T. meschniggi see ASSING & WUNDERLE (2001) and ASSING (2004 a), respectively. D i s t r i b u t i o n a n d b i o n o m i c s: ThetypelocalityissituatedintheOros Arénes, which is part of the Oros Grámos range, very close to the Albanian border. The specimens were sifted from grass roots, moss, and debris close to the edge of snowfields at an altitude of 1900 m (Fig. 32).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829252E976D48BFB1C883BFACF.taxon	materials_examined	Additional material examined: 22 exs., Makedhonía, Kavála, Pangéo, road from Akrovouni to ski resort, oak forest, 500 m, with Lasius fuliginosus, 28. V. 1999, leg. Assing (cAss); 1 ex., Fthiotis, ca. 20 km SSE Lamia, Oros Kallidromo, 38 ° 45 N, 22 ° 28 E, 940 m, 2. IV. 2001, leg. Assing (cAss); 1 ex., Pelopónnisos, Agios Petros, Vourvouni, 920 m, Quercus ilex forest, 22. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829252E977D48BFAD38BF7FE55.taxon	materials_examined	Additional material examined: Mainland Greece: 8 exs., Fthiotis, Oros Íti, W-slope, 38 ° 49 N, 22 ° 14 E, 1450 m, Abies forest, 10. IV. 2001, leg. Assing, Wunderle (cAss, cWun); 1 ex., Fthiotis, SSE Lamia, Oros Kallidromo, 38 ° 44 N, 22 ° 31 E, 1250 m, 7. IV. 2001, leg. Assing (cAss); 1 ex., Fthiotis, Parnassos Oros, ski resort, 38 ° 33 N, 22 ° 35 E, 1760 m, Abies wood, 15. IV. 2000, leg. Assing (cAss); 1 ex., Parnassos, Lilea, 20. V. 1997, leg. Wolf (cSch). Pelopónnisos: 3 exs., Panahaiko, above Ano Kastritsi, 38 ° 15 N, 21 ° 52 E, 1500 m, 28. III. 1997, leg. Assing (cAss); 5 exs., 10 km E Kalavrita, Aroania range, 38 ° 01 N, 22 ° 10 E, 1450 m, 29. III. 1997, leg. Assing, Wunderle (cAss, cWun); 1 ex., Killini range, S Trikalon, 37 ° 59 N, 22 ° 27 E, 1300 m, 31. III. 1997, leg. Assing (cAss); 2 exs., Killini, Ano Trikala, 1450 m, pine litter, 7. VI. 1996, leg. Wunderle (cWun); 2 exs., Erimanthos, above Kalendzi, 37 ° 57 N, 21 ° 47 E, 1500 m, 27. III. 1997, leg. Assing (cAss); 1 ex., Erimanthos, above Kalendzi, 37 ° 57 N, 21 ° 47 E, 1200 m, 27. III. 1997, leg. Assing (cAss); 1 ex., Erimanthos, Kalendzi, 1350 m, Abies forest, 27. III. 1997, leg. Wunderle (cWun); 5 exs., 20 km S Derveni, Sarandapiho, 38 ° 02 N, 22 ° 23 E, 1250 m, 3. IV. 1997, leg. Assing (cAss); 1 ex., Parnon range, 1100 - 1450 m, Pinus, Abies, 13. VI. 1996, leg. Wunderle (cWun). C o m m e n t s: The species had not been recorded since its original description, which is based on type material from the Pelopónnisos.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829257E973D48BFA078B0AF99D.taxon	materials_examined	Material examined: 1 ex. (det. Wunderle), Corfu, Zigos env., 400 m, Quercus ilex forest with moss, sifted, 24. IX. 1994, leg. Wunderle (cWun).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829257E973D48BFB0D882AFB60.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, 7 km NE Lálas, 700 m, Quercus, 4. X. 2004, leg. Schawaller (SMNS).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829257E973D48BFBB68A9FFBED.taxon	materials_examined	Material examined: 72 exs., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss); 7 exs., Pelopónnisos, Diafkoto, 26. III. - 4. IV. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829257E973D48BFCD08B19FC6A.taxon	materials_examined	Material examined: 2 exs., Makedhonía, Pieria range, above Skotina, 900 - 1000 m, mixed forest, sifted, 9. IV. 1998, leg. Assing (cAss); 1 ex., Pelopónnisos, Agios Petros, 1100 m, oak forest with Erica arborea, 22. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829257E973D48BFA8888F3FA1A.taxon	materials_examined	Material examined: 3 exs., Thessalia, Kato Olympos, E Kallipefki, 1300 m, under bark of Abies, 6. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829248E96CD48BFF7C883BFEB7.taxon	materials_examined	Material examined: 2 exs., Pieria range, Katafygi, 1450 m, pine forest, sifted, 9. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829248E96CD48BFDCC8B3BFC8D.taxon	materials_examined	Material examined: 2 exs. (det. A. Zanetti), Thásos, above Primos, 40 ° 44 N, 24 ° 39 E, 16. X. 1980, leg. Malicky (cAss); 1 ex. (det. A. Zanetti), Pelopónnisos, Erimanthos, above Kalendzi, 37 ° 57 N, 21 ° 47 E, 1200 m, 27. III. 1997, leg. Assing (cAss). C o mme n t: The species was previously known from Albania, Bulgaria, and Cyprus, but had not been recorded from Greece.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829248E96CD48BFEFB883BFE29.taxon	materials_examined	Material examined: 14 exs., Thessalia, Stómio, ca. 10 m, flood plain forest, 5. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829248E96CD48BFE7188EFFDAC.taxon	materials_examined	Material examined: 1 ex., Makhedonía, Kavála, Pangéo, near ski resort, 1700 m, beech forest, 24. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829248E96DD48BFCEC8AE7FDE3.taxon	materials_examined	Holotype ♀: Greece, Rhodes, Kolimbia, Loutani river, 10 mNN, 14.04.1994, leg. J. Frisch / Holotypus ♀ Aphaenostemmus rhodicus sp. n. det. V. Assing 2005 (MNHUB). Paratype ♀: same data as holotype (cAss). D e s c r i p t i o n: 3.8 - 4.0 mm; habitus as in Fig. 60. Coloration: head and pronotum castaneous; elytra yellowish brown; abdomen dark brown to blackish brown; legs and antennae brown. Head weakly transverse, approximately 1.05 times as wide as long; eyes large and bulging, about twice as long as postocular region in dorsal view; puncturation moderately coarse and dense, interstices in central dorsal area on average as wide as diameter of punctures, in lateral areas narrower; microsculpture absent (Fig. 62). Antenna as in Fig. 63. Pronotum approximately 1.10 times as wide as head and 1.20 times as wide as long, maximal width a short distance anterior to middle; posterior angles obtusely marked; puncturation coarser and denser than that of head, interstices distinctly narrower than diameter of punctures; microsculpture absent (Fig. 62). Elytra 1.35 - 1.40 times as wide and at suture about 1.8 times as long as pronotum; puncturation similar to that of pronotum (Fig. 61); microsculpture absent. Hind wings fully developed. Abdomen slightly wider than elytra; puncturation very fine, barely noticeable; integument with pronounced microreticulation rendering the surface almost completely matt; posterior margin of tergite VII with palisade fringe. 3: unknown. E t y m o l o g y: The name (Lat.: adj.) is derived from Rhódos, where the type locality is situated. C o m p a r a t i v e n o t e s: In the Palaearctic region sensu LÖBL & SMETANA (2004), the genus Aphaenostemmus PEYERIMHOFF 1914 previously included only five species (HERMAN 2001). Two species had become known from the Western Palaearctic: A. bordei PEYERIMHOFF 1914 (Algeria, Tunisia) and A. testaceus (KOCH 1936) (Egypt, Saudi Arabia, Oman). From the former, A. rhodicus is separated by the lighter coloration of the head, pronotum, legs, and antennae, by slightly smaller size, by the relatively narrower and smaller pronotum, by the shorter and more slender antennae, and by the much denser puncturation of the head and pronotum. The habitus of A. bordei is illustrated in ASSING (2005 f). Based on the details specified by KOCH (1936), the new species is separated from A. testaceus, of which I have seen no material, by the bicoloured body (in A. testaceus uniformly rufotestaceous), larger size, and by the denser puncturation of the forebody. D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from one locality situated in the Greek island Rhódos, off the southwest Anatolian coast. The types were apparently collected on a river bank; further bionomic data are not available.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFC0F887AFC63.taxon	materials_examined	Material examined: 6 exs., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFA0F89C6F9A1.taxon	materials_examined	Material examined: 2 exs., Pelopónnisos, Parnon, 1100 m, Abies, 26. IX. 2004, leg. Bense (SMNS, cAss); 1 ex., E Chelmós, 2 km S Zarúkhla, 1100 m, 23. IX. 2004, leg. Bense (SMNS). C o m m e n t: The species had been reported from Greece before, but is apparently very rare.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFB0F8B65FB63.taxon	materials_examined	Material examined: 2 exs., Fthiotis, ca. 30 km SW Lamia, bank of Inachos river near Perivoli, 38 ° 50 N, 22 ° 05 E, 470 m, 5. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFB8F8856FBE3.taxon	materials_examined	Material examined: 1 ex. (det. J. Frisch), Corfu, lake Korission, edge of pond, 28. IX. 1994, leg. Wunderle (cWun).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFA8F887AFAE3.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFD0F8855FD63.taxon	materials_examined	Material examined: 33 exs., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829249E96DD48BFC8F887AFCE3.taxon	materials_examined	Material examined: 2 exs., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924AE96FD48BFF7C8D3DFC2F.taxon	materials_examined	Holotype 3: GR. Pelopónnisos, 40 km SE Tripoli, SW Ag. Petros, 1160 m, 37 ° 18 ' 51 N, 22 ° 31 ' 10 E, 22. III. 1997, V. Assing / Holotypus 3 Xantholinus moreanus sp. n. det. V. Assing 2005 (cAss). Paratypes: 13: same data as holotype (cAss); 13, 3 ♀♀: same data, but 37 ° 18 ' 49 N, 22 ° 30 ' 37 E, 1100 m (cAss); 13: GR-Peloponnes, 40 km SO Tripoli, Ag. Petros-Karies, 1100 m, Quercus, 22.3.97, P. Wunderle (cWun); 13: GR: Peloponnes, Lakonia, 40 km SO Tripoli, Strasse zw. Agios Petros u. Karies, 1100 m, W-Hang, alte / Eichen, Erica arborea, Moos, 37 ° 18 ' 49 N, 22 ° 30 ' 37 O, 22.3.1979 [sic], leg. L. Zerche (DEI); 13, 1 ♀: GR: Peloponnes, Lakonia, 40 km SO Tripoli, Strasse zw. Agios Petros u. Karies, 1160 m, W-Hang, Tanne, / Eiche, Moos, 37 ° 18 ' 51 N, 22 ° 31 ' 10 O, 22.3.1979 [sic], leg. L. Zerche (DEI); 13: GR. Pelopónnisos, nördl. Taygetos, W Perivolia, 1300 m, 37 ° 08 ' 44 N, 22 ° 16 ' 03 E, 24. III. 1997, V. Assing (cAss); 13: GR: Peloponnes, Lakonía, Taigetos- Geb., Giátrissa bei Kastania, 1090 m, 21. IV. 1998 / 36 ° 51 ' 07 N, 22 ° 23 ' 12 E, unter Steinen, leg. Behne (DEI); 1 ♀: GR, S-Peloponnes, E Kardamíli, oberhalb Saidóna, Abies, 900 m, leg. W. Schawaller, 28. IX. 2004 (SMNS); 13: GR: Peloponnes, Arkadía, Parnon-Geb., SW Kastanitsa, 1380 m / 37 ° 14 ' 25 N, 22 ° 37 ' 16 E, 16. IV. 1998, Kiefern-Tannen-Wald, leg. Zerche (cAss); 233, 1 ♀: GR: Peloponnes, Arkadía, Parnon-Geb., SW Kastanitsa, 1385 m, 37 ° 12 ' 07 N, 22 ° 37 ' 52 E / 19. IV. 1998, Tannen-Kiefern-Wald, Schneefelder, leg. Zerche & Behne (DEI, cAss); 1 ♀: GR: Peloponnes, Lakonía, Parnon-Geb., Agios Pétros, 1150 m, 37.16.44 N, 22.35.03 O / 18. IV. 1998, Kiefern-Tannen-Wald, Schneefelder, leg. Zerche (DEI); 1 ♀: GR: Ahaïa, Panahaiko, oberh. Shuli, Plateau, 29. III. 2000, 1500 m, Igelsträucher, / Schneerand, 38 ° 11 ' N, 21 ° 51 ' E, leg. Zerche (DEI);. D e s c r i p t i o n: Species of intermediate size, 8.0 - 9.2 mm (abdomen fully extended). Coloration variable, ranging uniformly rufous to distinctly bicoloured, with the head and abdomen blackish brown to blackish, and pronotum and elytra rufous; legs rufous; antennae with the basal three antennomeres rufous, antennomeres IV-X reddish brown, and XI apically testaceous. Habitus of forebody as in Fig. 47 - 48. Head approximately 1.25 times as long as wide, weakly dilated posteriad; eyes very small, approximately 1 / 5 the length of postocular region; puncturation moderately sparse and relatively fine; microsculpture absent. Pronotum 0.9 times as wide as head and approximately 1.45 times as long as wide; dorsal rows of punctures composed of about 10 punctures; microsculpture absent. Elytra short, approximately 0.55 times as long as pronotum; puncturation relatively dense, rather coarse, and well-defined, interstices on either side of suture much narrower than diameter of punctures; microsculpture absent. Hind wings reduced; when fully unfolded, apices just protruding from under the elytra. Abdomen with fine and moderately sparse puncturation; microsculpture distinct, transverse everywhere; posterior margin of tergite VII without palisade fringe. 3: posterior segments without conspicuous modifications. Aedeagus basally with two series of distinctly scerotised spines, one with ca. 15 long spines, and one with more numerous smaller spines (Figs 49 - 53); long spines present also in anterior part of aedeagus (Fig. 54). E t y m o l o g y: The name (Lat., adj.) is derived from Morea, the Latin name of what is the Pelopónnisos today. C o m p a r a t i v e n o t e s: The geographically closest species of the subgenus Helicophallus is X. bulgaricus COIFFAIT from the Bulgarian Stara Planina, which, based on the original description, as well as on an examination of the holotype from the Coiffait collection (MNHNP) and a non-type male from the Stara Planina, is distinguished from the new species by the distinctly microsculptured elytra with almost indistinct puncturation, by the less slender (shorter and relatively broader) head and pronotum, by the smaller aedeagus (0.75 - 0.80 times the length of that of X. moreanus), and by the shorter and less massive proximal spines in the aedeagus. For an illustration of the aedeagus of X. bulgaricus see figure 98 A in COIFFAIT (1972). C o m m e n t: KORGE (1973) reports X. bulgaricus also from the Parnon range in the Pelopónnisos, stating he had found only negligible differences between the material from the Parnon and the original description of X. bulgaricus. As emphasised above, however, there are differences both in external morphology and in the internal structures of the aedeagus. In addition, there are other arguments suggesting that the populations from the Stara Planina and from the Pelopónnisos represent distinct species. First, aedeagal character divergence is generally low in the subgenus, so that striking differences like those in many other species groups of Xantholinus are not to be expected. Second, both X. bulgaricus and X. moreanus are micropterous and have small eyes, from which one can infer that dispersal power is low and the distributions are very likely to be restricted. Finally, despite much increased collecting activity in various parts of Greece in the past decade, no further records of Helicophallus have become known. Korge´s material was not examined, but it may hypothesised that it is conspecific with X. moreanus. It was not possible to examine the aedeagus of the – evidently slightly teneral – holotype of X. bulgaricus (" Bulg. St. planina, Midzur, 2150 - 2168 m, 26.10.1968, P. Beron leg. / Préparation microscopique / Holotype / Xantholinus (Helicophallus) bulgaricus H. Coiffait 1971 "), since it is apparently kepts separately (see labels). The transverse microsculpture of the head emphasised in the original description is visible only in the lateral parts; the dorsal median area is free of microsculpture. Fortunately, a male from the Stara Planina in the collections of the DEI was available for examination, which confirmed that X. bulgaricus and X. moreanus are indeed different species. The collection data of this specimen are: Stara Planina, Etropolska Planina, Murgana peak, 42 ° 42 N, 24 ° 03 E, 1600 - 1630 m, 11. V. 2001, leg. Zerche & Behne. D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from several localities in the Pelopónnisos (Map 1), where it is probably endemic, as can be inferred from the absence of records from other Greek localities, as well as from the reduced eyes and wings. It was collected in oak and mixed forests (oak, fir, maple, pine) at altitudes of 1100 - 1300 m.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924CE968D48BFF7C8A47FBA9.taxon	materials_examined	Holotype 3: GR. West-Kreta, Samariatal, 20. III. 1973, leg. Meybohm & Fülscher / Holotypus 3 Xantholinus creticus sp. n. det. V. Assing 2005 (cAss). D e s c r i p t i o n: Relatively small species, 6.8 mm (abdomen fully extended). Coloration: head blackish; pronotum dark brown, with the anterior and posterior margins diffusely paler; elytra brown; abdomen dark brown with the tergal margins reddish; legs and antennae rufous. Forebody as in Fig. 55. Head almost 1.3 times as long as wide, with subparallel lateral margins; eyes less than one third the length of postocular region; puncturation sparse and relatively fine; microsculpture absent. Pronotum 0.9 times as wide as head and approximately 1.45 times as long as wide; dorsal rows of punctures composed of 9 - 10 punctures; microsculpture absent. Elytra approximately 0.7 times as long as pronotum; puncturation relatively sparse and well-defined; microsculpture absent. Hind wings present, but possibly not fully developed. Abdomen with very fine and sparse puncturation; microsculpture distinct, on tergites III-VII transverse, on tergite VIII predominantly isodiametric; posterior margin of tergite VII with palisade fringe. 3: posterior segments without conspicuous modifications. Aedeagus with membranous tube, two large black spines, and two series of rather long brownish spines (Figs 56 - 59). E t y m o l o g y: The name (Lat., adj.) refers to Crete, where the species may be endemic. C o m p a r a t i v e n o t e s: Xantholinus creticus is distinguished from all other species of the genus by the internal structures of the aedeagus. In X. linearis (OLIVIER), which too has two long black spines in the internal sac, these spines are distinctly angled, the additional spines in the aedeagus are much shorter, and the head and pronotum have distinct transverse microsculpture. D i s t r i b u t i o n a n d b i o n o m i c s: The type locality, the Samaria valley, is located in the south of western Crete. Bionomic data are not available.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924CE968D48BFBF08881FA23.taxon	materials_examined	Material examined: Mainland: 13, Thessalia, Pilion, 5 km NE Vólos, Stagiate, 500 - 700 m, 15. V. 1996, leg. Schulz & Vock (cAss); 233, Voiotia, Oros Elikonas, Kiriaki, 38 ° 23 N, 22 ° 50 E, 900 m, 4. IV. 2001, leg. Assing (cAss). Evvoia: 13, Oros Lichas, 38 ° 52 N, 22 ° 53 E, 620 m, 9. IV. 2001, leg. Assing (cAss). Kérkira (Corfu): 333, Kérkira, 11. &. 13. X. 1987 (cAss); 13, Paleokastritsa, ruderal meadow, 14. X. 1987 (cAss). Pelopónnisos: 13, Agios Mammas, 36 ° 37 N, 22 ° 56 E, 3. X. 1980, leg. Malicky (cAss); 13, Olympia, 1. IV. 1986, leg. Assing (cAss); 13, pass 25 km S Kalávrita, 1100 m, 5. X. 2004, leg. Schawaller (cAss); 13, Sparta, stream bank, 2. IV. 1986, leg. Assing (cAss); 13, Parnon, 4.5 km W Karies, 37 ° 17 N, 22 ° 28 E, 930 m, oak litter sifted, 19. IV. 1999, leg. Behne (DEI). C o m m e n t: The species was previously considered to be endemic to the Greek island Kérkira (= Corfu) (BORDONI 1975, 1990), but is apparently widespread in southern and western Greece (Map 2).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924DE969D48BFB1D8AA8FA90.taxon	materials_examined	Material examined: 3 exs., Levkás, Sivota, beach debris, 27. IX. 1993, leg. Assing (cAss); 8 exs., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924DE969D48BFB86887AFB1D.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924DE969D48BFA98887AFA0A.taxon	materials_examined	Material examined: 35 exs., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924DE969D48BFA178AA8F98D.taxon	materials_examined	Material examined: 2 exs., Levkás, Sivota, beach debris, 27. IX. 1993, leg. Assing (cAss); 3 exs., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFF7C8B8CFE2D.taxon	materials_examined	Additional material examined: 2 exs., Fthiotis, ca. 20 km SSE Lamia, Oros Kallidromo, 38 ° 45 N, 22 ° 32 E, 1250 m, N-slope, under bark of Abies, 7. IV. 2001, leg. Assing (cAss); 22 exs., Pelopónnisos, Aroania, E Kalávrita, 1450 m, Abies forest, 29. III. 1997, leg. Assing (cAss); 6 exs., Pelopónnisos, E Chelmós, 2 km S Zarúkhla, Abies, Pinus, 1100 m, 23. IX. 2004, leg. Schawaller (SMNS, cAss); 4 exs., Chelmós, ski resort, 1800 m, leg. Bense (SMNS). Comment: The species is not listed for Greece by LÖBL & SMETANA (2004), but was reported from there by ASSING & WUNDERLE (2001).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFA7588F3F9AF.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Pieria range, above Skotina, 900 - 1000 m, mixed forest, sifted, 9. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFAFE883BFA35.taxon	materials_examined	Material examined: 2 exs., Thessalia, Stómio, ca. 10 m, flood plain forest, 5. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFE4C88EFFDA7.taxon	materials_examined	Material examined: 4 exs., Makedhonía, Sérrai, Vrontous, S Lailias, 1400 m, nest of Formica rufa, 25. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFDCB8B3AFC80.taxon	materials_examined	Material examined: 3 exs., Pelopónnisos, Chélmos, ski resort, Abies forest, 1600 m, 22. IX. 2004, leg. Schawaller (SMNS); 6 exs., same data, but 1800 m, leg. Bense (SMNS, cAss); 5 exs., Pelopónnisos, Taygetos, Neohóri, 1100 m, Platanus, Abies, 30. IX. 2004, leg. Bense (SMNS, cAss). Comment: The species was previously known neither from Greece nor from adjacent countries (LÖBL & SMETANA 2004).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFCEB8A9EFBFA.taxon	materials_examined	Material examined: 1 ex., Flórina, 40 km NE Flórina, Oros Voras, 40 ° 53 N, 21 ° 48 E, 1650 m, 22. IV. 2000, leg. Assing (cAss); 1 ex., Pella, 40 km NE Flórina, Oros Voras, 40 ° 54 N, 21 ° 50 E, 1950 m, 22. IV. 2000, leg. Assing (cAss); 1 ex., Thessalia, Pilion Oros, 4 km SW Zagora, 39 ° 25 N, 23 ° 05 E, 450 m, 3. IV. 1998, leg. Assing (cAss); 1 ex., Fthiótis, 40 km W Lamia, bank of Sperchios river, 38 ° 56 N, 21 ° 59 E, 16. IV. 2000, leg. Assing (cAss); 11 exs., Pelopónnisos, Taygetos, Manganiári source, 1000 m, Abies, Pinus, 29. IX. 2004, leg. Schawaller (SMNS, cAss). Comment: Quedius umbrinus had been reported from Greece before. Interestingly, the material from the Pelopónnisos is characterised by distinctly longer and larger elytra, as well as by darker coloration than material seen from other European regions.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924EE96AD48BFB268B8CFAB3.taxon	materials_examined	Additional material examined: 1 ex., Fthiotis, Oros Oxia, SW Paleochori, 38 ° 50 N, 21 ° 56 E, 1500 m, N-slope, 11. IV. 2001, leg. Assing (cAss). Comment: The species is not listed for Greece by LÖBL & SMETANA (2004), but was reported from there by ASSING & WUNDERLE (2001).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFB1388E9FA82.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, Erimanthos, above Kalendzi, 1200 m, Abies forest, 27. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFA64887AF9BF.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFC2E88B1FB94.taxon	materials_examined	Material examined: 1 ex., Fthiotis, ca. 30 km SW Lamia, bank of Inachos river near Perivoli, 38 ° 50 N, 22 ° 05 E, 470 m, 5. IV. 2001, leg. Assing (cAss); 1 ex., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss); 1 ex. (det. Wunderle), Crete, Preveli, 0 m, 16. X. 1991, leg. Wunderle (cWun).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFAEE8B2AFA04.taxon	materials_examined	Material examined: 17 exs. (det. Wunderle), W-Crete, Derés env., 400 m, edge of stream, 5. & 10. X. 1991, leg. Wunderle (cWun).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFDD48B03FD4F.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, Erimanthos, above Kalendzi, 1500 m, grass, moss, and leaf litter sifted, 27. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFD53883BFCC1.taxon	materials_examined	Material examined: 8 exs., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFF7C88DDFEDF.taxon	materials_examined	Material examined: 73 exs., Thessalia, Kato Olympos, E Kallipefki, 1300 m, under bark of Abies, 6. IV. 1998, leg. Assing (cAss); 1 ex., Pelopónnisos, E Chelmós, 2 km S Zarúkhla, 1100 m, 23. IX. 2004, leg. Bense (SMNS).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFEC388F3FE51.taxon	materials_examined	Material examined: 5 exs., Thessalia, Kato Olympos, E Kallipefki, 1300 m, under bark of Abies, 6. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFE5988F3FDD4.taxon	materials_examined	Material examined: 6 exs., Makedhonía, Olympos, NW Stavros refuge, 1450 m, under pine bark, 7. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782924FE96BD48BFB94887AFB0F.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, N Kalávrita, bank of Vouraikos river, 680 m, 30. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFD748B64FCAF.taxon	materials_examined	Material examined: 1 ex., Thessalia, Ossa Oros, NE Spilia, 39 ° 49 N, 22 ° 40 E, 1130 m, mixed Abies and Fagus forest, 22. VII. 2004, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFCF388EFFC49.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, northern Taygetos, E Perivolia, 1300 m, creek valley, 24. III. 1997, leg. Assing (cAss); 1 ex., Pelopónnisos, SE Zakháro, 6 km E Néa Figalia, 700 m, 2. X. 2004, leg. Bense (SMNS).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFC568855FBCC.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Dráma, Falakró, above ski resort, 1750 - 1850 m, 26. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFB2C882AFB47.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, Stimfalia, 800 m, lakeshore, 24. IX. 2004, leg. Schawaller (SMNS).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFAAB8D36FAFA.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss); 1 ex. (det. J. Vogel), Crete, Lefka Ori, Mt. Psári, 1700 m, VI. 2002, leg. Feldmann (cFel).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFA26887AFA64.taxon	materials_examined	Material examined: 1 ex., Thessalia, Ossa Oros, 6 km W Karitsa, 750 m, beech forest, 4. IV. 1998, leg. Assing (cAss); 1 ex., Thessalia, Ossa Oros, E-slope, 1150 - 1200 m, beech forest, 8. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFF7C8AA8FEC7.taxon	materials_examined	Material examined: 18 exs., Thessalia, Stómio, coast laguna, 5. IV. 1998, leg. Assing (cAss); 6 exs., Makedhonía, Kavála, 60 km W Kavála, beach of Ophrynia, laguna, 30. V. 1999, leg. Assing (cAss); 1 ex., Thessalia, Stómio, ca. 10 m, flood plain forest, 5. IV. 1998, leg. Assing (cAss); 1 ex., Pelopónnisos, 10 km NE Gíthio, beach debris, 23. & 26. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFD898BD6FD34.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Kavála, Pangéo, peak region, 1900 m, 24. V. 1999, leg. Assing (cAss); 2 exs., Pangéo, near ski resort, 1700 m, beech forest, 24. V. 1999, leg. Assing (cAss); 2 exs., same data, but 1650 m, 24. & 28. V. 1999, leg. Assing (cAss); 1 ex., Pangéo, road from Akrovouni to ski resort, beech forest, 1350 m, 24. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829240E964D48BFE2B8B60FE61.taxon	materials_examined	Material examined: 1 ex., Makedhonia, NW Kavála, Pangéo, road to ski resort, 1200 m, beech forest, 28. V. 1999, leg. Wunderle (cWun); 1 ex., Makedhonia, N Drama, Falakró, road to ski resort, 1000 m, N-slope, 26. V. 1999, leg. Wunderle (cWun).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFDCC8B67FD27.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Kavála, Pangéo, road from Akrovouni to ski resort, beech forest, 1350 m, 24. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFD4B88E9FCD9.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, Erimanthos, above Kalendzi, 1200 m, Abies forest, 27. III. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFCC68B65FC5C.taxon	materials_examined	Material examined: 1 ex., Fthiotis, ca. 30 km SW Lamia, bank of Inachos river near Perivoli, 38 ° 50 N, 22 ° 05 E, 470 m, 5. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFC5C8855FBD7.taxon	materials_examined	Material examined: 5 exs., Pelopónnisos, 30 km NE Pirgos, Koumani, oak forest, 630 m, 1. IV. 1997, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFEFB88F4FE29.taxon	materials_examined	Material examined: 1 ex. (det. J. Vogel), Ipiros, Peristeri range near Athohori, 1700 - 2200 m, 4. VI. 2000, leg. Marggi (cFel).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFF7C8B39FEB7.taxon	materials_examined	Material examined: 1 ex., Thessalia, Kato Olympos, E Kallipefki, 1450 - 1500 m, beech and fir forest, sifted, 6. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFE718855FDAC.taxon	materials_examined	Material examined: 1 ex., Makhedonía, Vermion, above Seli, 1500 m, mixed forest, sifted, 11. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFA3488E8FA6F.taxon	materials_examined	Material examined: 3 exs., Fthiotis, Oros Iti, 38 ° 49 N, 22 ° 14 E, 1400 m, N-slope, subalpine meadow, 10. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFBDB8A08FB72.taxon	materials_examined	Material examined: 2 exs., Thessalia, Stómio, ca. 10 m, flood plain forest, 5. IV. 1998, leg. Assing (cAss); 3 exs., Fokís, Oros Elikonas, road from Kiriaki to Elikonas, 38 ° 23 N, 22 ° 49 E, 900 m, Nslope, sifted Abies litter and under stones, 4. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829241E965D48BFABE8A20FAF4.taxon	materials_examined	Material examined: 11 exs., Makedhonía, Kavála, Pangéo, road from Akrovouni to ski resort, oak forest, 500 m, with Lasius fuliginosus, 28. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829242E966D48BFEFB8B65FE29.taxon	materials_examined	Material examined: 1 ex., Fthiotis, ca. 30 km SW Lamia, bank of Inachos river near Perivoli, 38 ° 50 N, 22 ° 05 E, 470 m, 5. IV. 2001, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829242E966D48BFB8D8BA1FA7B.taxon	description	Slope, 1900 m, 1. IV. 1995, leg. Assing, Wunderle (cAss, cWun). C o m m e n t s: The original description of Oxypoda corcyrica is explicitly based on a female holotype from " Ipsos auf der Insel Korfu " (SCHEERPELTZ 1958), not from Levkás, as indicated by LÖBL & SCHEERPELTZ (2004); that of O. bimaculata is based on two syntypes from Cyprus (BAUDI DI SELVE 1870). An examination of the holotype of O. corcyrica revealed that it is conspecific with O. bimaculata, which is widespread in the Eastern Mediterranean. Material of this species was seen from Cyprus, Greece (see above), and from Turkey (ASSING 2004 c, in press).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829242E966D48BFE718AF6FC6D.taxon	materials_examined	Material examined: Greece: 14 exs., Levkás, Vouno peak, 1050 m, 25. IX. 1993, leg. Assing (cAss, cWun); 1 ex., Fthiotis, 30 km W Lamía, W Kalithea, 38 ° 53 N, 22 ° 06 E, 800 m, oak forest, 16. IV. 2000, leg. Assing (cAss); 4 exs., N Larissa, Kato Olympos, above Goni, 39 ° 54 N, 22 ° 27 E, 550 m, roadside, nest of Messor sp., 6. IV. 1998, leg. Assing (cAss); 7 exs., N Larissa, Kato Olympos, N Larissa, Kato Olympos, E Kallipefki, 39 ° 58 N, 22 ° 29 E, 1500 - 1580 m, 6. & 12. IV. 1998, leg. Assing (cAss, cWun); 1 ♀, Makedhonia, NW Kavála, Pangéo, near ski resort, 1650 m, 24. V. 1999, leg. Assing (cAss); 1 ex., Pelopónnisos, Ahaia, Agios Nikolaos, IV. 1999, leg. Wachtel (cAss); 2 exs., Ikaría, Nas, 37 ° 37 N, 26 ° 03 E, 10 - 100 m, sifted from grass under Mastix shrubs, 26. IV. 2003, leg. Brachat & Meybohm (cAss). C o m m e n t s: The species was originally described from " Griechenland " (KRAATZ 1855), the locality is not specified. Recent records have not become known. Oxypoda micans is somewhat similar to, but not conspecific with O. referens MULSANT & REY from Corsica. The latter is of larger average body size, slightly paler coloration, and has shorter elytra. Most importantly, however, the median lobe of the aedeagus is of different morphology (shape of internal structures, smaller crista apicalis). These differences are not very pronounced, but constant. The record from the Pangéo is somewhat doubtful, since it is based on a single female; males are required to confirm the presence of the species in northeastern Greece.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829242E966D48BFF7C8D2FFEB7.taxon	materials_examined	Material examined: 1 ex., Pelopónnisos, 7 km SE Kalávrita, 1200 m, Abies, 21. IX. 2004, leg. Bense (cAss); 1 ex., Pelopónnisos, Chelmós, ski resort, 1800 m, Abies, 22. IX. 2004, leg. Bense (SMNS).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829243E967D48BFF7C8BC5F981.taxon	materials_examined	Holotype 3: N 35 ° 11 ', E 25 ° 31 ', GR Kreta Lassithi, Selia Afhin 1000 m, Meybohm 10.3.2001 / Holotypus 3 Oxypoda cretica sp. n. det. V. Assing 2005 (cAss). D e s c r i p t i o n: 3.5 mm. Coloration: whole body pale reddish brown, with the anterior areas of abdominal tergites V-VI infuscated; legs and antennae yellowish brown. Habitus as in Fig. 64. Head 1.2 times as wide as long; puncturation dense and distinct, interstices narrower than diameter of punctures and without microsculpture (Fig. 65); eyes moderately large (Fig. 66), slightly shorter than postocular region in dorsal view. Antennae relatively long and massive (Fig. 67), antennomeres 4 - 10 distinctly coniform, antennomere X approximately 1.5 times as wide as long, XI rather large. Preapical joint of maxillary palpus slightly more than twice as long as wide. Pronotum more than 1.4 times as wide as head and more than 1.3 times as wide as long (Fig. 65); maximal width approximately in the middle; puncturation very dense, denser and less well-defined than that of head; microsculpture absent. Elytra approximately as wide as pronotum and at suture 0.65 times as long as pronotum; near lateral margins shallowly impressed (Fig. 65); puncturation coarser than that of pronotum, very dense, asperate, and ill-defined; microsculpture absent. Hind wings reduced. Legs moderately slender; metatarsus almost as long as metatibia; metatarsomere I approximately as long as the combined length of metatarsomeres II-IV. Abdomen approximately as wide as elytra, widest at segments III / IV, moderately tapering posteriad (Fig. 64); puncturation fine and rather dense on tergites III-VI, sparser on tergites VII-VIII; microsculpture absent; posterior margin of tergite VII without palisade fringe. 3: posterior margin of sternite VIII obtusely angled in the middle; median lobe of aedeagus shaped as in Figs 68 - 69, ventral process apically bifid; apical lobe of paramere very long, with two long basal setae, short apical seta, and subapical seta of intermediate length (Fig. 70). E t y m o l o g y: The name (Lat., adj.) is derived from Creta, the name of the island where the type locality is situated. C o m p a r a t i v e n o t e s: Based on external and the male sexual characters, O. cretica is closely related to O. cingulum BERNHAUER from the Middle East and to O. hatayana ASSING from central southern Anatolia. It is distinguished from the former, whose male sexual characters are unknown, by the uniformly pale coloration of the forebody (in O. cingulum the head is darker than the pronotum) and from the latter by larger body size and by the shape of the aedeagus (apically straight ventral process in lateral view). From both species, it is additionally separated by the coarser puncturation of the forebody, the absence of microsculpture, the less transverse pronotum, the lateral impressions on the elytra, the relatively longer metatarsus, and the longer first metatarsomere. For illustrations of the habitus and the sexual characters of O. hatayana see ASSING (2004 c, in press). D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from one locality in eastern Crete. The reduced hind wings and palisade fringe at the posterior margin of tergite VII suggest that it is endemic to the island.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829244E960D48BFF7C8B1FF9A4.taxon	materials_examined	Material examined: 4 exs., Kato Olympos, E Kallipefki, 39 ° 58 N, 22 ° 29 E, 1500 - 1580 m, 6. IV. 1998, leg. Assing, Wunderle (cAss, cWun); 2 exs., Evritania, W Lamia, N Karpenisi, Timfristos, ski resort, 38 ° 57, 21 ° 48 E, 1900 m, 14. IV. 2000, leg. Assing, Wunderle (cAss, cWun); 1 ex., Fokis, 45 km SSW Lamia, Oros Giona, 38 ° 40 N, 22 ° 20 E, 1700 m, 17. IV. 2000, leg. Assing (cAss). C o m m e n t s: The species was only recently described from the Greek island Lesbos (ASSING 2005 g). A comparative study of the external and sexual characters of the above material and of the holotype male of O. lesbia revealed some differences between populations (microsculpture, puncturation, shape of median lobe of aedeagus and of spermatheca), but no convincing evidence that they should represent distinct species. The intraspecific variation of the aedeagus and of the – previously unknown – spermatheca are illustrated in Figs 72 - 74 and Figs 77 - 79. The species is closely related to O. brachyptera (STEPHENS), as can be inferred from the highly similar morphology of the aedeagus (shape and internal structures of median lobe, apically bifid ventral process of the median lobe, shape and chaetotaxy of the apical lobe of the paramere, shape of the spermatheca). For comparison, the median lobe of the aedeagus of O. brachyptera is illustrated in Fig. 71. All the specimens seen so far have hind wings of reduced length (approximately 1.5 times as long as elytra and distinctly projecting from under the posterior elytra margins when unfolded). However, the presence of a palisade fringe at the posterior margin of the abdominal tergite VII, as well as the wide distribution suggest that O. lesbia may be wing-dimorphic.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829245E962D48BFF7C885BFE30.taxon	materials_examined	Holotype 3: GR-Peloponnes, Erymanthos, Kalendzi, Pass, 1500 m, Rasen, Tanne, 27.3.97, P. Wunderle / Holotypus 3 Oxypoda afimbriata sp. n. det. V. Assing 2005 (cAss). Paratypes: 2 ♀♀: GR. Pelopónnisos, Erimanthos, oberh. Kalendzi, Sattel, 1500 m, 37 ° 56 ' 38 N, 21 ° 46 ' 30 E, 27. III. 1997, V. Assing (cAss, cWun). D e s c r i p t i o n: 2.5 - 2.7 mm. Coloration: whole body reddish yellow, with the central part of tergite VI and the anterior half of tergite VII infuscated; legs yellow. Habitus as in Fig. 84. Head indistinctly transverse; puncturation dense, but very shallow and somewhat illdefined; microsculpture very shallow (Fig. 85); eyes relatively small (Fig. 86), weakly projecting from lateral outline of head, distinctly shorter than postocular region in dorsal view. Antenna as in Fig. 87, antennomere X approximately 1.5 times as wide as long. Preapical joint of maxillary palpus somewhat dilated, slightly less than twice as long as wide. Pronotum 1.20 - 1.25 times as wide as head and 1.25 - 1.30 times as wide as long (Fig. 85); maximal width approximately in or a short distance anterior to middle; puncturation very dense, ill-defined, and asperate; surface almost matt. Elytra approximately as wide as pronotum and at suture 0.85 - 0.90 times as long as pronotum (Fig. 85); surface somewhat flattened; puncturation coarser than that of pronotum, very dense, rasp-like, and ill-defined; microsculpture present; surface almost matt. Hind wings reduced, slightly projecting from under elytral hind margins when unfolded (Fig. 85). Legs relatively short; metatarsomere I longer than the combined length of the two following metatarsomeres, approximately as long as the combined length of metatarsomeres II-IV or nearly so. Abdomen approximately as wide as elytra, widest at segments V / VI, segments VII-VIII weakly tapering (Fig. 84); puncturation fine and very dense on tergite III, gradually decreasing in density from tergites IV to VIII; microsculpture absent; posterior margin of tergite VII without palisade fringe. 3: posterior margin of sternite VIII obtusely angled in the middle (Fig. 80); median lobe of aedeagus shaped as in Figs 88 - 89, ventral process apically bifid; apical lobe of paramere of similar shape and chaetotaxy as in other species of the O. brachyptera group, very long, with two long basal and two short apical setae (Fig. 90). ♀: posterior margin of sternite VIII weakly convex (Fig. 81); spermatheca as in Fig. 82. E t y m o l o g y: The name (Lat., adj.: not fimbriate) refers to the absence of a palisade fringe at the posterior margin of tergite VII. C o m p a r a t i v e n o t e s: Oxypoda afimbriata belongs to the O. brachyptera group, as is suggested by the similar external characters and especially by the morphology of the median lobe of the aedeagus (general shape, internal structures, apically bifid ventral process), the shape and chaetotaxy of the apical lobe of the paramere, and by the morphology of the spermatheca. The new species is distinguished from the widespread O. brachyptera and O. lesbia by the paler coloration, which somewhat resembles that of some species of the subgenus Deropoda, by the much coarser and more asperate puncturation of the pronotum and the elytra, by the absence of a palisade fringe at the posterior margin of tergite VII, by the shape of the median lobe of the aedeagus, and by the shape of the spermathecal duct, which is longer and more slender than in O. lesbia (see Figs 77 - 79, 82). D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from the Erimanthos range in the northwestern Pelopónnisos, Greece (Map 3). The reduced hind wings and palisade fringe at the posterior margin of tergite VII suggest that it has a restricted distribution.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829246E963D48BFE788B21F9A7.taxon	materials_examined	Holotype ♀: GR. Pelopónnisos, Panahaiko-Geb., 1500 m, oberh. Ano Kastritsi, 38 ° 14 ' 58 N, 21 ° 51 ' 32 E, 28. III. 1997, V. Assing / Holotypus ♀ Oxypoda praecisa sp. n. det. V. Assing 2005 (cAss). Paratypes: 1 ♀: same data as holotype (cAss); 233, 1 ♀, 4 exs.: GR: Ahaïa, Panahaiko, oberh. Ano Kastrizi, N-Seite, 30. III. 2000, 1550 m, gefrorenes / Gras, Schneefeld, 38 ° 14 ' 46 N, 21 ° 51 ' 37 O, leg. Zerche & Behne (DEI, cAss); 433, 1 ♀, 2 exs.: GR: Peloponnes, Ahaïa, Panahaiko-Geb., N-Seite, 1610 m / 38 ° 11 ' 00 N, 21 ° 51 ' 41 E, 26. IV. 1998, Schneefelder, leg. Zerche (DEI, cAss); 1 ♀: GR: Peloponnes, Ahaïa, Panahaiko-Geb., N-Seite, 1610 m, 38.11.00 N, 21.51.41 O, 26. IV. 1998, Schneefelder, leg. Zerche (DEI); 13, 1 ♀, 2 exs.: GR: Peloponnes, Ahaïa, Panahaiko-Geb., N Shuli, Schnee auf Wiese, N-Hang / 1450 m, 38 ° 11 ' 03 N, 21 ° 51 ' 38 E, 29. IV. 1999, leg. Behne (DEI); 1 ♀: GR: Peloponnes, Ahaïa, Panahaiko, oberh. Shuli, N-Seite, 29. III. 2000, 1465 m, Igelsträucher / Schneerand, 38 ° 11 ' 00 N, 21 ° 51 ' 36 E, leg. Zerche & Behne (DEI); 3 exs., GR: Peloponnes, Ahaïa, Panahaiko-Geb., SO Ano Kastritsi, Schnee auf Wiese, NW- Hang, 1575 m, / 38 ° 14 ' 51 N, 21 ° 51 ' 32 O, 24. IV. 1999, leg. Zerche (DEI); 233, 2 ♀♀: GR: Peloponnes, Ahaïa, 10 km O Kalávrita, Aroánia, Strasse zum Chelmos, 1450 / m, Gesiebe im Tannenwald, 38 ° 01 ' 02 N, 22 ° 10 ' 21 O, 30.3.1997, leg. L. Zerche (DEI, cAss); 2 ♀♀: GR: Ahaïa, Chelmos, Strasse von Kalavrita zum Xerokambos (Ski-Center), 1450 m, 3. IV. 2000, Abies - Wald / Schneefel- der, 38 ° 01 ' 18 N, 22 ° 10 ' 45 O, leg. Zerche (DEI, cAss); 2 ♀♀: GR: Peloponnes, Korinthia, Killini- Geb., S Ano Trikala, Schnee im Rasen, O-Hang / 2010 m, 37 ° 55 ' 53 N, 22 ° 24 ' 06 O, 1. V. 1999, leg. Zerche & Behne (cAss); 1 ♀: GR: Peloponnes, Korinthia, S Ano Trikala, Abies, Juniperus, Schnee, N- / Hang, 1645 m, 37 ° 56 ' 24 N, 22 ° 25 ' 31 O, 27. IV. 1999, leg. Zerche (DEI); 233, 1 ♀: GR: Peloponnes, Korinthía, Killini-Geb., S Trikala, 1940 m, 37.55.48 N, 22 .. 24.45 O / 24. IV. 1998, Schneefeld mit Stachelsträuchern, leg. Zerche (DEI); 13, 2 ♀♀: GR: Arkadía, Melano (n), N-Hang am Ski-Center, 2. IV. 2000, 1670 m, Abies - Wald / Schneefelder, 37 ° 39 ' 06 N, 22 ° 15 ' 43 O, leg. Zerche (DEI, cAss). D e s c r i p t i o n: 2.5 - 3.0 mm. Coloration variable: body of almost uniformly dark colour, with the head and pronotum dark brown, the elytra slightly paler brown, and the abdomen blackish with paler tergal margins and paler apex, or bicoloured, with the pronotum and elytra reddish, the head slightly darker, and the abdomen reddish with tergite VI and anterior half of tergite VII infuscated; legs yellowish to yellowish brown; antennae brown, with antennomeres I-II yellowish to reddish yellow. Habitus as in Fig. 91. Head approximately 1.1 times as wide as long; puncturation dense, but usually very shallow, ill-defined, and indistinct; microsculpture shallow (Fig. 92); eyes relatively small (Fig. 93), not projecting from lateral outline of head, distinctly shorter than postocular region in dorsal view. Antenna as in Fig. 94, antennomere X approximately 1.5 times as wide as long. Preapical joint of maxillary palpus approximately twice as long as wide. Pronotum approximately 1.25 times as wide as head and 1.30 times as wide as long (Fig. 92); maximal width approximately in the middle; puncturation very dense, much coarser than that of head, and somewhat asperate; microsculpture very shallow; surface with subdued shine. Elytra approximately as wide as pronotum and at suture 0.70 times as long as pronotum (Fig. 92); puncturation coarser than that of pronotum, very dense, and rasp-like; microsculpture indistinct; surface with some shine. Hind wings reduced, not projecting from under elytral hind margins when unfolded (Fig. 85). Legs relatively short; metatarsomere I longer than the combined length of the two following metatarsomeres, but shorter than the combined length of metatarsomeres II-IV. Abdomen approximately as wide as elytra, widest at segments VI / VII, apically weakly tapering (Fig. 92); puncturation distinct and very dense on tergite III, decreasing in density from tergites IV to VIII; microsculpture absent, interstices with distinct shine; posterior margin of tergite VII without palisade fringe. 3: sternite VIII obtusely angled in the middle (Fig. 100); median lobe of aedeagus as in Figs 95 - 98, with pronounced crista apicalis, ventral process apically not bifid; apical lobe of paramere relatively short, with one very short apical and three long basal and subapical setae (Fig. 99). ♀: posterior margin of sternite VIII weakly convex, similar to that of O. afimbriata (cf. Fig. 81); spermatheca of highly distinctive, but somewhat variable shape (Figs 83, 101 - 107). E t y m o l o g y: The name (Lat., adj.: shortened) refers to the short duct of the spermatheca. I n t r a s p e c i f i c v a r i a t i o n: A comparison of specimens from different mountain ranges revealed some variability of the spermatheca. In the material from the Panahaiko and the Chelmos, the spermathecal duct tends to be shorter, whereas in females from the Killini and the Melanon it is somewhat longer and proximally more or less smoothly curved (Figs 83, 101 - 107). However, since convincing constant differences were found neither in the morphology of the aedeagus nor in external characters and, furthermore, since the shape of the spermatheca is subject to some variation even within populations, these differences are attributed to intra- rather than interspecific variation. C o m p a r a t i v e n o t e s: In external morphology, Oxypoda praecisa somewhat resembles species of the O. brachyptera group. However, the morphology of the apically unincised median lobe of the aedeagus, the shape and chaetotaxy of the apical lobe of the paramere, as well as the shape of the spermatheca suggest that it belongs to a different species group. From O. brachyptera and O. lesbia, the new species is readily distinguished by the coarser puncturation of the pronotum and elytra, the shorter elytra, the more pronounced shine especially of the abdomen, by the absence of a palisade fringe at the posterior margin of tergite VII, as well as by the completely different shape of the spermatheca. From O. afimbriata, it is separated by the darker coloration, the less reduced shine of the pronotum and elytra, the less dense puncturation of the pronotum, the shorter elytra, the more pronounced puncturation of the abdomen, and by the different shape of the spermatheca. D i s t r i b u t i o n a n d b i o n o m i c s: The species inhabits several mountain ranges (Panahaiko, Chelmos, Killini, Melanon) in the northern Pelopónnisos, Greece (Map 3). The reduced hind wings and palisade fringe at the posterior margin of tergite VII suggest that it has a restricted distribution. The type specimens were sifted from moss, grass, litter of coniferous trees and shrubs, often near snow, and the roots of grass and herbs at altitudes of 1450 - 2010 m.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829278E95DD48BFF7C8AC6FF73.taxon	materials_examined	Type material examined: Lectotype 3, here designated: ferruginea Er., Feodosia, Krim, Retowski / 4342 / longicollis Brh det. Bernhauer / c. Eppelsh. Steind. d. / Lectotypus 3 Oxypoda longicollis Bernhauer desig. V. Assing 2006 (NHMW). Paralectotypes: 1 ♀: ferruginea Er., Feodosia, Krim, Retowski / 4343 / longicollis Brh Type det. Bernhauer / c. Eppelsh. Steind. d. / Typus (NHMW); 1 ♀: Feodosia, Krim, Retowski, ex Doubl. Epp. / longicollis Brh. Russland Type. / Chicago NHMus. M. Bernhauer Collection (FMNH); 1 ♀: 3 [sic] / Graecia, Parnass / longicollis Brh., det. Bernhauer / Cotypus Oxypoda longicollis Bernhauer / ex coll. Skalitzky / ex coll. Scheerpeltz / Oxypoda sp. det. V. Assing (NHMW). Additional material examined: 1 ♀, Ukraine, Krim (FMNH). C o m m e n t s: The original description of this species is based on syntypes from the Krim peninsula, Ukraine (one male and two females), and from the Parnass range in Greece (one female). The type specimens from both localities are not conspecific, so that a lectotype designation is mandatory and the male from the Krim peninsula is here designated as the lectotype. The female from the Parnass belongs to an unidentified – possibly undescribed – species; its general appearance is similar to that of O. brachyptera, but the spermathecal duct is shorter and of different shape. The similar morphology of the median lobe of the aedeagus, the shape and chaetotaxy of the apical lobe of the paramere, as well as the morphology of the spermatheca suggest that O. longicollis is not affiliated with species of the O. brachyptera group, but rather closely related to O. praecisa. R e d e s c r i p t i o n: Small species, 2.3 - 2.7 mm; habitus of forebody as in Fig. 108. Coloration: Reddish brown, with abdominal segment VI and the anterior half of segment VII somewhat infuscate; legs testaceous; antennae pale reddish. Head as wide as long or weakly transverse, relatively large in relation to pronotum (Fig. 108); puncturation fine and indistinct; microsculpture shallow, but distinct; eyes relatively small (Fig. 110), little more than half the length of postocular region in dorsal view. Antennae as in Fig. 109. Penultimate joint of maxillary palpus of moderate length, about 2.5 - 3.0 times as long as wide Pronotum 1.20 - 1.25 times as wide as long and only 1.15 - 1.20 times as wide as head; maximal width in anterior half; puncturation shallow and dense, somewhat ill-defined, but more distinct than that of head; microsculpture shallow (Fig. 108). Elytra slightly wider than pronotum and at suture about 0.75 times as long as pronotum; puncturation dense and distinct, much more so than that of head and pronotum (Fig. 108). Hind wings reduced. Metatarsomere I slightly longer than the combined length of II-III. Abdomen about as wide as elytra; puncturation on anterior tergites dense and distinct, gradually decreasing in density towards apex of abdomen; microsculpture absent or indistinct; posterior margin of tergite VII without palisade fringe. 3: posterior margin of sternite VIII strongly convex, but not angled in the middle (Fig. 112); aedeagus with apically unincised and, in lateral view, somewhat angled ventral process (Figs 113 - 114); apical lobe of paramere as in Fig. 111. ♀: posterior margin of sternite VIII weakly concave in the middle (Fig. 115); spermatheca with rather short duct (Fig. 116). D i s t r i b u t i o n: The species is known only from the Krim peninsula, Ukraine.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829279E95DD48BFA2C8B4FF9BC.taxon	materials_examined	Material examined: 1 ♀, Pelopónnisos, Killini range, Mosia, 4. IV. 1992, leg. Frisch (cWun). C o m m e n t: The species was originally described based on a male and a female (LIKOVSKY 1973) – not two males, as indicated in the original description – from the Pelopónnisos, but has not been recorded since.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829279E95DD48BFB668AB4FACC.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Dráma, Falakró, above ski resort, 1750 - 1850 m, 26. V. 1999, leg. Assing (cAss); 1 ex., Falakró, below ski resort, 1500 m, pine forest, 26. V. 1999, leg. Assing (cAss); 1 ex., Makedhonía, Kavála, Pangéo, peak region, 1900 m, 28. V. 1999, leg. Assing (cAss); 1 exs., Pangéo, near ski resort, 1650 m, beech forest, 28. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829279E95DD48BFDEC8BDBFD07.taxon	materials_examined	Material examined: 2 exs., Makedhonía, Kavála, Pangéo, road from Akrovouni to ski resort, oak forest, 500 m, with Lasius fuliginosus, 28. V. 1999, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829279E95DD48BFD6B8AD2FB3A.taxon	materials_examined	Type material examined: Holotype ♀: Grecia nom Etol. - Akar., O. Váltou, 7. VI. 2003, 2 ° polje NW di Perdikáki, m 1115, Giachino & Vailati / Holotypus Aleochara hellenica n. sp. det. R Pace 2004 / Aleochara hellenica n. sp. det. R Pace 2004 / Haploglossa villosula (Stephens) det. V. Assing 2005 (private collection P. M. Giachino). C o m m e n t s: The description of Aleochara hellenica is based on a female holotype from " Etol. - Akar., O. Váltou, 2 ° polje NW di Perdikáki " deposited in the collection of Mauro Giachino (Torino). The details indicated in the original description, especially the illustration of the distinctive spermatheca, gave rise to the suspicion that A. hellenica was in fact a synonym of Haploglossa villosula. A subsequent examination of the holotype confirmed that it is doubtlessly conspecific with H. villosula; hence the synonymy proposed above. Haploglossa villosula has been recorded from Greece before (ASSING 2005 g, LÖBL & SMETANA 2004). The species is widespread and common in Europe; a recent record from China (ASSING 2005 e) shows that it is present also in the Eastern Palaearctic region.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E487829279E95DD48BFEBF88E6FD8D.taxon	materials_examined	Type material examined: Syntypes: 1 ♀: Kalávryta, Morea, Holtz / Moczarskii Brh. Typus / Chicago NHMus. M. Bernhauer Collection (FMNH); 1 ♀: Val de Ropa, Corfu / Moczarskii Brh. Cotypus / Chicago NHMus. M. Bernhauer Collection (FMNH). C o m m e n t s: The original description is explicitly based on two syntypes from the Pelopónnisos and Corfu (BERNHAUER 1906). Since both type specimens are females, a lectotype designation is refrained from until males become available from the vicinity of either of the two type localities.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782927AE95FD48BFF7C886CFB7A.taxon	materials_examined	Holotype 3: GR. N Larissa, No. 11, Kato Olympos, 1500 - 1580 m, E Kallipefki, 39 ° 57 ' 34 N, 22 ° 29 ' 15, 6. IV. 1998, V. Assing / Holotypus 3 Aleochara mortuisimilis sp. n. det. V. Assing 2005 (cAss). Paratypes: 433, 3 ♀♀: same data as holotype (cAss); 333, 1 ♀: same data, but leg. P. Wunderle (cWun). D e s c r i p t i o n: Relatively small and slender species, 4.3 - 5.3 mm (abdomen fully extended). Coloration variable. Light-coloured specimens: head brown; pronotum and elytra light brown to reddish brown; abdomen reddish brown with the central tergal areas infuscate; legs testaceous; antennae brown, with the basal 2 - 3 antennomeres paler. Darkcoloured specimens: head blackish; pronotum and elytra castaneous, with the elytra at least slightly paler than the pronotum; abdomen dark brown to blackish brown, with the tergal margins slightly paler; legs rufotestaceous; antennae brown, with the basal 2 - 3 antennomeres paler. Habitus as in Fig. 33. Head approximately as wide as long; eyes moderately large and protruding from lateral outline of head, postocular region approximately 1.5 times the length of eyes in dorsal view (Fig. 34); integument with pronounced and dense microreticulation, matt; puncturation fine, shallow, and relatively sparse; pubescence long and suberect. Antenna with antennomeres I-III approximately 2.5 times as long as wide and of subequal length, IV weakly transverse; IV-X of gradually increasing width and increasingly transverse; X approximately 1.5 times as wide as long; XI approximately as long as the combined length of IX and X (Fig. 33). Pronotum transverse, approximately 1.25 times as wide as long and 1.40 - 1.50 times as wide as head; maximal width approximately in the middle; posterior angles weakly marked, almost obsolete; microsculpture almost as pronounced as that of head; puncturation more distinct and somewhat denser than that of head; pubescence more or less depressed, along midline directed posteriad, on either side of midline laterad. Elytra approximately 1.1 (bachypterous morph) to 1.18 (macropterous morph) times as wide and at suture 0.80 - 0.85 times as long as pronotum; microreticulation similar to that of head and pronotum; puncturation of similar density as that of pronotum, but somewhat granulose; pubescence directed diagonally latero-caudad. Hind wing length dimorphic, either almost completely reduced or fully developed. Abdomen with segments III-V of subequal width, posterior segments (from segment VI) slightly tapering; tergites III-V with moderately deep anterior impressions, tergite VI with shallow impression; microsculpture very shallow, integument more shiny than that of forebody; puncturation relatively fine and sparse, in anterior impressions of tergites III-V very fine and very sparse; posterior margin of tergite VII with palisade fringe in both morphs; tergites III and VII with sexual dimorphism. 3: tergite III with weakly elevated median elevation (Fig. 35); tergite VII in the middle of posterior margin with broad, smoothly rounded (antero-dorsal view), and not very strongly elevated tubercle (Figs 36 - 38); posterior margin of tergite VIII convex and with very sparse setae posteriorly (Fig. 39); posterior margin of sternite VIII pointed and with moderately long marginal setae (Fig. 40); median lobe of aedeagus as in Figs 41 - 43. ♀: posterior margin of tergite VIII broadly convex and with more numerous short setae posteriorly (Fig. 44); posterior margin of sternite VIII broadly convex and with row of short, moderately stout, modified marginal setae (Fig. 45); spermatheca as in Fig. 46. I n t r a s p e c i f i c v a r i a t i o n: The species appears to be subject to lower intraspecific variation than most of its congeners; this conclusion, however, is based only on twelve specimens from one locality. As indicated above, coloration is rather variable, whereas variation in size is moderate. The male secondary sexual characters were present in all eight males, but especially the tubercle on tergite VII may be of reduced size. E t y m o l o g y: The name (Lat., adj.: seemingly dead) refers to the observation that the species displays thanatosis when moved or touched. C o m p a r a t i v e n o t e s: Aleochara mortuisimilis is readily separated from all other species of the subgenus by the following character combination: relatively small size; pronotum less than 1.3 times as wide as long; the pronounced microsculpture especially on the head, but also on the pronotum and elytra; the relatively short elytra; the dimorphism of the hind wings; the male secondary sexual characters, as well as the shapes of the aedeagus and the spermatheca. Two species of Ceranota have been recorded from Greece: Aleochara erythroptera GRAVENHORST and A. purkynei ROUBAL. From both, A. mortuisimilis is distinguished by distinctly smaller size, the pronounced microsculpture of the forebody, the shorter and less slender antennae, the less smaller and less transverse pronotum, the much sparser and more indistinct puncturation of the forebody, the fine and sparse puncturation of the anterior impressions of the abdominal tergites, the differently shaped male secondary sexual characters (especially the shape of the tubercle on the abdominal tergite VII), and by the shape of the genitalia. From A. erythroptera it is additionally separated by the different and more uniform coloration (in A. erythroptera the elytra are ferrugineous and distinctly contrasting with the dark pronotum) and from A. purkynei by the unmodified male tergite IV. For illustrations of the habitus and the male secondary sexual characters of A. purkynei see ROUBAL (1937). D i s t r i b u t i o n a n d b i o n o m i c s: The species is known only from the peak of the Kato Olympos, northeastern Thessalia. The types were collected by sifting moss and litter of beech and fir, partly near snow fields at an altitude of 1500 - 1580 m. Interestingly, the specimens displayed thanatosis when moved or touched. Aleochara mortuisimilis is apparently the first example of wing dimorphism to become known in the subgenus: two females are macropterous, the remaining eight males and two females are brachypterous.	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
03E48782927BE95FD48BFAA188F3FAFC.taxon	materials_examined	Material examined: 1 ex., Makedhonía, Olympos, NW Stavros refuge, 1500 m, under pine bark, 7. IV. 1998, leg. Assing (cAss).	en	Assing, V. (2006): New species and records of Staphylinidae from Greece, with two new synonymies (Insecta: Coleoptera). Linzer biologische Beiträge 38 (1): 333-379, DOI: 10.5281/zenodo.5433223
