taxonID	type	description	language	source
0DEEEC1347F05890B02BF2C79DAC3A73.taxon	diagnosis	Diagnosis. The main external distinctive feature of the species is the brownish-grey ground colour of forewings and hindwings. Lygephila lubrica differs from the externally somewhat similar Lygephila lubrosa by its characteristic brownish-grey ground color of the forewings; from Lygephila kazachkaratavika by more unicolorous forewings with a less-developed pattern; and from both related species by its brownish hindwings, which are generally ochreous in the two latter species. The differences in the genitalia structures among the three similar species are easily recognisable in both sexes. In males, the uncus dilation in Lygephila lubrica is wider than in Lygephila lubrosa, but narrower than in Lygephila kazachkaratavika, and the ampulla is more proximal, closer to the middle of the valve, than in the two other species; in the females, the cleft on the posterior margin of the antrum is U-shaped or V-shaped in Lygephila lubrica, whereas in Lygephila lubrosa it is evenly concave; in Lygephila kazachkaratavika the ostium cleft is deep, narrow, slit-like.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
0DEEEC1347F05890B02BF2C79DAC3A73.taxon	description	Description. Wingspan 37 - 50 mm, on average 42 - 48 mm. Head and body brownish grey; collar dark chocolate brown. Forewing brownish grey, sometimes dark brown; subbasal line indistinct; antemedial line arched, consisting of two elongated patches; medial fascia diffuse, wide and waved, with two costal patches; reniform stigma approximately triangular, dark brown, sometimes with sharp extension at inner corner and with satellite streak-like spots on outer margin; orbicular stigma as small white dot; postmedial line distinct; subterminal line with light fascia; terminal line a black sinuous stripe. Hindwing varies from brown to greyish brown; transverse line distinct; narrow discal spot present on underside; outer dark third with defuse inner margin; fringes as ground color. Male genitalia (Figs 21 - 32, 39 - 41). Uncus with short stem and dilated distal two thirds, apex with fine tip, anal tube membranous with characteristic oval hardening of tissue - scaphial crown on scaphium and sclerotized fig on subscaphium; valva elongated, relatively wide with parallel margins, valval apex rounded; clasper digitiform, slightly curved towards costa, situated rather far from apex. Aedeagus straight, long, tubular. Vesica globular, multidiverticulate, membranous; 1 st subbasal diverticulum small, adjacent to 2 nd terminal diverticulum; medial diverticulum large, tapering, with medium-large oblong chamber at base; 1 st terminal diverticulum large, more or less wedge shaped with one part densely scobinated and membranous, cauliflower-like, opposite part bears numerous small pockets; 2 nd terminal diverticulum tubular, scobinated on top; 3 rd terminal diverticulum irregular shaped with large rectangular scobinated basal part and membranous cylindrical extension; 4 th terminal diverticulum medium sized, situated between 1 st and 3 rd medial diverticulum; 2 nd subbasal diverticulum small, tubular, sometimes chili-pepper-like (Fig. 41), terminal tube membranous, as long as aedeagus, opening point of terminal tube located subbasally near carina. Female genitalia (Figs 46 - 57). Ovipositor relatively large, broad, papillae anales hairy with long setae on apical edges. Apophyses anteriores stout, apophyses posteriores thin, longer than apophyses anteriores. Ostium broad, antrum tapering, funnel shaped, posterior margin incised producing large U-shaped cleft; ductus bursae small, inflated with ventral sclerotized ribbon; appendix bursae small; corpus bursae membranous, ovoid.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
0DEEEC1347F05890B02BF2C79DAC3A73.taxon	distribution	Distribution. Siberian. Distributed from Zaporozhie region of Ukraine to Rostov, Samara, Povolzhie regions to Ural of Russia through Kazakhstan, Russian Altai and northern Mongolia.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
A7F00213DD9B54A2A159C87CA0A2E7AF.taxon	diagnosis	Diagnosis. Easily distinguishable from all other members of the species group by its unicolorous grey forewings. Comparing the genital structures of the taxa of the group, it differs from all related species by the narrow uncus without a real dilatation (only some slight thickening is present), the wide valva, and the subapically located, strong clasper with its tip reaching the valval edge (males); and by the funnel-shaped antrum bent dorsally at 45 degrees, being a unique female character for the whole genus (Fig. 60).	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
A7F00213DD9B54A2A159C87CA0A2E7AF.taxon	description	Description. Wingspan 42 - 46 mm. Head and body grey with some brownish scales; collar chocolate brown. Forewing almost unicolorous, wing pattern poorly developed; subbasal line hardly traceable, represented by groups of dark scales on veins; antemedial line semicircular; medial shade not expressed; reniform stigma small, indistinct, with one or two black dots basad; orbicular stigma small dot-like; postmedial and subterminal lines distinct; terminal line a row of black dots on veins. Hindwing pale ochreous; transverse line not discernible; outer dark third with sharply defined inner margin; fringes ochreous. Male genitalia (Figs 33, 34, 44, 45). Uncus long, stout, slightly thickened medially with skewed fine tip, anal tube membranous with oval hardening of tissue - scaphial crown on scaphium and sclerotized fig on subscaphium; valva elongated, wide, with parallel margins in the middle, valval apex rounded; clasper digitiform, strong, thickened with wide base, placed subapically, asymmetrical, left one shorter than the right one, both almost reach valval costa. Aedeagus tubular with narrow, long, acute sclerotised lamina on ventral side of carina. Vesica globular, multidiverticulate, membranous; basal diverticulum small; medial diverticulum large with small lateral hemispherical bulging; 1 st terminal diverticulum large, more or less wedge shaped, membranous with cauliflower-like part bearing numerous small pockets; 2 nd terminal diverticulum large, cone shaped, scobinated on top; 3 rd terminal diverticulum medium-sized, bifurcated, Y-shaped; 4 th terminal diverticulum large, bilobate, located oppositely to the 3 rd medial diverticulum; terminal tube membranous as long as aedeagus, opening point of terminal tube located near to carina. Female genitalia (Figs 58 - 61). Ovipositor relatively large, broad, papillae anales hairy with long setae on apical edges. Apophyses anteriores stout, apophyses posteriores thin, longer than apophyses anteriores. Antrum funnel shaped, bent dorsally at 45 degrees, boomerang shaped from lateral view; ostium bursae broad, posterior margin U-shaped; ductus bursae medium sized; appendix bursae small; corpus bursae membranous, bevelled oval.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
427A763DE68B5DD5B904453896BC2203.taxon	diagnosis	Diagnosis. Differs from Lygephila lubrosa lubrosa by its smaller size and better marked reniform stigma. In the male genitalia, ssp. orbonaria differs from ssp. lubrosa by its narrower uncus without a medial thickening, and the narrower upper part of valva with more expressed asymmetry.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
427A763DE68B5DD5B904453896BC2203.taxon	description	Description. Wingspan 34 - 43 mm. The external features, with the exception of the smaller size and somewhat roundish forewing, match those of the nominate subspecies. Male genitalia (Figs 35, 36). Uncus long, stout, sabre-like, anal tube membranous with oval hardening of tissue - scaphial crown on scaphium and sclerotized fig on subscaphium; valva elongated, wide, with parallel margins in the middle, tapering to apex; clasper digitiform, strong, thickened with wide base, placed subapically, somewhat asymmetrical, left one short, right one longer, almost reaches costa. Aedeagus tubular with narrow, long, acute sclerotised lamina on ventral side of carina. Vesica globular, multidiverticulate, membranous; basal diverticulum small; medial diverticulum large with small lateral hemispherical bulging; 1 st terminal diverticulum large, more or less wedge shaped, membranous with cauliflower-like part bearing numerous small pockets; 2 nd terminal diverticulum large, cone shaped, scobinated on top; 3 rd terminal diverticulum medial sized, bifurcated, Y-shaped; 4 th terminal diverticulum large, bilobate, located opposite to 3 rd medial diverticulum; terminal tube membranous, as long as aedeagus, opening point of terminal tube located near carina. Female genitalia. Unknown.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
427A763DE68B5DD5B904453896BC2203.taxon	distribution	Distribution. SW Tajikistan, Pianj river valley.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
36218B37B8B5639984E19A6B12A8B961.taxon	diagnosis	Diagnosis. Easily separable from Lygephila lubrica and Lygephila lubrosa by the very contrasting, well-developed pattern on the forewings. In the male genitalia, it differs from all close relatives by its wider dilatation of the uncus, and the more acute apex of the valva (males); the longer antrum with a deep slit-like cleft on the posterior margin is diagnostic for females.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
36218B37B8B5639984E19A6B12A8B961.taxon	description	Description. Wingspan 41 - 44 mm. Head and body brownish grey; collar dark chocolate brown. Forewing contrastingly marked, variable in coloration from mottled light greyish brown to uniform dark brown; noctuid pattern well developed; subbasal line traceable; antemedial line arched, consisting of three elongated patches edged by light fascia basally; medial shade waved, bifurcated from below cell to anal margin, with two costal patches; reniform stigma somewhat triangular, black, sometimes with satellite streak-like spots on outer margin; orbicular stigma absent; postmedial line distinct; subterminal line with light fascia; terminal line a row of black dots. Hindwing ochreous; transverse line distinct; small discal spot present on underside; border between pale proximal part and dark outer third diffuse; fringes ochreous. Male genitalia (Figs 37, 38, 42, 43). Uncus stem short, stout, distal part dilated, terminated in fine tip; anal tube membranous with oval hardening of tissue - scaphial crown on scaphium and with sclerotized fig on subscaphium; valva elongated, relatively wide with parallel margins in the middle and convergent basally and distally; clasper digitiform, undulate, placed subapically, not reaching costa. Aedeagus straight, long, tubular, with small sclerotized fig on ventral side of carina. Vesica globular, multidiverticulate, membranous; basal diverticulum small; medial diverticulum large, cupola shaped with two hemispherical chambers medially and basally; 1 st terminal diverticulum large, more or less wedge shaped, with one part densely scobinated and membranous cauliflower-like, opposite part bearing numerous small pockets; 2 nd terminal diverticulum tubular, elongated, scobinated on top; 3 rd terminal diverticulum medium sized with lateral bulging; 4 th terminal diverticulum large, conical, situated opposite to 3 rd medial diverticulum; terminal tube membranous, as long as aedeagus, opening point of terminal tube located subbasally near carina. Female genitalia (Figs 62, 63). Ovipositor relatively large, broad, papillae anales hairy with very long setae on apical edges. Apophyses anteriores long and thin, apophyses posteriores thin, somewhat longer than apophyses anteriores. Ostium broad, antrum tapering, funnel shaped, posterior margin deeply incised producing slit-like cleft; ductus bursae small; appendix bursae small; corpus bursae membranous, ovoid.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
36218B37B8B5639984E19A6B12A8B961.taxon	distribution	Distribution. Kazakhstan, Kyrgyzstan.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
FD93720C601D98E343E55A2DFDA5B152.taxon	diagnosis	Diagnosis. The distinctive features of Lygephila lupina and Lygephila vulcanea (Fig. 66) are given in the works of Sviridov (1990) and Kononenko (1996) (under the names Lygephila vulcanea and Lygephila mirabilis). The main external differences between the two species are found in the colouration and shape of the forewing: Lygephila lupina is broader winged and paler in colouration, usually ochreous brown to buff coloured, whereas Lygephila vulcanea is darker, deep brown to claret brown, most often with a clearly visible violaceous shade and the forewing apex is somewhat more pointed. In the majority of the specimens the reniform stigma of Lygephila lupina is stronger, sharper, and more distinctly marked against the paler background. The female genitalia differ from those of Lygephila vulcanea (Fig. 69) by the shallower incision of the posterior margin of the antrum.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
FD93720C601D98E343E55A2DFDA5B152.taxon	description	Description. Wingspan 44 - 49 mm. Head and body brownish grey; collar dark chocolate brown. Forewing brownish grey with sparse dark brown irroration; subbasal line indistinct; antemedial line arched with costal patch; reniform stigma large, dark brown, consists of 5 or 6 streak-like spots; orbicular stigma absent; postmedial and subterminal lines distinct; terminal line a row of black dots. Hindwing brownish; small discal spot present on underside; outer third dark brown; fringes as ground color. Female genitalia (Figs 67, 68). Ovipositor long, papillae anales large, hairy with long setae on apical edges. Apophyses anteriores relatively short, apophyses posteriores thin, longer than apophyses anteriores. Antrum long, narrow anteriorly, dilated posteriorly, ostium broad, posterior margin with small U-shaped cleft. Corpus bursae membranous, ovoid.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
FD93720C601D98E343E55A2DFDA5B152.taxon	distribution	Distribution. Russian Far East, China, Korea.	en	Pekarsky, Oleg (2014): Taxonomic studies of the Lygephila lubrica (Freyer, 1842) species complex with notes on other species in the genus (Lepidoptera, Erebidae, Toxocampinae). ZooKeys 452: 107-129, DOI: http://dx.doi.org/10.3897/zookeys.452.8152, URL: http://dx.doi.org/10.3897/zookeys.452.8152
