taxonID	type	description	language	source
03D955344E1C257D59DEF82C2DFD98AC.taxon	description	1976 Rumohra recentior (UNGER) BARTHEL, p. 457, text-fig. 6 a – d, pls 80, 81. M a t e r i a l. Fragments and incomplete sterile fern fronds, impressions PA 1618, PA 1619, PA 1620, PA 1621 – 28, PA 1625, NM-G 12803, NM-G 12804, NM-G 12806. D e s c r i p t i o n. Incomplete sterile fern fronds, apex with a lateral pinna and a detached pinna, preserved length up to 100 mm, rachis strong, surface smooth, pinna almost perpendicular, pinnules ovate, slightly asymmetrical, deeply irregularly dentate, up to 15 mm long and 5 mm wide, segments broadly attached, admedially oriented, shallowly dissected, secondary veins alternate, 3 times forked, the lowermost admedially reaching the margin. R e m a r k s. These fragments are up to now the first records of this fern in the Oligocene of the České středohoří Mts. and Saxony. From the Palaeogene of North Bohemia, it is only known from the Doupovské hory Mts., the locality Valeč (Bůžek et al. 1990) and from the České středohoří Mts., localities Kučlín (Kvaček and Teodoridis 2011) and Holý Kluk (Radoň et al. 2006). The new finds from Ludvíkovice morphologically fully correspond to the above mentioned material from the Czech Republic. For more detailed analysis of morphology and affinities see Barthel (1976).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E1B257D5B06FBA128119BC0.taxon	description	1850 Daphnogene cinnamomifolia (BRONGN.) UNGER, p. 424. M a t e r i a l. 14 incomplete simple leaves and fragments, impressions PA 1623, PA 1638, PA 1663 A, PA 1665, PA 1673, PA 1678, PA 1679, NM-G 12797, NM-G 12802, NM-G 12805, NM-G 12807, NM-G 12812, NM-G 12815. D e s c r i p t i o n. Leaves lanceolate to widely oval, up to 120 mm long and 52 mm wide, apex not preserved probably shortly acuminate and blunt, base cuneate with fragmentary petiole, margin entire, venation suprabasal acrodromous, midrib strong, moderate, straight or curved, lateral veins thinner, alternate, running along the margin, usually connecting with secondaries at 2 / 3 of the blade length, secondary veins thinner, alternate, originating at 40 ° to 50 °, curved and looping by margin or between midrib and lateral veins, tertiary veins alternate perpendicular, straight to sinuous, often forked, venation of the higher orders poorly preserved. R e m a r k s. The leaves of Daphnogene cinnamomifolia are known from most Oligocene localities of Europe and obviously represent lineal to oval lanceolate “ sun ” and larger oval “ shade ” leaves distinguished as two foliage forms “ cinnamomifolia ” and “ lanceolata ” (e. g., Kvaček and Walther 1995). In volcanic floras this element usually occurs abundantly (lacking at Bechlejovice). In Hammerunterwiesenthal (Krušné hory / Erzgebirge Mts.) and Valeč (Doupovské hory Mts.) it occurs in lacustrine sediments with volcaniclastic admixture as a possible “ pioneer ” plant (Kvaček and Walther 1995).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E1B257D59C1FF4A28A29FDE.taxon	description	M a t e r i a l. 12 incomplete simple leaves and fragments, impressions PA 1623, PA 1650, PA 1662, PA 1665, PA 1679, NM-G 12808. D e s c r i p t i o n. Leaves lanceolate to elongate, up to 69 mm long and up to 18 mm wide, apex not preserved probably shortly attenuate to acute or blunt, base cuneate with fragmentary petiole (up to 7 mm), margin entire, venation brochidodromous, midrib straight, secondary veins thin, alternate, at 40 ° to 60 °, looping, tertiary veins alternate perpendicular, straight to sinuous, venation of the higher orders regular polygonal reticulate, poorly preserved. R e m a r k s. The lauroid leaves with entire margin and brochidodromous venation are typical lanceolate to elongate in outline and in gross morphology match slender leaves of Laurophyllum cf. acutimontanum, which are common elements of late Eocene to Oligocene floras in Central Europe, e. g., Staré Sedlo, Bechlejovice, Kundratice, Suletice-Berand, Holý Kluk, Seifhennersdorf (Kvaček and Walther 1995, 1998, 2004, Knobloch et al. 1996, Radoň et al. 2006). However, the lack of epidermal structure does not allow for a clear assignment because the foliage of Laurophyllum pseudoprinceps WEYLAND et KILPPER and Laurophyllum medimontanum BŮŽEK, HOLÝ et KVAČEK have similar leaf morphology and co-occurred with Laurophyllum acutimontanum at Suletice-Berand, Kundratice and Bechlejovice (Kvaček and Walther 1995, 1998, 2004).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E1A257F5944FB522DF49A84.taxon	description	1840 Phyllites furcinervis ROSSM., p. 33, pl. 6, fig. 25, pl. 7, figs 26 – 31. 1989 Eotrigonobalanus furcinervis (ROSSM.) H. WALTHER et KVAČEK in Kvaček and Walther, p. 581, text-figs 3 – 10, pls 33 – 36, 38 – 46, pl. 47, figs 1 – 3. M a t e r i a l. Incomplete simple leaves, impressions PA 1645, PA 1661. D e s c r i p t i o n. Leaves simple, incomplete, oblong to elliptic, 70 mm and 102 mm long, 22 mm and 24 mm wide, base cuneate without petiole, apex not preserved probably shortly attenuate to acute, margin entire, venation eucamptodromous, midrib thick, almost straight, secondary veins thinner, slightly bent and looping along the margin, regularly spaced, alternate to subopposite, originating at 40 °, tertiary veins percurrent, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided. R e m a r k s. The leaves morphologically clearly match Eotrigonobalanus furcinervis, but the identification is tentative due to the lack of leaf epidermal anatomy. The leaves of this extinct fagaceous element are common in the late Eocene fluviatile sandstones of the Staré Sedlo Fm. (Knobloch et al. 1996, Teodoridis et al. 2012), Kučlín (Kvaček 2002, 2010, Kvaček and Teodoridis 2011) and other sites of this kind in Germany and elsewhere in Europe (Kvaček and Walther 1989). The occurrence of Eotrigonobalanus in Seifhennersdorf (Walther and Kvaček 2007) is the oldest evidence of this genus in volcanic floras of Central Europe during the early Oligocene. Regionally, this element is also known from the late Oligocene at Kleinsaubernitz (Walther 1999).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E1A257C5E2EFF4429439F78.taxon	description	Pl. 2, Figs 4 – 6 M a t e r i a l. 2 complete leaves and 1 incomplete leaf, impressions PA 1632, PA 1639, PA 1646. D e s c r i p t i o n. Leaves widely elliptic, incomplete leaf obovate, 50 mm to 61 mm long, 34 to 39 mm wide, apex acute, base widely cuneate, margin simple dentate, teeth acute, sinus rounded, venation craspedodromous, secondary veins thinner, slightly curved, alternate, originating at 40 ° to 60 °, tertiary veins alternate to opposite percurrent, straight to sinuous, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided, veinlets lacking. R e m a r k s. The acute to short spiny teeth show an affinity to Quercus, however the widely elliptic outline is not typical for Fagaceae and may also resemble Betulaceae (see below). Relatively common leaves described as Quercus lonchitis UNGER (typified in Sotzka – Unger 1850, also occurring in Seifhennersdorf, Suletice, Lochočice and Stadice – Walther and Kvaček 2007) show similar features of the margin, however, the leaf outline is more elongate. Generally, the studied leaves also resemble the stratigraphically younger morphotypes of younger Quercus drymeja UNGER and Q. mediterranea UNGER. The incomplete obovate leaf (PA 1646) may morphologically resemble the basal part of “ roburoid ” oaks.	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E19257F5B3DFA4F289B9E5B.taxon	description	Walter and Kvaček 2007 / Carya quadrangula (KIRCHH.) J. - F. LEROY, 1955 Pl. 3, Figs 6 – 9, Pl. 4, Fig. 1 1825 Phyllites fragiliformis STERNB., p. 42, index iconum, pl. 50, fig. 1. 1936 Caryojuglans quadrangular KIRCHH., pp. 82 – 85, text-fig. 13, pl. 12, fig. 36 a – l. 1955 Carya quadrangular (KIRCHH.) J. - F. LEROY, pp. 162 – 163, text-fig. 81. 2007 Carya fragiliformis (STERNB.) KVAČEK et H. WALTHER in Walter and Kvaček, p. 110, text-fig. 6 b, pl. 11, figs 1 – 3, pl. 23, figs 8 – 10. M a t e r i a l. Incomplete leaflets and one incomplete endocarp, impressions PA 1627, PA 1644, PA 1659, PA 1660, PA 1682 (fruit). D e s c r i p t i o n. Detached incomplete leaflets, lanceolate to oblanceolate, 35 to 63 mm long, 12 to 26 mm wide, apex acute to acuminate, base cuneate with fragmentary petiolule up to 5 mm, often asymmetrical, margin irregularly serrate, teeth triangular, sharp, bent admedially, venation craspedodromous, to semicraspedodromous, midvein straight or curved, secondaries opposite in the basal part then alternate, originating at 40 ° to 60 °, tertiary veins percurrent, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided. Incomplete endocarp widely elliptic in outline, apex acute, upper surface and distinct longitudinal ribs are not visible, base not preserved, 22 mm long and 18 mm wide, thickness of the nutshell 2.8 mm, the primary and secondary septum fragmentary. Seed body (kernel) oval incomplete, 2 lateral symmetric parts preserved. R e m a r k s. The material corresponds to the holotype of this fossil species based on a single specimen from Žichov (Kvaček and Straková 1997, Walther and Kvaček 2007). The hickory leaves and / or leaflets described as C. fragiliformis or C. serrifolia (GÖPP.) KRÄUSEL are relatively common elements from Oligocene localities in Central Europe, known from e. g., Bechlejovice (Kvaček and Walther 2004), Suletice-Berand (Kvaček and Walther 1995), Seifhennersdorf (Walther and Kvaček 2007) and Matrý (Kvaček et al. 2018). According to Walther and Kvaček (2007: 110), C. serrifolia, which in most cases was assigned to Oligocene – early Miocene material from Central Europe, may represent a descendant of this lineage in the late Neogene. Koutecký et al. (2019) associated this type of foliage with fruits of C. quadrangula (KIRCHH.) J. - F. LEROY and wood of Eucaryoxylon crystallophorum MÜLL. - STOLL et MÄDEL following the whole plant concept. The fruit find from Ludvíkovice morphologically corresponds to C. quadrangula. On the other hand, some poorly preserved leaf material may show affinity to Engelhardia orsbergensis (P. WESSEL et C. O. WEBER) JÄHNICHEN, MAI et H. WALTHER.	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E19257F5AAAFE5E2CAE9EB5.taxon	description	1840 Phyllites rhamnoides ROSSM., p. 35, pl. 8, figs 30, 31. 1988 Trigonobalanopsis rhamnoides (ROSSM.) KVAČEK et H. WALTHER, p. 405, text-figs 2 – 4, pl. 2, figs 1 – 8, pls 50 – 54, pl. 55, figs 2 – 7, pl. 56, figs 1 – 4, pl. 57. M a t e r i a l. 2 complete and 2 incomplete leaves, impressions PA 1651, PA 1665, PA 1667, NM-G 12813. D e s c r i p t i o n. Leaves oblong to ovate, 32 to 86 mm long, 11 to 30 mm wide, base cuneate, apex shortly attenuate and blunt, margin entire, venation brochidodromous, midrib straight, slightly curved in its upper third, secondaries very regularly spaced, subopposite, originating at 30 ° to 40 °, venation of higher order not preserved. R e m a r k s. This extinct element of the Fagaceae has usually been determined mainly according to its rhamnoid (brochidodromous) venation. Occurrences in the late Eocene Staré Sedlo Fm. are rare (Knobloch et al. 1996) as well as in Kučlín (Kvaček and Teodoridis 2011). The early Oligocene flora of Bechlejovice contains two leaves described as cf. Quercus bavarica (KNOBLOCH et KVAČEK) KVAČEK, which may show a morphological affinity to Trigonobalanopsis. In the volcanic facies Trigonobalanopsis was usually poorly represented (Rott, Kleinsaubernitz, Lower Rhine Basin), while it became a dominant element in the Miocene mastixioid floras (e. g., Kvaček and Walther 1988, Walther 1999, Winterscheid 2018).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E19257E595DFA612C87992F.taxon	description	1856 Corylus rhenana P. WESSEL et C. O. WEBER, p. 134, pl. 22, fig. 5. 2014 Alnus rhenana (P. WESSEL et C. O. WEBER) WINTERSCHEID et KVAČEK, p. 22, pl. 4, figs 1 – 5, pl. 11, figs 4 – 6. M a t e r i a l. Several fragments and incomplete leaves, impressions PA 1628, PA 1629, PA 1632, PA 1633, PA 1635 – 38, PA 1640, PA 1642, PA 1643, PA 1650, PA 1655 – 56, PA 1657, PA 1658, PA 1663 A, NM-G 12814. D e s c r i p t i o n. Incomplete leaves and fragments are ovate to obovate, 23 to 85 mm long, 17 to 75 mm wide, apex acute to shortly attenuate, base rounded or slightly cordate, margin finely widely serrate, venation craspedodromous to semicraspedodromous, midrib thick, secondaries, subopposite to alternate, widely spaced, curved, originating at 30 ° to 50 °, tertiary veins percurrent, straight, curved or forked, venation of the higher orders regular polygonal reticulate, areolation well-developed, 4 sided. R e m a r k s. The foliage in the samples from Ludvíkovice is very similar to the leaf material described from Matrý and Žichov as Alnus rhenana, which is typically long petiolate and variable in shape (Ettingshausen 1869, Winterscheid and Kvaček 2014). In addition to A. rhenana, narrow elliptic leaves of Alnus gaudinii (HEER) ERW. KNOBLOCH et KVAČEK known from Bechlejovice, Kundratice (Kvaček and Walther 1998, 2004), Holý Kluk (Radoň et al. 2006), Hrazený (Kvaček et al. 2015) and Seifhennersdorf (Walther and Kvaček 2007), are also a very common Oligocene elements of North Bohemia and Saxony.	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E18257E597DFAEC29C69324.taxon	description	1996 Dicotylophyllum rossmaessleri ERW. KNOBLOCH et KVAČEK in Knobloch et al., p. 114, pl. 8, fig. 2, text-fig. 59 f. M a t e r i a l. Almost complete leaf, impression PA 1663 B. D e s c r i p t i o n. Almost complete leaf, linear-ovate in outline, 110 mm long and 27 mm wide, base cuneate, apex long attenuate, margin entire, venation eucamptobrochidodromous, midrib straight, secondary veins alternate, originating at 30 ° to 40 °, tertiary veins perpendicular higherorder venation regular polygonal reticulate, areolation welldeveloped. R e m a r k s. The specific type of venation in combination with the almost linear character of the leaf lamina and an extremely attenuate apex allow an assignment to Dicotylophyllum rossmaessleri without any doubt. The element is typified in the late Eocene flora of Staré Sedlo and probably has an affinity to evergreen Fagaceae such as Eotrigonobalanus (Knobloch et al. 1996).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E18257E5B3EF81E2EFE9993.taxon	description	M a t e r i a l. 2 incomplete leaves / leaflets, impressions PA 1628. D e s c r i p t i o n. Leaves / leaflets incomplete, lamina elliptic and obovate, 41 and 93 mm long, 19 and 21 mm wide, base cuneate, apex shortly attenuate, margin entire or? finely crenulate in apical part, venation brochidodromous or semicraspedodromous, midrib straight, secondary veins subopposite to alternate, numerous, originating at 40 ° to 60 °, looping, intersecondaries parallel, venation of higher order not preserved. R e m a r k s. The leaves / leaflets resemble Carya leaflets based on the lamina shape, size and numerous secondary veins, however a typical margin is not preserved. On the other hand, an affinity to Lauraceae or Leguminosae cannot be excluded due to the resemblance in shape, a relatively distinct midrib and the “ evergreen ” character of the leaf / leaflet lamina.	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E18257E59F4FD192EAB9E26.taxon	description	M a t e r i a l. Incomplete leaf, impression PA 1650. D e s c r i p t i o n. Incomplete leaf, elliptic, 49 mm long, 21 mm wide, base not preserved, apex incomplete probably blunt or shortly attenuate, margin entire, venation brochidodromous, midrib straight, secondary veins subopposite to alternate, originating at 40 ° to 60 °, 9 pairs preserved, looping by the margin, venation of higher order not preserved. R e m a r k s. The preserved fragmentary leaf lamina seems to have a coriaceous character, which may also be proven by the entire margin and should indicate an evergreen character of the element. The spacing of secondary veins shows an affinity to Fagaceae rather than to Lauracaeae, therefore an assignment to Trigonobalanus or Quercus bavarica (KNOBLOCH et KVAČEK) KVAČEK may be acceptable but cannot be proven without knowledge of the epidermal structure.	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
03D955344E18257E5A9CFD702C4D9DA1.taxon	description	1869 Quercus artocarpites ETTINGSH., p. 63, pl. 55, figs 19 – 19 a. 2001 Sloanea artocarpites (ETTINGSH.) KVAČEK et HABLY in Kvaček et al., p. 117. M a t e r i a l. Incomplete leaves and fragments, impressions PA 1630, PA 1631, PA 1634, PA 1641, PA 1654, PA 1666, PA 1667, PA 1671, PA 1677. D e s c r i p t i o n. Leaves incomplete, without petiole, lamina elliptic to ovate, 61 to 161 mm long and 38 to 76 mm wide, base widely cuneate to rounded, apex not preserved, probably acute or attenuate, margin subentire, with widely spaced minute teeth, venation camptosemicraspedodromous, midrib straight, secondaries subopposite, curved, originating at 40 ° to 60 °, looping very near margin, with abmedial veinlets entering teeth, tertiary veins percurrent, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided. R e m a r k s. The leaves are characterized by a specific marginal venation of campto-semicraspedodromous type which allows the unambiguous identification of this mesophytic and thermophilous element. Sloanea artocarpites was widespread at most sites of the České středohoří Mts. such as Suletice-Berand (Kvaček and Walther 1995), Holý Kluk (Radoň et al. 2006) and Matrý (Kvaček et al. 2018) and Seifhennersdorf in Saxony (Walther and Kvaček 2007). Similar foliage is known from the early Oligocene sites of Hungary and Slovenia accompanied by fruits of the Sloanea - type (Kvaček et al. 2001).	en	Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav, Bruch, Angela A. (2020): A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic). Fossil Imprint 76 (2): 297-314, DOI: 10.37520/fi.2020.024, URL: http://dx.doi.org/10.37520/fi.2020.024
