taxonID	type	description	language	source
03D887EDFF84814873E1A8E273B9F49B.taxon	type_taxon	(type species: M. austinianus LeConte, 1874, fixed by monotypy)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF84814873E1A8E273B9F49B.taxon	description	(Figs 1 – 29, 31 – 42)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF84814873E1A8E273B9F49B.taxon	description	Redescription. Head, pronotum and abdomen slightly convex; elytra flattened, slightly or strongly broadened posteriad. Body shiny, with dense coriaceous microsculpture on head and pronotum or without it (M. puncticollis); sometimes elytra subrugosely sculptured (M. rogersi); body length varying from 2.60 to 5.50 mm. Coloration brown to reddish-brown or black. Head and pronotum with fine and moderately dense punctation; punctation of elytra distinctly denser, larger and deeper than that on pronotum; abdomen with distinct, regular, dense and fine punctation. Head subtrapezoidal, with postocular parts usually widened and relatively long, with distinctly elevated part between bases of antennae and infraorbital portions; supra-antennal protuberances narrow, distinctly elevated, diagonally stretching from base of antennae to about level of apical margins of eyes; medioapical depression deep and wide, slightly or strongly narrowing posteriad toward level of anterior margins of eyes; interocellar depression shallow or well-distinct and moderately deep; anteocellar furrows (grooves in front of ocelli) narrow or very deep and moderately wide, diagonally stretching anteriad toward level of anterior third of eyes. Eyes relatively small, convex. Ocelli convex, small, but well visible. Labrum transverse, with slightly and widely concave apical margin and slightly elongate basolateral projections, with several very long setae in latero-apical and middle portions (Figs 6 – 7, and Fig. 398 in Watanabe (1990 )). Mandibles with wide basal part and elongate, strongly curved apical portion; inner margin of each mandible with two elongate teeth (Figs 8 – 9, and Fig. 396 in Watanabe (1990 )). Maxillae relatively wide, with elongate galea and maxillary palpomeres; apical maxillary palpomere somewhat elongate, narrow, distinctly more than twice narrower than markedly widened preceding segment; lacinia moderately short, with several elongate preapical teeth (Figs 10 – 11). Labium narrow; labial palpomere 2 about as long as broad, preapical palpomere distinctly longer than broad, apical palpomere narrow, distinctly longer than preceding segment (Figs 12 – 13 and Fig. 399 in Watanabe (1990 )); labial palpomere 2 slightly elongate (Fig. 12) or transverse (Fig. 13). Mentum narrow, transverse, as in Fig. 399 in Watanabe (1990). Gular sutures narrowly (see Fig. 400 in Watanabe (1990 )) or relatively widely separated from each other at level of posterior margins of eyes, strongly divergent posteriad. Antennae long, exceeding middle or apical third length of elytra when reclined, with antennomeres 4 – 10 slightly or distinctly elongate. Pronotum convex, distinctly transverse, widest in anterior portion, not or distinctly protruding anteriad (M. fenderi), gradually or strongly narrowed posteriad, narrowly or widely bordered laterally and sometimes distinctly explanate and reflexed (M. fenderi); mediobasal portion sometimes with shallow transverse impression. Prothorax short, strongly transverse, distinctly convex in middle, with long and acute intercoxal process reaching about middle of front coxae. Mesoventrite moderately short, strongly convex, with acute elongate process, reaching apex of relatively long and narrow intercoxal process extending to about middle of mesocoxae. Scutellum large, triangular, without punctation. Metaventrite transverse, convex, with very deep intercoxal cavities and angular intercoxal process extending to mesosternal process. Elytra wide, slightly or strongly broadened posteriad, with narrow or wide and distinctly reflexed (M. fenderi) lateral margins, elongate or moderately short, reaching apical margin of abdominal tergite III to V. Hind wings fully developed. Legs relatively slender, moderately long, with transverse procoxae and metacoxae, and large suboval mesocoxae; protarsi of males with distinctly dilated protartsomeres 1 – 4; metatarsus more than twice shorter than metatibiae; apical antennomere slightly shorter than preceding four metatarsomeres. Abdomen slightly or distinctly broader than elytra, with two oval or transverse tomentose spots in the middle of abdominal tergite IV; apical margin of abdominal tergite VII with narrow palisade fringe. Aedeagus elongate or moderately short (M. kastcheevi sp. n., M. schilenkovi), with small or very large phallobase, gradually narrowed apicad; parameres narrow, reaching or exceeding apex of median lobe; internal sac narrow, without or with elongate and sclerotized structures, sometimes with flagellum. Female genital segment with narrow and very long gonocoxites, with moderately short and narrow styli, each with long apical setae (Figs 29, 42). Immature stages unknown. Comparative notes. Based on the general shape of the body and the aedeagus, and the presence of an interocellar depression, the genus Microedus is related to genera of the Hygrogeus complex, distributed in the Holarctic Region (Zerche 1992, Shavrin 2017). Regarding the general shape of the narrow apical segment of the maxillary palpi, the ground sculpture of the forebody, and the shape of the transverse pronotum, Microedus is similar to Hygrodromicus Tronquet, 1981 and Philydrodes Bernhauer, 1929. It can be distinguished from Hygrodromicus by the narrower apical and somewhat broader preapical segments of the maxillary palpi, longer temples, and the general shape of the aedeagus (basal portion of aedeagus of Hygrodromicus usually small and rotated inside the abdomen in lateral position (90 °), as in Philydrodes too). It can be distinguished from Philydrodes by the shape of the labrum, broader mandibles, the shorter and broader preapical and longer apical segments of the maxillary palpi, shorter temples, and the significantly broader pronotum. From all genera listed above and other genera of the Hygrogeus complex it can be distinguished by the external and internal morphology of the aedeagus. A more detailed analysis of relations of these genera within Hygrogeus complex will be published in detail in a forthcoming revision of the genus Trichodromeus Luze, 1903 and related genera of the Palaearctic Region (in prep.).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF84814873E1A8E273B9F49B.taxon	distribution	Distribution. Species of the genus Microedus are distributed in the Nearctic (Canada, USA) and Palaearctic regions (Russia (Altai, Russian Far East), Kazakhstan (Altai and Dzhungar mts.), India (Himalayas), and Japan (Hokkaido, Honshu). Bionomics. Species of Microedus inhabit sand or gravel banks of rivers, streams and lakes, in moss and litter, and can be found frequently in the mountains (Hatch 1957, Newton et al. 2000, present study).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF84814873E1A8E273B9F49B.taxon	discussion	Remarks. The genus Altaiodromicus was originally described based on a single species (A. schilenkovi Zerche, 1992) from Altai Mts., Russia. Based on the shape of the body (Fig. 4), it was compared with Hygrodromicus and Geodromicus Redtenbacher, 1857 and distinguished by the narrow apical maxillary palpomere and different morphology of the aedeagus. During the study of Omaliinae material collected by Dr. Kastcheev, I found a new, similar species from Kazakhstan. This new species can be clearly distinguished from A. schilenkovi by the presence of sclerotized elongate structures within the internal sac (Fig. 22) and some other morphological features (see below). Besides that, I studied the holotype of Geodromicus brevitarsis Champion, 1925 from northern India (Fig. 2). The apical maxillary palpomere of this species is also narrow, and based on the slightly elongate antennomeres 4 – 10, G. brevitarsis is somewhat similar to the two species listed above, but differs by the smaller and narrower body, with markedly narrower pronotum, and different morphology of the aedeagus. Based on the external morphological features, one unnamed species from Kashmir is also similar to G. brevitarsis (see below). Several years ago, with the assistance of the late A. Smetana, I received several Nearctic species of Microedus from Canada and USA. During the study of mouthparts and aedeagi of these species, I found that they are similar to species of Altaiodromicus (Figs 6 – 13) and G. brevitarsis. The general shapes of the body (Fig. 1) and the aedeagus (Fig. 14) of the type species of Microedus (M. austinianus) are somewhat similar to these of M. brevitarsis (Fig. 16). The shapes of antennomeres 4 – 10 of M. austinianus are also similar to these in all Asian species listed above. Other Nearctic species of Microedus studied by me (M. fenderi Hatch, 1957, M. laticollis (Mannerheim, 1843) and M. porterae Hatch, 1957) have more elongate antennomeres 4 – 10. Besides that, all these species have similar ground sculpture of the forebody and shape of the female genital segment (Fig. 29). Thus, I decided to synonymize the genus Altaiodromicus with Microedus and transfer G. brevitarsis to this genus. Species of Liophilydrodes, known from Japan and Far Eastern Russia have a narrow apical segment of maxillary palpi, similar general shape of the body and mouthparts (see Figs 396 – 399 in Watanabe (1990 )) as in species from Middle Asia and the Nearctic Region. All species of Liophilydrodes have elongate antennomeres 4 – 10. Some species have similar shapes of aedeagi as those from Middle Asia, with the presence of elongate structures within the internal sac (Fig. 34), sometimes with flagellum as in Figs. 21 – 26 in Shavrin & Makarov (2019). Thus, the genus Liophilydrodes is synonymized with Microedus. The Palaearctic species of Microedus vary in proportions of the antennomeres, the shape of the interocellar foveae, the pronotum and the aedeagus as in some other taxa of the Hygrogeus group (e. g. Shavrin (2022 b )). Species of the Nearctic Region are in need of a comprehensive taxonomic revision due to significant morphological differences between species (e. g. M. fenderi).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814A73E1AA72754AF711.taxon	description	(Figs 1, 6, 8, 10, 12)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814A73E1AA72754AF711.taxon	materials_examined	Material examined. CANADA: QUEBEC: 1 ♂: Mont Albert, Parc Gaspesie. 1000 m a. s. l. 09.08.1972. J. M. & B. A. Campbell leg. (CNC); 2 ♂♂: 6 mi. S Riviere-a-Claude. 1000 m a. s. l. 18.07.1972. J. M. Campbell leg. (cSh); USA: NEW HAMPSHIRE: 1 ♂: Glen Boulder Trail, White Mts. 2500 m a. s. l. 17.08.1976. J. M. & B. A. Campbell leg. (CNC).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814A73E1AA72754AF711.taxon	discussion	Remarks. Microedus austinianus was originally described from “ White Mountains, Vancouver Island ”. This species is widely distributed in Canada and USA (Herman (2001), Newton et al. (2000), etc.). Habitus as in Fig. 1. Labrum as in Fig. 6. Mandibles as in Fig. 8. Maxilla as in Fig. 10. Labium as in Fig. 12. Aedeagus as in Figs 14 – 15. The aedeagus of this species is illustrated for the first time.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814473E1ACB87005F4DB.taxon	description	(Figs 2, 16 – 17)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814473E1ACB87005F4DB.taxon	materials_examined	Type material examined. Holotype of Geodromicus brevitarsus Champion, 1925 ♂ (Fig. 2; dissected): ‘ ♂ ’ <printed>, ‘ Shelshel, | N Kumaon, India | 15750 ft. H. G. C. ’ <printed>, ‘ Type | H. T. ’ <round label with red margin, printed>, ‘ Geodromicus | brevitarsis, | Champ. ’ <printed>, ‘ E. M. M. 1925. | det. G. C. C. ’ <printed>, ‘ G. C. Champion | Brit. Mus. | 1925 - 42. ’ <printed>, ‘ Microedus | brevitarsis Cam. [handwritten] | Shavrin A. V. det. 2016 ’ <printed> (BMNH).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814473E1ACB87005F4DB.taxon	description	Redescription. Measurements: HW: 0.55; HL: 0.35; OL: 0.12; LT: 0.07; AL: 1.83; PL: 0.50; PWMax: 0.62; PWMin: 0.42; ESL: 0.87; EW: 0.95; MTbL: 0.72; MTrL: 0.25 (MTrL 1 – 4: 0.15; MTrL 5: 0.10); AW: 1.00; AedL: 0.67; BL: 3.00. Habitus as in Fig. 2. Body reddish-brown, with slightly paler elytra; mouthparts, antennae and legs yellow (femora slightly darker). Head with dense microreticulation, isodiametric in middle and transverse on infraorbital portions, finer in portions between antennal bases and anterior margin of eyes; neck with large isodiametric meshes; pronotum with very dense and fine isodiametric sculpture, finer in mediobasal portion; scutellum with wide transverse meshes; abdominal tergites with dense, transverse microreticulation, finer on abdominal tergites VI – VIII. Head 1.5 times as broad as long, with slightly elevated infraorbital portions and widely U-shaped depression between eyes; medioapical depression relatively deep and wide; portion between ocelli and neck distinctly depressed; anteocellar foveae distinct, narrow and long, diagonally stretching posteriad toward level of anterior margins of eyes; temples distinctly shorter than longitudinal length of eyes. Eyes moderately small, slightly convex. Ocelli moderately large, located slightly behind level of posterior margins of eyes, distance between ocelli about one and half times as long as distance between ocellus and posterior margin of eye. Punctation very fine and dense, slightly deeper on infraorbital portions. Preapical segment of maxillary palpi significantly widened apicad, about as long as preceding segment; apical palpomere very narrow, about twice as long as preapical segment. Antennae reaching about middle length of elytra when reclined; basal antennomere moderately wide, about three times as long as broad, antennomere 2 slightly shorter and narrower than basal antennomere, 3 slightly longer and narrower than 2, 4 – 6 slightly shorter and wider than 3, 7 – 8 slightly wider than 6, 9 – 10 slightly shorter than 8, apical antennomere about 1.7 times as long as 10, from apical third gradually narrowed toward subacute apex. Pronotum slightly convex, 1.2 times as broad as long, 1.1 times as broad as head, widest in anterior portion, gradually rounded toward widely rounded anterior angles and sharply narrowed posteriad toward rounded posterior angles; anterior margin widely rounded, slightly longer than indistinctly concaved posterior margin; laterobasal portions distinctly depressed. Punctation very fine and dense, indistinct in middle. Elytra somewhat flattened, slightly broader than long, 1.7 times as long as pronotum, slightly broadened posteriad. Punctation dense, moderately deep and regular, denser around scutellum, finer and sparser along suture. Abdomen slightly broader than elytra, with two large and transverse tomentose spots in the middle of abdominal tergite IV. Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus elongate, narrow, from widest point at basal bulb gradually narrowed toward preapical part of median lobe, with rounded apex; parameres slightly exceeding apex of median lobe, with four moderately short apical setae; internal sac without any visible structures (Fig. 16). Lateral aspect of the aedeagus as in Fig. 17. Female unknown. Comparative notes. Microedus brevitarsis can be distinguished from all Palaearctic species of the genus by the distinctly smaller pronotum slightly broader than the head, and the morphology of the narrow aedeagus without sclerotized structures of the internal sac. Based on the slightly elongate antennomeres 4 – 10, it is somewhat similar to the Middle Asian M. kastcheevi sp. n. and M. schilenkovi, from which it can be distinguished by the presence of distinct interocellar foveae, narrower pronotum, and the morphology of the aedeagus.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF80814473E1ACB87005F4DB.taxon	distribution	Distribution. Microedus brevitarsis is known only from the type locality in Kumaon, Uttarakhand, India. Bionomics. The type specimen was collected at an elevation of about 4800 m a. s. l. Detailed bionomical data are unknown	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8F814573E1A8AA7039F4D8.taxon	description	(Figs 3, 22 – 23, 30)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8F814573E1A8AA7039F4D8.taxon	materials_examined	Type material examined. Holotype ♂ ‘ Kazakhstan | Narym Riv., Maimer | 06.08.1986 | V. Kastcheev’ <printed>, ‘ HOLOTYPE | Microedus | kastcheevi sp. n. | Shavrin A. V. des. 2024 ’ <red, printed> (ZIN). Paratypes: 9 ♂♂ (one specimen without head and pronotum; one specimen without head), 5 ♀♀: same data as the holotype (1 ♀: cSch; 4 ♂♂, 2 ♀♀: cSh; 5 ♂♂, 2 ♀♀: ZIN); 13 ♂♂ (two specimens without abdomen), 10 ♀♀: ‘ SE Kazakhstan, | Dzhungar Mts., Sarkand Riv. | 15.08.2006 | V. Kastcheev’ (3 ♂♂, 2 ♀♀: cSh; 1 ♂, 1 ♀: NME; 1 ♂, 1 ♀: NMW: 1 ♂, 1 ♀: ZMM; 7 ♂♂, 5 ♀♀: ZIN); 2 ♂♂, 4 ♀♀: ‘ SE Kazakshtan | Chylik Riv., ur. Sarybastau | 14.06.1988 | V. Kastcheev’ (1 ♂, 1 ♀: cSh; 1 ♂, 3 ♀♀: ZIN). All paratypes with additional red printed label: ‘ PARATYPE | Microedus | kastcheevi sp. n. | Shavrin A. V. des. 2024 ’.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8F814573E1A8AA7039F4D8.taxon	description	Redescription. Measurements (n = 44): HW: 0.65 – 0.78; HL: 0.42 – 0.56; OL: 0.13 – 0.20; LT: 0.10 – 0.15; AL (holotype): 2.10; PL: 0.55 – 0.74; PWmax: 0.85 – 1.02; PWmin: 0.70 – 0.88; ESL: 0.92 – 1.25; EW: 1.35 – 1.50; MTbL (holotype): 0.85; MTrL (holotype): 0.33 (MTrL 1 – 4: 0.16; MTrL 5: 0.17); AW: 1.20 – 1.61; AedL: 0.85 – 1.00; BL: 3.40 – 4.95 (holotype: 3.95). Habitus as in Fig. 3. Body brown to reddish-brown (some paratypes with distinctly paler elytra and darkened mediobasal portion). Head with moderately dense isodiametric or somewhat transverse microsculpture, distinctly finer in middle and coarser on infraorbital portions (holotype and several paratypes without visible meshes between anteocellar foveae); neck with dense transverse microreticulation; pronotum with dense transverse or isodiametric microsculpture, finer and sometimes indistinct in middle. Head 1.3 – 1.5 times as broad as long, with medioapical depression strongly narrowing posteriad; temples long, but distinctly shorter than longitudinal length of eyes. Ocelli located distinctly below level of posterior margins of eyes, about as long as distance between ocellus and posterior margin of eyes or slightly shorter. Punctation fine, denser and larger in middle and on infraorbital portions. Apical segment of maxillary palpi 1.4 – 1.6 times as long as preapical segment. Antennomeres 4 – 10 slightly elongate; antennomere 3 slightly longer than 2, 4 slightly shorter and broader than 3, 5 – 7 slightly longer and broader than 4, 8 – 10 slightly shorter and broader than 7, apical antennomere distinctly longer than 10, from about middle gradually narrowed toward subacute apex. Pronotum 1.2 – 1.3 times as broad as long, 1.3 times as broad as head, from widest anterior portion gradually narrowed posteriad. Punctation dense and fine, sometimes finer and sparser in middle; several paratypes without punctures in mediobasal third. Elytra 1.2 – 1.4 times as broad as long, 1.6 times as long as pronotum. Interstices between punctures in middle about as long as distance of one-two nearest punctures, portions of each elytron along suture with finer and sparser or sometimes with very dense punctation. Abdomen distinctly broader than elytra, with two transverse tomentose spots in the middle of abdominal tergite IV. Male. Apical margin of abdominal tergite VIII somewhat straight. Apical margin of abdominal sternite VIII widely concave. Aedeagus long, with very wide basal portion, strongly narrowed toward moderately wide median lobe, from apical third gradualy narrowed toward small rounded apex; parameres narrow, slightly broadened apically, distinctly exceeding apex of median lobe, with one short apical and two preapical setae; internal sac very long and narrow, with several very long sclerotized structures in basal portion (Fig. 22). Lateral aspect of the aedeagus as in Fig. 23. Female. Apical margins of abdominal tergite VIII and sternite VIII straight or somewhat rounded. Comparative notes. Altaiodromicus kastcheevi sp. n. can be distinguished from A. schilenkovi by the slightly broader head with slightly shorter temples, finer punctation of the middle portion of the head, and more distinct punctation of the pronotum. It can be easily distinguished from A. schilenkovi by the broader median lobe of the aedeagus, the lack of short setae along inner margin of each paramere, and the presence of large sclerotized structures in basal portion of the internal sac. Based on the slightly elongate antennomeres 4 – 10, it is somewhat similar to the Himalayan M. brevitarsis, from which it can be distinguished by the shallow interocellar foveae, broader pronotum, and the morphology of the aedeagus.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8F814573E1A8AA7039F4D8.taxon	distribution	Distribution. Altaiodromicus kastcheevi sp. n. is known from three localities in Altai Mts. and Dzhungar Mts., Kazakhstan (Fig. 30). Bionomics. Detailed bionomical data are unknown.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8F814573E1A8AA7039F4D8.taxon	etymology	Etymology. The species is named in memory of V. A. Kastcheev (20.12.1953 – 14.07.2012), collector of the type specimens.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8C814073E1A8AA75CEF49F.taxon	description	(Figs 4, 7, 9, 11, 13, 18 – 21, 24 – 30)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8C814073E1A8AA75CEF49F.taxon	materials_examined	Type material examined. Holotype ♂: ‘ Алтай, хр. Коргон | верх. руч. Мохнатый | прит. Кумира. 1750 - | 2000 м, 10 - 24. У [V] II. | В. Шиленков 1984 [Altai, Korgon Mts., upper flow of Mokhnatyi Stream, tributary of Kumir River […] V. G. Shilenkov leg.] ’ <photo label>, ‘ Коргон | 1950 м | берег оЗера [Korgon, 1950 m a. s. l., bank of the lake] ’ <handwritten by blue pen>, ‘ Holotypus | Altaiodromicus [handwritten] | schilenkovi [handwritten] | Zerche’ <red, printed> (ZIN). Paratypes 2 ♀♀: same data as that of holotype, except red printed label: ‘ Paratypus | Altaiodromicus [handwritten] | schilenkovi [handwritten] | Zerche’ <red, printed> (ISU, ZIN). Material examined. RUSSIA: ALTAI: 1 ♂, 2 ♀♀: same data as the type material (ISU); 1 ♀: Altaiskiy Nature Reserve, Akitchu River. 10.07.2002. A. V. Matalin & S. S. Demidov leg. (MSPU); 2 ♂♂, 2 ♀♀: same Nature Reserve and collectors, Teletskoye Lake, kordon Kokchin. 05.07.2002 (cSh, ZMM); 1 ♂, 1 ♀: SE slope of Aygulakskiy Mts., lower flow of Bel`gabash River. 1250 m a. s. l., under stones. 29.06.2000. S. S. Demidov leg. (cSh, MSPU); 1 ♂, 1 ♀: same data, middle flow of Bel`gabash River. 1700 m a. s. l., stones near river. 02 - 03.07.2000. K. V. Makarov leg. (cSh, ZMM); KAZAKHSTAN: EAST KAZAKHSTAN: 6 ♂♂, 5 ♀♀: Altai, connections of Kara-Koba and Tauchilik rivers. 15.08.1988. V. A. Kastcheev leg. (cSh, NME, ZIN); 14 ♂♂, 16 ♀♀: same data. 06.08.1989 (cSh, ZIN); 7 ♂♂, 4 ♀♀: same data. 16.08.1988 (cSh, ZIN); 1 ♂, 1 ♀: Karakoba River, 2 nd bridge. 16.08.1988. V. A. Kastcheev leg. (ZIN); 3 ♂♂, 2 ♀♀: Sorvenok River near Aksubas Mt. 12 - 14.08.1989. V. A. Kastcheev leg. (ZIN); 6 ♂♂, 2 ♀♀: Prokhodnoy Belok, Marchikha River. 12.07.1989. V. A. Kastcheev leg. (ZIN); 4 ♂♂, 5 ♀♀: same data. 06.07.1991 (cSh, ZIN); 27 ♂, 13 ♀♀: same data. 11.08.1989 (cSh, ZIN); 16 ♂♂, 20 ♀♀: Marchikha River. 12.07.1989. V. Kastcheev leg. (cSh, ZIN); 1 ♀: Markakol`, Sorvenok River. 14.08.1989. V. Kastcheev leg. (ZIN); 1 ♂: same data, 18.06.1988 (cSh); 3 ♂♂, 3 ♀♀: same data, 12.08.1989 (cSh); 1 ♀: Sarymsakty River, Katon-Karagai. 28.08.2010. V. Kastcheev leg. (NMW); 5 ♂♂, 5 ♀♀: Burkhat Pass. 06.08.1989. V. Kastcheev leg. (cSh, ZIN).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8C814073E1A8AA75CEF49F.taxon	description	Redescription. Measurements (n = 180): HW: 0.60 – 0.83; HL: 0.37 – 0.50; OL: 0.17 – 0.19; LT: 0.10 – 0.11; AL (averaged): 1.99; PL: 0.55 – 0.75; PWmax: 0.80 – 1.02; PWmin: 0.63 – 0.80; ESL: 0.92 – 1.28; EW: 1.22 – 1.41; MTbL (averaged): 0.87; MTrL (averaged): 0.32 (MTrL 1 – 4: 0.17; MTrL 5: 0.15); AW: 1.30 – 1.72; AedL: 0.45 – 0.77; BL: 3.15 – 4.60. Habitus as in Fig. 4. Body reddish-brown, sometimes with darker head and abdomen; mouthparts, antennae and legs yellow-brown; apical segment of maxillary palpi and tarsi yellowish. Head with dense and coarse isodiametric microreticulation, usually finer in middle and slightly coarser on infraorbital portions; neck with dense and coarse mesh-like sculpture; pronotum with very dense and coarse isodiametric microsculpture, somewhat similar to that on head, sometimes finer in medioapical portion; scutellum with fine isodiametric meshes; abdominal tergites with very dense isodiametric microreticulation, usually coarser on abdominal tergites III – IV. Head 1.6 times as broad as long; medioapical depression slightly or strongly narrowing posteriad; temples about 1.7 times as long as longitudinal length of eyes. Ocelli located slightly below level of posterior margins of eyes, distance between ocelli 1.2 – 1.4 times as long as distance between ocellus and posterior margin of eyes. Punctation of head very sparse and fine, indistinct in middle portion, usually larger and denser on infraorbital portions. Labrum as in Fig. 7. Mandibles as in Fig. 9. Maxilla as in Fig. 11; apical segment of maxillary palpi about 1.3 times as long as preapical segment. Labium as in Fig. 13. Antennomeres 4 – 10 slightly elongate; basal antennomere about three times as long as broad, 2 slightly narrower and distinctly shorter than basal antennomere, 3 slightly broader than 2, 4 moderately small, distinctly shorter than 3, 5 – 6 distinctly longer than 4, 7 – 8 slightly shorter than 6, 9 – 10 slightly broader than 8, apical antennomere about as long as two preceding antennomeres, from about middle gradually narrowed toward rounded or subacute apex. Pronotum 1.3 – 1.4 times as broad as long, 1.2 – 1.3 times as broad as head, gradually or strongly narrowed posteriad toward obtuse posterior angles; anterior margin rounded, about as long as straight posterior margin or slightly shorter. Punctation indistinct; longitudinal band of pronotal disc usually without punctation. Elytra distinctly broader than long, 1.6 – 1.7 times as long as pronotum, with widely rounded apical margins. Punctation distinctly finer around scutellum and along suture. Abdomen distinctly broader than elytra, with two transverse tomentose spots in the middle of abdominal tergite IV. Male. Apical margin of abdominal tergite VIII somewhat straight (Fig. 24). Apical margin of abdominal sternite VIII concave (Fig. 25). Genital segment as in Fig. 26. Aedeagus with wide basal portion, strongly narrowed toward narrow and moderately short median lobe with small rounded apex; parameres narrow, distinctly exceeding apex of median lobe, with two short apical setae and two additional short setae on inner margin of apical half of each paramere; internal sac narrow and moderately long, without additional sclerotized structures (Figs 18, 20, 21; Fig. 1 in Zerche (1992 )). Lateral aspect of the aedeagus as in Fig. 19. Female. Apical margins of abdominal tergite VIII (Fig. 27) and sternite VIII (Fig. 28) straight. Genital segment as in Fig. 29. Comparative notes. Altaiodromicus schilenkovi can be distinguished from A. kastcheevi sp. n. by the slightly broader head with slightly longer temples, somewhat coarser punctation of the middle portion of the head, and less distinct punctation of the pronotum. It can be reliably distinguished from A. kastcheevi sp. n. by the external and the internal structure of the aedeagus, with narrower median lobe, the presence of short setae on inner margin of each paramere and the lack of large sclerotized structures of the internal sac. Based on the slightly elongate antennomeres 4 – 10, it is somewhat similar to the Himalayan M. brevitarsis, from which it can be distinguished by the shallow interocellar foveae, broader pronotum, and the morphology of the aedeagus.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8C814073E1A8AA75CEF49F.taxon	distribution	Distribution. Altaiodromicus schilenkovi is known from several localities in Altai mountain range, Altai Republic (Russia) and north-eastern part of Kazakhstan (Fig. 30) Bionomics. Specimens from Russia were collected under stones near rivers and lakes at elevations from 1250 to 1950 m a. s. l. Detailed bionomical data for the specimens collected in Kazakhstan are unknown.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8B815C73E1ACD371C2F677.taxon	description	(Figs 31 – 42)	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8B815C73E1ACD371C2F677.taxon	materials_examined	Type material examined. Lectotype (here designated) of Anthophagus subtilis Sharp, 1889 ♀ (Figs 31 – 32): ‘ Anthophagus | subtilis. | Type D. S. | Iwake. Japan’ <handwritten on a card with the specimen, in black India ink>, ‘ Type’ <round label with red margin, printed>, ‘ Japan [underlined by yellow] | G. Lewis. ’ <printed>, ‘ Sharp Coll. | 1905 - 313. ’ <printed>, ‘ NHMUK 015009854 ’ <printed label, with BMNH barcode on rigth side of the label>, ‘ LECTOTYPE | Anthophagus | subtilis Sharp, 1889 | Shavrin A. V. des. 2024 ’ <red, printed>, ‘ Microedus | subtilis (Sharp, 1889) | Shavrin A. V., det. 2024 ’ <printed> (BMNH). Paralectotype ♀ (left apical antennomere, left middle tibia and tarsus and right elytron are absent): ‘ Anthophagus | subtilis. D. S. ’ <handwritten on a card with the specimen, in black India ink>, ‘ Japan. [underlined by yellow] | G. Lewis | 1910 - 320. ’ <printed>, ‘ Iwake’ <handrwitten>, ‘ Philydrodes [handwritten] | subtilis Sh. [handwritten] Det. C. Koch’ <printed>, ‘ Microedus | subtilis (Sharp, 1889) | Shavrin A. V., det. 2024 ’ <printed> (BMNH). Additional material examined: JAPAN: HONSHU: 7 ♂♂, 6 ♀♀: ‘ JAPAN: Honshu B. M. 1980 - 492 P. M. Hammond’, ‘ Gumma Pref. Mt, Hotaka (foot) ca 1300 m 14 - 15. viii. [19] 80 ’ (BMNH).	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8B815C73E1ACD371C2F677.taxon	description	Redescription. Measurements (n = 15): HW: 0.72 – 0.87; HL: 0.47 – 0.63; OL: 0.15 – 0.21; LT: 0.12 – 0.14; AL (lectotype): 2.96; PL: 0.67 – 0.78; PWmax: 0.90 – 1.03; PWmin: 0.75 – 0.87; ESL: 1.02 – 1.32; EW: 1.30 – 1.57; MTbL (lectotype): 1.05; MTrL (lectotype): 0.37 (MTrL 1 – 4: 0.17; MTrL 5: 0.20); AW: 1.35 – 1.68; AedL: 0.75 – 0.80; BL: 2.90 – 5.30 (lectotype: 4.70). Habitus as in Fig. 33. Body reddish-brown, usually with darker head and abdomen and paler elytra (some specimens with darker mediobasal portion of elytra; lectotype and paralectotype somewhat paler); antennae brown, sometimes with paler antennoimeres 1 – 2; mouthparts and legs yellow-brown to brown; apical maxillary palpomere and tarsi yellowish. Head with very dense isodiametric microreticulation, more transverse in middle of vertex, finer and sometimes indistinct in middle, and coarser on infraorbital portions, mediobasal portion of head with dense and fine isodiametric meshes; neck with dense and coarse isodiametric sculpture; pronotum with very dense and coarse isodiametric microsculpture, distinctly finer and sometimes indistinct in middle; scutellum with very fine transverse or isodiametric meshes; abdominal tergites with very dense and coarse isodiametric microreticulation. Head 1.3 – 1.5 times as broad as long; medioapical depression slightly or strongly narrowing posteriad toward level of apical margins of eyes; temples 1.2 – 1.5 times as long as longitudinal length of eyes. Ocelli located distinctly below level of posterior margins of eyes, distance between ocelli 1.3 – 1.6 times as long as distance between ocellus and posterior margin of eyes. Punctation of head sparse and irregular, sparser and finer (sometimes indistinct) in middle and slightly larger, deeper and denser on infraorbital portions. Antennomeres 4 – 10 elongate; basal antennomere slightly more than three times as long as broad, 2 slightly narrower and distinctly shorter than basal antennomere, 3 – 5 distinctly longer than 2, 6 – 7 slightly longer and indistinctly broader than 5, 8 – 10 slightly shorter than 7, apical antennomere 1.4 – 1.7 times as long as 10. Pronotum 1.3 times as broad as long, 1.1 – 1.2 times as broad as head, from widest anterior portion slightly or strongly narrowed posteriad, laterobasal margins in front of hind angles sometimes straight and parallel-sided. Punctation dense and fine, sparser and sometimes indistinct in middle. Elytra 1.1 – 1.2 times as broad as long, 1.5 – 1.6 times as long as pronotum. Punctation dense and regular, distinctly larger and deeper than that on pronotum, finer in parascutellar portion and along suture. Male. Apical margin of abdominal tergite VIII truncate (Fig. 37). Apical margin of abdominal sternite VIII widely concave (Fig. 38). Genital segment as in Fig. 39. Aedeagus with moderately narrow basal portion, gradually narrowed toward long and narrow median lobe, slightly (Fig. 34) or strongly (Fig. 35) narrowed apicad; parameres narrow, distinctly exceeding apex of median lobe, sometimes sligthly broadened apically, with two short apical and two preapical setae; internal sac narrow and long, with two narrow elongate and sclerotized structures in middle (sometimes these structures invisible). Lateral aspect of the aedeagus as in Fig. 36. Female. Apical margins of abdominal tergite VIII (Fig. 40) and sternite VIII (Fig. 41) somewhat straight. Genital segment as in Fig. 42. Comparative notes. Based on the shape of the slightly flattened pronotum, the presence of coriaceous ground sculpture on head and pronotum, and the general shape of the aedeagus, M. subtilis is similar to the Japanese M. yamanakai Watanabe, 1990, known from Honshu (Watanabe 1990), from which it can be distinguished by the larger eyes, the finer punctation of head and pronotum, and the distinctly narrower median lobe of the aedeagus.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8B815C73E1ACD371C2F677.taxon	distribution	Distribution. Microedus subtilis is known from Japan (Honshu) and Russia (Iturup Island, Kurile islands). Bionomics. Some specimens were collected under stones near streams and rivers. Detailed bionomical data are uknown.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
03D887EDFF8B815C73E1ACD371C2F677.taxon	discussion	Remarks. Anthophagus subtilis was originally descibed from “ … Iwakisan … ” based on “ … two mutilated examples ”. I studied two syntypes (females) from BMNH, and one of them, with additional label ‘ Type’, I designated here as the lectotype in order to fix the name. Nakane & Sawada (1956) regarded it as a species of Philydrodes (Lioplax). Watanabe (1990) redescribed it and transferred it to the genus Liophilydrodes, and provided additional material from northeastern Honshu and illustrations of the aedeagus. Philydrodes (Lioplax) troglophila was originally described based on two specimens from “ Nippara cave, Okutama, Tokyo ”. Watanabe (1990) redescribed it, transferred it to the genus Liophilydrodes, provided new material from Honshu and illustrations of the aedeagus. Based on the external morphology of the body, and the shape of the aedeagus, this species corresponds with other specimens of M. subtilis. I could not find sufficient morphological differences between the type and additional specimens. The apex of the median lobe is variable and can be narrow or somewhat wide (Figs 34 – 35 and Figs 407, 414 in Watanabe (1990 )). It should be noted that the width of the apical portions of the median lobe and the parameres is variable in many species of Anthophagini (e. g. Shavrin 2022 b). Thus, this species is synonymized with M. subtilis. Liophiydrodes subtilis iturupensis was originally described based on one female from Iturup Island (“ Konservnaya bay ”), Kurile islands. Based on the original description, features of the body of this taxon correspond with morphological features of specimens of M. subtilis. Thus, L. subtulis iturupensis is synonymyzed with M. subtilis.	en	Shavrin, Alexey V. (2024): Microedus LeConte, 1874 (Coleoptera: Staphylindae: Omaliinae: Anthophagini), a new genus for the Palaearctic Region. Zootaxa 5443 (2): 205-223, DOI: 10.11646/zootaxa.5443.2.4, URL: http://dx.doi.org/10.11646/zootaxa.5443.2.4
