identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DB557C912B415AE0C1A238FCD74097.text	03DB557C912B415AE0C1A238FCD74097.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Endecous (Endecous) painensis Castro-Souza & Junta & Reira 2020	<div><p>Endecous (Endecous) painensis n. sp.</p><p>(Figures 5–10, 11–18, 19, 20–25, 26–30, 31–32, Table 2)</p><p>Depository. Holotype and 12 Paratypes of E. (E.) painensis n. sp. (10 ♂♂ and 3 ♀♀), were deposited in the “Coleção de Invertebrados Subterrâneos de Lavras” (ISLA), Centro de Estudos em Biologia Subterrânea, Setor de Zoologia Geral, Departamento de Biologia, Universidade Federal de Lavras (Federal University of Lavras), Minas Gerais, Brazil.</p><p>Sc.d, endophallic sclerite duct; End.Sc.p, endophallic sclerite posterior portion</p><p>Material examined: Holotype, ♂, code ISLA 59121, Brazil, Minas Gerais, municipality of Pains, Brega cave (20° 25’ 5.25” S; 45°46’ 20.67”O), 20.i.2018, R. A. Castro-Souza, leg. Holotype condition: right tegmen were detached and maintained in holotype’s tube. Paratypes, 9 ♂♂ (ISLA 59122; 59123; 59124; 59125; 59126; 59127; 59128; 59129; 59130) and 3 ♀♀ (ISLA 59131; 59132; 59133), same locality of holotype., 20.i.2018, 03.ii.2018, 14.ix.2018, R. A. Castro-Souza, leg. Individuals examined, municipality of Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Mastodonte cave</a> (20° 25’ 34.44”S; 45°37’53.17”O), 30.i.2018, 17.iii.2018, R. A. Castro-Souza, leg., 2 ♂ (ISLA 59133; 59134); Pains, Macacos III cave (20° 24’ 28.44”S; 45° 40’ 18.82”O), 02. iii.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15381); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Loca D’Água de Baixo cave</a> (20° 25’ 25.04” S; 45°41’34.11” O), 10.ii.2016, R. A. Castro-Souza, leg., 1 ♂ (ISLA 59135); Pains, Loca dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Negros</a> cave (20° 26’ 7.66”S; 45° 39’ 34.70”O), 21.iv.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15349); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Éden cave</a> (20°23’5.24”S; 45°40’1.66”O), 06.x.2018, R. A. Castro-Souza, leg., 1 ♂ (ISLA 62610); Pains, Tabocas III cave (20°19’36.30”S; 45°41’24.97”O), 03.iii.2017, R. L. Ferreira, leg., 1 ♂ (ISLA 62611); Pains, Cornélio I cave (20°18’8.21”S, 45°40’9.76”O), 28.i.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15364); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Zizinho Beraldo cave</a> (20°21’22.69”S; 45°50’2.71”O), 21.i.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15380); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">São Lourenço cave</a> (20°19’34.40”S, 45°41’29.38”O), 20.iv.2018, R.A. CastroSouza, leg., 1 ♂ (ISLA 62612); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Cerâmicas cave</a> (20°24’15.46”S, 45°35’51.16”O), 03.iv.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 15346); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Vicente Amargoso cave</a> (20°23’17.69”S, 45°37’33.92”O), 26.vi.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 15394); Pains, Tio Rafa II cave (20°24’47.44”S, 45°39’52.88”O), 24.i.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 16082); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Paranoá cave</a> (20°21’54.23”S, 45°40’8.46”O), 27.i.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15411); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Torre cave</a> (20°25’25.85”S, 45°36’3.65”O), 03.ii.2017, R. A. Castro-Souza, leg., 1 ♂ (ISLA 62613); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Ponte Velha cave</a> (20°20’17.22”S, 45°48’37.59”O), 04.i.2017, R. A. Castro-Souza, leg., 1 ♂ (ISLA 64048); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Capoeirão cave</a> (20°20’25.42”S, 45°46’57.13”O), 22.i.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 16085); Pains, Dimas I cave (20°27’58.98”S, 45°39’41.49”O), 26.i.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 15412); Pains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Fisico cave</a> (20°24’3.72”S, 45°40’9.64”O), 01.iv.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 15352); Pains, Manada II cave (20°22’20.28”S, 45°40’26.40”O), 26.iii.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 15362); municipality of Doresópolis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Milagres cave</a> (20°20’36.24”S, 45°51’49.68”O), 13.i.2009, R. L. Ferreira, leg., 1 ♂ (ISLA 15382); Doresópolis, Bar-reado II cave (20°20’44.50”S, 45°51’30.03”O), 20.i.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 16076); Doresópolis, B 082 cave (20°18’16.78”S; 45°54’45.54”O,) 12.xi.2002, R. L. Ferreira, leg., 1 ♂ (ISLA 64047); Doresópolis, Helinho I cave (20°18’34.73”S, 45°50’47.35”O), 02.vii.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 62614); Doresópolis, Buraco dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Curiós</a> cave (20°18’19.44”S, 45°54’21.14”O), 04.vii.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 62615); Doresópolis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Peixe cave</a> (20°17’10.54”S, 45°47’40.18”O), 29.xii.2017, R. L. Ferreira, leg., 1 ♂ (ISLA 62616); Doresópolis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.364956&amp;materialsCitation.latitude=-20.843163" title="Search Plazi for locations around (long 45.364956/lat -20.843163)">Ninfeta de Baixo cave</a> (20°20’16.19”S, 45°36’53.42”O), 25.i.2009, R. A. Zampaulo, leg., 1 ♂ (ISLA 62617); municipality of Lagoa da Prata, Bicho que Foi cave (20° 4’33.19”S, 45°34’44.29”O), 03.v.2003, R. L. Ferreira, leg., 1 ♂ (ISLA 15385); municipality of Campo Belo, 52A cave (20°50’35.38”S; 45°21’53.84”O), 24.iii.2012, H. Nadolny, 1 ♂ (ISLA 16071) .</p><p>Distribution. This species is distributed in caves in the municipalities of Arcos, Campo Belo, Córrego Fundo, Doresópolis, Lagoa da Prata and Pains, many of them associated with the Arcos-Pains-Doresópolis speleological unit, which comprises the region with the highest density of caves known for South America (Fig. 1 A and B).</p><p>Etimology. Specific epithet “ painensis ” refers to the municipality Pains, type locality of the new species. This name originates in language, which cite the family “ Paim Pamplona ” around the 18th century, where was it usual pronounce the phrase “Let’s go to the farm of Paim ”, region where the name of the municipality originated.</p><p>Diagnosis. Combination of the following characters: phallic complex slightly more elongated than wide in dorsal or ventral view (Figs 5 and 6); pseudepiphallic dorsal branch underdeveloped, elongated, rounded at apex, with the presence of a small angulation, leaning in greater degree to inwards of the phallic complex (Figs 5, 7 and 9, Ps. db); pseudepiphallic paramere well developed, discoidal and concave (in dorsal, diagonal and front view) (Figs 5–9, Ps. P); a small region forming an arc partially sclerotized U-shaped (Fig. 5, blue arrow) between the bases of the pseudepiphallic parameres (Ps.P) and below the endophallic sclerite posterior portion (End.Sc.p); endophallic sclerite anterior portion well developed and sclerotized, with a groove (in ventral view); a dilated and elongated apodeme is evident after the groove (Figs 6 and 7, End.Sc.a, yellow arrow).</p><p>Description, male holotype ♂. Body color: head, pronotum and right tegmen brown color in dorsal view (Figs 12–14, 31), slightly light yellowish brown ventrally, as well as the abdomen (Fig. 15); legs slightly light yellowish brown at base and medium brown in the extremities, cerci pubescent and brown (Figs 17 and 18). Head: slightly pubescent, elongated in front view (4.65 and 3.43 mm, length and width respectively), vertex brownish, with a central band reaching the occiput; gena, clypeus, frons and labrum light yellowish brown, mandibles brown whitish and sclerotized in all its extension; first two maxillary palpomeres of similar proportion, smaller than the third and fourth that have the same proportion, fifth palpomere greater than the fourth, slightly elongated, claviform format, light yellowish brown and curved at apex; first, second and third labial palpomeres crescent in size, slightly pubescent, third claviform; scape, pedicel and antennomeres medium brown and pubescent, the region between the antennas has well developed bristles; compound eyes developed, slightly surrounded by depigmented ommatidium, and with a small depigmented dorsal field where scape fits; ocelli absent (Figs 11 and 12). Thorax: pronotum pubescent, with long bristles around the anterior and posterior extremities, a lighter vertical band crosses the center of the structure, lateral lobes rounded (Figs 13 and 15). Leg I: femur and tibia of similar size; oval auditory tympanum in the ventralproximal region of the tibia (Fig. 19–I, (t) highlighted in yellow); tibia armed with a pair of ventro-apical spurs of the same-size; first tarsomere serrulated, greater than the second and third tarsomeres (Fig. 19 –I–II). Leg II: tibia with a pair of internal and external apical spurs, dorsal smaller (Fig. 16 –III–IV, μ and λ highlighted in yellow). Leg III: femur developed; tibia serrulated of proportional size to the femur, armed with a sequence of three internal subapical spurs (Fig. 19–V, α, β and γ highlighted in yellow) and four outer side (Fig. 19 –VI, w, x, y and z highlighted in yellow), distal spur smaller (Fig. 19 –VI, z); four apical spurs at inner (Fig. 19–V, d, e, f and g highlighted in yellow) and and three at outer side (Fig. 19 –VI, a, b and c highlighted in yellow); three tarsomeres present, first serrulated distally and more elongated than the second and third, with a pair of apical spurs, the inner longer (Fig. 19 –V–VI, θ and δ highlighted in yellow). Right tegmen developed and sclerotized, covering the first three abdominal tergites (Fig. 15); mirror oval with three crosse-vein and fourth cells, first caudal vein has two small discontinuous branches and a small secondary cell; harp with six cross-veins, the last three caudal cells connected to a diagonal vein; lateral field with two large vertical cells and numerous accessory veins, from those are projected numerous irregular vein, some of these may be efurcate; basal field vein 1A, 2A, and 3A not furcate, main route leading up to the meeting of the 1A and 2A presents two small secondary cells (Figs 11 and 17); stridulatory file with 91 teeth. Abdomen: cerci elongated and pubescent, mainly on the basis; supranal plate short, rounded apex, lateral projections short (Figs 16 and 18); subgenital plate short, subtriangular apex (Fig. 17).</p></div>	https://treatment.plazi.org/id/03DB557C912B415AE0C1A238FCD74097	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Castro-Souza, Rodrigo Antônio;Junta, Vitor Gabriel Pereira;Reira, Rodrigo Lopes Fer-	Castro-Souza, Rodrigo Antônio, Junta, Vitor Gabriel Pereira, Reira, Rodrigo Lopes Fer- (2020): Description and ecology of a new species of the cricket genus Endecous (Orthoptera: Grylloidea: Phalangopsidae) in the speleological province of Arcos-Pains-Doresópolis, Southeastern Brazil. Zootaxa 4821 (2): 305-332, DOI: 10.11646/zootaxa.4821.2.4
