identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DB87E7FFA09D2362D9FD3F6BF1222E.text	03DB87E7FFA09D2362D9FD3F6BF1222E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Centromedon Sars 1891	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Centromedon Sars, 1891</p>
            <p> Centromedon Sars, 1891: 99 –101, pl. 34, fig. 2. — Barnard, 1962: 35. — Olerod, 1980: 50, 51. — Barnard &amp; Karaman, 1991: 474. — Lowry &amp; Stoddart, 1995: 13. </p>
            <p> Type species.  Anonyx pumilus Lilljeborg, 1865 , original designation. </p>
            <p> Diagnosis. Lateral cephalic lobes triangular, subacute to acute. Eyes apparently absent. Callynophore well-developed in both males and females. Epistome flattened and separated from upper lip by a sinus or notch; upper lip rounded and projecting in front of epistome. Inner lobes of lower lip weak or absent. Mandibular molar conicolaminate with vestigial triturative area, palp attached level with molar. Maxilla 1, inner plate stout with two pappose setae on apex (one in  C. pavor ); outer plate with 7–11 setal-teeth in 7–4 crown arrangement (reduced to 4–3 crown in  C. pavor and 6–3 crown in  C. pumilus ); palp two–articulate, reaching beyond outer plate. Maxilla 2, outer plate longer than inner plate. Maxilliped, outer plate not reaching article 3 of palp, suboval, with long robust setae apically, small and sparsely placed robust nodular setae on straight inner margin; article 4 of palp normal to vestigial. Gnathopod 1 variable from subchelate to simple; coxa 1 variable from strongly to weakly tapered, (not tapered in  C. typhlops ); ischium short (1: &lt;1.13). Gnathopod 2 subchelate to chelate. Pereopod 7 with distal articles shortened. Epimeron 3 posteroventrally produced into an acute tooth. Urosomite 1 with dorsal carina. Outer ramus of uropod 3 two-articulate, article 2 long (&gt;50% article 1 length). Telson longer than wide, deeply cleft. </p>
            <p> Species composition.  Centromedon pumilus (Lilljeborg, 1865) ;  Centromedon productus (Goës, 1866) ;  Centromedon typhlops (Sars, 1879) ;  Centromedon calcaratus (Sars, 1879) ;  Centromedon laevis (Bonnier, 1896) (new combination);  Centromedon mediator (Barnard, 1962) (new combination);  Centromedon pavor Barnard 1966 ; and  Centromedon zoe sp. nov.</p>
            <p> Remarks.  Uristes typhlops mediator Barnard, 1962 was originally described as a subspecies of  Centromedon typhlops (Sars, 1879) . An examination of the type material of both  Uristes typhlops mediator and  Centromedon typhlops shows that they differ by characters of the gnathopod 2, epimeron 3 and telson. There is also a clear size difference between the two species, with mature females of  C. typhlops reaching 17 mm in length while those of  C. mediator are 6 mm.  Uristes typhlops mediator conforms to the diagnosis of  Centromedon and is hereby transferred to this genus as  C. mediator . </p>
            <p> Orchomenella laevis Bonnier, 1896 was described from a single adult male (5 mm) taken in the Bay of Biscay (44°17' N, 02°18' W) at 950 m. It was transferred to  Tryphosella by Barnard, 1964 but it has a non–dominant epistome, reduced distal articles of pereopod 7 and otherwise conforms to the new diagnosis of  Centromedon , except for the outer plate of the maxilliped, which does reach article 3 of the palp. It is hereby transferred to this genus as  C. laevis . </p>
            <p> The distinctions between the genus  Uristes and  Centromedon are unclear but clarification awaits a thorough redescription of  Uristes gigas Dana, 1849 , the type species of  Uristes and a re-examination of the species currently included in that genus, an analysis which is beyond the scope of this paper. The location of Dana’s material is unknown. It is very likely that the type material was part of a collection on loan from the Smithsonian Institution to William Stimpson, at the Chicago Academy of Science, at the time of the Chicago fire in 1871. All of Stimpson’s material was lost in the fire and it is known that he had on loan 10,000 jars of Crustacea, many of which contained type material described by James Dana (Evans 1967). Type material is not held at the Natural History Museum, London (Thurston &amp; Allen 1969) and is presumed to be no longer extant. It is generally accepted, however, that  Tryphosa antennipotens Stebbing, 1888 is synonymous with  Uristes gigas (see Stebbing 1899). A full redescription of this species and designation of a neotype is needed to clarify the status of the genus. It is probable that a large number of the species currently included in the genus (22+) will be transferred to other genera following a critical analysis. </p>
            <p> Centromedon should be retained for those species characterised by: sub-acute to acute head lobes; tapered coxa 1; laminar, unridged molar with vestigial triturative area; maxilla 1 outer plate with 7-4 crown setal-tooth arrangement; shortened distal articles of pereopod 7 and acutely produced epimeron 3. </p>
            <p> The similarity of  Centromedon to  Galathella Barnard &amp; Karaman, 1987 was noted by Lowry &amp; Stoddart (1995), who pointed out that these are the only two genera in the  Uristidae with a tapering first coxa and 7-4 crown setal-tooth arrangement. It was also noted that both genera show a range of gnathopod 1 morphologies with species characterised by a simple, weak structure through to others with a strongly subchelate gnathopod 1. The only differences between the two genera appear to be the sub-acute lateral cephalic lobe (rounded in  Galathella ), the lack of proximal A3 setae on the mandibular palp, the less well-developed molar, and the acutely produced posteroventral corner of epimeron 3 in  Centromedon . Contrary to Lowry &amp; Stoddart (1995), most species of  Centromedon (  C. productus ,  C. typhlops ,  C. zoe and  C. mediator ) have callynophores in the female. There is a degree of variability within the genus in that  C. pumilus and  C. pavor have a reduced number of setal-teeth on the outer plate of maxilla 1, a reduced article 4 of the maxilliped palp and less well-developed callynophore and gnathopod 1 (Olerod, 1980). </p>
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	https://treatment.plazi.org/id/03DB87E7FFA09D2362D9FD3F6BF1222E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Horton, Tammy;Thurston, Michael	Horton, Tammy, Thurston, Michael (2011): Centromedon zoe (Crustacea: Amphipoda: Lysianassoidea: Uristidae), a new deep-water scavenger species from the North Atlantic, with a key to the genus Centromedon. Zootaxa 2869: 54-62, DOI: 10.5281/zenodo.277530
03DB87E7FFA39D2662D9FC346FAD222E.text	03DB87E7FFA39D2662D9FC346FAD222E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Centromedon zoe	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Centromedon zoe sp. nov.</p>
            <p>(Figures 1–3)</p>
            <p>Type material. Holotype: female, 6.5 mm and 9 slides, NHMUK 2011.925. Allotype: male 6.3 mm (and one slide mount of antennae) NHMUK 2011.946. Paratypes: 12 mature females, average 6.5 mm (4.9–7.0 mm), NHMUK 2011.926 – 935; 12 mature males, average 6.1 mm (5.5–7.3 mm), NHMUK 2011.936 – 945; 5 mature females and 5 mature males (AM P.85244); collected at the Mid-Atlantic Ridge in the northern Atlantic Ocean, RRV James Cook, (cruise 011), station number JC011/098; freefall, acoustically-released, baited trap, deployed at 1313 hrs (UTC) 9 August 2007, 54° 04.08' N, 34° 09.43' W at 2500 m, released at 1125 hrs, 11 September 2007; bottom time, 46 hrs.</p>
            <p>Additional material. 678 specimens, Mid-Atlantic Ridge, JC011/079, 05–07/08/2007, 53°56.44'N, 36°11.56'W, 2564 m; 1861 specimens, Mid-Atlantic Ridge, JC011/114, 12–13/08/2007, 54°02.31'N, 34°09.60'W, 2453 m; 860 specimens, Mid-Atlantic Ridge, JC011/098, 09–11/08/2007, 54°04.08'N, 34°09.43'W, 2500 m. 879 specimens, base of Sedlo Seamount, Azores, Stn. 56319#1, 40° 11.43' N, 26° 33.99' W, 2655 m; 574 specimens, 56354#1, 39° 50.17' N, 26° 17.82' W, 2876 m. 21 specimens, Stn. 51403#1, Benthic Net 1.5/ 3m, 51° 37.7' N, 12° 59.8' W, 1292–1314 m; 6 specimens, Stn. 51403#2, Benthic Net 1.5/ 3m, 51° 37.4' N, 12° 59.2' W, 1317–1325 m; 1 specimen, Stn. 51403#3, Benthic Net 1.5/ 3m, 51° 36.8' N, 12° 59.1' W, 1319–1325 m; 11 specimens, Stn. 51403#4 Benthic Net 1.5/ 3m, 51° 36.7' N, 12° 59.6' W, 1319–1333 m; 8 specimens, Stn. 51403#5, Benthic Net 1.5/ 3m, 51° 37.8' N, 12° 58.9' W, 1289–1297 m; 11 specimens, Stn. 51420#1, Benthic Net 1.5/ 3m, 51° 37.3' N, 12° 58.6' W, 1326–1328 m; 11 specimens, Stn. 51420#3, Benthic Net 1.5/ 3m, 51° 38.3' N, 12° 58.9' W, 1293–1298 m; 6 specimens, Stn. 51420#4, Benthic Net 1.5/ 3m, 51° 37.9' N, 12° 59.5' W, 1302–1319 m – all from the ‘ Pheronema’ ground in the Porcupine Seabight.</p>
            <p>Diagnosis. Lateral cephalic lobes subacute, not projecting; mandibular palp with 8 distolateral A2 setae; maxillipedal palp article 4 well-developed; coxa 1 weakly tapered, anterior margin concave; gnathopod 1 subchelate, propodus subrectangular; basis anterior margin fully setose; gnathopod 2 minutely subchelate; propodus rectangular; pereopods 3 and 4 dactyli long, subequal to propodus; epimeron 3 tooth acute, not upcurved; urosome 1 with distinct triangular acute carina.</p>
            <p>Description. Holotype female 6.5 mm. Head: exposed, as long as deep; lateral cephalic lobe large, narrow, subacute; Eyes: apparently absent. Antenna 1: short; peduncular article 1 short; peduncular article 2 short, length 0.3 x article 1; peduncular article 3 short, length 0.3 x article 1; primary flagellum nine- articulate; accessory flagellum long, length 0.45 x primary flagellum, three-articulate, forming cap, partially covering callynophore; callynophore strong, two-field; calceoli absent. Antenna 2: short, length 1.17 x antenna 1; peduncle without brush setae; peduncular article 1 not greatly enlarged; weakly geniculate between peduncular articles 3–4; article 3 long, length 0.8 x article 4; flagellum well-developed, nine-articulate.</p>
            <p>Epistome and upper lip: separate, upper lip dominant, weakly produced. Lower lip lobes: widely separated, inner lobes present. Mandible: incisor ventral margin smooth with small hook; lacinia mobilis present only on left mandible, a stemmed irregularly cusped peg; left and right accessory setal rows with simple robust setae and plumose setae; molar laminar, unridged with vestigial triturative area at tip; palp attached level with molar; article 1 short, length 0.8 x breadth; article 2 slender, length 5 x breadth, with eight distolateral A2 setae; article 3 slender, blade-like, with two long B3 setae, ten D3 setae and three E3 setae. Maxilla 1: inner plate narrow, with 2 apical pappose setae; outer plate with setal-teeth in 7/4 crown arrangement, outer row with ST1–7 large and slender, ST1 four-cuspidate, ST2 three-cuspidate, ST3 four-cuspidate, ST4 six-cuspidate, ST5 seven-cuspidate, ST6 six-cuspidate, ST7 displaced from ST6, five-cuspidate, STA-D large, slender, STA four-cuspidate, STB five-cuspidate, STC six-cuspidate, STD six-cuspidate; palp large, two-articulate, article 1, short, article 2 with seven robust terminal setae, one flag seta and one sub-terminal simple seta. Maxilla 2: inner plate narrow, tapering distally, shorter than outer plate, with two sub-parallel, medial setal rows; outer plate with short robust and plumose setae. Maxilliped: inner plate large, suboval with three robust nodular setae, setal row strong, with eight pappose setae reaching apical margin, and two marginal setae; outer plate medium, subrectangular, with six simple robust setae apically and ten robust nodular setae medially; palp large, four-articulate, article 2 slender, length 2.5 x breadth, article 3 long, slen- der, length 1.9 x breadth, article 4 well-developed, with one subterminal seta.</p>
            <p>Gnathopod 1: subchelate; coxa large, slightly shorter than coxa 2, weakly tapered distally, anterior margin concave, anteroventral corner broadly rounded, basis slender, long, length 4.4 x breadth, strongly setose anteriorly; ischium short, length 1.0 x breadth; carpus subrectangular, lacking posterior lobe, subequal in length to propodus, length 1.9 x breadth; propodus subrectangular, margins subparallel, palm oblique. Gnathopod 2: minutely subchelate; coxa large, subequal to coxa 3; ischium long, length 2.5 x breadth; carpus length 2.4 x breadth; propodus subrectangular; dactylus inserted at anterior corner of propodus, reaching palm edge. Pereopod 3: coxa large, subrectangular; basis slender, straight, margins subparallel, dactylus long, straight. Pereopod 4: coxa deeper than wide with posteroventral lobe broadly rounded, posterior margin straight. Pereopod 5: coxa posterior lobate (no lateral ridge); basis expanded, posterior margin weakly convex, posterior lobe narrowly rounded. Pereopod 6: coxa small, posterior lobate; basis expanded, posterior margin weakly convex, posterodistal lobe narrowly rounded. Pereopod 7: coxa small; basis broadly expanded and rounded, posterodistal lobe broadly rounded; distal articles shortened, together 0.76 x length of basis.</p>
            <p>Pleonites 1 to 3: not carinate. Epimeron 2: posteroventral corner convex, broadly rounded. Epimeron 3: posteroventral corner produced acutely. Urosome: urosomite 1 with distinct carina, forming a posteriorly directed acute ‘tooth’. Uropod 1: peduncle, length 1.24 x inner ramus, with four lateral robust setae and six medial robust setae; outer ramus subequal to inner ramus; inner ramus with one medial robust seta and two lateral robust setae; outer ramus with one lateral robust seta and two medial robust setae. Uropod 2: peduncle, length 0.86 x inner ramus, with four lateral robust setae; outer ramus subequal to inner ramus; inner ramus with two medial robust setae and one lateral robust seta; outer ramus with two lateral robust setae. Uropod 3: peduncle, length 0.8 x inner ramus, with four apicolateral robust setae, four apicomedial robust setae and two medial simple slender setae; inner ramus shorter than outer ramus, length 0.84 x outer ramus, with one medial slender seta, one medial robust seta and one lateral robust seta; outer ramus two-articulate, article 2 length 0.77 x article 1, article 1 with one medial robust seta and three lateral robust setae. Telson: long, lobes tapering, length 1.56 x breadth, cleft 81%, with two dorsal robust setae per lobe; apices incised, each with apical notch and one robust seta.</p>
            <p>Male. As for female except Antenna 1: peduncular article 2, length 0.23 x article 1; peduncular article 3, length 0.18 x article 1; primary flagellum 11-articulate; accessory flagellum, length 0.65 x primary flagellum, calceoli present. Antenna 2: long, length 2.3 x antenna 1; peduncle with brush setae; calceoli present; flagellum 22- articulate.</p>
            <p>Distribution. North-east Atlantic Ocean, 1289–2876 m; Mid-Atlantic Ridge, Azores seamounts, Porcupine Seabight.</p>
            <p> Etymology. The species is named in honour of the first author’s first daughter,  Zoë , noun in apposition. </p>
            <p> Remarks. This small scavenging species has been captured in great numbers in baited traps set at around 2500 m at the Mid-Atlantic Ridge. It resembles most closely the species  C. typhlops ,  C. mediator and  C. laevis . It can be distinguished from  C. typhlops by the acute boss on urosome 1, the longer dactyls of the pereopods, the rectangular form of the propodus of gnathopod 2 (suboval in  C. typhlops ), details of the mouthparts and antennae, and the much smaller mature female size (6 mm in  C. zoe vs. 17.5 mm in  C. typhlops ). It can be distinguished from  C. mediator by the form of gnathopod 2 (chelate in  C. mediator ), the number of articles in the antennae, the less welldeveloped molar, the number of A2 setae on the mandibular palp, and the less produced epimeron 3 tooth. It can be distinguished from  C. laevis by the fully setose anterior margin of the basis of gnathopod 1 (only two setae in  C. laevis ), the length of the dactyls on pereopods 3 and 4 (subequal to propodus in  C. zoe , longer than propodus in  C. laevis ), the anterior margin of coxa 1 (concave in  C. zoe , convex in C.  laevis ) and the shape of coxa 4 posterior lobe (more rounded in  C. zoe ). </p>
            <p> Tryphosella abyssi Norman, 1900 , included under  Uristes by Barnard &amp; Karaman (1991) has been recorded only once. A single 6 mm male specimen was collected at 1071 m in the ‘cold area of the Faroe Channel’ by Sir John Murray, aboard HMS  Triton , in 1882 (Station 7: 60°19' N, 7°10' W). Type material of this species is not in the Norman Collection at the Natural History Museum. The description and illustrations given by Norman (1900), particularly of the sub-acute lateral cephalic lobes, the dominant upper lip, and the acute urosome boss, place it close to the new species  Centromedon zoe . However, the Norman species is distinguished by the rounded posterodistal lobe of epimeron 3. Without type material, or further material that can be linked with this species it is hereby referred to nomen dubium. </p>
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	https://treatment.plazi.org/id/03DB87E7FFA39D2662D9FC346FAD222E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Horton, Tammy;Thurston, Michael	Horton, Tammy, Thurston, Michael (2011): Centromedon zoe (Crustacea: Amphipoda: Lysianassoidea: Uristidae), a new deep-water scavenger species from the North Atlantic, with a key to the genus Centromedon. Zootaxa 2869: 54-62, DOI: 10.5281/zenodo.277530
03DB87E7FFA69D2662D9FC336BF124E1.text	03DB87E7FFA69D2662D9FC336BF124E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Centromedon	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to the species of  Centromedon</p>
            <p> 1. Gnathopod 1 simple; pereopod 7, basis with prominent postero-distal spur...........  Centromedon calcaratus (Sars, 1879)</p>
            <p>- Gnathopod 1 subchelate; pereopod 7, basis lacking prominent postero-distal spur................................... 2</p>
            <p> 2. Lateral cephalic lobe strongly projecting, acute; epimeron 3 with postero-distal corner produced into an acute strongly upcurved tooth...........................................................  Centromedon productus (Goës, 1866)</p>
            <p>- Lateral cephalic lobe subacute; epimeron 3 produced into an acute tooth, not strongly upcurved........................ 3</p>
            <p>3. Coxa 1 tapering strongly, sub-triangular; gnathopod 1 propodus sub-triangular, palmar angle obscure................... 4</p>
            <p>- Coxa 1 tapering weakly; gnathopod 1 propodus sub-rectangular, palmar angle distinct............................... 5</p>
            <p> 4. Lateral cephalic head lobe short, sub-triangular; dactyls of pereopods 3 and 4 longer than propodus; article 4 of maxillipedal palp vestigial.............................................................  Centromedon pavor (Barnard, 1966)</p>
            <p> - Lateral cephalic head lobe produced, sub-acute; dactyls of pereopods 3 and 4 shorter than propodus; article 4 of maxillipedal palp short............................................................  Centromedon pumilus (Lilljeborg, 1865)</p>
            <p>5. Urosomite 1 with distinct triangular acute carina............................................................ 6</p>
            <p> - Urosomite 1 lacking distinct triangular acute carina...............................  Centromedon typhlops (Sars, 1879)</p>
            <p> 6. Gnathopod 2 chelate....................................................  Centromedon mediator (Barnard, 1962)</p>
            <p>- Gnathopod 2 minutely subchelate......................................................................... 7</p>
            <p> 7. Gnathopod 1 basis setose anteriorly; pereopods 3 and 4 dactyls subequal in length to propodus; coxa 1 concave anteriorly...................................................................................  Centromedon zoe sp. nov.</p>
            <p> - Gnathopod 1 basis with two setae anteriorly; pereopods 3 and 4 dactyls longer than propodus; coxa 1 convex anteriorly..............................................................................  Centromedon laevis (Bonnier, 1896)</p>
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	https://treatment.plazi.org/id/03DB87E7FFA69D2662D9FC336BF124E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Horton, Tammy;Thurston, Michael	Horton, Tammy, Thurston, Michael (2011): Centromedon zoe (Crustacea: Amphipoda: Lysianassoidea: Uristidae), a new deep-water scavenger species from the North Atlantic, with a key to the genus Centromedon. Zootaxa 2869: 54-62, DOI: 10.5281/zenodo.277530
