taxonID	type	description	language	source
03DD6822FFA6326BFF32FE59FA4F627A.taxon	type_taxon	Type species. Attus redii Audouin, 1826.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326BFF32FE59FA4F627A.taxon	discussion	Remarks. The genus includes 43 nominal species (WSC 2020), many of them poorly studied and known from one sex only. One of the most important diagnostic characters, the shape of the embolus, in particular its base, is documented for only a few species. The majority of Langona representatives are known from Eurasia and Africa. The major papers on relationships and distributions of Oriental Langona are: Hęciak & Prószyński (1983), Prószyński (1992), Song et al. (1999), Peng et al. (2008), Caleb et al. (2015), Li & Lin (2016) and Caleb (2020).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	description	Figs 1 – 20	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	etymology	Etymology. The species is named after the type locality.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	diagnosis	Diagnosis. In the similar L. davidi (Caleb, Mungkung & Mathai, 2015), the anterior tegular outgrowth is oriented retrolaterally vs prolaterally in L. pattayensis sp. nov. The embolus (Fig. 10) is much longer than in L. davidi (see Caleb 2020: fig. 17 I), coiling more than 360 ° vs 180 º in L. davidi. The shape of the epigyne (Fig. 18) is most similar to that of L. pallida Prószyński, 1993, but the pockets are deeper, and the spermathecae (Figs 19 – 20) have more distinctive accessory glands.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	description	Description. Male (Holotype, UUME UPSZTY 155662, Figs 1 – 12). Carapace rather low and flat. Thoracic part almost twice as long as cephalic part, with 2 longitudinal stripes of white setae. Clypeus high, light brown, cheeks covered with white scales. Endites and labium light brown, with light tips. Sternum brown. Legs light brown, dorsally darker, legs I and IV with brown joint areas. Leg spination not distinctive. Dorsal opisthosoma with delicate scutum making up 57 % of opisthosoma length, colouration light brown, centrally lighter with long protruding anterior white and brown setae. Spinnerets not distinctive, brown. Ventral opisthosoma light brown, with light setae. Palp as in Figs 7 – 12. Ventral tibial apophysis in form of round outgrowth, retrolateral apophysis wide at base, distally pointed, ventrally bent and accompanied by long, thick, brown bristles. Tegulum elongate, with posterior pointed lobe and apical outgrowth. Embolus long, proximal part hidden by tegulum, coiling ca. 540 º, visible distal part needle-like. Dimensions: CL 2.90, CW 2.08, CH 1.39, clypeus height 0.27, EFL 0.99, AEW 1.57, AME diameter 0.47, PEW 1.39, OL 2.75, OW 1.80, legs: I 4.81 (0.30, 1.45, 0.93, 0.89, 0.65, 0.59), II 4.47 (0.28, 1.36, 0.87, 0.81, 0.53, 0.62), III 6.40 (0.32, 2.03, 1.03, 1.06, 1.32, 0.64), IV 6.19 (0.32, 1.70, 1.02, 1.08, 1.37, 0.70). Female (Paratype, UUME UPSZTY 155663, Figs 13 – 20). Carapace brown, with darker eye field, covered with brown setae and white scales, latter forming two thoracic stripes. Clypeus light brown, cheeks covered with white scales. Chelicerae light brown, palps light brown with white setae. Endites and labium light brown, with lighter tips, sternum elongate, yellowish. Legs light brown, with brown patches and annulations. Leg spination not distinctive. Opisthosoma poorly preserved, dorsally light brown, covered with brown setae. Ventral opisthosoma light grey. Spinnerets not distinctive, light brown. Epigyne as illustrated in Figs 18 – 20: with 2 merging posterior pockets, copulatory openings crescent-shaped, oriented towards each other, copulatory ducts C-shaped, membranous, spermathecae thick-walled, strongly sclerotized, multi-chambered, with distinctive accessory glands. Dimensions: CL 3.06, CW 2.08, CH 1.52, clypeus height 0.10, EFL 1.05, AEW 1.60, AME diameter 0.31, PEW 1.47, OL 4.15, OW 2.92, legs: I 4.88 (0.38, 1.57, 0.97, 0.84, 0.61, 0.51), II 4.61 (0.38, 1.43, 0.94, 0.71, 0.57, 0.58), III 6.14 (0.44, 1.17, 1.09, 1.07, 1.21, 0.63), IV 6.09 (0.47, 1.68, 0.84, 1.09, 1.40, 0.61).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	materials_examined	Type material. Holotype ³: THAILAND: Chonburi Province: Pattaya, leg. Å. Holm, 2 – 15. XII. 1979 (amongst grass near seashore) (UUME UPSZTY 155662). Paratypes: Same data as holotype, 13 ³, 1 ♀ (UUME UPSZTY 155663).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	distribution	Distribution. Known only from the type locality (Fig. 34).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA6326FFF32FC74FCC0609F.taxon	discussion	Remarks. The copulatory organs are very uniform within the genus. The new species is most similar to L. albolinea Caleb & Mathai, 2015, L. alfensis Hęciak & Prószyński, 1983, L. davidi, L. hongkong Song, Xie, Zhu & Wu, 1997 and L. simoni Hęciak & Prószyński, 1983, but the true relationships may not reflect these similarities; all species listed are known from south and southeast Asia.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA2326FFF32FDA8FA46629C.taxon	type_taxon	Type species. Attus membrosus Simon, 1868.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA2326FFF32FDA8FA46629C.taxon	discussion	Remarks. The body form and structure of the copulatory organs are similar to those in related Bianor Peckham & Peckham, 1886, Harmochirus Simon, 1885, Sibianor Logunov, 2001 and Modunda Simon, 1901. All species of Neaetha are known from Africa and southern Europe, except N. tomkovichi Logunov, 2019, which is found in India.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	description	Figs 21 – 34	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	etymology	Etymology. The species is named in honour of Professor Wanda Wesołowska (Wrocław University, Poland), distinguished Polish arachnologist and world authority of salticid taxonomy.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	diagnosis	Diagnosis. The palps are similar to those of N. absheronica Logunov & Guseinov, 2001, N. tomkovichi and N. membrosa (Simon, 1868), but the embolus is crescent shaped versus almost straight, narrower at the base, and the tegulum more elongate in the new species (Figs 24 – 27). The retrolateral tibial apophysis is long, dorsally oriented, more slender than in the three species mentioned above. The epigyne is similar to that of N. tomkovichi, but the pockets are relatively larger, and the spermathecae are distant from one another versus touching to one other. The copulatory openings are ovoid versus crescent shaped in most species. The copulatory ducts are much shorter than those of other species (Figs 32 – 33).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	description	Description. Male (Holotype, UUME UPSZTY 155664, Figs 21 – 27). Specimen in poor condition. Carapace brown, robust, rather high, widest behind PLE, thoracic slope distinctive, fovea just behind PLE. Entire surface covered with white scales, more numerous on sides and near eyes. Clypeus brown, with light scales along the lower margin. Chelicerae brown, with 2 promarginal teeth and single retromarginal tooth. Endites and labium brown, with lighter tips, sternum light brown. Legs brown, legs I thickest, tibia swollen, femur, patella and tibia with feathery ventral setae. Leg spination not distinctive. Opisthosoma ovoid, dorsally with 3 transverse, wide, brown stripes disconnected medially. Setae and scales rather numerous, whitish and brown, anterior setae longer. Ventral opisthosoma light grey, with darker spots. Spinnerets light brown, not distinctive. Palps as in Figs 24 – 27. Retrolateral tibial apophysis long, gradually narrowing, tip pointed and oriented dorsally. Tegulum ovoid, longer than wide, sperm duct not meandering. Embolus crescent-shaped, originating at 9 o’clock position. Dimensions: CL 1.49, CW 1.30, CH 0.65, clypeus height 0.10, EFL 0.56, AEW 0.91, AME diameter 0.30, PEW 0.95, OL 1.20, OW 0.71, legs: I 3.15 (0.21, 0.96, 0.62, 0.66, 0.42, 0.28), II? (0.18, 0.61, 0.33,?), III 3.13 (0.24, 1.06, 0.52, 0.55, 0.39, 0.37), IV 2.40 (0.18, 0.70, 0.36, 0.40, 0.42, 0.34). Female (Paratype, UUME UPSZTY 155665, Figs 28 – 33). Carapace rather high, thoracic slope distinctive, fovea short, just behind PLE. Colouration brown, eye surroundings black. Entire surface covered with white scales and fewer brown setae. Clypeus brown, with white scales and longer setae centrally. Chelicerae brown, endites light brown, labium darker. Sternum ovoid, light brown. Legs light brown, femora III with proximal and distal brown rings. Legs I thickest, covered with numerous whitish scales and brown setae. Leg spination as in male, not distinctive. Dorsal opisthosoma yellowish, with darker spots, covered with light and brown scales. Ventral opisthosoma yellowish, with irregular darker spots, covered with white and fewer brown setae. Spinnerets light brown. Epigyne with large, central anterior pocket, copulatory openings ovoid, copulatory ducts very short, membranous, spermathecae strongly sclerotized, as long as wide, separated by ca. one spermathecal radius (Figs 32 – 33). Dimensions: CL 0.86, CW 0.74, CH 0.50, clypeus height 0.13, EFL 0.47, AEW 0.65, AME diameter 0.38, PEW 0.71, OL 0.97, OW 0.72, legs: I 3.22 (0.27, 0.91, 0.64, 0.63, 0.46, 0.31), II 2.40 (0.21, 0.73, 0.50, 0.37, 0.29, 0.30), III 3.83 (0.30, 1.24, 0.64, 0.69, 0.49, 0.47), IV 2.76 (0.26, 0.84, 0.40, 0.44, 0.42, 0.40).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	materials_examined	Type material. Holotype ³: THAILAND: Kanchanaburi Province: Ban Say Yok, Kwai River, 100 m a. s. l., leg. Å. Holm, 24 – 25. XI. 1979 (secondary rain forest) (UUME UPSZTY 155664). Paratype ♀: THAILAND: Chonburi Province: Pattaya, leg. Å. Holm, 27. XI. 1979 (on shrubs) (UUME UPSZTY 155665).	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	distribution	Distribution. The easternmost record of Neaetha species (Fig. 34); known only from the type locality.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
03DD6822FFA23262FF32FC11FDAD644B.taxon	discussion	Remarks. The copulatory organs are similar to those of N. absheronica from Azerbaijan, Crete, Greece, North Macedonia, Turkey, N. tomkovichi from India and N. membrosa occurring from the western Mediterranean to Germany. The great distance between the localities of the aforementioned species and N. wesolowskae sp. nov. suggests that the reconstruction of intrageneric relations should be based on genetic rather than morphological data. Both sexes are from different localities, and the collection of a male and a female from the same locality will help confirm their conspecific affiliation.	en	Żabka, Marek, Patoleta, Barbara M. (2020): New species of Langona Simon, 1901 and Neaetha Simon, 1884 (Araneae Salticidae) from Thailand. Zootaxa 4899 (1): 374-383, DOI: 10.11646/zootaxa.4899.1.19
