taxonID	type	description	language	source
03DD4C47FFCEFFE6FF33B858FD03F936.taxon	description	(Fig. 1)	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCEFFE6FF33B858FD03F936.taxon	materials_examined	Type material. Holotype: ISEA MA 2013 - 1, Burmese amber, Myanmar; Cenomanian, late Cretaceous. Syninclusion. The second damaged specimen of this species is not included in the type series.	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCEFFE6FF33B858FD03F936.taxon	etymology	Etymology. The epithet of this new species is in honour of Evgeny E. Perkovsky, a specialist in fossil insect.	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCEFFE6FF33B858FD03F936.taxon	description	Description. Body black, convex dorsally, covered with short suberect setae. Forehead convex, broad, densely punctate and granulate. Eyes well protruded, finely facetted, rounded. Antennae elevensegmented, slightly serrate, without club. First antennomere about 1.8 times as long as broad at apex. Second antennae subequal in length and width, about 0.4 times as long and about 0.8 times as narrow as the first. Third antennomere about 1.3 times as long and wide at apex, about 1.7 times as long and about 1.3 times as wide as second antennomere. Fourth antennomere 1.4 times as long as broad at apex, 1.4 times as long as and about 1.3 times as broad as third antennomere. Fifth and sixth antennomeres equal to fourth antennomere. Seventh antennomere about 1.8 times as long as wide at apex, about 1.2 times as long as and equal in width to sixth antennomere. Eighth antennomere about 1.9 times as long as wide at apex, as long as and slightly narrower than seventh antennomere. Ninth antennomere about 1.9 times as long as wide at apex, about 0.9 times as long and about 0.8 times as narrow as eighth antennomere. Tenth antennomere 2.5 times as long as wide at apex, about 1.1 times as long as ninth antennomere. Eleventh antennomere about 3.2 times as long as broad at middle, about 1.3 times as long and broad as tenth antennomere. Pronotum almost bell-shaped, about 1.3 times as long as wide at apex, about 0.9 times as long as wide at middle, about 0.8 times as long as wide at base, without lateral pronotal carinae. Disc densely punctate. Elytra oval, broadest at humeri, about 3.4 times as long as broad at base, about 1.5 times as long as broad at middle, about 1.6 times as long as broad at apical fourth, about 2.2 times as long as pronotum, with convex humeri. Elytral striae absent. Metaventrite weakly convex, about 2.9 times as long as length of mesocoxal cavity. Metepisternum narrow. Abdomen convex. First ventrite about 1.4 times as long as length of mesocoxal cavity. Second ventrite equal in length to first ventrite. Third ventrite about 0.6 times as long as second ventrite. Fourth ventrite equal in length to third ventrite. Fifth ventrite about 1.7 times as long as fourth ventrite, covered by elytra. Legs long. Femora weakly clavate. Tibiae apically expanded, with two spurs. Tarsi long, five-segmented. Tarsomeres first-third long conical. Fourth tarsomere weakly bilobed. Claws weakly toothed. Body length 3.1 mm. Comparison. The new species differs from Granulobium whitei Li, Philips et Cai, 2023 in the punctate pronotum, the larger eyes, densely punctate elytra, and the larger body size. The following is a list of the Mesozoic Ptinidae:	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE0FF33BEE0FC37FA98.taxon	discussion	Discussion Representatives of two subfamilies, Eucradinae and Anobiinae, have been identified from the Cretaceous (Peris et al. 2015, 2020; Molino-Olmedo 2017; Li et al. 2023; Háva & Zahradník 2023). Mesernobius anawrahtai Engel, 2010, from Burmese amber, was described as a primitive anobiid beetle (Engel 2010) but was subsequently placed in the family Scirtidae (Peris et al. 2020). Benemerita burmitica Molino­Olmedo, 2024, a species described from Burmese amber, was classified as a new genus in the subfamily Dryophilinae of the family Ptinidae (Molino­Olmedo 2024). However, it has been determined that Benemerita Molino­Olmedo, 2024, syn. nov. is synonymous with Amberophytum Yu et al., 2019 and it belongs to the family Cerophytidae, because it is characterised by the broad, acute posterior angles of the pronotum, the simple claws and the hypognathous head. The earliest documented record indicates the presence of an undescribed species from Barremian Lebanese amber (Kirejtshuk & Azar 2013), and a species of the recent Californian genus Actenobius has been described from early Albian San Just amber (Peris et al. 2015). The most diverse fauna, with five species, is found in Burmese amber. Acknowledgements The authors are grateful to the anonymous reviewers who helped to improve the manuscript.	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE0FF33BEE0FC37FA98.taxon	description	https: // doi. org / 10.59893 / bjc. 24 (2). 009 Engel, M. S. (2010) A primitive anobiid beetle in mid-Cretaceous amber from Myanmar (Coleoptera: Anobiidae). Alavesia, 3, 31 – 34. Háva, J. & Zahradník, P. (2023) Nicobium cretaceum sp. nov. (Coleoptera: Bostrichoidea: Ptinidae), a new species from mid-Cretaceous Burmese amber. Studies and Reports. Taxonomical Series, 19 (2), 289 – 292. Kirejtshuk, A. G. & Azar, D. (2013) Current knowledge of Coleoptera (Insecta) from the Lower Cretaceous Lebanese amber and taxonomical notes for some Mesozoic groups. Terrestrial Arthropod Reviews, 6, 103 – 134. https: // doi. org / 10.1163 / 18749836 - 06021061 Lawrence, J. F., Beutel, R. G., Leschen, R. A. B., Ślipiński, S. A. (2010) Chapter 2. Glossary of Morphological Terms. Handbook of Zoology. Arthropoda: Insecta. Tb. 40: Coleoptera (Beetles). Vol. 2: Morphology and Systematic (Elateroidea, Bostrichformia, Cucujiformia partim), P. 9 – 20. Li, Y. - D., Philips, T. K., Huang, D. - Y. & Cai, C. - Y. (2023) Earliest fossil record of Eucradinae in mid-Cretaceous amber from northern Myanmar (Coleoptera: Ptinidae). Bulletin of Geosciences, 98 (2), 171 – 180. https: // doi. org / 10.3140 / bull. geosci. 1876 Molino-Olmedo, F. (2017) Descripción de Molinernobius fuentesi gen. et n. sp. del ámbar cretácico de Myanmar (Coleoptera, Ptinidae, Ernobiinae). Lambillionea, 117, 151 – 154. Molino-Olmedo, F. (2024) Description of a new Ptinidae Latreille, 1802 (lnsecta: Coleoptera) from Cretaceous amber of Myanmar. Paleontological Journal, 58 (11), 1226 – 1231. https: // doi. org / 10. ll 34 / S 0031030124601099 Peris, D., Bao, T., Maehler, B. & Philips, T. K. (2020) A morphologically unique species of Ptinidae (Coleoptera) and the first found in mid-Cretaceous Kachin amber (Myanmar). Journal of Systematic Palaeontology, 18, 873 – 883. https: // doi. org / 10.1080 / 14772019.2019.1695291 Peris, D., Philips, T. K. & Delclòs, X. (2015) Ptinid beetles from the Cretaceous gymnosperm-dominated forests. Cretaceous Research, 52, 440 – 452. https: // doi. org / 10.1016 / j. cretres. 2014.02.009 Philips, T. K. (2002) 70. Anobiidae Fleming 1821. American beetles. Polyphaga: Scarabaeoidea through Curculionoidea. Arnett, R. H., Thomas, M. C., Skelley, P. E., Frank, J. H. (eds.). CRC Press, Boca Raton, London, New York, Washington, D. C., pp. 245 – 260. Shi, G., Grimaldi, D. A., Harlow, G. E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q. & Li, X. (2012) Age constraint on Burmese amber based on U – Pb dating of zircons. Cretaceous Research, 37, 155 – 163. https: // doi. org / 10.1016 / j. cretres. 2012.03.014 White, R. E. (1969) On Crichtonia, described as a new fossil genus of Anobiidae (Coleoptera). Proceedings of the Entomological Society of Washington, 71, 597. Zahradník, P. (2018) Ptinomorphus kratkyi sp. nov. – a new species from Europe (Coleoptera: Bostrichoidea: Ptinidae). Studies and Reports. Taxonomical Series, 14, 497 – 502. Zahradník, P. & Háva, J. (2014) Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia). Zootaxa, 3754, 301 – 352. https: // doi. org / 10.11646 / zootaxa. 3754.4.1	en	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
