identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DD4C47FFCEFFE6FF33B858FD03F936.text	03DD4C47FFCEFFE6FF33B858FD03F936.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Granulobium perkovskyi Legalov & Legalov 2025	<div><p>Granulobium perkovskyi A. Legalov, sp. nov.</p><p>https://zoobank.org/ urn:lsid:zoobank.org:act: B6C8617A-F6C6-46A4-8D9F-F529BAFCEFF4</p><p>(Fig. 1)</p><p>Type material. Holotype: ISEA MA 2013-1, Burmese amber, Myanmar; Cenomanian, late Cretaceous .</p><p>Syninclusion. The second damaged specimen of this species is not included in the type series.</p><p>Etymology. The epithet of this new species is in honour of Evgeny E. Perkovsky, a specialist in fossil insect.</p><p>Description. Body black, convex dorsally, covered with short suberect setae. Forehead convex, broad, densely punctate and granulate. Eyes well protruded, finely facetted, rounded. Antennae elevensegmented, slightly serrate, without club. First antennomere about 1.8 times as long as broad at apex. Second antennae subequal in length and width, about 0.4 times as long and about 0.8 times as narrow as the first. Third antennomere about 1.3 times as long and wide at apex, about 1.7 times as long and about 1.3 times as wide as second antennomere. Fourth antennomere 1.4 times as long as broad at apex, 1.4 times as long as and about 1.3 times as broad as third antennomere. Fifth and sixth antennomeres equal to fourth antennomere. Seventh antennomere about 1.8 times as long as wide at apex, about 1.2 times as long as and equal in width to sixth antennomere. Eighth antennomere about 1.9 times as long as wide at apex, as long as and slightly narrower than seventh antennomere. Ninth antennomere about 1.9 times as long as wide at apex, about 0.9 times as long and about 0.8 times as narrow as eighth antennomere. Tenth antennomere 2.5 times as long as wide at apex, about 1.1 times as long as ninth antennomere. Eleventh antennomere about 3.2 times as long as broad at middle, about 1.3 times as long and broad as tenth antennomere. Pronotum almost bell-shaped, about 1.3 times as long as wide at apex, about 0.9 times as long as wide at middle, about 0.8 times as long as wide at base, without lateral pronotal carinae. Disc densely punctate. Elytra oval, broadest at humeri, about 3.4 times as long as broad at base, about 1.5 times as long as broad at middle, about 1.6 times as long as broad at apical fourth, about 2.2 times as long as pronotum, with convex humeri. Elytral striae absent. Metaventrite weakly convex, about 2.9 times as long as length of mesocoxal cavity. Metepisternum narrow. Abdomen convex. First ventrite about 1.4 times as long as length of mesocoxal cavity. Second ventrite equal in length to first ventrite. Third ventrite about 0.6 times as long as second ventrite. Fourth ventrite equal in length to third ventrite. Fifth ventrite about 1.7 times as long as fourth ventrite, covered by elytra. Legs long. Femora weakly clavate. Tibiae apically expanded, with two spurs. Tarsi long, five-segmented. Tarsomeres first-third long conical. Fourth tarsomere weakly bilobed. Claws weakly toothed. Body length 3.1 mm.</p><p>Comparison. The new species differs from Granulobium whitei Li, Philips et Cai, 2023 in the punctate pronotum, the larger eyes, densely punctate elytra, and the larger body size.</p><p>The following is a list of the Mesozoic Ptinidae:</p></div>	https://treatment.plazi.org/id/03DD4C47FFCEFFE6FF33B858FD03F936	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCDFFE7FF33BBABFA94FF51.text	03DD4C47FFCDFFE7FF33BBABFA94FF51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Granulobium Li, Philips et Cai 2023	<div><p>Genus Granulobium Li, Philips et Cai, 2023</p><p>Granulobium whitei Li, Philips et Cai, 2023 – Burmese amber (the earliest Cenomanian); Myanmar Granulobium perkovskyi A. Legalov, sp. nov. – Burmese amber (the earliest Cenomanian); Myanmar</p></div>	https://treatment.plazi.org/id/03DD4C47FFCDFFE7FF33BBABFA94FF51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCDFFE6FF33BB24FA9AF8EF.text	03DD4C47FFCDFFE6FF33BB24FA9AF8EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Molinernobius Molino-Olmedo 2017	<div><p>Genus Molinernobius Molino-Olmedo, 2017</p><p>Molinernobius fuentesi Molino-Olmedo, 2017 – Burmese amber (the earliest Cenomanian); Myanmar</p></div>	https://treatment.plazi.org/id/03DD4C47FFCDFFE6FF33BB24FA9AF8EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE7FF33BD4BFAE9FE06.text	03DD4C47FFCCFFE7FF33BD4BFAE9FE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actenobius Fall 1905	<div><p>Genus Actenobius Fall, 1905</p><p>Actenobius magneoculus Peris, Philips et Delclòs, 2015 – San Just amber (early Albian); Spain</p></div>	https://treatment.plazi.org/id/03DD4C47FFCCFFE7FF33BD4BFAE9FE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE7FF33BDD2FA9AFD81.text	03DD4C47FFCCFFE7FF33BDD2FA9AFD81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nicobium LeConte 1861	<div><p>Genus Nicobium LeConte, 1861</p><p>Nicobium cretaceum Háva et Zahradník, 2023 – Burmese amber (the earliest Cenomanian); Myanmar</p></div>	https://treatment.plazi.org/id/03DD4C47FFCCFFE7FF33BDD2FA9AFD81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE7FF33BE59FB35FD1B.text	03DD4C47FFCCFFE7FF33BE59FB35FD1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stegobium Motschulsky 1860	<div><p>Genus Stegobium Motschulsky, 1860</p><p>Stegobium raritanensis Peris, Philips et Delclòs, 2015 –Raritan amber (Turonian); USA</p></div>	https://treatment.plazi.org/id/03DD4C47FFCCFFE7FF33BE59FB35FD1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
03DD4C47FFCCFFE0FF33BEE0FC37FA98.text	03DD4C47FFCCFFE0FF33BEE0FC37FA98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cretasernus Peris et Philips 2020	<div><p>Genus Cretasernus Peris et Philips, 2020</p><p>Cretasernus spinosus Peris et Philips, 2020 – Burmese amber (the earliest Cenomanian); Myanmar</p><p>Discussion</p><p>Representatives of two subfamilies, Eucradinae and Anobiinae, have been identified from the Cretaceous (Peris et al. 2015, 2020; Molino-Olmedo 2017; Li et al. 2023; Háva &amp; Zahradník 2023). Mesernobius anawrahtai Engel, 2010, from Burmese amber, was described as a primitive anobiid beetle (Engel 2010) but was subsequently placed in the family Scirtidae (Peris et al. 2020) . Benemerita burmitica Molino­Olmedo, 2024, a species described from Burmese amber, was classified as a new genus in the subfamily Dryophilinae of the family Ptinidae (Molino­Olmedo 2024) . However, it has been determined that Benemerita Molino­Olmedo, 2024, syn. nov. is synonymous with Amberophytum Yu et al., 2019 and it belongs to the family Cerophytidae, because it is characterised by the broad, acute posterior angles of the pronotum, the simple claws and the hypognathous head. The earliest documented record indicates the presence of an undescribed species from Barremian Lebanese amber (Kirejtshuk &amp; Azar 2013), and a species of the recent Californian genus Actenobius has been described from early Albian San Just amber (Peris et al. 2015). The most diverse fauna, with five species, is found in Burmese amber.</p><p>Acknowledgements</p><p>The authors are grateful to the anonymous reviewers who helped to improve the manuscript.</p><p>References</p><p>Abdullah, M. &amp; Abdullah, A. (1967) Crichtonia macleani, a new genus and species of the Hedobiini (Coleoptera: Anobiidae) from the Baltic amber. Entomological News, 28, 23–27.</p><p>Alekseev, V. &amp; Bukejs, A. (2024) A new extinct genus of the tribe Hedobiini (Coleoptera: Ptinidae) from Baltic amber. Baltic Journal of Coleopterology, 24 (2), 223–230.</p><p>https://doi.org/10.59893/bjc.24(2).009</p><p>Engel, M.S. (2010) A primitive anobiid beetle in mid-Cretaceous amber from Myanmar (Coleoptera: Anobiidae). Alavesia, 3, 31–34.</p><p>Háva, J. &amp; Zahradník, P. (2023) Nicobium cretaceum sp. nov. (Coleoptera: Bostrichoidea: Ptinidae), a new species from mid-Cretaceous Burmese amber. Studies and Reports. Taxonomical Series, 19 (2), 289–292.</p><p>Kirejtshuk, A.G. &amp; Azar, D. (2013) Current knowledge of Coleoptera (Insecta) from the Lower Cretaceous Lebanese amber and taxonomical notes for some Mesozoic groups. Terrestrial Arthropod Reviews, 6, 103–134. https://doi.org/10.1163/18749836-06021061</p><p>Lawrence, J.F., Beutel, R.G., Leschen, R.A.B., Ślipiński, S.A. (2010) Chapter 2. Glossary of Morphological Terms. Handbook of Zoology. Arthropoda: Insecta. Tb. 40: Coleoptera (Beetles). Vol. 2: Morphology and Systematic (Elateroidea, Bostrichformia, Cucujiformia partim), P. 9–20.</p><p>Li, Y.-D., Philips, T.K., Huang, D.-Y. &amp; Cai, C.-Y. (2023) Earliest fossil record of Eucradinae in mid-Cretaceous amber from northern Myanmar (Coleoptera: Ptinidae). Bulletin of Geosciences, 98 (2), 171–180. https://doi.org/10.3140/bull.geosci.1876</p><p>Molino-Olmedo, F. (2017) Descripción de Molinernobius fuentesi gen. et n. sp. del ámbar cretácico de Myanmar (Coleoptera, Ptinidae, Ernobiinae). Lambillionea, 117, 151–154.</p><p>Molino-Olmedo, F. (2024) Description of a new Ptinidae Latreille, 1802 (lnsecta: Coleoptera) from Cretaceous amber of Myanmar. Paleontological Journal, 58 (11), 1226–1231.</p><p>https://doi.org/10.ll34/S0031030124601099</p><p>Peris, D., Bao, T., Maehler, B. &amp; Philips, T.K. (2020) A morphologically unique species of Ptinidae (Coleoptera) and the first found in mid-Cretaceous Kachin amber (Myanmar). Journal of Systematic Palaeontology, 18, 873–883. https://doi.org/10.1080/14772019.2019.1695291</p><p>Peris, D., Philips, T.K. &amp; Delclòs, X. (2015) Ptinid beetles from the Cretaceous gymnosperm-dominated forests. Cretaceous Research, 52, 440–452. https://doi.org/10.1016/j.cretres.2014.02.009</p><p>Philips, T.K. (2002) 70. Anobiidae Fleming 1821. American beetles. Polyphaga: Scarabaeoidea through Curculionoidea. Arnett, R.H., Thomas, M.C., Skelley, P.E., Frank, J.H. (eds.). CRC Press, Boca Raton, London, New York, Washington, D.C., pp. 245–260.</p><p>Shi, G., Grimaldi, D.A., Harlow, G.E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q. &amp; Li, X. (2012) Age constraint on Burmese amber based on U–Pb dating of zircons. Cretaceous Research, 37, 155–163. https://doi.org/10.1016/j.cretres.2012.03.014</p><p>White, R.E. (1969) On Crichtonia, described as a new fossil genus of Anobiidae (Coleoptera). Proceedings of the Entomological Society of Washington, 71, 597.</p><p>Zahradník, P. (2018) Ptinomorphus kratkyi sp. nov. – a new species from Europe (Coleoptera: Bostrichoidea: Ptinidae). Studies and Reports. Taxonomical Series, 14, 497–502.</p><p>Zahradník, P. &amp; Háva, J. (2014) Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia). Zootaxa, 3754, 301–352. https://doi.org/10.11646/zootaxa.3754.4.1</p></div>	https://treatment.plazi.org/id/03DD4C47FFCCFFE0FF33BEE0FC37FA98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Legalov, Andrei A.;Legalov, Vladislav A.	Legalov, Andrei A., Legalov, Vladislav A. (2025): A new species of the genus Granulobium Li et al., 2023 (Coleoptera: Ptinidae) from Cretaceous Kachin amber. Ecologica Montenegrina 82: 69-73, DOI: 10.37828/em.2025.82.3, URL: https://doi.org/10.37828/em.2025.82.3
