taxonID	type	description	language	source
03DDA36B7D5FFFE2FF09FC4201B9FCAA.taxon	description	(Fig. 1 A)	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5FFFE2FF09FC4201B9FCAA.taxon	materials_examined	TYPE MATERIAL. — The single remaining syntype of Deltocyathus magnificus is deposited at the BMNH (uncataloged). TYPE LOCALITY. — 5 ° 49 ’ S, 132 ° 14 ’ E (off Kei Islands, Banda Sea), 236 m. NEW RECORDS. — HALIPRO 1, stn CP 877, 1 specimen (MNHN-Scl. 2008 - 001); 1 specimen (USNM 1114164).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5FFFE2FF09FC4201B9FCAA.taxon	distribution	DISTRIBUTION. — New Caledonia waters (23 ° 03.51 ’ S, 166 ° 59.20 ’ E, 464 - 480 m). Previous records: east China sea, 88 - 422 m (Cairns 1994); western Pacific from Japan to southeastern Australia, 88 - 1500 m (Cairns & Zibrowius 1997); including Vanuatu region, 408 - 433 m (Cairns 1999); Sulu Archipelago, Moluccas, 118 - 1500 m; Philippines, Indonesia, 8 - 522 m (Cairns & Parker 1992); western Australia, 250 m (Cairns 2004).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5FFFE2FF09FC4201B9FCAA.taxon	description	DESCRIPTION Corallum discoidal to hexagonal with a slightly concave base. Largest specimen examined (HALIPRO 1, stn CP 877) 25.5 mm in calicular diameter and 7 mm in height, with a depression of 6 mm in diameter in centre of base. Costae equal thin ridges, finely dentate, being separated by wide furrows and extending up to 1 mm beyond calicular margin. C 3 - 5 not present at centre of base. All costae laterally spinose especially near calicular edge, where intercostal furrows are deeper. Corallum white. Septa hexamerally arranged in 5 cycles (S 1 ≥ S 2> S 3> S 4 - 5), but any specimen examined display a complete fifth cycle, denoting their juvenile stage. S 1 only independent septa, 6 mm exsert, joining columella through a wide palus that sometimes are divided at the upper margin in 3 paliform lobes. S 2 equal to slightly less wide than S 1, also joining columella through a wide and tall pali. S 3  ⁄  width of S 3, less exsert and bear a paliform lobe that fuses to P 2 near columella. S 4 half size of S 3 and less exsert, joining P 3 through a typical deltocyathid chevron arrangement. S 5 less exsert but equal in width to S 4, also bearing a paliform lobe that fuses P 4. Fossa very shallow, aligned with the 2 principal and opposite S 1, containing a papillose columella variable in size, shape, and number of rods.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5FFFE2FF09FC4201B9FCAA.taxon	discussion	REMARKS Among all species of the genus Deltocyathus that do not present coastal spines, only four can reach five cycles of septa in the adult stage: D. sarsi, D. magnificus, D. suluensis, and D. rotulus. Of these species, D. sarsi is known only from off Vanuatu islands (Gardiner & Waugh 1938), and at 80 m in western Australia (Cairns 2004). Among the other three species, all were collected off New Caledonia, and D. magnificus is easily distinguished by having a more robust corallum and a flat to concave base, and usually is whiter than D. rotulus and D. suluensis which normally present a reddish brown pigment in some parts of the coralla.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5EFFE1FD42FCFC033FF903.taxon	materials_examined	TYPE MATERIAL. — The holotype of Trochocyathus rotulus is presumed to be deposited at the IM. Six syntypes of Deltocyathus fragilis are deposited at the ZMA (Coel. 1188). TYPE LOCALITY. — North Maldive Atoll, 1408 - 1756 m (Trochocyathus rotulus). 7 ° 24 ’ S, 118 ° 15.2 ’ E (Flores Sea), 794 m (Deltocyathus fragilis). NEW RECORDS. — BATHUS 3, stn CP 842, 2 specimens (MNHN-Scl. 2008 - 0002); stn DW 786, 1 specimen (MNHN-Scl. 2008 - 0003). — BATHUS 4, stn CP 950, 62 specimens (MNHN-Scl. 2008 - 0004); 10 specimens (USNM 1114141).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5EFFE1FD42FCFC033FF903.taxon	distribution	DISTRIBUTION. — New Caldonia waters from 20 ° 31 ’ S to 23 ° 54 ’ S, 699 - 830 m. Elsewhere: Indian Ocean off Durban, off Mozambique, off Zanzibar (Cairns & Keller 1993), 510 - 1986 m, including Maldive islands (Alcock 1898; Gardiner & Waugh 1938), Gulf of Aden (Gardiner & Waugh 1938), off Sri Lanka (Van der Horst 1931), and off Tanzania (Gardiner & Waugh 1938); Pacific Ocean off southeastern Japan (Cairns 1994), 799 - 1187 m, off Philippines, Indonesia (Flores and Celebes Sea), Malaysia (Alcock 1902 a, c; Cairns & Zibrowius 1997), 210 - 1719 m, Vanuatu and Wallis and Futuna region (Cairns 1999), 1050 - 1160 m, Australia region off Queensland (Cairns 2004), 143 - 1192 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5EFFE1FD42FCFC033FF903.taxon	description	DESCRIPTION Corallum circular to slightly elliptical with lancetted calicular margin. Each apex correspond to those C 4 flanked by C 5. Base normally slightly conical and pointed in centre. However, largest specimen examined (BATHUS 3, stn CP 842), with 19 mm in calicular diameter and 4 mm in height, shows a flat base with upturned edges. Costae more prominent near calicular edge as low serrated ridges. C 1 - 3 originates in centre of the base, C 4 present only near calicular edge, and C 5 slightly wider than C 4. Corallum white to reddish-brown. Septa hexamerally arranged in 5 cycles (S 1 - 2> S 3> S 4> S 5). S 1 - 2 do not fuse to any adjacent septa, each bearing a palus about 1 - 2.5 mm wide. S 3 less exert with irregularly shaped paliform lobe, sometimes P 3 wider than P 1 - 2, but some specimens present P 3 very small. Adjacent to the P 3, near columella, sometimes 2 or 3 small paliform teeth are present. S 4 less wide and exsert than S 3 but bear a wide and tall palus, producing a prominent palar crown encircling columella. Each P 4 fuses to the flanked P 3 near columella. S 5 fuses to the axial edge of adjacent P 4 close to basal theca through a long and porous lamella. Fossa extremely shallow containing a welldeveloped papillose columella composed of several interconnected rods.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5EFFE1FD42FCFC033FF903.taxon	discussion	REMARKS Among the Deltocyathus species from New Caledonia with five septal cycles, D. rotulus is easily distinguished from D. suluensis by having a scalloped calicular edge, a pigmented columella and paliform lobes, and paliform lobes that reach the same high of the septa. For comparation with D. magnificus see previous remarks.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5DFFE0FD42FF7A003CFDB7.taxon	materials_examined	TYPE MATERIAL. — The types of D. suluensis are presumed to be deposited at the IM. The holotype (CO 266) of D. formosus and one paratype (CO 228) are deposited at the MoNZ. Additional paratypes are deposited at NZOI (station numbers: stn K 826 [1], K 828 [2], and K 870 [1]), and at the USNM (94171 [3], 94172 [1], and 94173 [6]). TYPE LOCALITY. — 5 ° 43 ’ N, 119 ° 40 ’ E- 6 ° 11 ’ N, 120 ° 37 ’ E (Sulu Archipelago), 450 - 522 m (D. magnificus var. suluensis). 29 ° 13.96 ’ S, 177 ° 52.84 ’ W (Kermadec Ridge), 530 - 567 m (D. formosus). NEW RECORDS. — BATHUS 3, stn DW 786, 3 specimens (MNHN-Scl. 2008 - 0005). — BATHUS 4, stn CP 889, 2 specimens (MNHN-Scl. 2008 - 0006). — NORFOLK 2, stn DW 2097, 3 specimens (MNHN-Scl. 2008 - 0007); stn DW 2104, 1 specimen (MNHN-Scl. 2008 - 0008).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5DFFE0FD42FF7A003CFDB7.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 21 ° 00.83 ’ S, 164 ° 27.34 ’ E to 24 ° 44 ’ S, 169 ° 49.15 ’ E, 416 - 752 m. Previous records: Indian Ocean off western Australia from Dampier Land to Port Hedland (Cairns 1998; 2004), 401 - 530 m; Pacific Ocean from Philippines and Indonesian region (Cairns & Zibrowius 1997), 204 - 540 m to southern Norfolk Ridges, New Zealand (Cairns 1995), 142 - 565 m, including Wallis and Futuna region and Vanuatu region (Cairns 1999), 400 - 650 m, and off Queensland, Australia (Cairns 2004), 246 - 1050 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5DFFE0FD42FF7A003CFDB7.taxon	description	DESCRIPTION Corallum circular with base flat to slightly convex. Calicular margin serrate but not lancetted. Largest specimen examined (BATHUS 3, stn DW 786) 22 mm in calicular diameter and 7 mm in height. Costae present as round granular ridges separated by well-defined same width intercostal furrows. Only C 1 - 2 originates at centre of base. Septa hexamerally arranged in 5 complete cycles (S 1 ≥ S 2> S 3> S 4 >> S 5). S 1 only independent septa and extend half distance to columella, being separated from their pali by a moderately wide notch. Higher septal cycles progressive less wide. S 1 - 4 and P 1 - 4 arranged in a typical Deltocyathus fashion, fusing through porous processes. S 5 rudimentary. Fossa shallow containing a well-developed papillose columella.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5DFFE0FD42FF7A003CFDB7.taxon	discussion	REMARKS See Remarks of D. magnificus and D. rotulus.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5CFFEEFF27FDFE0058FE36.taxon	materials_examined	TYPE MATERIAL. — The holotype of Levipalifer orientalis is deposited at the USNM (19391). TYPE LOCALITY. — Off Bosyu, Japan, depth unknown (Levipalifer orientalis). NEW RECORDS. — BATHUS 4, stn DW 914, 1 specimen (USNM 1114140). — NORFOLK 2, stn DW 2037, 1 specimen (MNHN-Scl. 2008 - 0009).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5CFFEEFF27FDFE0058FE36.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 18 ° 48.79 ’ S, 163 ° 15.23 ’ E to 23 ° 40 ’ S, 167 ° 41 ’ E, 517 - 616 m. Previous records: Japan, off Honshu and Kyushu (Yabe & Eguchi 1932; Cairns 1994), Sagami Sea (Ogawa 2006), 88 - 1097; Philippines and Indonesia regions (Cairns & Zibrowius 1997), 247 - 807 m; Vanuatu region (Cairns 1999), 585 - 919 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5CFFEEFF27FDFE0058FE36.taxon	description	DESCRIPTION Corallum circular with base flat to slightly convex bearing a small epicentre boss. Largest specimen examined (NORFOLK 2, stn DW 2037) 21 mm in calicular diameter and 9.1 mm in height. Costae rounded, granular and separated by intercostal furrows deeper at calicular edge. Near calicular margin, lateral faces of costae bears numerous pointed granules. All costae extend to base as a row of low and round granules, except from those of C 3 and C 4, which do not reach the centre of base. Corallum white. Septa hexamerally arranged in 4 complete cycles (S 1> S 2> S 4> S 3) for a total of 48 septa. S 1 highly exsert and only independent septa, almost reaching columella and bearing a wide lamellar paliform lobe that fuses to the columella. S 2 less exsert and wide than S 1, each bearing a paliform lobe (P 2) recessed from columella moreso than P 1. S 3 smallest septa, bearing a wide pali. S 4 as exsert as S 3, slightly wider and often bear a narrow paliform lobe, which fuses to enclosed P 3. If P 4 are absent, inner edge of S 4 fuses to P 3. All septal and palar faces show sparsely pointed granules. Fossa shallow containing and elongate columella composed by small interconnected, sometimes granulated papillae.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D5CFFEEFF27FDFE0058FE36.taxon	discussion	REMARKS Among the three New Caledonia Deltocyathus species that do not have spines at the calicular edge and have four septal cycles (D. vaughani, D. cameratus, D. crassiseptum), D. vaughani is distinguished by having: a more robust corallum, a non-jagged calicular edge, all septa reaching the same distance Deltocyathus (Scleractinia, Caryophylliidae) from New Caledonia from calicular margin, dimorphic S 4 (those S 4 near flanking S 1 are larger than S 3 and those flanking S 2 are equal in width to S 3) and the presence of P 4. According to Cairns (1994) the presence of P 4 can be used to distinguish this species from all other Deltocyathus with four septal cycles.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEEFF1AFE780189FC2B.taxon	materials_examined	TYPE MATERIAL. — The holotype is deposited at the BMNH. TYPE LOCALITY. — Baie du Santal, Lifu, Loyalty Islands, 73 m. NEW RECORDS. — BATHUS 4, stn DW 887, 4 specimens (MNHN-Scl. 2008 - 0010); stn DW 902 (MNHN- Scl. 2008 - 0012); 1 specimen (USNM 1114163); stn DW 903, 1 specimen (MNHN-Scl. 2008 - 0011). — NOR- FOLK 2, stn DW 2092, 4 specimens (USNM 1114145); stn DW 2117, 5 specimens (MNHN-Scl. 2008 - 0013).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEEFF1AFE780189FC2B.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 18 ° 59.93 ’ S, 163 ° 13.55 ’ E to 24 ° 45 ’ S, 168 ° 07 ’ E, Loyalty islands (Gardiner 1899), 73 - 400 m. Previous records: Australia off Queensland (Cairns 1995), 315 - 360 m; Wallis and Futuna region, and Vanuatu region (Cairns 1999), 295 - 360 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEEFF1AFE780189FC2B.taxon	description	DESCRIPTION Corallum circular, lancetted and shaped as a shallow bowl with upturned edges. Base flat to slightly concave, sometimes with small scar at centre. C 1 - 2 equal and always reach centre of base, being separated from adjacent costae by moderately deep intercostal furrows. C 3 most developed costae projecting up to 1.5 mm beyond calicular edge as 12 granular spines. C 4 do not reach centre of base, but fuses adjacent C 3 beyond calicular margin producing 12 well-developed apexes. Corallum white. Septa hexamerally arranged in 4 cycles (S 1> S 2> S 4> S 3). S 1 only independent septa, extending ¾ the distance to columella, being separated from their dimorphic paliform lobe (as in D. crassiseptum) by a deep notch. S 2 slightly narrower than S 1 and bear a small palus. S 3 are smallest septa (⅓ of S 2) but bears largest paliform lobes which fuse P 2 near columella. S 4 fuses outer edge of flanked P 3. Fossa shallow containing an elliptical papillose columella aligned with the two principal S 1. Columella consists of small, short, fused tuberculate papillae.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEEFF1AFE780189FC2B.taxon	discussion	REMARKS Among the five-spined species of Deltocyathus, three occur in New Caledonian waters: D. ornatus, D. heteroclitus, and D. corrugatus. Deltocyathus ornatus and D. corrugatus are easily separated from D. heteroclitus using the number of spines present: 12 versus 6 - 8 in D. heteroclitus, and the P 1 dimorphism: dimorphic in D. ornatus and D. corrugatus and not dimorphic in D. heteroclitus. However, the differences between the other two species with 12 spines that occur in New Caledonia region are very subtle: D. corrugatus usually possesses a circular basal scar, C 3 very proeminent, and longer and slender costal spines (also see Cairns 1999).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEDFD4BFC7D0011FAEF.taxon	materials_examined	TYPE MATERIAL. — The holotype and 5 paratypes are deposited at the USNM (71849, 71850, and 71851). TYPE LOCALITY. — Navaka River, Espiritu Santo, Vanuatu (Late Pleistocene). NEW RECORDS. — BATHUS 4, stn DW 887, 2 specimens (MNHN-Scl. 2008 - 0015); stn DW 894, 2 specimens (MNHN-Scl. 2008 - 0016); stn DW 898, 1 specimen (MNHN-Scl. 2008 - 0017); 1 specimen (USNM 1114160). DISTRIBUTION. — New Caledonia waters from 20 ° 15.77 ’ S, 163 ° 50.21 ’ E to 21 ° 06.67 ’ S, 164 ° 27.62 ’ E, 245 - 600 m. Previous records: Wallis and Futuna region, 349 - 355 m, and Vanuatu region (Cairns 1999), 208 - 210 m, including late Pleistocene of Espiritu Santo (Wells 1984).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEDFD4BFC7D0011FAEF.taxon	description	DESCRIPTION Corallum polygonal and lancetted in calicular margin, each lancet corresponding to each P 3 and P 4. Largest specimen examined (BATHUS 4, stn DW 894) 11.9 mm in calicular diameter and 4.1 mm in height. Base conical with a pointed centre, but one specimen (BATHUS 4, stn DW 887) shows a large depression in the middle to the base. All costae well developed and equal sized, extending from calicular margin to centre of the base, being separated by shallow intercostal furrows. However, when flanked, C 3 is broader and most prominent costae, extending up to 1 mm beyond the calicular edge forming robust costal spines, not reaching the centre of the base. All costae bear several granules near calicular edge, sometimes becoming a row near the middle of the base. Septa hexamerally arranged in four cycles (S 1> S 2> S 4> S 3), however, no specimen examined have a complete fourth cycle, usually resulting in 39 - 42 septa. Number of spines (C 3) dependent of number of flanked S 3. S 1 only independent septa extending ½ to ¾ distance to columella, being separated from a lamellar palus by a slender notch. S 2 slightly smaller than S 1 but bear a wider palus. S 3 dimorphic in size: those unflanked by S 4 slightly smaller than S 2, being fused by axial edges to adjacent P 2; however, those S 3 flanked by S 4 are the smallest septa, but bear the widest and tallest paliform lobe, and sometimes 2 or 3 small paliform teeth near columella. P 3 high granular, sinuous in upper edge, and are the most recessed pali from columella. S 4 fuse to adjacent P 3 through a solid lamella at the level of S 3 - P 3 notch. All septal faces granulated and slightly sinuous in upper edge. Fossa shallow containing an elliptical papillose columella composed by several granular elements.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D52FFEDFD4BFC7D0011FAEF.taxon	discussion	REMARKS See identification key and Remarks of D. ornatus.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D51FFECFF1EFA2603FDFEB4.taxon	materials_examined	TYPE MATERIAL. — The holotype is at the NZOI (H 689), and 10 paratypes are deposited at the USNM (94169 [5], and 94170 [5]). TYPE LOCALITY. — 28 ° 54 ’ 36 ’’ S, 167 ° 44 ’ 12 ’’ E (Norfolk Islands), 390 m. NEW RECORDS. — BATHUS 4, stn DW 898, 8 specimens (MNHN-Scl. 2008 - 0019); stn CP 883, 2 specimens (MNHN-Scl. 2008 - 0020); 1 specimen (USNM 1114161); stn DW 903, 2 specimens (MNHN-Scl. 2008 - 0021); stn CP 885, 1 specimen (MNHN-Scl. 2008 - 0022); stn CP 889, 3 specimens (MNHN-Scl. 2008 - 0023); 3 specimens (USNM 1114162); stn CP 967, 3 specimens (MNHN-Scl. 2008 - 0018).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D51FFECFF1EFA2603FDFEB4.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 18 ° 59.93 ’ S, 163 ° 13.55 ’ E to 23 ° 02.53 ’ S, 166 ° 58.18 ’ E, including Loyalty Islands (Cairns 1995), 250 - 600 m. Previous records: from southern Great Barrier Reef, Australia (Wells 1984) to Norfolk Ridge, New Zealand (Cairns 1995), 73 - 390 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D51FFECFF1EFA2603FDFEB4.taxon	description	DESCRIPTION Corallum circular. Calice lancetted, lancets correspond to each C 3 flanked by C 4. Largest specimen examined (BATHUS 3, stn CP 847) 15 mm in calicular diameter and 4.2 mm in height. Corallum usually white but sometimes slightly reddish-brown pigmented. Base flat to slightly convex, having a small scar of previous attachment (during early stages) in centre. All costae well developed. C 1 - 2 more prominent near calicular edge, becoming a row of granules near centre of base. C 3 thickest and tallest costae, extending up to 2 mm and normally upturned slightly beyond calicular edge as granulated costal spines. C 4 and C 3 form a broad three-step ridge. All costae granulated specially near calicular edge, and a small low granule ridge is present beside each C 1 - 2. Septa hexamerally arranged in 4 complete cycles (S 1 ≥ S 2> S 4> S 3). S 1 independent, extending half distance to columella and bearing a wide lamellar palus, which is often fused to columella. S 2 equal to or slightly less wide than S 1, also bearing a pali of same size of P 1, forming a crown more recessed from columella than the crown formed by P 1. S 3 smallest septa, bearing the tallest pali which fuse to inner edge of near P 2 by a porous lamella. S 4 slightly wider than S 3, and have a spinose margin. Each S 4 fuses the adjacent P 3 through a long porous lamella. All pali equal sized (P 1 sometimes dimorphic) and septa and palar faces bear pointed granules. Fossa shallow with an elliptical papillose columella.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D51FFECFF1EFA2603FDFEB4.taxon	discussion	REMARKS See Remarks of D. ornatus.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFECFF08FEFB01B1FB8C.taxon	materials_examined	TYPE MATERIAL. — The holotype and 78 paratypes are deposited at the MNHN (uncatalogued), and 39 at the USNM (98681 [2], 98682 [9], 98683 [13], 98684 [8], 98685 [2], 98686 [5]). TYPE LOCALITY. — 19 ° 21 ’ S, 169 ° 25 ’ E (Tanna), 433 - 450 m. NEW RECORDS. — BATHUS 3, stn CP 818, 4 specimens (MNHN-Scl. 2008 - 0024). — BATHUS 4, stn CP 889, 1 specimen (MNHN-Scl. 2008 - 0026). — HALIPRO 1, stn CP 877, 1 specimen (MNHN-Scl. 2008 - 0025). — NORFOLK 2, stn DW 2041, 2 specimens (MNHN-Scl. 2008 - 0027); stn DW 2024, 6 specimens (MNHN-Scl. 2008 - 0028); 6 specimens (USNM 1114122); stn DW 2025, 1 specimen (MNHN-Scl. 2008 - 0029); stn DW 2097, 2 specimens (MNHN-Scl. 2008 - 0030); stn DW 2117, 2 specimens (MNHN-Scl. 2008 - 0031).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFECFF08FEFB01B1FB8C.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 21 ° 00.83 ’ S, 164 ° 27.34 ’ E to 24 ° 44 ’ S, 168 ° 06 ’ E, 370 - 583 m. Previous records: Wallis and Futuna region (Cairns 1999), 420 - 510 m; Vanuatu region (Cairns 1999), 413 - 536 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFECFF08FEFB01B1FB8C.taxon	description	DESCRIPTION Corallum circular to slightly elliptical shaped as a shallow bowl. Theca thick. Base flat to slightly convex, usually with a scar, and sometimes protuberant. Largest specimen examined (BATHUS 3, stn CP 877) 14.2 mm in calicular diameter and 4.6 mm in height. However, smaller coralla show a CD: HT ratio near 2. Costae unequal and more prominent at calicular edge where they are separated by intercostal grooves bisected by a row of low and rounded granules that sometimes resemble a small ridge in the valley formed by the adjacent costae. C 1 - 2 broader than other costae. Corallum white to reddish-brown. Septa hexamerally arranged in four cycles (48 septa) with septal formula: S 1> S 2> S 3> S 4. However, in larger coralla up to 3 pairs of S 5 are present resulting in 56 septa. S 1 thick and most exsert septa, extending half distance to columella and bear a dimorphic lamellar paliform lobe. Those paliform lobes before the two principal S 1 are narrower than the other four P 1, but all reach the columella. S 2 equal to slightly smaller and less exsert than S 1, also bearing a paliform lobe that reaches columella by a small paliform teeth. S 3 about ¾ width of S 2, bearing a tall recessed pali that fuses P 2 in a chevron arrangement typical of the genus. S 4 smallest septa and in some coralla composed by a row of spines. All septal and palar faces very granular. Fossa shallow to moderately deep, containing a papillose columella.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFECFF08FEFB01B1FB8C.taxon	discussion	REMARKS First time reported since the original description, the specimens of D. crassiseptum are distinguished from the New Caledonia congeners by having: a slighty thicker S 1 - 2 in relation to the other septa, corallum shaped as a bowl, absence of costal spines, well-arranged crowns formed by the paliform lobes, and paliform lobes and columellar elements that terminate below the septa upper edges. Different pigmentation patterns can be found in New Caledonia specimens of D. crassiseptum: some have the entire calicular edge and septa reddish-brown, other specimens present only the S 3 and paliform lobes pigmented, some are completely pigmented and some are enterely white.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFEBFD4BFBC3032EFA20.taxon	materials_examined	TYPE MATERIAL. — The holotype and 38 paratypes are deposited at the MNHN (uncatalogued), and 15 at the USNM (98687 [1], 98688 [4], 98689 [1], 98690 [1], 98691 [3], 98692 [1], 98693 [4]). TYPE LOCALITY. — 18 ° 52 ’ S, 168 ° 52 ’ E (Erromango, Vanuatu), 720 - 830 m. NEW RECORDS. — NORFOLK 2, stn DW 2086, 2 specimens (MNHN-Scl. 2008 - 0032); stn DW 2103, 2 specimens (MNHN-Scl. 2008 - 0033); 2 specimens (USNM 1114123).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFEBFD4BFBC3032EFA20.taxon	distribution	DISTRIBUTION. — New Caledonia waters from 23 ° 57 ’ S, 167 ° 44 ’ E to 24 ° 56 ’ S, 168 ° 22 ’ E, 707 - 777 m. Previous records: known only in the Pacific Ocean from Wallis and Futuna region, 305 - 1010 m, and Vanuatu region (Cairns 1999), 512 - 1175 m to Lord Howe Rise, including Elizabeth Reef and Britannia (Cairns 2004), 419 - 1078 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFEBFD4BFBC3032EFA20.taxon	description	DESCRIPTION Calice circular with lancetted margin. Lancets correspond to each C 3 and C 4 that project outward. All costae ridged and covered with low rounded granules. Intercostal grooves more prominent at calicular edge and coarsely pointed spines are present in this region. Between C 3 and C 4, intercostal grooves are slightly shallower than others, and just C 1 and C 2 originate at centre of base. C 4 smaller than C 3. Base slightly conical. Corallum white but reddish-brown in well-preserved coralla. Septa hexamerally arranged in 4 complete cycles (S 1> S 2> S 4> S 3). S 1 only independent septa and extend half distance to columella. Each S 1 bears a lamellar paliform lobe, which encircles the columella forming the first palar crown. S 2 less wide and exsert than S 1, bearing a palus more recessed from columella than P 1. S 3 smallest septa, but posses the tallest palus which fuses S 2 in a well-developed lamellar structure. P 3 form outermost crown. S 4 less exsert but slightly wider than S 3. Each pair of S 4 fuses the flanked S 3 with a porous lamella that extend lower in theca almost to columella. All septa and palar faces coarsely granulated. Fossa nonextant containing a well-developed papillose columella formed by 10 - 15 granular rods.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D50FFEBFD4BFBC3032EFA20.taxon	discussion	REMARKS Among the other four non-spined species of Deltocyathus from New Caledonia with four septal cycles, D. cameratus is distinguished by the conical base with pointed centre, calicular margin lancetted, and the well-developed lamellar fusions of the S 4 to P 3 and S 3 to P 2.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	description	(Fig. 1 J)	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	materials_examined	HOLOTYPE. — NORFOLK 2, stn DW 2097 (MNHN- Scl. 2008 - 0034). PARATYPES. — NORFOLK 2, stn DW 2025, 2 specimens (MNHN-Scl. 2008 - 0035, MNHN-Scl. 2008 - 0036); stn DW 2034, 1 specimen (MNHN-Scl. 2008 - 0037); 1 specimen (USNM 1114137); stn DW 2086, 1 specimen (MNHN-Scl. 2008 - 0038); stn DW 2087, 1 specimen (MNHN-Scl. 2008 - 0039); 1 specimen (USNM 1114138); stn DW 2097, 1 specimen (MNHN-Scl. 2008 - 0040). TYPE LOCALITY. — Bank Kaimon Maru (24 ° 44 ’ S, 168 ° 06 ’ E), 580 - 583 m.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	etymology	ETYMOLOGY. — The species name inusitatus (from Latin inusitatus, meaning unusual, extraordinary, strange, rare) refers to the unusual septal arrangement presented in this species.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	distribution	DISTRIBUTION. — New Caledonia region from 23 ° 27 ’ S, 167 ° 51 ’ E to 24 ° 56 ’ S, 168 ° 22 ’ E (Bank Brachiopode, Bank Stylaster, Bank Éponge, and Bank Kaimon Maru).	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	description	DESCRIPTION Corallum free and patellate, with flat to very slightly convex base. Well-preserved coralla show a small and usually convex epicentre boss in base. Specimens range from 11 to 13.4 mm in GCD and 2.9 to 4 mm in HT. Holotype measures 13.3 × 13.2 mm in CD and 3.6 mm in HT. Well-preserved specimens have a circular calice, and worn specimens appear to have a polygonal margin. Calicular margin not lancetted or serrate, projection of all septa being equal. Costae granular being more conspicuous near calicular edge, where they are separated by shallow intercostal grooves. Granules small and low, becoming more slender and taller near calicular edge. All specimens analyzed show a reddish-brown to purple calicular edge, including all septal edges and approximatelly 3 mm of base edge in direction to centre. Septa always hexamerally arranged in 5 incomplete cycles (S 1 ≥ S 2> S 3 ≥ S 5> S 4), normally having 69 to 72 septa. All systems present two dimorphic half systems, one composed of 1 S 1, 1 S 2, 1 S 3, 2 S 4, and 4 S 5, and the other composed of 1 S 1, 1 S 2, 1 S 3 and 2 S 4. All specimens contain 3 half-systems. S 5 oriented to the right of half systems with less septa, and 3 to left. S 1 about 1.8 mm exsert and only independent septa, extending half distance to columella. P 1 well developed and separated from S 1 by a wide notch which contains a slender paliform tooth. S 2 similar in shape and exserteness, but sometimes slightly less wide than S 1. Notch separating S 2 from their paliform lobes less wide than those of S 1, and sometimes bearing a small paliform tooth. At the site that S 3 join S 2 there are usually 1 to 3 paliform teeth. S 3 dimorphic in width, those in the half-system with no S 5 equal to slightly less wide than the flanking S 4 (specially S 4 near S 1), extending ¼ to  ⁄  distance to columella. P 3 as wide as S 3, being separated by a small notch. However, those S 3 in the half-system with S 5 are bigger than the flanking S 4, extending about ⅓ distance to columella. Before joining columella, all P 3 usually bear 1 to 4 paliform teeth. Axial edges of each pair of S 5 solidly fused as a thick lamella to outer edge of adjacent P 4. Usually S 5 beside a primary or secondary septa is wider than other S 5. P 4 of crowded half system join P 3 at approximately  ⁄  distance from columella and in half systems without S 5, P 4 fuse P 3 as thick lamella. P 3 join P 2 near columella and axial eges of P 1 and P 2 are fused to columella. All septa and palar faces bear sparse, pointed granules. Fossa extremely shallow to nonextant, containing a well-developed papillose columella, consisting of 7 to 20 often fused granular rods.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
03DDA36B7D57FFEAFF73FA6702CCFA00.taxon	discussion	REMARKS Among the other 24 Recent species belonging to the genus Deltocyathus, D. inusitatus can most easily be distinguished by its unusually dimorphic septal cycles, one half system composed of 1 S 1, 1 S 2, 1 S 3, 2 S 4, and 4 S 5, and the other composed of 1 S 1, 1 S 2, 1 S 3 and 2 S 4. All specimens consist of 3 half-systems with S 5 oriented to the right of the half systems with less septa, and 3 to the left. Other distinctive characters are its flat base, all septa extending the same distance beyond the calicular margin, and usually a small paliform tooth present between S 1 and P 1. Two coralla from NORFOLK 2 expedition, stn DW 2087 showed evidence of calicular regeneration.	en	Kitahara, Marcelo V., Cairns, Stephen D. (2009): A revision of the genus Deltocyathus Milne Edwards & Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species. Zoosystema 31 (2): 233-248
